taxonID	type	description	language	source
05B3683E83175DC39567AA5195E1784D.taxon	description	Figures 1, 2, 3, 4, 5, 6, 7, 8	en	Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo, Gallao 2, Jonas E. (2021): Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species. ZooKeys 1075: 33-75, DOI: http://dx.doi.org/10.3897/zookeys.1075.67459, URL: http://dx.doi.org/10.3897/zookeys.1075.67459
05B3683E83175DC39567AA5195E1784D.taxon	etymology	Etymology. The species epithet is a derivative form of the Greek noun, σπήλαιον (Latin: caverna), which means cave, in reference to the subterranean habitat where Tityus spelaeus has an established population. It is a noun in apposition.	en	Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo, Gallao 2, Jonas E. (2021): Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species. ZooKeys 1075: 33-75, DOI: http://dx.doi.org/10.3897/zookeys.1075.67459, URL: http://dx.doi.org/10.3897/zookeys.1075.67459
05B3683E83175DC39567AA5195E1784D.taxon	diagnosis	Diagnosis. (Based on female). This species belongs to the Tityus trivittatus species-group (Figs 2, 3). Among members of the group distributed in Brazil (T. carvalhoi Mello-Leitao, 1945; T. charreyroni Vellard, 1932; T. confluens Borelli, 1899; T. fasciolatus Pessoa, 1935; T. jeanvellardi Lourenco, 2001; T. karaja Lourenco, 2016; T. rupestre Lourenco, 2019; T. sylviae Lourenco, 2005, and T. trivittatus Kraepelin, 1898), Tityus spelaeus sp. nov. can be readily recognized. Tityus spelaeus sp. nov.; T. carvalhoi; T. charreyroni; T. confluens; T. fasciolatus; T. rupestre, and T. trivittatus share a subaculear tubercle small, and acute, pointing towards the tip of the aculeus (Fig. 12 A). In contrast, T. jeanvellardi; T. karaja, and T. sylviae exhibit a small and coarse subaculear tubercle that points either towards the tip of the aculeus (T. sylviae) or towards the middle of the aculeus (T. jeanvellardi and T. karaja). On the other hand, Tityus spelaeus sp. nov. and T. sylviae share a very large glandular region occupying beyond the anterior two thirds of the medial region of the pectinal basal piece of female pectines (Figs 6 E, F, 11). In T. carvalhoi; T. charreyroni; T. confluens; T. fasciolatus, and T. trivittatus (females of T. jeanvellardi; T. karaja and T. rupestre are unknow) the glandular region occupies a large area of anterior two thirds of the anteriomedian region of the pectinal basal piece of female pectines (e. g., Fig. 5 A, B). Finally, Tityus spelaeus sp. nov. and T. karaja can be readily distinguished from T. carvalhoi; T. charreyroni; T. fasciolatus; T. jeanvellardi; T. rupestre; T. sylviae, and T. trivittatus, based on the presence of residual spots on tergites (Figs 7, 13), and having the carapace (Figs 7, 8 A) and chericeral manus immaculate (Fig. 8 A). In contrast, T. charreyroni; T. confluens; T. fasciolatus; T. jeanvellardi; T. rupestre; T. sylviae, and T. trivittatus have the carapace and tergites moderately covered with brownish spots and the cheliceral manus with reticulations (except T. jeanvellardi that exhibit a cheliceral manus immaculate).	en	Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo, Gallao 2, Jonas E. (2021): Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species. ZooKeys 1075: 33-75, DOI: http://dx.doi.org/10.3897/zookeys.1075.67459, URL: http://dx.doi.org/10.3897/zookeys.1075.67459
05B3683E83175DC39567AA5195E1784D.taxon	description	Description. Based on the female holotype (MZSP 74633). Male unknown. Total length. Female: 53.52 mm (measurements in Table 4).	en	Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo, Gallao 2, Jonas E. (2021): Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species. ZooKeys 1075: 33-75, DOI: http://dx.doi.org/10.3897/zookeys.1075.67459, URL: http://dx.doi.org/10.3897/zookeys.1075.67459
