taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619807	https://doi.org/10.3897/zookeys.1075.67459.figure1	Figure 1. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI). Maximum likelihood tree (Log-likelihood = - 12896.086), showing species-groups and subgenera. Values on nodes correspond to ultrafast-bootstrap (Ubst) values.	Figure 1. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI). Maximum likelihood tree (Log-likelihood = - 12896.086), showing species-groups and subgenera. Values on nodes correspond to ultrafast-bootstrap (Ubst) values.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619808	https://doi.org/10.3897/zookeys.1075.67459.figure2	Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold.	Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619809	https://doi.org/10.3897/zookeys.1075.67459.figure3	Figure 3. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, Cytochrome c Oxidase I), showing distribution of the characters states exhibited by the female basal pectinal piece (orange: GR absent; grey: medium-sized GR; turquoise: relatively large GR; blue: very large GR) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold. Abbreviations = GR, glandular region.	Figure 3. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, Cytochrome c Oxidase I), showing distribution of the characters states exhibited by the female basal pectinal piece (orange: GR absent; grey: medium-sized GR; turquoise: relatively large GR; blue: very large GR) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold. Abbreviations = GR, glandular region.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619810	https://doi.org/10.3897/zookeys.1075.67459.figure4	Figure 4. Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I-IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm.	Figure 4. Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I-IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619811	https://doi.org/10.3897/zookeys.1075.67459.figure5	Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm.	Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619812	https://doi.org/10.3897/zookeys.1075.67459.figure6	Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm.	Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619813	https://doi.org/10.3897/zookeys.1075.67459.figure7	Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm.	Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619814	https://doi.org/10.3897/zookeys.1075.67459.figure8	Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm.	Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619818	https://doi.org/10.3897/zookeys.1075.67459.figure12	Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D).	Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D).	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619817	https://doi.org/10.3897/zookeys.1075.67459.figure11	Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E).	Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E).	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619819	https://doi.org/10.3897/zookeys.1075.67459.figure13	Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket.	Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619815	https://doi.org/10.3897/zookeys.1075.67459.figure9	Figure 9. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), right pedipalp segments A femur, dorsal view B, C patella B dorsal view C external view. Scale bars: 1.5 mm.	Figure 9. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), right pedipalp segments A femur, dorsal view B, C patella B dorsal view C external view. Scale bars: 1.5 mm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619816	https://doi.org/10.3897/zookeys.1075.67459.figure10	Figure 10. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), chela A external view B dorsal view C ventral view. Scale bar: 3 mm.	Figure 10. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), chela A external view B dorsal view C ventral view. Scale bar: 3 mm.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
05B3683E83175DC39567AA5195E1784D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/619820	https://doi.org/10.3897/zookeys.1075.67459.figure14	Figure 14. Habitat of Tityus (Tityus) spelaeus sp. nov. in the Russao cave A inside landscape of the cave B females on the cave walls.	Figure 14. Habitat of Tityus (Tityus) spelaeus sp. nov. in the Russao cave A inside landscape of the cave B females on the cave walls.	2021-12-07	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.		Zenodo	biologists	Moreno-Gonzalez 1, Jairo A.;Pinto-da-Rocha 1, Ricardo;Gallao 2, Jonas E.			
