identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D82A3EBD6A11493F5DFA5EFDD7FB0E.text	03D82A3EBD6A11493F5DFA5EFDD7FB0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola Sharp 1874	<div><p>Genus Sagola Sharp, 1874</p><p>Sagola Sharp, 1874: 506 . Broun 1880: 134; Reitter 1882: 199; Brendel 1888: 302; Schaufuss 1888: 64; Raffray 1890: 84, 1893: 2, 1904: 492; Casey 1894: 434; Hudson 1923: 365, 1934: 183; Jeannel 1961: 47, 1967: 442; Newton 1985: 195; Newton and Chandler 1989: 18; Kuschel 1990: 48; Klimaszewski et al. 1996: 147; Chandler 2001: 50; Newton and Thayer 2005a; Nomura and Leschen 2006: 241.</p><p>Type Species. Sagola misella Sharp (designated by Oke 1928: 5).</p><p>Diagnosis. The members of the genus Sagola can be distinguished from other genera of Pselaphinae by the following combination of characters: habitus elongate parallel, dorsoventrally slightly flattened; antenna gradually clavate, not abruptly clubbed; dorsum of head with anterior or posterior frontal fovea or both; prosternum with median procoxal fovea and lateral procoxal fovea (Fig. 59p); mesoventrite with promesocoxal fovea (Fig. 59q); metaventrite with lateral metasternal fovea (Fig. 59q); abdominal ventrites IV–VI with basolateral fovea (Fig. 59r); abdominal tergite VI slightly larger than V and smaller than VII (Fig. 59r).</p><p>Comments. Males of many species possess secondary sexual characters such as an enlarged triangular head (Figs. 69, 70z), enlarged hind tarsal claw (Fig. 56k), and/or semicircular depression on the fore femur (Fig. 4u: arrow). Males of other species lack obvious characters that distinguish them from females. Most keys provided here are based on male characters, so recognizing the sex of specimens is important to evaluate secondary sexual characters used in the keys. Male abdominal ventrite IX is usually fragile and partially concealed by ventrite VIII, rendering it simple and reduced in appearance (Fig. 2; male). Females possess a more robust, triangular abdominal ventrite IX bearing a pair of long setae (Fig. 2; female) that are usually visible in ventral view.</p><p>Antennal shape is usually unique within speciesgroups and individual species. However, apparent variations may result from viewing at inconsistent orientations among specimens. Characterizing antennomeres is difficult, but still useful in comparing types or specimens in series.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6A11493F5DFA5EFDD7FB0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6C114A3C99FB34FB7FF998.text	03D82A3EBD6C114A3C99FB34FB7FF998.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola SHARP	<div><p>KEY TO THE SPECIES- GROUPS OF SAGOLA SHARP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Hind-tarsal claw enlarged relative to fore and mid-claw (Fig. 56k)................ ........................ notabilis species-group</p><p>1′. Hind-tarsal claw simple, similar to fore and mid-claw.......................................2</p><p>2(1′). Abdominal tergite V distinctly depressed, bearing processes from anterior and posterior edges and long, dense setae between processes (Fig. 54f)............... ...................... excavata species-group</p><p>2′. Abdominal tergite V simple, unmodified ............................................................... 3</p><p>3(2′). Frontal sulcus reversed Y- or V-shaped, divided with each posterior branch reaching vertexal fovea (Fig. 52p, r: arrows) ......................... furcata species-group</p><p>3′. Frontal sulcus not divided...................4</p><p>4(3′). Fore femur with semicircular depression (Fig. 4u: arrow)...................................5</p><p>4′. Fore femur without semicircular depression....................................................... 13</p><p>5(4). Head transverse (Figs. 25g, 27e) ........6</p><p>5′. Head round or bluntly triangular ........7</p><p>6(5). Anterior frontal fovea small, round, but not externally observable owing to concealment by connected frontal rostrum (Fig. 27e: arrow) ................................... .......................... flavipes species-group</p><p>6′. Anterior frontal fovea elongate, partially covered by frontal rostrum (Fig. 25g) ............................ incisa species-group</p><p>7(5′). Eye relatively more prominent due to depressed frontal gena (Fig. 23c) ......... ............................ valida species-group</p><p>7′. Eye normal and frontal gena not depressed .................................................................... 8</p><p>8(7′). Antennomeres 5–6 enlarged (Fig. 21e) ............................ parva species-group</p><p>8′. Antennomeres not enlarged.................9</p><p>9(8′). Ventral surface of head with pair of processes laterally (Fig. 19j: arrow)....... ........................... pulchra species-group</p><p>9′. Ventral surface of head without processes laterally...................................10</p><p>10(9′). Gular area of head with transverse depression and medial process (Fig. 16i: arrow).............. baiknami species-group</p><p>10′. Gular area of head with small, round depression without medial process......11</p><p>11(10′). Gular area of head with bean-shaped opening (Fig. 8j: arrow); ventral surface of head with pair of short carinae from posterior point of eyes, as long as eye (Fig. 8j).......... rugifrons species-group</p><p>11′. Gular area of head with round opening; ventral surface of head without carinae .........................................................12</p><p>12(11′). Ventral surface of head with large round depression (Fig. 12m: arrow) ................ ............................. bifida species-group</p><p>12′. Ventral surface of head with transverse depression (Fig. 4s: arrow) ................... ........................... hirtalis species-group</p><p>13(4′). Head with dense setae around posterior frontal fovea (Fig. 29a–b) ..................... ........................ helmorei species-group</p><p>13′. Head with sparse setae throughout...14</p><p>14(13′). Ventral surface of head distinctly convex, with heart-shaped depression (Fig. 35d) ......... sharpi species-group</p><p>14′. Ventral surface of head slightly convex or flat, without heart-shaped depression ................................................... 15</p><p>15(14′). Frontal sulcus deep and long, reaching between behind eye to occiput (Fig. 38j–k)........ auripila species-group</p><p>15′. Frontal sulcus not exceeding posterior margin of eye ....................................16</p><p>16(15′). Ventral surface of head with a pair of cone-shaped temporal depressions and small triangular medial processes (Fig. 50c: short arrow) .......................... ........................ owensae species-group</p><p>16′. Ventral surface of head without coneshaped temporal depressions and medial processes............................................17</p><p>17(16′). Head triangular, ventral temple of head depressed and projecting (Figs. 64f, 66n, 70y–z, 75d).......................................... 18</p><p>17′. Head round, ventral temple of head simple.................................................21</p><p>18(17). Posterior frontal fovea large and rounded, as large as eye (Fig. 75d) ........................ ............................. genalis species-group</p><p>18′. Posterior frontal fovea small.............19</p><p>19(18′). Ventral surface of neck with patch of long, dense setae anteriorly (Fig. 66o–t, 69)..... ................................................................ 20</p><p>19′. Ventral surface of neck without patch of setae anteriorly.................................. ................. moehauensis species-group</p><p>20(19). Genitalia robust and convex (Fig. 66g –l) ............................... major species-group</p><p>20′. Genitalia broad and flat (Fig. 70a–x) ...................... insignis species-group</p><p>21(17′). Ventral surface of head with process behind mouthparts bearing dense setae along posterior margin (Figs. 41f, 43d, 77b)............................................... 22</p><p>21′. Ventral surface of head without process behind mouthparts.............................24</p><p>22(21). Ventral surface of head with W-shaped process (Fig. 77b: arrow) ...................... .................... opercularis species-group</p><p>22′. Ventral surface of head with round process (Figs. 41f, 43d)..........................23</p><p>23(22′). Ventral surface of head with distinct triangular process; eye one-third length of temple (Fig. 43d)............................. ........................... strialis species-group</p><p>23′. Ventral surface of head with round process; eye large, as long as temple (Fig. 41f)..... arboricola species-group</p><p>24(21′). Anterior frontal fovea open anteriorly (Fig. 34c: arrow) ................................... ........................ gourlayi species-group</p><p>24′. Anterior frontal fovea closed anteriorly ......................................................... 25</p><p>25(24′). Ventral surface of head convex with setose gula.........................................26</p><p>25′. Ventral surface of head simple..........27</p><p>26(25). Larger, 2.3–2.9 mm total length; posterior frontal fovea elongate (Fig. 31j) .......................... rustica species-group</p><p>26′. Smaller, 1.7–1.9 mm total length; posterior frontal fovea round (Fig. 33a)..... ......................... lescheni species-group</p><p>27(25′). Antennomere 1 approximately 2.0 times longer than wide, with dull surface (Fig. 4q) ............... gilae species-group</p><p>27′. Antennomere 1 approximately 1.5– 2.0 times longer than wide, with surface at least slightly shining (Fig. n) ....................................................... 28</p><p>28(27′). Frontal rostrum rectangular, lobes continuous, reaching posterior frontal fovea (Fig. 59o)........... misella species-group</p><p>28′. Frontal rostrum round, lobes not continuous, not reaching eye if continuous ........................................................ 29</p><p>29(28′). Elytra subquadrate; hind wings reduced to small pads (Fig. 76a–b) .................... .................. flemingensis species-group</p><p>29′. Elytra rectangular; hind wings fully developed...........................................30</p><p>30(29′). Median lobe of genitalia divided vertically, major lobe partially covers minor lobe (Fig. 47f–j)..... socia species-group</p><p>30′. Median lobe of genitalia not divided (Fig. 45f–k)..... anisarthra species-group</p></div>	https://treatment.plazi.org/id/03D82A3EBD6C114A3C99FB34FB7FF998	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6F11443F7EF9BEFD44FDD2.text	03D82A3EBD6F11443F7EF9BEFD44FDD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hirtalis Broun, 1893 b: 1050	<div><p>1. hirtalis species-group (16 species)</p><p>Diagnosis. Members of the hirtalis speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.0–4.0 mm; gular surface of male head with deep, round depression; male antennomeres longer than those of female, antennomere 1 at least 2 times longer than wide, with dull surface; prosternum with lateral procoxal foveae; male fore femur with semicircular depression (Fig. 4u: arrow); abdominal tergites IV–VI with discal carinae; abdominal ventrites IV–VI with basolateral foveae; parameres distinctly asymmetrical and finely punctured (Fig. 4a–p); present on North Island, not known from South Island (Figs. 5–7).</p><p>KEY TO SPECIES OF THE HIRTALIS SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Hind wings reduced to small pads; abdominal tergite IV without patches of microtrichia.....................................2</p><p>1′. Hind wings well-developed; abdominal tergite IV with pair of transverse patches of microtrichia reaching middle............3</p><p>2(1). Smaller body size, length 2.4 mm; antennomere 1 approximately 2 times longer than wide, 4–5 subquadrate; mid-tibia bent (Fig. 12o: arrow) ........ ..................... S. brookesi new species</p><p>2′. Larger body size, length 2.5–2.9 mm; antennomere 1 approximately 2.5 times longer than wide, 4–5 longer than wide; mid-tibia straight ................................... .................. S. hunuaensis new species</p><p>3(1′). Larger body, length 3.2–4.0 mm; antennomere 5 elongate, at least 1.5 times longer than wide.................................... 4</p><p>3′. Smaller body, length 2.0– 3.3 mm; antennomere 5 elongate, not exceeding 1.5 times longer than wide ................... 5</p><p>4(3). Apical lobe of genitalia bird-head-shaped; left paramere with two minor slender lobes, originating from base (Fig. 4b) .................................. S. terricola Broun</p><p>4′. Apical lobe of genitalia shaped differently; left paramere with one lobe originating anteriorly (Fig. 4a).............. ................................... S. hirtalis Broun</p><p>5(3′). Smaller, body length, 2.0– 2.5 mm; antennomere 1 not exceeding 2.5 times longer than wide...............................6</p><p>5′. Larger, body length, 2.4–3.2 mm; antennomere 1 approximately 3 times longer than wide.................................... 8</p><p>6(5). Phallobase of median lobe symmetrical and elongate (Fig. 4c) ........................... ................................. S. convexa Broun</p><p>6′. Phallobase of median lobe asymmetrical and triangular.................................7</p><p>7(6′). Eye small, one-third length of temple; mid-femur and tibia normal; apical lobe of genitalia curved; left paramere rectangular (Fig. 4k).................................... ....................... S. thayerae new species</p><p>7′. Eye large, as long as temple; mid-femur with semicircular depression; mid-tibia bent; apical lobe of genitalia straight; left paramere L-shaped (Fig. 4n) .......... ................ S. gisbornensis new species</p><p>8(5′). Apical lobe of genitalia bulbous and semicircular..........................................9</p><p>8′. Apical lobe not bulbous and not semicircular ...............................................12</p><p>9(8). Process, either slender or triangular, extending from middle of apical lobe of genitalia .........................................10</p><p>9′. Middle of apical lobe of genitalia lacking ...............................................11</p><p>10(9′). Slender process extending from middle of apical lobe of genitalia (Fig. 4e) ...... ............ S. northlandensis new species</p><p>10′. Broad, triangular process extending from middle of apical lobe of genitalia (Fig. 4h).......... S. newtoni new species</p><p>11(9′). Slender process extending from base of apical lobe of genitalia; minor lobes of left paramere folded (Fig. 4j)................ ....................... S. dugdalei new species</p><p>11′. Apical lobe of genitalia lacking process; minor lobes of left paramere weakly bent (Fig. 4i).... S. thorpei new species</p><p>12(8′). Slender and short process extending from apical lobe of genitalia; left paramere lacking minor lobe (Fig. 4d)........ ......... S. coromandelensis new species</p><p>12′. Apical lobe of genitalia lacking process; left paramere divided, forming two lobes ...........................................13</p><p>13(12′). Apical lobe of genitalia deeply divided into two lobes (Fig. 4g) ........................ ....................... S. kuscheli new species</p><p>13′. Apical lobe of genitalia not divided.....................................................14</p><p>14(13′). Apical lobe of genitalia faucet-shaped; minor lobe of left paramere crescentshaped (Fig. 4f)..................................... ............... S. taranakiensis new species</p><p>14′. Apical lobe of genitalia triangular; minor lobe of left paramere divided but not crescent-shaped....................15</p><p>15(14′). Minor lobes of left paramere divided and U-shaped; right paramere rectangular (Fig. 4m) .......................................... ................ S. waikatoensis new species</p><p>15′. Minor lobes of left paramere divided and V-shaped; right paramere L-shaped (Fig. 4o)............... S. weiri new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD6F11443F7EF9BEFD44FDD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6111463F6FFF31FDA1FA82.text	03D82A3EBD6111463F6FFF31FDA1FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hirtalis Broun, 1893 b: 1050	<div><p>Sagola hirtalis Broun, 1893</p><p>(Figs. 3a, 4a, 4q–s, 5)</p><p>Sagola hirtalis Broun, 1893b: 1050 . Raffray 1904: 498, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola cilipes Broun, 1921a: 491 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Type Material Examined. Holotype: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1876. ♂.” [white label, handwritten]; “Howick” [ white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola hirtalis ” [white label, handwritten]. The original label indicates this specimen is a male, but it is female. Syntypes of S. cilipes: 2♂♂ (BMNH), glued on rectangular card, “3999. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Waitakerei. 26.10.1914.” [white label, handwritten]; “ sagola cilipes . ♂ ” [white label, handwritten] .</p><p>Additional Material (n = 7: 4 males; 3 females). NEW ZEALAND: Auckland: 1♂ (slide-mounted), Waitakere Range, 260m, Nohoanga Scenic Res., 8 XII 1984 – 25 I 1985, hdwd.-podocarp forest, A. Newton, M. Thayer, FIT ; 1♀, Titirangi, Waitakere Range, 26 X 1914, A. Brookes ; Northland: 2♂♂ 2♀♀ (1♀ slide-mounted), Waipoua State Forest, 0.8km nw Wairau Summit, 350m, 27 XI 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 689, berl., leaf &amp; log litter forest floor ; 1♂, Waipoua State Forest, Wairau Summit, 400m, 27 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 687, window trap 687 .</p><p>Diagnosis. Specimens of S. hirtalis can be distinguished from the other species of the hirtalis species-group by the following combination of characters: large body size, 3.2–4.0 mm; antennomere 1 elongate, approximately 3 times longer than wide; 4–5 antennomeres elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Redescription. Length 3.2–4.0 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown (Fig. 3a). Head: As long as wide, widest across eyes (Fig. 4r). Antennomere 1 approximately 3 times longer than wide, 2–3 longer than wide, 4–5 elongate, 6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes (Fig. 4r). Anterior fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple (Fig. 4r). Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3a). Hind wings well-developed. Meso-and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle in male and more limited in female. Aedeagus: Median lobe longer than parameres, bearing 2 basal processes (Fig. 4a). Phallobase of median lobe asymmetrical and triangular (Fig. 4a). Left paramere larger than right with 1 slender branch apically (Fig. 4a).</p><p>Type Locality. Near Howick, Auckland, New Zealand .</p><p>Distribution. Auckland, Northland (Fig. 5: black circles).</p><p>Habitat. Specimens of this species were collected using window traps and flight intercept traps, or by sifting leaf or log litter in hardwood and podocarp forests.</p><p>Comments. Specimens of S. hirtalis are difficult to distinguish from those of other species externally, but the shape and size of the antennomeres, size of the eye, presence or absence of hind wings, and genitalia are diagnostic. The type specimens of S. cilipes share these diagnostic characters, and this species has been collected at or near the type locality. For these reasons, we have placed S. cilipes in synonymy with S. hirtalis .</p></div>	https://treatment.plazi.org/id/03D82A3EBD6111463F6FFF31FDA1FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6311473D48FABBFDD0FB79.text	03D82A3EBD6311473D48FABBFDD0FB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola terricola Broun Desig. Park and Carlton 2013	<div><p>Sagola terricola Broun, 1886</p><p>(Figs. 3b, 4b, 4t –u, 5)</p><p>Sagola terricola Broun, 1886: 832 . Raffray 1893: 34, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Type Material. Lectotype. NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1480. ♂.” [white label, handwritten]; “Waitakerei” [red label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola terricola ” [white label, handwritten]; “ LECTOTYPE Sagola terricola Broun Desig. Park and Carlton 2013 ” [white label, printed]. Paralectotype (1 male). NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1480. ♂.” [white label, handwritten]; “Waitakerei” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined (Broun, 1886: 832). This designation will fix the identity of the species and facilitate its recognition by future workers.</p><p>Additional Material (n = 6: 5 males; 1 female). NEW ZEALAND: Northland: 3♂♂ (1♂ slidemounted), Waipoua State Forest, Toatoa Track, 270m, 12–15 IV 1080, toatoa-kauri-podocarpbroadlf, A. Newton, M. Thayer, pitfall trap, berl. leaf and log litter, forest floor; 1♂, Waipoua SF, Ricker Tr., 100m, 11–15 IV 1980, kauri-podocarpbroadleaf, A. Newton, M. Thayer, pitfall trap 619; 1♂, Waipoua SF, Toronui Track, 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd, A. Newton, M. Thayer 685, FIT, window trap; 1♀, SH12, Waipoua SF, 20 IX 1977, D. W. Helmore, rotten stumps and trees 77/102.</p><p>Diagnosis. Specimens of S. terricola can be distinguished from the other species of the hirtalis species-group by the following combination of characters: larger body size, 2.8–3.5 mm; antennomere 1 elongate, approximately 3 times longer than wide; antennomeres 4–5 elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Redescription. Length 2.8–3.5 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown (Fig. 3b). Head: As long as wide, widest across eyes (Fig. 3b). Antennomere 1 approximately 3 times longer than wide, 2 subquadrate, 3 longer than wide, 4–5 elongate, 6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3b). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide (Fig. 4t). Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with bird-head-shape of apical lobe (Fig. 4b). Phallobase of median lobe asymmetrical and triangular (Fig. 4b). Left paramere with 2 slender branches from base, right paramere longer than left (Fig. 4b).</p><p>Type Locality. Waitakerei Range, Auckland, New Zealand.</p><p>Distribution. Auckland, Northland (Fig. 5: triangles).</p><p>Habitat. Specimens of this species were collected using pitfall, window, and flight intercept traps or by sifting leaf or log litter in kauri, broadleaf, hardwood, and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6311473D48FABBFDD0FB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6211403CBCFAE0FC84FC85.text	03D82A3EBD6211403CBCFAE0FC84FC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola convexa Broun 1886	<div><p>Sagola convexa Broun, 1886</p><p>(Figs. 3c, 4c, 5)</p><p>Sagola convexa Broun, 1886: 889 . Raffray 1893: 41, 1904: 498, 1911: 6, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Sagola basalis Broun, 1911: 496 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola indiscreta Broun, 1915: 288 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1583. ♂.” [white label, handwritten]; “Tuakau” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola convexa . ♂.” [white label, handwritten]. Holotype of Sagola basalis: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3368. ♂ ” [white label, handwritten]; “Howick” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola basalis ” [white label, handwritten]. The original label indicates this specimen is a male, but it is a female. Holotype of Sagola indiscreta: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3703. ♂ ” [white label, handwritten]; “Howick” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola indiscreta . ♂ ” [white label, handwritten]. The original label indicates this specimen is a male, but it is female.</p><p>Additional Material (n = 60: 27 males; 33 females). NEW ZEALAND: Auckland: 2♂♂ 2♀♀, Waitakere Ranges Reg. Park, 1.3km N Huia, Parau Track, 133m, 20 III 2010, D.S. Chandler, sift kauri, Cyathea &amp; etc. leaf litter ; 1♂, Waitakere Ra., Cascade kauri Park, up Kauri Tr., 170m, 23 XI–8 XII 1984, kauri-podo-hdwd, A. Newton, M. Thayer 680, FIT &amp; window trap ; 2♀♀, Waitakere Ra., Cascade kauri Park, up Kauri Tr., 170m, 8 XII 1984 – 25 I 1985, kauripodo-hdwd, A. Newton, M. Thayer 680, FIT &amp; window trap ; 1♂ 3♀♀, Manurewa, Murphy’ s Bush, 2 V 1981, G. Kuschel, rotten wood ; 1♂ 2♀♀, Goldie Bush Scenic Res., Mokoroa Falls Track, 6km W Waitakere, 28 III 2910, 106m, D.S. Chandler, rotten wood &amp; branch debris ; 1♂ 1♀, Waitakere Ranges Reg. Park, Upper Huia Reser. Track, 31 I 2010, 340m, 4km SW Waiatarua, D.S. Chandler, sift rotten wood ; 1♂, Hunua Ranges Reg. Park, Kohukohunui Track nr. Mine Rd, 475m, 37″02.396′S, 175″11.251′E, 17 XI–28 XII 2005, broadleaf-podocarp forest, numerous trees ferns, FMHD#2005-001, FIT, A. Newton &amp; M. Thayer, et al., ANMT site 1140 ; 1♂, Waipipi Scen. Res., 2km wsw Waipipi, 40m, 37.14′S 174.40′E, 22 II 1987, native hardwood forest in gully, A. Newton &amp; M. Thayer 835, FMHD#87-269, berl., leaf &amp; log litter, forest floor ; 1♂, Clevedon Scenic Reser., 0.5km N Clevedon, 20m, 19 III 2010, D.S. Chandler, sift forest litter by stream ; 1♀, Flat bush, Murphy’ s bush, 20 VIII 1977, J.C. Watt, podocarp forest litter ; Bay of Plenty: 1♂ 1♀, Tapapa Tukorehe Res., 200m, 25 III 1978, S. &amp; J. Peck, berl. litter ; Northland: 2♂♂ 5♀♀, Parahaki Park, 5 XI 1981, G. Kuschel, litter &amp; decayed wood ; 2♂♂ 1♀, Waipoua SF, Lookout, 29 X 1980, G. Kuschel, decayed wood &amp; litter 80/94 ; 1♂ 3♀♀, Ngaiotonga Res., Walkway, 3 XI 1981, G. Kuschel, litter &amp; decayed wood 81/118 ; 3♂♂ 4♀♀ (1♂ 1♀, slidemounted), Waipoua For., Te Matua Ngahere, 370m, 19 III 1978, S. &amp; J. Peck, berl., rotten kauri log, kauri forest ; 2♂♂ 1♀, Waipoua For., Te Matua Ngahere, 400m, 19 II 1978, berl., under bark of fallen tree with mushroom, S. Peck ; 2♂♂ 1♀, Waipoua SF, 25 XI 1980, G. Kuschel, decayed wood 80/120 ; 2♀♀, Waipoua SF, Toronui Track, 20 X 1980, G. Kuschel, sifted wood &amp; litter 80/95 ; 1♂ 1♀, Waipoua SF, Waikohatu Br., 290m, 11–14 IV 1980, Agathis -podocarp-broadlf, A. Newton, M. Thayer, berl., leaf &amp; log litter, forest floor ; 1♂, Mangamuka stm, 386m, 13 XII 1976, V. A. May, litter 76/109 ; 1♂, Waipoua SF, Yakas Tk, 27 VII–2 VIII 1998, R. Leschen, R. Hoare, FIT2 236, 35′37S 173′32E ; 1♂, Waipoua SF, vic. Wairau Summit, 460m, 27 XI–4 XII 1984, hdwd-podocarp forest, A. Newton, M. Thayer 683, FIT &amp; window trap ; 2♂ (1♂, slide-mounted), Waipoua State Forest, Toronui Track, 120m, 2 XII 1984, kauri-podocarp-hdwd, A. Newton, M. Thayer 685 ; 1♀, Waipoua SF, Toronui Tr., 150m, 13 IV 1980, kauri-podocarp-broadlf-nikau palm forest, A. Newton, M. Thayer, under bark with slime molds ; 1♀, Waipoua SF, Toatoa Tr., 270m, 12–15 IV 1980. toatoa-kauri-podocarp-broadlf, A. Newton, M. Thayer, berl. leaf &amp; log litter ; Wellington: 1♀, Balance Bridge Res., 3 I 1975, J.C. Watt, litter 75/23 .</p><p>Diagnosis. Specimens of S. convexa can be distinguished from the other species of the hirtalis species-group by the following combination of characters: smaller body size, 2.0– 2.4 mm; antennomere 1 elongate, approximately 3 times longer than wide; antennomeres 4–5 elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-half length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Redescription. Length 2.0– 2.4 mm. Body yellowish brown, elytra, legs, and maxillary palpi paler (Fig. 3c). Head: As long as wide, widest across eyes (Fig. 3c). Antennomere 1 approximately 3 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 7–10 subquadrate. Frontal sulcus reaching midpoint of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3c). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe shorter than right paramere with triangular apical lobe (Fig. 4c). Phallobase of median lobe symmetrical and elongate (Fig. 4c). Left paramere shorter than right, setae present along edge (Fig. 4c).</p><p>Type Locality. Tuakau, Auckland, New Zealand .</p><p>Distribution. Auckland, Bay of Plenty, Northland, Wellington (Fig. 5: black squares).</p><p>Habitat. Most specimens of this species were collected using window and flight intercept traps, or by sifting moss or leaf litter in Nothofagus Blume (Nothofagaceae), kauri, broadleaf, hardwood, and podocarp forests. A few specimens were found under bark with mushrooms or slime molds.</p><p>Comments. Specimens of S. convexa are difficult to distinguish from those of other species externally, but the shape and size of the antennomeres, size of eye, presence or absence of hind wings, and genitalia are diagnostic. The type specimens of S. basalis and S. indiscreta share these diagnostic characters, and these species have been collected at or near the type locality of S. convexa . For these reasons, we have placed S. basalis and S. indiscreta as junior synonyms of S. convexa .</p></div>	https://treatment.plazi.org/id/03D82A3EBD6211403CBCFAE0FC84FC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6511413E9FFCB9FDE9FD1A.text	03D82A3EBD6511413E9FFCB9FDE9FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola coromandelensis Park and Carlton	<div><p>Sagola coromandelensis Park and Carlton, new species</p><p>(Figs. 3d, 4d, 5)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: CL: Coromandel Ra. nr Summit, 19 I 1972 G.W. Ramsay, Litter 72/35”, “ HOLOTYPE Sagola coromandelensis Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), Mt. Te Aroha, 2.5km SE Te Aroha, 390m, Tui Road at Tui Creek, Mt. Domain Track, II-15-2010, D.S. Chandler, sift rotten wood .</p><p>Etymology. The species name is based on the biotic region of the type locality, Coromandel.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea elongate; eye small, one-fourth length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description of Male. Length 2.6–3.2 mm.Body yellowish brown, elytra, legs, and maxillary palpi paler (Fig. 3d). Head: As long as wide, widest across eyes (Fig. 3d). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-fourth length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3d). Hind wings well-developed. Meso-and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, bearing one branch apically (Fig. 4d). Phallobase of median lobe asymmetrical and triangular (Fig. 4d). Right paramere divided into 2 branches from base, and minor branch divided into 2 branches at apex (Fig. 4d).</p><p>Distribution. Bay of Plenty, Coromandel (Fig. 5: stars).</p><p>Habitat. Specimens of this species were collected by sifting leaf and rotten wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6511413E9FFCB9FDE9FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6411413CFDFD58FB46FEDB.text	03D82A3EBD6411413CFDFD58FB46FEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola northlandensis Park and Carlton	<div><p>Sagola northlandensis Park and Carlton, new species</p><p>(Figs. 3e, 4e, 5)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand: ND: Omahuta S.F., Kauri Res., litter 74/81 10 Oct 1974, JC Watt”, “ HOLOTYPE Sagola northlandensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the type locality, Northland.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis group by the following combination of characters: antennomere 1 elongate, approximately 3 times longer than wide; antennomere 4 elongate; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description of Male. Length 3.0 mm. Body brown and elytra, legs, maxillary palpi yellowish brown (Fig. 3e). Head: As long as wide, widest across eyes (Fig. 3e). Antennomere 1 approximately 3 times longer than wide, 2–3 longer than wide, 4 elongate, approximately 1.5 times longer than wide, 5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3e). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with semicircular apical lobe (Fig. 4e). Phallobase of median lobe asymmetrical and triangular (Fig. 4e). Right paramere divided into 2 branches from base, and minor branch divided into 2 long branches (Fig. 4e).</p><p>Distribution. Northland (Fig. 5: white circle).</p><p>Habitat. The holotype and only known specimen of this species was collected by sifting litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6411413CFDFD58FB46FEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6411413E90FE9FFB46F99D.text	03D82A3EBD6411413E90FE9FFB46F99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola taranakiensis Park and Carlton	<div><p>Sagola taranakiensis Park and Carlton, new species</p><p>(Figs. 3f, 4f, 5)</p><p>Type Material. Holotype. NEW ZEALAND: Taranaki: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: TK: Awakino Gorge 23 I 1972, G.W. Ramsay Litter 72/30”, “ HOLOTYPE Sagola taranakiensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the type locality, Taranaki.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; antennomere 4 elongate; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description of Male. Length 2.8 mm. Body yellowish brown, elytra, legs, and maxillary palpi paler (Fig. 3f). Head: As long as wide, widest across eyes (Fig. 3f). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4 elongate, approximately 1.5 times longer than wide, 5–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, onethird length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3f). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, with faucet-shaped apical lobe (Fig. 4f). Phallobase of median lobe asymmetrical and triangular (Fig. 4f). Right paramere divided into 2 branches, with minor branch crescent-shaped (Fig. 4f).</p><p>Distribution. Taranaki (Fig. 5: white square).</p><p>Habitat. The holotype and only known specimen of this species was collected by sifting litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6411413E90FE9FFB46F99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6411423EEAF9C1FDC0FA82.text	03D82A3EBD6411423EEAF9C1FDC0FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola kuscheli Park and Carlton	<div><p>Sagola kuscheli Park and Carlton, new species</p><p>(Figs. 3g, 4g, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, ND Unuwhao, 270m 25 Nov 1982 G. Kuschel ”, “sifted litter and decayed wood 82/125”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola kuscheli Park and Carlton 2013 ”. Paratype (1 male). NEW ZEALAND: Northland : 1♂ (NZAC), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype and world renowned beetle specialist, Guillermo Kuschel.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 3 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description. Length 2.4–2.7 mm. Body yellowish brown, elytra, legs,and maxillary palpi paler (Fig. 3g). Head: As long as wide, widest across eyes (Fig. 3g). Antennomere 1 approximately 3 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching hind-point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3g). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, bearing one branch from base (Fig. 4g). Phallobase of median lobe asymmetrical and triangular (Fig. 4g). Left paramere divided into 2 branches from base, with minor branch divided into 2 branches apically (Fig. 4g).</p><p>Distribution. Northland (Fig. 6: black circle).</p><p>Habitat. Specimens of this species were collected by sifting decayed wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6411423EEAF9C1FDC0FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6711433CF4FAB0FDAFFE36.text	03D82A3EBD6711433CF4FAB0FDAFFE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola newtoni Park and Carlton	<div><p>Sagola newtoni Park and Carlton, new species</p><p>(Figs. 3h, 4h, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: ND Waipoua State Forest, 0.8km nw Wairau Summit 350m, 27. xi. 1984 hdwd.-podocarp forest A.Newton / M.Thayer 689 ”, “berl., leaf &amp; log litter, forest floor”, “ HOLOTYPE Sagola newtoni Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Auckland: 1♂ (DSC), Mt. Auckland, Atuanui Scenic Reserve, 250m, 2.5km NE Glorit, 2 II 2010, D.S. Chandler, sift Nikau palm litter .</p><p>Etymology. This species is named for one of the collectors of the holotype and world renowned beetle specialist, Alfred F. Newton.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description. Length 2.4–2.7 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown (Fig. 3h). Head: As long as wide, widest across eyes (Fig. 3h). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4 longer than wide, 5–7 subquadrate, 8–10 transverse. Frontal sulcus deep and reaching hind-point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3h). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with semicircular apical lobe (Fig. 4h). Phallobase of median lobe asymmetrical and triangular (Fig. 4h). Left paramere longer than right (Fig 4h).</p><p>Distribution. Auckland, Northland (Fig. 6: triangles).</p><p>Habitat. Specimens were collected by sifting nikau palm, hardwood, and podocarp forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6711433CF4FAB0FDAFFE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6611433EEBFE4FFB93F95B.text	03D82A3EBD6611433EEBFE4FFB93F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola dugdalei Park and Carlton	<div><p>Sagola dugdalei Park and Carlton, new species</p><p>(Figs. 3j, 4j, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND, Omahuta S.F. 22 Apr 1975 J.S. Dugdale ”, “Liverworts &amp; mosses 75/141”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola dugdalei Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Northland: 1♂ (NZAC), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype, John S. Dugdale.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 3 times longer than wide; antennomeres 4–5 elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description. Length 2.4–3.2 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 3j). Head: As long as wide, widest across eyes (Fig. 3j). Antennomere 1 approximately 3 times longer than wide, 2–3 longer than wide, 4–5 elongate, 6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind-point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3j). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with semicircular apical lobe, bearing small branch from base (Fig. 4j). Phallobase of median lobe asymmetrical and triangular (Fig. 4j). Right paramere divided into 2 branches from base and minor branch divided into 2 branches apically (Fig. 4j).</p><p>Distribution. Northland (Fig. 6: star).</p><p>Habitat. Specimens of this species were collected by sifting liverwort and moss litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6611433EEBFE4FFB93F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD6611433CDAFE25FBE7FE0B.text	03D82A3EBD6611433CDAFE25FBE7FE0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola thorpei Park and Carlton	<div><p>Sagola thorpei Park and Carlton, new species</p><p>(Figs. 3i, 4i, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (AMNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, CL Cuvier I., Old Radar Station, 180m, 10–18.xi.1999, J.W. Early, S.E. Thorpe ”, “ In forest, Yellow pan trap, L8170”, “ AMNZ 30285 AUCKLAND MUSEUM NEW ZEALAND ”, “ HOLOTYPE Sagola thorpei Park and Carlton 2013 ” . Paratypes (n = 3: 2 males; 1 female). NEW ZEALAND: Coromandel: 1♂ (AMNZ), Great Barrier I, Little Windy Hill, 220m, 22 V –7 VII 2002, P. Sutton, forest edge malaise trap ; 1♂ (AMNZ), Great Barrier I, Mt. Hobson, 500m, pitfall trap, 9 XI–17 XII 2003, J. W. Early ; 1♀ (AMNZ), slide-mounted, Great Barrier I, Little Windy Hill, 20 XI–19 XII 2002, K. Parsons, in forest pitfall trap .</p><p>Etymology. This species is named for one of the collectors of the holotype, Stephen E. Thorpe.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 3 times longer than wide; antennomere 4 elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description. Length 2.7–3.2 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 3i). Head: As long as wide, widest across eyes (Fig. 3i). Antennomere 1 approximately 3 times longer than wide, 2–3 subquadrate, 4 elongate, 5–6 longer than wide, 7–9 subquadrate, 10 transverse. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea oval, posterior frontal fovea oval. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3i). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse, reaching middle, patches of microtrichia. Aedeagus: Median lobe longer than parameres with semicircular apical lobe (Fig. 4i). Phallobase of median lobe asymmetrical and triangular (Fig. 4i). Left paramere divided into 2 branches from base, with minor branch divided into 2 branches apically (Fig. 4i).</p><p>Distribution. Coromandel (Fig. 6: black squares).</p><p>Habitat. Specimens were collected using yellow pan, Malaise, and pitfall traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD6611433CDAFE25FBE7FE0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD79115C3E86FE52FC12F99D.text	03D82A3EBD79115C3E86FE52FC12F99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hunuaensis Park and Carlton	<div><p>Sagola hunuaensis Park and Carlton, new species</p><p>(Figs. 3l, 4l, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: AK: Hunua Ra., Kohukohunui, 550m, 30III1974 G. Kuschel, litter 74/19”, “ HOLOTYPE Sagola hunuaensis Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Auckland: 1♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named after the type locality, Hunua, Auckland.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings reduced to small pads; shape of genitalia unique to species.</p><p>Description. Length 2.5–2.9 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown (Fig. 3l). Head: As long as wide, widest across eyes (Fig. 3l). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at onethird length. Elytra rectangular (Fig. 3l). Hind wings absent. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without microtrichia. Aedeagus: Median lobe longer than parameres, apical lobe bird-head-shaped (Fig. 4l). Phallobase of median lobe asymmetrical and triangular (Fig. 4l). Left paramere elongate and right paramere short and rounded (Fig. 4l).</p><p>Distribution. Auckland (Fig. 6: white square).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD79115C3E86FE52FC12F99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD79115C3CE8FF31FC45FEEC.text	03D82A3EBD79115C3CE8FF31FC45FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola thayerae Park and Carlton	<div><p>Sagola thayerae Park and Carlton, new species</p><p>(Figs. 3k, 4k, 6)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), “ NEW ZEALAND: ND: Waipoua SF, Waikohatu Br. 290m, 11–14.iv.80 Agathis-podocarp-broadlf. A.Newton, M. Thayer ”, “berl, leaf &amp; log litter, forest floor”, “ HOLOTYPE Sagola thayerae Park and Carlton 2013 ” . Paratypes (n = 7: 6 males; 1 female). NEW ZEALAND: Northland: 1♂ (slide-mounted; FMNH), Waipoua Forest, Waipoua Stream, 70m, 16–21 II 1978, berl. litter, kauri forest, S. Peck, J, Peck ; 1♂ (DSC), Waipoua State Forest, Toronui Track, 120m, 26 XI–4 XII 1984, kauri-podocarphdwd, A. Newton, M. Thayer 685 ; 1♂ (FMNH), Waipoua State Forest, Kauri Ricker Track, 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd., A. Newton, M. Thayer 684 ; 1♂ (DSC), Waipoua State Forest, 0.9km e Forest Headquarters, 120m, 26 XI-4 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 686 ; 1♂ (NZAC), Waipoua S.F., Te Matua Ngahere, 4 II 1975, J.C. Watt, litter 75/94; Waipoua S.F., Lookout, 29 XI 1980, G. Kuschel, moss 80/99 (NZAC) ; 1♀ (NZAC), Waipoua Stream, 100m, 19 III 1978, S. B. Peck, litter .</p><p>Etymology. This species is named for one of the collectors of the holotype and world renowned beetle specialist, Margaret K. Thayer.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: smaller body size, 2.1–2.6 mm; antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings welldeveloped; shape of genitalia unique to species.</p><p>Description. Length 2.1–2.6 mm. Body yellowish brown, elytra, legs, and maxillary palpi paler (Fig. 3k). Head: As long as wide, widest across eyes (Fig. 3k). Antennomere 1 approximately 2.5 times longer than wide, 2 subquadrate, 3 longer than wide, 4 elongate, 5–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3k). Hind wings welldeveloped. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres (Fig. 4k). Phallobase of median lobe asymmetrical and triangular (Fig. 4k). Right paramere divided into 2 branches from base, with minor, slender branch bearing setae (Fig. 4k).</p><p>Distribution. Northland (Fig. 6: white circle).</p><p>Habitat. Specimens of this species were collected by sifting leaf or log litter in kauri, broadleaf, hardwood, Agathis Salisb. (Araucariaceae), and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD79115C3CE8FF31FC45FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD79115D3EABF9C1FE7DFA82.text	03D82A3EBD79115D3EABF9C1FE7DFA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola waikatoensis Park and Carlton	<div><p>Sagola waikatoensis Park and Carlton, new species</p><p>(Figs. 3m, 4m, 7)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “NEW ZEALAND WO Hapuakohe Ra Maungakawa, 500m 3 Jan 1984 J.C. Watt ”, “ Sifted wood mould 84/3 ”, “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “HOLOTYPE Sagola waikatoensis Park and Carlton 2013 ”. Paratypes (n = 5: 1 male; 4 females). NEW ZEALAND: Waikato: 1♂ 4♀♀ (1♀, slide-mounted; NZAC), same data as holotype.</p><p>Etymology. The species name is based on the biotic region of the type locality, Waikato.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea oval; eye small, one-third length of temple; hind wings welldeveloped; shape of genitalia unique to species.</p><p>Description. Length 2.7–3.3 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 3m). Head: As long as wide, widest across eyes (Fig. 3m). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye small and prominent, onethird length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3m). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, bearing small process from apex (Fig. 4m). Phallobase of median lobe asymmetrical and triangular (Fig. 4m). Right paramere divided into 2 branches with an additional minor branch divided into 2 branches apically (Fig. 4m).</p><p>Distribution. Waikato (Fig. 7: black circle).</p><p>Habitat. Specimens of this species were collected by sifting wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD79115D3EABF9C1FE7DFA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD78115E3C81FAB0FE78FE5C.text	03D82A3EBD78115E3C81FAB0FE78FE5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola gisbornensis Park and Carlton	<div><p>Sagola gisbornensis Park and Carlton, new species</p><p>(Figs. 3n, 4n, 7)</p><p>Type Material. Holotype. NEW ZEALAND: Gisborne: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: GB: Lake Waikaremoana 17 I 1972, G.W. Ramsay Litter 72/20”, “ HOLOTYPE Sagola gisbornensis Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Waikato: 1♂ (NZAC), Okauia, 28 XII 1947, Leaf-mould, A. E. Brookes .</p><p>Etymology. The species name is based on the biotic region of the type locality, Gisborne.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: smaller body size, 2.0– 2.5 mm; antennomere 1 elongate, approximately 2 times longer than wide; antennomeres 4–5 subquadrate, approximately 1.5 times longer than wide; posterior frontal fovea oval; eye large, as long as temple; hind wings well-developed; mid-femur with semicircular depression; mid-tibia bent; shape of genitalia unique to species.</p><p>Description of Male. Length 2.0 – 2.5 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 3n). Head: As long as wide, widest across eyes (Fig. 3n). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from end of frontal rostral lobes. Anterior frontal fovea oval, posterior frontal fovea oval. Eye large and prominent, as long as temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular and at least 1.5 times longer than wide (Fig. 3n). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Midfemur with semicircular depression. Mid-tibia bent. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres (Fig. 4n). Phallobase of median lobe asymmetrical and triangular (Fig. 4n). Left paramere L-shaped and longer than right. Right paramere divided into 3 branches from base (Fig. 4n).</p><p>Distribution. Gisborne, Waikato (Fig. 7: triangles).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD78115E3C81FAB0FE78FE5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7B115E3E81FF31FC74FAD9.text	03D82A3EBD7B115E3E81FF31FC74FAD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola brookesi Park and Carlton	<div><p>Sagola brookesi Park and Carlton, new species</p><p>(Figs. 3p, 4p, 7)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ Fairburn Mt. Te Aroha 30-3-51”, “ A.E.Brookes Collection ”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola brookesi Park and Carlton 2013 ”.</p><p>Etymology. This species is named for the collector of the holotype and world renowned insect specialist, Albert E. Brookes.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: smaller body size, 2.4 mm; antennomere 1 elongate, approximately 2 times longer than wide; antennomeres 2–10 subquadrate; posterior frontal fovea oval; eye large, one-half length of temple; hind wings reduced to small pads; mid-tibia bent; shape of genitalia unique to species.</p><p>Description of Male. Length 2.4 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 3p). Head: As long as wide, widest across eyes (Fig. 3p). Antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from end of frontal rostral lobes. Anterior frontal fovea oval, posterior frontal fovea oval. Eye large and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3p). Hind wings absent. Meso- and metathorax trapezoidal, as long as wide. Mid-tibia bent. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe longer than parameres. Phallobase of median lobe asymmetrical and triangular (Fig. 4p). Left paramere divided into 2 branches from base and minor branch long and bent (Fig. 4p). Right paramere L-shaped and as long as left (Fig. 4p).</p><p>Distribution. Waikato (Fig. 7: star).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7B115E3E81FF31FC74FAD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7B115E3CFAFE02FDD2F95B.text	03D82A3EBD7B115E3CFAFE02FDD2F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola weiri Park and Carlton	<div><p>Sagola weiri Park and Carlton, new species</p><p>(Figs. 3o, 4o, 7)</p><p>Type Material. Holotype. NEW ZEALAND: Taranaki: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: TK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=174.78334&amp;materialsCitation.latitude=-38.933334" title="Search Plazi for locations around (long 174.78334/lat -38.933334)">Waitaanga Forest</a> Campsite 38°56S 174°47E, Punky wood, berlese, 11 Mar 2000 C. Carlton, A Weir, #039”, “ HOLOTYPE Sagola weiri Park and Carlton 2013 ”.</p><p>Etymology. This species is named for one of the co-collectors of the holotype, Alex Weir.</p><p>Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; antennomere 4 elongate, approximately 1.5 times longer than wide; posterior frontal fovea oval; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.</p><p>Description of Male. Length 3.0 mm. Body brown, elytra, legs, maxillary palpi paler (Fig. 3o). Head: As long as wide, widest across eyes (Fig. 3o). Antennomere 1 approximately 2.5 times longer than wide, 2 subquadrate, 3 longer than wide, 4 elongate, 5–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from end of frontal rostral lobes. Anterior frontal fovea oval, posterior frontal fovea oval. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 3o). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, bearing 1 branch at apex (Fig. 4o). Phallobase of median lobe asymmetrical and triangular (Fig. 4o). Left paramere divided into 2 branches from base and minor branch divided into 2 branches apically (Fig. 4o). Right paramere L-shaped and longer than left (Fig. 4o).</p><p>Distribution. Taranaki (Fig. 7: black square).</p><p>Habitat. Specimens of this species were collected by sifting punky wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7B115E3CFAFE02FDD2F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7B11583F75FA8BFD45FEFE.text	03D82A3EBD7B11583F75FA8BFD45FEFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola rugifrons Broun Desig. Park and Carlton 2013	<div><p>2. rugifrons species-group</p><p>(4 species)</p><p>Diagnosis. The members of the rugifrons species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.1–3.4 mm; antennomere 1 at least 2 times longer than wide with dull surface; gular region of male head with bean-shaped opening and transversely open inside (Fig. 8j: arrow); ventral surface of male head with pair of short carinae from posterior margin of eyes, as long as eye (Fig. 8j); male fore femur with semicircular depression; male mid-femur with shallow depression; male mid-tibia bent; abdominal tergites IV–VI with discal carinae; present on North Island, not known from South Island (Figs. 9–10).</p><p>KEY TO SPECIES OF THE RUGIFRONS SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Body large, length 3.0– 3.4 mm; head weakly transverse; frontal rostrum with distinctive punctures; densely setose around posterior frontal fovea.................................. ................................ S. brouni new species</p><p>1′. Body smaller, length 2.1–2.6 mm; head as long as wide; not densely setose around posterior frontal fovea; frontal rostrum without punctures .................................... 2</p><p>2(1′). Antennomeres 4–8 longer than wide; major process of median lobe of genitalia slender .................................................................... 3</p><p>2′. Antennomeres 4–8 subquadrate; major process of median lobe of genitalia broad and rectangular (Fig. 8h) .................................... ......................... S. plentyensis new species</p><p>3(2). Apical major process of median lobe of genitalia arrowhead-shaped (Fig. 8b)........ ................... S. waipouaensis new species</p><p>3′. Apical major process of median lobe of genitalia triangular (Fig. 8a)...................... ................................... S. rugifrons Broun</p></div>	https://treatment.plazi.org/id/03D82A3EBD7B11583F75FA8BFD45FEFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7D11593D4DFE62FED8F95B.text	03D82A3EBD7D11593D4DFE62FED8F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola rugifrons Broun Desig. Park and Carlton 2013	<div><p>Sagola rugifrons Broun, 1895</p><p>(Figs. 8a, 8e, 9)</p><p>Sagola rugifrons Broun, 1895: 73 . Raffray 1904: 498, 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola puncticeps Broun, 1911: 489 . Hudson 1923: 365, 1923: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material Examined. L e c t o t y p e o f Sagola rugifrons: NEW ZEALAND: Waikato: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “2722. ♂ ” [white label, printed]; “ Waikato ” [white label]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola rugifrons ” [white label, handwritten]; “ LECTOTYPE Sagola rugifrons Broun Desig. Park and Carlton 2013 ” [red label, printed]. Paralectotype (1 female) . NEW ZEALAND: Waikato: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2722. ♀ ” [white label, printed]; “ Waikato ” [white label]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola rugifrons ” [white label, handwritten]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined (Broun 1895: 73). This designation will fix the identity of the species and facilitate its recognition by future workers. Holotype of Sagola puncticeps: NEW ZEALAND: Coromandel: 1♂ (BMNH), “3361. ♂ ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Tairua. Auckland.” [white label]; “ Sagola puncticeps ” [white label, handwritten] .</p><p>Additional Material (n = 23: 14 males; 9 females). NEW ZEALAND: Auckland: 1♂ 2♀♀, Hunua Range, Vining Reserve, Mangatangi Tr., 240m, 37″07.747′S, 175″13.024′E, 17 XI 2005, Nothofagus - Agathis -Phyllocladus trichomomoides, etc, FMHD #2005-008, berl., leaf &amp; log litter, A. Newton, S. Solodovnikov, et al., ANMT site 1141; 1♂, Waitakere Ra., Cascade-Kauri Park, Anderson Tr., 85m, 23 XI 1984, hdwd,-podocarp, A. Newton, M. Thayer 681; 1♂, Bethells, Matuku Res., 25 II 1992, G. Hall, litter 92/5; 1♂, Mt. Dome, 15 III 1978, S.B, Peck, litter; 1♀, Kirks Bush, Papakura, 26 V 1992, J. Nunn; Bay of Plenty: 1♂, Okauia, 30 X 1921, A.E. Brookes collection; 1♂, Mt. Te Aroha, 2.5km SE Te Aroha, 390m , Tui Rd. at Tui Creek, Mt. Domain Tr ., 15 II 2010, D.S. Chandler, sift forest leaf litter; Northland: 2♂♂ (1♂, slide-mounted), Mangamuka Walk, 28 VII–1 VIII 1998, R. Leschen, R. Hoare, FIT 223, 36 ′11S, 173′28E; 1♂, Mangamuka Gorge SR, 6.6km nw Mangamuka, 70m , 25 XI–5 XII 1984, hdwd.- podocarp forest, A. Newton, M. Thayer 682, window trap; 1♂, Puketi Forest, 18 XII 1951, moss; 1♀, Tangihua Range, 17 X 1936, F. Fairburn; 1♀, Waipoua SF, Yakas Tree tr., 350m , 11–14 IV 1980, broadleaf.-podocarp, A. Newton, M. Thayer, berl, litter at bases of Metrosideros robusta trunks; 1♀, Mangamuka Summit, 386m , 13 XII 1976, V. A. May, litter 76/109; 1♀, Waipoua SF, 1km E Headquarters, 90m , 15 IV 1980, J.C. Watt, litter80/50; Waikato: 1♂, Pirongia Forest Park, Tirohanga Tr. (above end Corcoran rd.), 500m , 37″58.409′S 175″07.173′E, 20 XI 2005, broadleaf forest, FMHD#2005-015, berl., forest litter, D. Clarke, A. Solodovnikov, ANMT site 1146; 1♂, Piripiri Caves Reserve, 25 V 1986, C.L. Lyal, leaf litter CL 480; 1♂, Mt. Karioi near base, II X1981, C.F. Butcher, litter 81/96; 1♂, Waitomo, Maori Tumutumu Rd ., 24 V 1983, J.C. Watt, litter and soil 83/60; 1♀, Te Mata Bridal, Veil Falls, 19 IX 1981, C.F. Butcher, litter 81/88; 1♀, Hapuakohe Ra ., 29 I 1984, J.C. Watt, sieved leaf litter 84/8.</p><p>Diagnosis. This species can be distinguished from the other species of the rugifrons speciesgroup by the following combination of characters: smaller body, length 2.3–2.6 mm; frontal rostrum with distinctive punctures; antennomere 1 approximately 2.5 times longer than wide, 2–8 longer than wide; posterior frontal fovea surrounded by dense setae; ventral surface of male head with weak pair of short carinae behind eyes; shape of genitalia unique to species.</p><p>Redescription. Length 2.3–2.6 mm. Body brown, antennae, legs, and maxillary palpi paler (Fig. 8a). Head: Round, widest across eyes (Fig. 8a). Frontal rostrum with distinctive punctures. Ventral surface of male head with weak pair of short carinae behind eyes. Antennomere 1 approximately 2.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea oval. Posterior frontal fovea round. Male eye large and prominent, two-thirds length of temple. Female eye small one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular in male (Fig. 8a), shorter in female. Hind wings welldeveloped in male, reduced to small pads in female. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe deeply divided into 2 lobes (Fig. 8e). Phallobase of median lobe nearly symmetrical and rounded (Fig. 8e). Right paramere broader than left (Fig. 8e).</p><p>Type Locality. Mount Pirongia, Waikato, New Zealand .</p><p>Distribution. Auckland, Bay of Plenty, Northland, Waikato (Fig. 9: black circles).</p><p>Habitat. Most specimens of this species were collected in flight intercept traps and sifting leaf litter or logs in broadleaf, hardwood, Nothofagus, or podocarp forests.</p><p>Comments. Specimens of S. rugifrons can be distinguished from other species externally by the frontal rostrum with distinctive punctures; female hind wings reduced to small pads; and the female eye and antenna size and shape. The type specimens of S. puncticeps share these diagnostic characters, and this species has been collected at or near the type locality of S. rugifrons . For these reasons, we have placed this species in synonymy with S. rugifrons .</p><p>(black circles).</p></div>	https://treatment.plazi.org/id/03D82A3EBD7D11593D4DFE62FED8F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7C115A3E90FF31FDB2FCCF.text	03D82A3EBD7C115A3E90FF31FDB2FCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola waipouaensis Park and Carlton	<div><p>Sagola waipouaensis Park and Carlton, new species</p><p>(Figs. 8b, 8f, 8i–j, 10)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: ND: Waipoua SF, Yakas Tree Tr. 350m 11–14.iv.1980 broadlf.-podocarp A. Newton, M. Thayer ”, “berl, litter at bases of Metrosideros rubusta trunks”, “ HOLOTYPE Sagola waipouaensis Park and Carlton 2013 ” . Paratypes (n = 11: 9 males; 2 females). NEW ZEALAND: Auckland: 1♂ (NZAC), Whangateau, 27 XII 1951, P625; Northland: 5♂♂ 1♀ (2♂♂, DSC; 3♂♂ 1♀, FMNH), Waipoua SF, Toronui Tr., 150m, 13 IV 1980, kauri-podocarp-broadlf.-nikau palm forest, A. Newton, M. Thayer, berl., leaf &amp; log litter, forest floor; 1♂ (DSC), Waipoua SF, 0.9km e Forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 686, leaf &amp; log litter, forest floor; 1♂ (slide-mounted; FMNH), Waipoua SF, 0.9km e forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 686, FIT &amp; window trap ; 1♂ (DSC), Waipoua SF, 1.7km nw Waikohatu Stream Bridge, 380m, 28 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 691, FIT, window trap ; 1♂ (FMNH), Waipoua SF, Kauri Ricker Track, 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd., A. Newton, M. Thayer 684, berl., leaf &amp; log litter, forest floor; 1♀ (FMNH), Waipoua SF, kauri Ricker Tr., 100m, 11–15 IV 1980, kauri-podocarp-broadleaf, A. Newton, M. Thayer, berl., leaf &amp; log litter, forest floor.</p><p>Etymology. This species is named after the type locality, Waipoua, Auckland.</p><p>Diagnosis. This species can be distinguished from the other species of the rugifrons speciesgroup by the following combination of characters: smaller body, length 2.1–2.6 mm; frontal rostrum with distinctive punctures; antennomere 1 approximately 2.5 times longer than wide, 4–8 longer than wide; round posterior frontal fovea without surrounding dense setae; ventral surface of male head with distinctive pair of short carinae behind eyes; shape of genitalia unique to species.</p><p>Description. Length 2.1–2.6 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 8b). Head: As long as wide, widest across eyes (Fig. 8i). Frontal rostrum with distinctive punctures. Ventral surface of male head with distinctive pair of short carinae behind eyes (Fig. 8j). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea round. Eye small and prominent, one-half length of temple. Female eye small one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular in male (Fig. 8b), subquadrate in female. Hind wings present in male, absent in female. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe shorter than parameres, divided in 2 lobes. Apical lobe of major lobe triangular (Fig. 8f). Minor lobe slender and forked into 3 branches (Fig. 8f). Phallobase of median lobe nearly symmetrical and rounded (Fig. 8f).</p><p>Distribution. Auckland, Northland (Fig. 10: black circles).</p><p>Habitat. Specimens of this species were collected using window and flight intercept traps, or by sifting leaf and log litter in kauri, broadleaf, hardwood, nikau palm, and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7C115A3E90FF31FDB2FCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7F115A3E89FF31FBC3FA67.text	03D82A3EBD7F115A3E89FF31FBC3FA67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola brouni Park and Carlton	<div><p>Sagola brouni Park and Carlton, new species</p><p>(Figs. 8c, 8g, 10)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “19.ix.51 La Roy’ s Bush Birkenhead N.I. P282”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola brouni Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Coromandel: 1♂ (NZAC), Little Barrier Island, 31 XII 1951, rotten wood, P622 .</p><p>Etymology. This species is named for the most influential pioneer of the study of New Zealand Coleoptera, including Faronitae, Thomas Broun.</p><p>Diagnosis. This species can be distinguished from the other species of the rugifrons speciesgroup by the following combination of characters: larger body, length 3.0– 3.4 mm; weakly transverse head; antennomere 1 approximately 2.5 times longer than wide, 2–7 subquadrate, 8–10 transverse; elongate posterior frontal fovea surrounded by dense setae; ventral surface of male head with weak pair of short carinae behind eyes; shape of genitalia unique to species.</p><p>Description of Male. L e n g t h 3. 0 –3.4 m m. Body brown, elytra, legs, and maxillary palpi paler (Fig. 8c). Head: Slightly transverse, widest across eyes (Fig. 8c). Ventral surface of male head with weak pair of short carinae behind eyes. Antennomere 1 approximately 2.5 times longer than wide, 2–7 subquadrate, 8–10 transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea elongate, surrounded by dense setae. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 8c). Hind wings welldeveloped. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, forked into 3 lobes, major apical lobe bulbous with 9 branches apically (Fig. 8g). Phallobase of median lobe symmetrical and rounded (Fig. 8g).</p><p>Distribution. Auckland (Fig. 10; triangles).</p><p>Habitat. Specimens of this species were collected by sifting rotten wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7F115A3E89FF31FBC3FA67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7F115B3EB2F9C1FE4FFA07.text	03D82A3EBD7F115B3EB2F9C1FE4FFA07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola plentyensis Park and Carlton	<div><p>Sagola plentyensis Park and Carlton, new species</p><p>(Figs. 8d, 8h, 10)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: BP: Ikitiki Stream Mamaku Plateau Horohoro S.F. 24 VII 1976, J.S. Dugdale Litter 76/48”, “HOLOTYPE Sagola plentyensis Park and Carlton 2013 ”. Paratypes (4 males). NEW ZEALAND: Bay of Plenty: 2♂♂ (NZAC), same data as holotype; 1♂ (NZAC), Tapapa, 300m, 25 III 1978, S.B. Peck, litter; Waikato: 1♂ (NZAC), Kaimai Ranges, Matamata, A. E. Brookes, V 1920.</p><p>Etymology. The species name is based on the biotic region of the type locality, Bay of Plenty.</p><p>Diagnosis. This species can be distinguished from the other species of the rugifrons speciesgroup by the following combination of characters: smaller body, length 2.1–2.6 mm; frontal rostrum with distinctive punctures; antennomere 1 approximately 2 times longer than wide, 2–8 subquadrate, 9–10 transverse; round posterior frontal fovea not surrounded by dense setae; ventral surface of male head with weak pair of short carinae behind eyes; shape of genitalia unique to species.</p><p>Description of Male. Length 2.1–2.6 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 8d). Head: As long as wide, widest across eyes (Fig. 8d). Frontal rostrum with distinctive punctures. Ventral surface of male head with weak pair of short carinae behind eyes. Antennomere 1 approximately 2 times longer than wide, 2–8 subquadrate, 9–10 transverse. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea round. Eye small and prominent, onehalf length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 8d). Hind wings well-developed. Meso-and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres and divided, major lobe rectangular (Fig. 8h). Phallobase of median lobe slightly symmetrical and rounded (Fig. 8h).</p><p>Distribution. Bay of Plenty, Waikato (Fig. 10: black squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7F115B3EB2F9C1FE4FFA07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7E11543D52FA3BFD44FE26.text	03D82A3EBD7E11543D52FA3BFD44FE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola bifida Broun 1915	<div><p>3. bifida species-group</p><p>(11 species)</p><p>Diagnosis. The members of the bifida speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.4–3.2 mm; antennomere 1 approximately 2 times longer than wide with dull surface; ventral surface of male head depressed transversely, at least one-half covered with tubular structures on each side (Fig. 12m), with dense setae along edge; hind wings reduced to small pads; male fore femur with semicircular depression; male mid-femur with shallow depression (Fig. 12n: arrow); male mid-tibia bent (Fig. 12o: arrow); abdominal tergites IV–VI with discal carinae; present on North Island, not known from South Island (Figs. 13–14).</p><p>KEY TO SPECIES OF THE BIFIDA SPECIES- GROUP</p><p>The key is based on genitalia because most specimens are indistinguishable based on external morphology.</p><p>1. Median lobe of genitalia deeply divided, major lobe broad and covering minor (Fig. 12f)............. S. mayae new species</p><p>1′. Median lobe of genitalia not divided...2</p><p>2(1′). Median lobe of genitalia shorter than both parameres; right paramere twisted............................................................3</p><p>2′. Median lobe of genitalia longer than at least one of parameres; right paramere not twisted .............................................4</p><p>3(2). Apex of right paramere broad (Fig. 12c) .............................. S. crawi new species</p><p>3′. Apex of right paramere simple (Fig. 12a) ....................................... S. bifida Broun</p><p>4(2′). Right paramere with long minor branch ...............................................................5</p><p>4′. Right paramere without branch............7</p><p>5(4). Apical lobe of genitalia bent ventrally (Fig. 12d).............. S. earlyi new species</p><p>5′. Apical lobe of genitalia straight...........6</p><p>6(5′). Median lobe of genitalia cylindrical, longer than right paramere (Fig. 12b) .... ................................. S. latistriata Broun</p><p>6′. Median lobe of genitalia broad vertically, shorter than right paramere (Fig. 12e) .................. S. embersoni new species</p><p>7(4′). Left paramere with U-shaped depression apically (Fig. 12g)................................... .................... S. ranatungae new species</p><p>7′. Left paramere without depression.........8</p><p>8(7′). Median lobe of genitalia shorter than left paramere, with simple apical lobe (Fig. 12i)...... S. tekoauensis new species</p><p>8′. Median lobe of genitalia longer than parameres with modified apical lobe....9</p><p>9(8′). Apical lobe of genitalia broad with acute lobe behind apex (Fig. 12h) ................... ........................ S. baylessae new species</p><p>9′. Apical lobe of genitalia round and blunt .......................................................... 10</p><p>10(9′). Apical lobe of genitalia with semicircular depression; phallobase of genitalia transverse (Fig. 12k) ....................................... ........................ S. fairburni new species</p><p>10′. Apical lobe of genitalia with branch; phallobase of genitalia as long as wide (Fig. 12j)............... S. ferroi new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD7E11543D52FA3BFD44FE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7111543D5FFE05FB58FEEC.text	03D82A3EBD7111543D5FFE05FB58FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola bifida Broun 1915	<div><p>Sagola bifida Broun, 1915</p><p>(Figs. 11a, 12a, 13)</p><p>Sagola bifida Broun, 1915: 290 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3706. ♂.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Woodhill Waikakerei.” [white label, printed]; “ Sagola bifida . ♂.” [white label, handwritten]. The original label indicates this specimen is a male, but it is a female.</p><p>Additional Material (4 males). N E W ZEALAND: Auckland: 2♂♂, Mt. Auckland, Atuanui Scenic Reserve, 250m, 2.5km NE Glorit, 2 II 2010, D.S. Chandler, sift Nikau palm litter; 1♂, Waitakere Ranges Pk., Upper Huia Reser. Tr., 28 II 2010, 4.3km SW Waiatarua, 277m, D.S. Chandler, sift leaf litter at base of Cyathea; 1♂, Ahuroa, 240m, nr Woodcocks, 5 I 1983, J.C. Watt, litter 83/6 .</p><p>Diagnosis. This species is similar in appearance to other members of this species- group, but can be distinguished by the shape of the antennae and genitalia.</p><p>Redescription. Length 2.6–3.2 mm. Body brown, antennae, legs,and maxillary palpi paler, elytra yellowish brown (Fig. 11a). Head: As long as wide, widest across eyes (Fig. 11a). Antennomere 1 approximately 2 times longer than wide, 2–4 longer than wide, 5–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe shorter than parameres, apical lobe simple (Fig. 12a). Phallobase of median lobe asymmetrical and triangular (Fig. 12a). Parameres asymmetrical, right paramere twisted (Fig. 12a).</p><p>Type Locality. Woodhill, Kaipara Railway, Auckland, New Zealand .</p><p>Distribution. Auckland (Fig. 13: black circles).</p><p>Habitat. Specimens of this species were collected by sifting leaf and nikau palm tree litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7111543D5FFE05FB58FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7111563F5AFE55FE78FB64.text	03D82A3EBD7111563F5AFE55FE78FB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola latistriata Broun 1911	<div><p>Sagola latistriata Broun, 1911</p><p>(Figs. 11b, 12b, 12l–o, 13)</p><p>Sagola latistriata Broun, 1911: 495 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3367. ♂.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Ligar’ s Bush. Papakura.” [white label, printed]; “ Sagola latistriata .” [white label, handwritten]. The original label indicates this specimen is a male, but it is a female.</p><p>Additional Material (2 males). N E W ZEALAND: Auckland: 1♂, Kohukohunui, Hunua Ra., 600m, 30 III 1974, G. Kuschel, litter 74/20 ; 1♂, Kohukohunui, Hunua Ra., 550m, 30 III 1974, G. Kuschel, 74/19 .</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Redescription. Length 2.4–2.8 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11b). Head: As long as wide, widest across eyes (Fig. 12l). Antennomere 1 approximately 2 times longer than wide, 2–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11b). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres, apical lobe simple (Fig. 12b). Phallobase of median lobe asymmetrical and triangular (Fig. 12b). Parameres asymmetrical, right paramere divided, minor lobe slender with setae (Fig. 12b).</p><p>Type Locality. Ligar’ s Bush, Papakura, New Zealand .</p><p>Distribution. Auckland (AK) (Fig. 13: triangles).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7111563F5AFE55FE78FB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7311573CF9FADBFEE8FA6F.text	03D82A3EBD7311573CF9FADBFEE8FA6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola crawi Park and Carlton	<div><p>Sagola crawi Park and Carlton, new species</p><p>(Figs. 11c, 12c, 13)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND Waipoua SF Tanemahuta 29 Oct 1985 R.C.Craw &amp; ”, “C.Lyal Sifted litter 85/48”, “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ”, “ HOLOTYPE Sagola crawi Park and Carlton 2013 ” . Paratypes (n = 4: 3 males, 1 female). NEW ZEALAND: Northland: 1♂ (DSC), Waipoua SF, 0.9km e Forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.- podocarp forest, A. Newton, M. Thayer 686, leaf &amp; log litter, forest floor ; 1♂ (FMNH), Waipoua SF, 0.8km nw Wairau Summit, 350m, 27 XI 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 689, berl., leaf &amp; log litter, forest floor; 1♂ (NZAC), Waipoua SF, Four Sisters, 4 II 1975, S.E. Nicols, litter 75/93 ; 1♀ (FMNH), Waipoua SF, 1.7km nw Waikohatu Stream Bridge, 380m, 28 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer, FIT &amp; window trap .</p><p>Etymology. This species is named for the collector of the holotype, Robin C. Craw.</p><p>Diagnosis. This species is similar in appearance to other members of this species- group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 2.6–3.0 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 11c). Head: As long as wide, widest across eyes (Fig. 11c). Antennomere 1 approximately 2 times longer than wide, 2–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11c). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe shorter than parameres, apical lobe simple (Fig. 12c). Phallobase of median lobe asymmetrical and triangular (Fig. 12c). Parameres asymmetrical, right paramere twisted, broad apically (Fig. 12c).</p><p>Distribution. Northland (Fig. 13: black square).</p><p>Habitat. Specimens of this species were collected using window and flight intercept traps or by sifting leaf and log litter in hardwood and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7311573CF9FADBFEE8FA6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7211573CE2F9E4FC54FBDE.text	03D82A3EBD7211573CE2F9E4FC54FBDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola earlyi Park and Carlton	<div><p>Sagola earlyi Park and Carlton, new species</p><p>(Figs. 11d, 12d, 13)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (AMNZ), “ NEW ZEALAND CL Great Barrier I Mt. Hobson, 500m pitfall trap 9.xi.-17.xii.2003 J.W. Early ”, “ AMNZ86364 AUCKLAND MUSEUM NEW ZEALAND ”, “ HOLOTYPE Sagola earlyi Park and Carlton 2013 ”. Paratypes (n = 4; 2 males; 2 females). NEW ZEALAND: Coromandel : 2♂♂ 2♀♀ (1♂, slide-mounted; NZAC), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype, John W. Early.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group, but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 2.7–3.0 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11d). Head: As long as wide, widest across eyes (Fig. 11d). Antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea elongate. Eye small and prominent, onethird length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11d). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres, apical lobe bent to ventral (Fig. 12d). Phallobase of median lobe asymmetrical and triangular (Fig. 12d). Parameres asymmetrical, right paramere divided, minor lobe slender with setae (Fig. 12d).</p><p>Distribution. Coromandel (Fig. 13: star).</p><p>Habitat. Specimens of this species were collected by pitfall traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7211573CE2F9E4FC54FBDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7211503EA0FB9DFE8AFCCF.text	03D82A3EBD7211503EA0FB9DFE8AFCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola embersoni Park and Carlton	<div><p>Sagola embersoni Park and Carlton, new species</p><p>(Figs. 11e, 12e, 13)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, BP Orete Forest Te Puia Hut 230m NZMS 260 Y 14 484819 29.I.1993 R. M. Emberson ”, “ EAST CAPE INSECT SURVEY 1992 /3”, “ex litter from upland broadleaf/ podocarp forest LCNZ 93 /4”, “ HOLOTYPE Sagola embersoni Park and Carlton 2013 ” . Paratypes (2 males). NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), Orete Forest, Te Puia Hut, 13 IV 1992, G. Hall, litter 92/43 ; 1♂ (NZAC), Motu River, V 1920, A.E. Brookes .</p><p>Etymology. This species is named for the collector of the holotype, Rowan M. Emberson.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description of Male. L e n g t h 2.5 –2.8 m m. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11e). Head: As long as wide, widest across eyes (Fig. 11e). Antennomere 1 approximately 2 times longer than wide, 2–7 subquadrate, 8–10 weakly transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11e). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe shorter than right paramere, apical lobe blunt (Fig. 12e). Phallobase of median lobe asymmetrical and triangular (Fig. 12e). Parameres asymmetrical, right paramere divided, minor lobe with setae (Fig. 12e).</p><p>Distribution. Bay of Plenty (Fig. 13: white circle).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter in broadleaf and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7211503EA0FB9DFE8AFCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7511503CFDFC8CFC12FD1A.text	03D82A3EBD7511503CFDFC8CFC12FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola mayae Park and Carlton	<div><p>Sagola mayae Park and Carlton, new species</p><p>(Figs. 11f, 12f, 13)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND BP Kaimai Ra 427m Leyland Tramline 22 Sep 1981 B.M.May ”, “Litter 81/92”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola mayae Park and Carlton 2013 ” . Paratypes (5 males). NEW ZEALAND: Bay of Plenty: 2♂♂ (NZAC), Kaimai, 4 XII 1941, A.E. Brookes (NZAC); Coromandel : 1♂ (NZAC), Coromandel Ra., nr Summit, 19 I 1972, G.W. Ramsay, litter 72/36 ; 1♂ (NZAC), Kirikiri Saddle, 488m, 21 X 1976, J.S. Dugdale, litter 76/82 ; 1♂ (NZAC), Kopu Rd., 27 IX 1967, R. A. Cumber, leaf litter .</p><p>Etymology. This species is named for the collector of the holotype, Brenda M. May.</p><p>Diagnosis. This species is similar in appearance to other members of this group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description of Male. Length 2.8–3.2 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11f). Head: As long as wide, widest across eyes (Fig. 11f). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and reaching hind-point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11f). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres, divided from base, major lobe broad and covering minor lobe (Fig. 12f). Phallobase of median lobe asymmetrical and triangular (Fig. 12f). Parameres asymmetrical, right paramere with setae (Fig. 12f).</p><p>Distribution. Bay of Plenty, Coromandel (Fig. 13: white squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7511503CFDFC8CFC12FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7511513EB1FD58FEA1FEBD.text	03D82A3EBD7511513EB1FD58FEA1FEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola ranatungae Park and Carlton	<div><p>Sagola ranatungae Park and Carlton, new species</p><p>(Figs. 11g, 12g, 14)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (AMNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: ND: North Cape Area Taputaputa Bay 30IV1968, J.L. Bertrand ”, “ HOLOTYPE Sagola ranatungae Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Northland: 1♀ (AMNZ), same data as holotype .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Dhahara Ranatunga. She assisted with specimen access through loans and during the first author’ s visit to the Auckland War Memorial Museum, Auckland, New Zealand.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 2.4–2.8 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11g). Head: As long as wide, widest across eyes (Fig. 11g). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11g). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe weakly bent and longer than left paramere (Fig. 12g). Phallobase of median lobe asymmetrical and triangular (Fig. 12g). Parameres asymmetrical, left paramere approximately twice larger than right, divided apically (Fig. 12g).</p><p>Distribution. Northland (Fig. 14: black circle).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7511513EB1FD58FEA1FEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7411513CD2FEBAFBC0FC9F.text	03D82A3EBD7411513CD2FEBAFBC0FC9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola baylessae Park and Carlton	<div><p>Sagola baylessae Park and Carlton, new species</p><p>(Figs. 11h, 12h, 14)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND Mair Park Whangarei 17 Aug 1977 G. Kuschel ”, “Litter+wood”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola baylessae Park and Carlton 2013 ”.</p><p>Etymology. This species is named for LSAM Curator, President of The Coleopterists Society (2013–2014), and enthusiastic supporter of this study, Victoria M. Bayless.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description of Male. Length 2.8 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11h). Head: As long as wide, widest across eyes (Fig. 11h). Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3–7 subquadrate, 8–10 weakly transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11h). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres, apical lobe blunt with sharp branch (Fig. 12h). Phallobase of median lobe asymmetrical and triangular (Fig. 12h). Parameres asymmetrical, left paramere triangular and right round (Fig. 12h).</p><p>Distribution. Northland (Fig. 14: triangles).</p><p>Habitat. The specimen of this species was collected by sifting leaf and wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7411513CD2FEBAFBC0FC9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7411523EE8FCDCFE78FEEF.text	03D82A3EBD7411523EE8FCDCFE78FEEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola tekoauensis Park and Carlton	<div><p>Sagola tekoauensis Park and Carlton, new species</p><p>(Figs. 11i, 12i, 14)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, BP Te Koau, 140m Twin Puriri’ s 31 Jan 1993 J.S. Dugdale ”, “Litter 93/10”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola tekoauensis Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Bay of Plenty: 1♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named after the type locality, Te Koau, Bay of Plenty.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 2.5–2.8 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11i). Head: As long as wide, widest across eyes (Fig. 11i). Antennomere 1 approximately 2 times longer than wide, 2–4 subquadrate, 5–10 transverse. Frontal sulcus deep and reaching hind-point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11i). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe shorter than left paramere, apical lobe simple (Fig. 12i). Phallobase of median lobe asymmetrical and triangular (Fig. 12i). Parameres asymmetrical, left paramere triangular, right broad and round depression apicolaterally (Fig. 12i).</p><p>Distribution. Bay of Plenty (Fig. 14: black square).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7411523EE8FCDCFE78FEEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7711523CE8FA0EFC74FB95.text	03D82A3EBD7711523CE8FA0EFC74FB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola fairburni Park and Carlton	<div><p>Sagola fairburni Park and Carlton, new species</p><p>(Figs. 11k, 12k, 14)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “Tangihua Range N. Auckland. 1800’-2000’”. “ Coll. E.Fairburn 27.10.1926 ”, “ A.E. Brookes Collection”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola fairburni Park and Carlton 2013 ”. Paratype (1 female). NEW ZEALAND: Northland: 1♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype, Edwin Fairburn.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 2.7 mm. Body brown, antennae, elytra, maxillary palpi, and legs yellowish brown (Fig. 11k). Head: As long as wide, widest across eyes (Fig. 11k). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11k). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres with semicircular depression (Fig. 12k). Phallobase of median lobe asymmetrical and triangular (Fig. 12k). Parameres asymmetrical, left paramere blunt and right triangular (Fig. 12k).</p><p>Distribution. Northland (Fig. 14: white circle).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7711523CE8FA0EFC74FB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7711523CF9FE64FDC0FA41.text	03D82A3EBD7711523CF9FE64FDC0FA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola ferroi Park and Carlton	<div><p>Sagola ferroi Park and Carlton, new species</p><p>(Figs. 11j, 12j, 14)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, ND Te Paki Trig 23 Nov 1982 G. Kuschel ”, “Sifted litter and decayed wood 82/ 116”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola ferroi Park and Carlton 2013 ”.</p><p>Etymology. This species is named for LSAM alumnus, North American faronite specialist, and enthusiastic supporter of this study, Michael L. Ferro.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description of Male. Length 2.8 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 11j). Head: As long as wide, widest across eyes (Fig. 11j). Antennomere 1 approximately 2 times longer than wide, 2–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep and reaching hind point of eye from frontal rostral lobes. Anterior frontal fovea an inverted triangle. Posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 11j). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without microtrichial patch. Aedeagus: Median lobe longer than parameres, apical lobe bent with branch from apex (Fig. 12j). Phallobase of median lobe asymmetrical and triangular (Fig. 12j). Parameres asymmetrical, left paramere long and bent, right paramere diamondshaped (Fig. 12j).</p><p>Distribution. Northland (Fig. 14: star).</p><p>Habitat. The specimen of this species was collected by sifting decayed wood litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7711523CF9FE64FDC0FA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7711533F75FBCFFDABFC43.text	03D82A3EBD7711533F75FBCFFDABFC43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>4. baiknami species-group</p><p>(7 species)</p><p>Diagnosis. Members of the baiknami speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.2–3.1 mm; antennomere 1 approximately 2 times longer than wide with dull surface; gular region of male head with transverse depression and with blunt process in middle (Fig. 16i: arrow); fore femur with semicircular depression; abdominal tergites IV–VI with discal carinae; present on North Island, not known from South Island (Figs. 17–18).</p><p>KEY TO SPECIES OF THE BAIKNAMI SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Body large,&gt; 2.9 mm; mid-tibia not bent; median lobe of genitalia S-shaped (Fig. 16a) ............................. S. baiknami new species</p><p>1′. Body smaller, &lt;2.8 mm; mid-tibia bent; median lobe of genitalia straight or bent at 90°.......................................................2</p><p>2(1′). Antennomeres 4–11 bearing tubercles; fore tibia with semicircular depression ..... ................................. S. watti new species</p><p>2′. Antennomeres without tubercles; fore tibia without depression...................................3</p><p>3(2′). Apical lobe of genitalia bird-head- or umbrella-shaped.......................................4</p><p>3′. Apical lobe of genitalia blunt.................5</p><p>4(3). Apical lobe of genitalia bird-head-shaped; right paramere without minor lobe (Fig. 16d) ............................ S. taupoensis new species</p><p>4′. Apical lobe of genitalia umbrella-shaped; right paramere with minor lobe (Fig. 16e) ............................... S. nunni new species</p><p>5(3′). Median lobe slender, approximately 3 times longer than wide; left paramere U-shaped (Fig. 16g) ........... S. choeunae new species</p><p>5′. Median lobe slender, at least 4 times longer than wide; left paramere L-shaped........... 6</p><p>6(5′). Right paramere with 2 acute lobes (Fig. 16c) ............................ S. butcherae new species</p><p>6′. Right paramere with arm-shaped lobe and 2 acute lobes (Fig. 16f) ............................ ........................ S. sunsookae new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD7711533F75FBCFFDABFC43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD7611533C96FBF7FC3DFB79.text	03D82A3EBD7611533C96FBF7FC3DFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola baiknami Park and Carlton	<div><p>Sagola baiknami Park and Carlton, new species</p><p>(Figs. 15a, 16a, 17)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), “ NEW ZEALAND: ND: Mangataipa Scen. Res., near Mangamuka Bridge, 20m, 29 xi-5 xii 1984, mixed hardwood forest, A. Newton / M. Thayer 695, berl., leaf &amp; log litter, forest floor”, “ HOLOTYPE Sagola baiknami Park and Carlton 2013 ” . Paratypes (n = 14: 12 males; 2 females). NEW ZEALAND: Northland: 2♂♂ 1♀ (1♂ 1♀, slide-mounted; FMNH), Mangamuka Summit, 12km nw Mangamuka, 400m, 25 XI–5 XII 1984, hdwd.- podocarp forest, A. Newton, M. Thayer 683, window trap ; 2♂♂ 1♀ (NZAC), Waipoua SF, 100m, 17 III 1978, S. &amp; J. Peck, under Agathis log on bark ; 3♂♂ (</p><p>FMNH), Waipoua SF, Waipoua Stream, 16–21 III 1978, 70m, S. &amp; J. Peck, Kauri forest, frass under bark; 3♂♂ (NZAC), Te Paki Trig, 23 XI 1982, G. Kuschel, sifted litter and decayed wood 82/116 ; 1♂ (FMNH), Waipoua SF, Wairau Summit, 40m, 27 XI–6 XII 1984, hdwd-. podocarp forest, A. Newton, M. Thayer 687, FIT &amp; window trap ; 1♂ (FMNH), Waipoua SF, Kauri Ricker tr. 100m, 11–15 IV 1980, kauripodocarp-broadleaf, A. Newton, M. Thayer, pyrethrin-fogging fungusy log.</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Baik-Nam Park. He is the first author’ s father and has provided a lifetime of support and encouragement.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: larger body, length 2.9–3.1 mm; antennomeres 4–10 with tubercles; triangular head with flat ventral temporal lobe; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.9–3.1 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 15a). Head: Male head triangular, widest across temples (Fig. 15a), temple lobed and flat ventrally. Female head rectangular, widest across eyes. Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3 subquadrate, 4 elongate, 5–10 subquadrate, 4–10 with tubercles. Frontal sulcus reaching midpoint of eye from frontal fovea. Anterior frontal fovea round. Posterior frontal fovea round. Eye small and prominent, one-third length of temple in male, one-fourth length of temple in female. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15a). Meso- and metathorax trapezoidal, longer than wide. Hind wings well-developed. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, S-shaped (Fig. 16a). Phallobase of median lobe symmetrical and rectangular (Fig. 16a). Parameres asymmetrical, right paramere elongate (Fig. 16a).</p><p>Distribution. Northland (Fig. 17: black circles).</p><p>Habitat. Most specimens of this species were collected using window and flight intercept traps or by sifting leaf and log litter in kauri, broadleaf, hardwood, and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD7611533C96FBF7FC3DFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD76116E3E84FAFDFBB1FC6E.text	03D82A3EBD76116E3E84FAFDFBB1FC6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola watti Park and Carlton	<div><p>Sagola watti Park and Carlton, new species</p><p>(Figs. 15b, 16b, 17)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: ND: Omahuta S.F., Kauri Res. 10 X 1974, J.C. Watt litter 74/81”, “ HOLOTYPE Sagola watti Park and Carlton 2013 ” . Paratypes (n = 33: 22 males; 11 females). NEW ZEALAND: Northland: 2♂♂ (FMNH, DSC), Waipoua SF, Kauri Ricker Tr., 120m, 26 XI–4 XII 1984, kauripodocarp-hdwd., A. Newton, M. Thayer 684, berl., leaf &amp; log litter, forest floor; 1♂ (FMNH), Waipoua SF, Kauri Ricker Tr., 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd., A. Newton, M. Thayer 684, berl., FIT &amp; window trap ; 1♂ 1♀ (FMNH), c) S. butcherae, d) S. taupoensis, e) S. nunni, f) S. sunsookae, g) S. choeunae . Scale bars = 1 mm.</p><p>Waipoua SF, 1.7km nw Waikohatu Stream bridge, 380m, 28 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer; 1♂ (FMNH), Waipoua SF, Wairau Summit 400m, 27 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer, FIT &amp; window trap ; 1♀ (FMNH), Waipoua SF, Toronui Tr., 150m, 13 IV 1980, kauri-podocarpbroadlf-nikau palm forest, A. Newton, M. Thayer, berl., leaf &amp; log litter, forest floor ; 2♂♂ (NZAC), Te Paki Trig, 23 XI 1982, G. Kuschel, sifted litter and decayed wood 82/116 ; 1♂ (NZAC), Waipoua SF, Wairau Stm, 390m, 15 IV 1980, J.C. Watt, sifted litter 80/ 51 ; 1♂ (NZAC), Waipoua Stm, 100m, 19 III 1978, S.B. Peck, litter ; 1♂ (DSC), Waipoua SF, Toronui Tr., 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd., A. Newton, M. Thayer 685, berl., leaf &amp; log litter, forest floor ; 1♂ (FMNH), Waipoua SF, 0.8km nw Wairau Summit, 350m, 27 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 689, FIT &amp; window trap ; 1♂ (FMNH), Waipoua SF, Wairau Summit, 387m, 11–14 IV 1980, podocarp-broadleaf, A. Newton. M. Thayer, pitfall trap ; 1♂ (FMNH), Waipoua SF, Toatoa Tr., 270m, 12–15 IV 1980, toatoa-kauri-podocarp-broadlf., A. Newton, M. Thayer, berl., leaf &amp; log litter, forest floor ; 1♂ (NZAC), Mangamuka Walk, 28 VII–1 VIII 1998, R. Leschen, R. Hoare, FIT 223, 36 ′11S, 173′28E ; 1♂ (NZAC), Above Hihi, 6 I 1969, K. Wise, juvenile Kahikatea ; 1♂ (NZAC), Puketi SF, 21 I 1972, G.W. Ramsay, litter 72/56 ; 1♂ (NZAC), Waipoua SF, 7 XII 1961, G. Kuschel, litter 61/13. Auckland : 1♂ (FMNH), Waitakere Range, 260m, Nohoanga Scenic Reserve, 23 XI–8 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 679, FIT &amp; window trap ; 1♂ 2♀♀ (1♀, slide-mounted; 1♂ 1♀, FMNH; 1♀, DSC), Waitakere Range, 260m, Nohoanga Scenic Reserve, 8 XII 1984 – 25 I 1985, hdwd.- podocarp forest, A. Newton, M. Thayer 679, FIT &amp; window trap ; 1♀ (FMNH), Waitakere Range, 260m, Nohoanga Scenic Reserve, 23 XI 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 679, berl., leaf &amp; log litter, forest floor ; 3♀♀ (FMNH), Waitakere Range, Cascade-Kauri Park, up. Kauri Tr., 170m, 8 XII 1984 – 25 I 1985, kauri-podo-hdwd., A. Newton, M. Thayer, FIT &amp; window trap ; 1♂ (NZAC), Huia, 150m, Twin Peaks Ridge, 1 V 1981, B.M. May, at roots of Metrosideros robusta; 1♂ (DSC), Goldie Bush Scenic Res., Mokoroa Falls Tr., 6km W Waitakere, 28 III 2010, 106m, D.S. Chandler, kauri &amp; podocarp leaf litter ; 1♂ (NZAC), Kohukohunui, Hunua Ra, 600m, 30 III 1974, G. Kuschel, litter 74/20 ; 1♀ (FMNH), Waitakere Ra., Nohoanga Scenic Res., 260m, 36.57′S, 174.35′E, 23 II 1987, hardwood-podocarp forest, A. Newton, M. Thayer 836, FMHD#87-271, berl., frond litter at base of tree ferns; Waikato : 1♀ (slide-mounted; FMNH), Pirongia Forest Park, Mahaukura Track (above end Grey Rd.), 270m, 37″58.218′S, 175″06.523′E, 18 XI–27 XII 2005, mixed broadleaf forest, FMHD#2005-009, FIT, A. Newton, M. Thayer, et al., ANMT site 1142 ; 1♀ (NZAC), Waitomo, Maori L, Tumutumu Rd, 24 V 1983, J.C. Watt, litter and soil 83/60 .</p><p>Etymology. This species is named for the collector of the holotype, John C. Watt.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.2–2.5 mm; antennomeres 4–10 with tubercles; male head bluntly triangular, ventral surface of head with short carinae from posterior point of eyes; male fore tibia with semicircular depression and mid-tibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.2–2.5 mm. Body brown, antennae and maxillary palpi paler, legs and elytra yellowish brown (Fig. 15b). Head: Male head as long as wide, widest across eyes, ventral surface with short carina from hind point of eyes, as long as eye (Fig. 16h). Antennomere 1 approximately 2 times longer than wide, 2–3 transverse, 4 longer than wide, 5 subquadrate, 9–10 transverse, 4–10 with tubercles. Frontal sulcus deep, reaching beyond eye from frontal fovea (Fig. 16h). Anterior frontal fovea round (Fig. 16h). Posterior frontal fovea elongate (Fig. 16h). Eye large and prominent, one-half length of temple in male (Fig. 16h), one-third length of temple in female. Thorax: Prosternum as long as wide, widest at midpoint. Male fore tibia with semicircular depression. Elytra rectangular (Fig. 15b). Meso- and metathorax trapezoidal, longer than wide. Hind wings well-developed. Male mid-tibia bent. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided deeply (Fig. 16b). Phallobase of median lobe asymmetrical and triangular (Fig. 16b). Parameres asymmetrical, right paramere larger than left (Fig. 16b).</p><p>Distribution. Auckland, Northland, Waikato (Fig. 17: triangles).</p><p>Habitat. Most specimens of this species were collected using window, flight intercept, and pitfall traps or by sifting leaf and log litter in kauri, broadleaf, hardwood, and podocarp forests. A few specimens were found at the bases of Metrosideros robusta A. Cunn. (Myrtaceae) and tree ferns.</p></div>	https://treatment.plazi.org/id/03D82A3EBD76116E3E84FAFDFBB1FC6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4B116F3E86FBECFDCEFC3E.text	03D82A3EBD4B116F3E86FBECFDCEFC3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola butcherae Park and Carlton	<div><p>Sagola butcherae Park and Carlton, new species</p><p>(Figs. 15c, 16c, 17)</p><p>Type Material. Holotype. NEW ZEALAND: Taupo: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND TO Waituhi Saddle 16 Nov 1983 C.F. Butcher ”, “Leaf litter, moss and rotten wood 83/112”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola butcherae Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Gisborne: 1♂ (FMNH), Urewera N.P., Waikaremoana Rd., S end Matanunui Ridge, 720m, 38″44.404′S, 177″05.806′E, 22 XII 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp, FMHD#2005-118, berl., leaf &amp; log litter, M. Thayer, A. Solodovnikov, ANMT site 1149 .</p><p>Etymology. This species is named for the collector of the holotype, Clare F. Butcher.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.3–2.8 mm; male head bluntly triangular, widest across temples, ventral temporal lobe flat; male mid-femur weakly depressed at base and mid-tibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. L e n g t h 2.3– 2.8 m m. Body brown, antenna, elytra, legs, and maxillary palpi paler (Fig. 15c). Head: Bluntly triangular, widest across temples, ventral temporal lobe flat (Fig. 15c). Antennomere 1 approximately 2 times longer than wide, 2–7 subquadrate, 8–10 weakly transverse. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea elongate. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15c). Meso- and metathorax trapezoidal, longer than wide. Male mid-femur weakly depressed at base. Male mid-tibia bent. Abdomen: Tergite IV with pair of oval or transverse patches of microtrichia. Aedeagus: Median lobe blunt (Fig. 16c). Phallobase of median lobe asymmetrical and triangular (Fig. 16c). Parameres asymmetrical, left paramere L-shaped, right paramere with thick setae apically (Fig. 16c).</p><p>Distribution. Gisborne, Taupo (Fig. 17: black squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf, moss, rotten wood, and log litter in broadleaf and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4B116F3E86FBECFDCEFC3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4A116F3C88FC3CFCA8FC10.text	03D82A3EBD4A116F3C88FC3CFCA8FC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola taupoensis Park and Carlton	<div><p>Sagola taupoensis Park and Carlton, new species</p><p>(Figs. 15d, 16d, 17)</p><p>Type Material. Holotype. NEW ZEALAND: Taupo: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND TO Waituhi Saddle 16 Nov 1983 C.F. Butcher ”, “Leaf litter, moss and rotten wood 83/112”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola taupoensis Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Gisborne: 1♂ (NZAC), Urewera NP, Waikaremoana Rd., 4 end Matanunui Ridge, 720m, 38″44.404′S. 177″05.806′E, 22 XI–23 XII 2005, mixed broadleaf (incl. Nothofagus fusca)- podocarp, FMHD#2005-028, FIT, M. Thayer, A. Solodovnikov, ANMT site 1149 .</p><p>Etymology. The species name is based on the biotic region of the holotype, Taupo.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.4–2.7 mm; male head bluntly triangular, widest across temples, ventral temporal lobe flat; male mid-femur weakly depressed at base and mid-tibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.4– 2.7 mm. Body brown, antenna, elytra, legs, and maxillary palpi paler (Fig. 15d). Head: Bluntly triangular, widest across temples, ventral temporal lobe flat (Fig. 15d). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea elongate. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15d). Meso- and metathorax trapezoidal, longer than wide. Male midfemur weakly depressed at base. Male mid-tibia bent. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres, with bird-head-like apical lobe (Fig. 16d). Phallobase of median lobe asymmetrical and elongate (Fig. 16d). Parameres asymmetrical, left paramere elongate, right paramere blunt (Fig. 16d).</p><p>Distribution. Gisborne, Taupo (Fig. 17: stars).</p><p>Habitat. Specimens of this species were collected using flight intercept traps or by sifting leaf, moss, and rotten wood litter in broadleaf and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4A116F3C88FC3CFCA8FC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4A11683EFFFC47FE17FCF5.text	03D82A3EBD4A11683EFFFC47FE17FCF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola nunni Park and Carlton	<div><p>Sagola nunni Park and Carlton, new species</p><p>(Figs. 15e, 16e, 18)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND Whangae Stm Kawakawa, 14.11.94”, “ HOLOTYPE Sagola nunni Park and Carlton 2013 ” The original label does not mention who collected this specimen, but the collector is J. T. Nunn . Paratypes (2 males). NEW ZEALAND: Northland: 1♂ (NZAC), Waipoua SF, Four Sisters, 4 II 1975, S.E. Nicols, litter 75/93; 1♂ (AMNZ), Waima Forest, Hauturu Tr., 220–360m, 25 III 1993, J.W. Early, kauribroadleaf-nikau forest, screen sweep .</p><p>Etymology. This species is named for the collector of the holotype, John T. Nunn, who also provided many valuable specimens for this study.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.5–2.8 mm; male head bluntly triangular, widest across eyes, ventral surface of head with short carina from hind-point of eyes; male mid-femur weakly depressed at base and mid-tibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.5–2.8 mm. Body brown, antenna, elytra, legs, and maxillary palpi paler (Fig. 15e). Head: As long as wide, widest across eyes, ventrally with short carina from hind point of eyes, as long as eye (Fig. 15e). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4 elongate, 5–9 longer than wide, 9–10 subquadrate. Frontal sulcus reaching midpoint of eye from end of frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea elongate. Eye large and prominent, onethird length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15e). Hind wings well-developed. Meso-and metathorax trapezoidal, longer than wide. Male mid-femur weakly depressed at base. Male mid-tibia bent. Abdomen: Tergite IV with a pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe elongate with acute minor lobe at base (Fig. 16e). Apical lobe of genitalia umbrella-shaped (Fig. 16e). Phallobase of median lobe asymmetrical and triangular. Parameres asymmetrical, right paramere with minor lobe apically (Fig. 16e).</p><p>Distribution. Northland (Fig. 18: black circles).</p><p>Habitat. Specimens of this species were collected by sweeping and sifting leaf litter in kauri, broadleaf, and nikau forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4A11683EFFFC47FE17FCF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4D11683EBCFF31FB75F95B.text	03D82A3EBD4D11683EBCFF31FB75F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sunsookae Park and Carlton	<div><p>Sagola sunsookae Park and Carlton, new species</p><p>(Figs. 15f, 16f, 18)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND WO Mt Karioi near base 11 Oct 1981 C.F. Butcher ”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “Litter 81–96”, “ HOLOTYPE Sagola sunsookae Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Auckland: 1♂ (FMNH), Hunua Range Reg. Park, Kohukohunui Tr. nr Mine Rd., 475m, 37″02.396′S 175″11.251′E, 17 XI–28 XII 2005, broadleaf-podocarp forest, numerous tree fern, FMHD#2005-001, FIT, A. Newton, M. Thayer et al., ANMT site 1140 .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Sun-Sook Choi. She is the first author’ s mother and has provided a lifetime of support and encouragement.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.4–2.7 mm; male head bluntly triangular, widest across eyes, ventral surface of head with pair of short carinae on temple; male mid-femur weakly depressed at base; midtibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.4–2.7 mm. Body brown, antenna, elytra, legs, and maxillary palpi paler (Fig. 15f). Head: As long as wide, widest across eyes (Fig. 15f), ventral surface with pair of short carinae on temple. Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea oval. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15f). Meso- and metathorax trapezoidal, longer than wide. Male mid-femur weakly depressed at base. Male mid-tibia bent. Abdomen: Tergite IV with pair of oval or transverse patches of microtrichia. Aedeagus: Median lobe blunt (Fig. 16f). Phallobase of median lobe asymmetrical and triangular (Fig. 16f). Parameres asymmetrical, left paramere L-shaped, right paramere with 3 lobes with thick setae apically (Fig. 16f).</p><p>Distribution. Waikato, Auckland (Fig. 18: triangles).</p><p>Habitat. Specimens of this species were collected using flight intercept traps or by sifting litter in broadleaf and podocarp forests with tree ferns.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4D11683EBCFF31FB75F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4C11693CD2FF31FEA1FA52.text	03D82A3EBD4C11693CD2FF31FEA1FA52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola choeunae Park and Carlton	<div><p>Sagola choeunae Park and Carlton, new species</p><p>(Figs. 15g, 16g, 18)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “Maraenui Bay of Plenty.”, “ Coll. A.E.Brookes. May1928.”, “ A.E.Brookes Collection ”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola choeunae Park and Carlton 2013 ”.</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Cho-Eun Jeong. She is the first author’ s wife and has always cheered him up.</p><p>Diagnosis. This species can be distinguished from the other species of the baiknami speciesgroup by the following combination of characters: smaller body, length 2.4 mm; male head triangular, widest across temples, ventral temporal lobe flat; male mid-femur weakly depressed at base and mid-tibia bent; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.4 mm. Body brown, antenna, elytra, legs, and maxillary palpi yellowish brown (Fig. 15g). Head: Head triangular, widest across temples, ventral temporal lobe flat (Fig. 15g). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea oval. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 15g). Meso- and metathorax trapezoidal, longer than wide. Male mid-femur weakly depressed at base. Male mid-tibia bent. Abdomen: Tergite IV with pair of oval or transverse patches of microtrichia. Aedeagus: Median lobe rectangular and blunt (Fig. 16g). Phallobase of median lobe asymmetrical and triangular (Fig. 16g). Parameres asymmetrical, left paramere U-shaped, right paramere divided, minor lobe with tubercle and thick setae apically (Fig. 16g).</p><p>Distribution. Bay of Plenty (Fig. 18: black square).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4C11693CD2FF31FEA1FA52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4C11693D48FA0EFB45FA9E.text	03D82A3EBD4C11693D48FA0EFB45FA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola pulchra Broun 1880	<div><p>5. pulchra species-group</p><p>(4 species)</p><p>Diagnosis. The members of the pulchra species-group can be distinguished from other Sagola species-groups by the following combination of characters: large body length 3.2–4.2 mm; antennomere 1 approximately 2.5 times longer than wide with dull surface; gular region of male head with transverse depression (Fig. 19j); ventral surface of male head with a pair of processes laterally (Fig. 19j: arrow); hind wings reduced to small pads; fore and mid-femur with semicircular depression (Fig. 19k); male mid-femur at least twice as broad as tibia (Fig. 19k); Male tergite IV without microtrichial patch; abdominal tergites IV–VI with discal carinae; female abdominal ventrite VIII triangularly produced posteriorly (Fig. 19l), ventrite IX deeply emarginate posteriorly (Fig. 19m); present on North Island, not known from South Island (Fig. 20).</p><p>KEY TO SPECIES OF THE PULCHRA SPECIES- GROUP</p><p>The key is based on genitalia because most specimens are indistinguishable based on external morphology.</p><p>1. Median lobe of genitalia without minor lobe; broader paramere without branch from base (Fig. 19e)...... S. pulchra Broun</p><p>1′. Median lobe of genitalia with minor lobe; broader paramere with 1 or 2 branches from base .................................................2</p><p>2(1′). Branches of apical lobe of genitalia approximately equal............................................... 3</p><p>2′. Apical lobe of genitalia with longer major branch and shorter minor branch (Fig. 19h) .......................... S. poortmani new species</p><p>3(2). Apical lobe of genitalia with U-shaped emargination; right paramere with 2 branches from base (Fig. 19f) .................. ......................... S. keejeongi new species</p><p>3′. Apical lobe of genitalia with V-shaped emargination; left paramere with 1 branch from base (Fig. 19g) ................................. ............................ S. ramsayi new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD4C11693D48FA0EFB45FA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4C116B3F54FADDFDF3FCF5.text	03D82A3EBD4C116B3F54FADDFDF3FCF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola pulchra Broun 1880	<div><p>Sagola pulchra Broun, 1880</p><p>(Figs. 19a, 19e, 19l–m, 20)</p><p>Sagola pulcher Broun, 1880: 137 . Schaufuss 1888: 85; Raffray 1893: 31, 1904: 497, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola robusta Broun, 1893b: 1420 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material Examined. Holotype: NEW ZEALAND: Coromandel: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “250” [white label, printed]; “Tairua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola pulcher . ♂.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female. Holotype of Sagola robusta: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “2468. ♂ ” [white label, handwritten]; “Hunua” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ sagola robusta . ♂ ”[white label, handwritten].</p><p>Additional Material (n = 16: 7 males; 9 females). NEW ZEALAND: Bay of Plenty: 1♂, Mt. Te Aroha, 2.5km SE Te Aroha, 390m, Tui Road at Tui Creek, Mt. Domain Tr., 25 II 2010, D.S. Chandler, sift forest litter &amp; old epiphyte clumps ; 1♂, Mt. Te Aroha, 500–1000m, 14 X 1979, J.S. Dugdale ; 2♂♂, Kaimai, 18 I 1931, A.E. Brookes collection ; 1♂ 5♀♀, Tapapa Tukorehe Res., 300m, 25 III 1978, S. &amp; J. Peck, berl., litter ; 1♀, Waiaroho, 17 IX 1992, G. Hall, R. C. Henderson, sifted rotten wood 92/54 ; 1♀, Waenga, NZMS 260 Y14 652913, 28 I 1993, R. M. Emberson, mixed broadleaf nikau forest litter LCNZ 93 /3 ; 1♀, Waioeka Gorge, 14 IV 1987, J. Nunn, with Mesoponera; Coromandel: 1♀, Coromandel, 350m, 23 III 1977, G. Kuschel, decayed wood ; Waikato: 1♂, Maungatautari, 17 XII 2007 – 20 I 2008, D.H. Watts, Pitfall trap .</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Redescription. Length 3.5–4.1 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 19a). Head: Transverse, widest across temples (Fig. 19a). Antennomere 1 approximately 2.5 times longer than wide, 2–3 longer than wide, 4–5 elongate, 6–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep and wide, reaching midpoint of eye from frontal fovea. Anterior frontal fovea extremely small. Posterior frontal fovea round. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 19a). Meso-and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe longer than parameres (Fig. 19e). Phallobase of median lobe symmetrical and rectangular (Fig. 19e). Parameres asymmetrical, right paramere broader and with slender branch apically (Fig. 19e).</p><p>Type Locality. Tairua, New Zealand .</p><p>Distribution. A u c k l a n d, B a y o f P l e n t y, Coromandel, Waikato (Fig. 20: black circles).</p><p>Habitat. Most specimens of this species were collected using pitfall traps or by sifting leaf litter or rotten wood in kauri or broadleaf forests. One specimen was collected with ants of the genus Mesoponera Emery.</p><p>Comments. Specimens of S. pulchra are difficult to distinguish from those of other species externally, but the shape and size of the antennomeres and genitalia are diagnostic. The type specimens of S. robusta share these diagnostic characters, and this species has been collected at or near the type locality. For these reasons, we have placed S. robusta in synonymy with S. pulchra .</p></div>	https://treatment.plazi.org/id/03D82A3EBD4C116B3F54FADDFDF3FCF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4E11643EE8FF31FDDFFF68.text	03D82A3EBD4E11643EE8FF31FDDFFF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola keejeongi Park and Carlton	<div><p>Sagola keejeongi Park and Carlton, new species</p><p>(Figs. 19b, 19i–j, 20)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, BP Te Koau, 120m 29 Apr 1993 J.S. Dugdale Litter 93/101”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola keejeongi Park and Carlton 2013 ” . Paratypes (n = 8: 5 males; 3 females). NEW ZEALAND: Bay of Plenty: 1♂ (slide mounted, NZAC), Te Koau, Hovells, Watching Dog, 240m, 24 X 1992, J.S. Dugdale, litter 92/78 ; 1♂ (NZAC), Hicks Bay, 20 IX 1992, G.M. Barker, J.S. Dugdale, under rotten logs and stones ; 1♂ (NZAC), Lottin Pt Rd., Waenga Bush, 16 IX 1992, R. C. Henderson, sifted litter 92/51 ; 1♀ (slide-mounted; NZAC), Lottin Pr Rd., Waenga Bush, 29 I 1993, J.I. Townsend, sifted litter 93/7 ; 1♂ (JTN), Te Araroa, 28 III 1991, J. Nunn ; 1♀ (JTN), Te Koau, Te Araroa, 4 II 1993, J.I. Townsend, J. Nunn ; 1♀ (NZAC), Te Koau, Hovells, Watching Dog tr., 260m, 31 I 1993, J.S. Dugdale, litter 93/11 ; 1♂ (AMNZ), Te Koau, track to Hovells Watching dog, 160m, 31 I–4 II 1993, J.W. Early, Puriri /nikau forest, yellow pan trap .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study and world staphylinid specialist, Kee-Jeong Ahn.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 3.2–3.5 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 19b). Head: Transverse, widest across temples (Fig. 19i). Antennomere 1 approximately 2.5 times longer than wide, 2–3 longer than wide, 4–5 elongate, 6–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep and wide, reaching midpoint of eye from frontal fovea (Fig. 19i). Anterior frontal fovea extremely small (Fig. 19i). Posterior frontal fovea round (Fig. 19i). Eye small and prominent, one-third length of temple (Fig. 19i). Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 19b). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Apical lobe of genitalia U-shaped (Fig. 19f). Phallobase of median lobe symmetrical and rectangular (Fig. 19f). Parameres asymmetrical, right paramere with 3 branches, 2 from base, 1 apically (Fig. 19f).</p><p>Distribution. Bay of Plenty (Fig. 20: triangles).</p><p>Habitat. Most specimens of this species were collected using yellow pan traps, by sifting leaf and log litter, or were from under logs and stones. One specimen was found with ants.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4E11643EE8FF31FDDFFF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4111643F06FCE6FB7FFA48.text	03D82A3EBD4111643F06FCE6FB7FFA48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola parva	<div><p>6. parva species-group</p><p>(2 species)</p><p>Diagnosis. The members of parva group can be distinguished from other Sagola groups by the following combination of characters: body length 2.3–3.8 mm; antennomere 1 approximately 2.5 times longer than wide with dull surface, male antennomeres 5–6 enlarged (Fig. 21e); gular region of male head with round depression (Fig. 21g); hind wings well-developed; fore-femur with semicircular depression; abdominal tergites IV–VI with discal carinae; present on North Island, not known from South Island (Fig. 22).</p><p>KEY TO SPECIES OF THE PARVA SPECIES- GROUP</p><p>The key is based on genitalia because most specimens are indistinguishable based on external morphology.</p><p>1. Head bluntly triangular with large eyes (Fig. 21a, f); antennomeres 5–6 enlarged, without tubercles................. S. parva Sharp</p><p>1′. Head transverse with small eyes (Fig. 21b); antennomeres 4–6 enlarged, with distinctive tubercles ........................ S. denticollis Broun</p></div>	https://treatment.plazi.org/id/03D82A3EBD4111643F06FCE6FB7FFA48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4111643CA1FAFDFC74FD70.text	03D82A3EBD4111643CA1FAFDFC74FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola poortmani Park and Carlton	<div><p>Sagola poortmani Park and Carlton, new species</p><p>(Figs. 19d, 19h, 20)</p><p>Type Material. Holotype. NEW ZEALAND: Gisborne: 1♂ (AMNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, GB Hikurangi Hut, 260m 4.v.1996 P.Poortman L4459”, “ AMNZ 35222 AUCKLAND MUSEUM NEW ZEALAND ”, “ HOLOTYPE Sagola poortmani Park and Carlton 2013 ”.</p><p>Etymology. This species is named for the collector of the holotype, Peter Poortman.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 3.5 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 19d). Head: Transverse, widest across temples (Fig. 19d). Antennomere 1 approximately 2.5 times longer than wide, 2–3 longer than wide, 4 elongate, 5–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and wide, reaching midpoint of eye from frontal fovea. Anterior frontal fovea extremely small. Posterior frontal fovea round. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 19d). Meso-and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe with blunt lobe at middle (Fig. 19h). Phallobase of median lobe symmetrical and rectangular (Fig. 19h). Parameres asymmetrical, right paramere with 1branch from base (Fig. 19h).</p><p>Distribution. Gisborne (Fig. 20: star).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4111643CA1FAFDFC74FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4111643CF4FEEFFE78FB79.text	03D82A3EBD4111643CF4FEEFFE78FB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola ramsayi Park and Carlton	<div><p>Sagola ramsayi Park and Carlton, new species</p><p>(Figs. 19c, 19g, 20)</p><p>Type Material. Holotype. NEW ZEALAND: Gisborne: 1♂ (NZAC), “ New Zealand: GB: Lake Waikaremoana 17 I 1972, G.W. Ramsay Litter 72/17”, “ HOLOTYPE Sagola ramsayi Park and Carlton 2013 ” . Paratypes (3 males). NEW ZEALAND: Gisborne: 3♂♂ (NZAC), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype, Graeme W. Ramsay.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description. Length 3.2–3.5 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 19c). Head: Transverse, widest across temples (Fig. 19c). Antennomere 1 approximately 2.5 times longer than wide, 2–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep and wide, reaching midpoint of eye from frontal fovea. Anterior frontal fovea extremely small. Posterior frontal fovea round. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 19c). Meso-and metaventrite trapezoidal, as long as wide. Aedeagus: Apical lobe of genitalia V-shaped (Fig. 19g). Phallobase of median lobe symmetrical and rectangular (Fig. 19g). Parameres asymmetrical, left paramere with 2 branches from apex and base (Fig. 19g).</p><p>Distribution. Gisborne (Fig. 20: black square).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4111643CF4FEEFFE78FB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4111603F77FA0EFE4DFF68.text	03D82A3EBD4111603F77FA0EFE4DFF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola parva Sharp 1874	<div><p>Sagola parva Sharp, 1874</p><p>(Figs. 21a, 21c, 21e–g, 22)</p><p>Sagola parva Sharp, 1874: 508 . Broun 1880: 136; Schaufuss 1888: 85; Raffray 1893: 25, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen, 2006: 243.</p><p>Sagola fovealis Broun, 1886: 886 . Raffray 1893: 37, 1904: 498, 1911: 6, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola brevicornis Raffray, 1893: 27 . Raffray 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola insolens Broun, 1893b: 1051 . Raffray 1904: 498, 1911: 6; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen, 2006: 242. New synonymy.</p><p>Sagola rectipes Broun, 1893b: 1051 . Raffray 1904: 498, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola punctata Broun, 1893b: 1052 . Raffray 1904: 498, 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola concolorata Broun, 1915: 292 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola rotundiceps Broun, 1915: 289 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material Examined. Holotype of Sagola parva: NEW ZEALAND: 1♀ (BMHM), glued on rectangular card, “249” [green label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola parva Cotype. Sharp. ” [white label, handwritten] . Holotype of Sagola concolorata: NEW ZEALAND: Coromandel: 1♀ (BMHM), glued on rectangular card, “Type” [red label, printed]; “3709. ♂ ” [white label, handwritten]; “Tairua. Auckland.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ concolorata.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola fovealis: NEW ZEALAND: Auckland: 1♂ (BMHM), glued on rectangular card, “Type” [red label, printed]; “1577. ♂ ” [white label, handwritten]; “Waitakerei” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “sagola ♂ fovealis.”[white label, handwritten] . Syntypes of Sagola insolens: NEW ZEALAND: Auckland: 1♀ (BMHM), glued on rectangular card, “Type” [red label, printed]; “1879” [white label, handwritten]; “ ♀ “ [white label, handwritten]; “ Howick ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola insolens ”[white label, handwritten]. 1♂, glued on rectangular card, “1879.” [white label, handwritten]; “ ♂ “ [white label, handwritten]; “ Howick ” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola insolens ”[white label, handwritten] . Holotype of Sagola punctata: NEW ZEALAND: Coromandel: 1♀ (BMHM), glued on rectangular card, “Type” [red label, printed]; “1880.” [white label, handwritten]; “Tairua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola punctata .” [white label, handwritten] . Holotype of Sagola rectipes: NEW ZEALAND: Coromandel: 1♀ (BMHM), glued on rectangular card, “Type” [red label, printed]; “1878.” [white label, handwritten]; “Tairua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola rectipes ” [white label, handwritten] . Holotype of Sagola rotundiceps: NEW ZEALAND: Taranaki: 1♀ (BMHM), glued on rectangular card, “Type” [red label, printed]; “3704. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Stratford. Taranaki.” [white label, handwritten]; “ Sagola ♂ rotundiceps.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female. Two of three syntypes of Sagola brevicornis: NEW ZEALAND: Auckland: 2♂ ♀ (MNHN), glued on rectangular card, “ MP / S. brevicornis AR ” .</p><p>Additional Material (n = 88: 50 males; 38 females). NEW ZEALAND: Auckland: 1♂ 3♀♀, Clevedon Scenic Res., 0.5km N Clevedon, 20m, 19 III 2010, D.S. Chandler, sift forest litter by stream ; 1♀, Manurewa, Murphy’ s Bush, 2 III 1981, G. Kuschel, rotten wood 81/59 ; 1♀, Grafton, Gulley, 6 III 2002, J. Nunn ; 1♀, Riverhead SF Res., 17–26 XI 1983, P.A. Maddison, Malaise trap ; 1♀, Waitakere Ra., Cascade-Kauri Park, up. Kauri tr., 170m, 8 XII 1984 – 25 I 1985, kauripodo-hdwd., A. Newton, M. Thayer 680, FIT &amp; window trap; 1♂, Symond St. Cemetery, Auckland, 40m, 2–6 IV 2010, L. Masner, yellow pan traps; 1♂, Matuku Res., 45m, 6.5km W Waitakere end of Jonker’ s Rd., 2–5 IV 2010, L. Masner, yellow pan traps ; 1♂, Symonds St. Cemetery, Auckland, 40m, 23 II 2010, D.S. Chandler, sift leaf litter in ravine; 1♂, Hunua Ranges Reg. Park, 7km W Kaiaua Workman Tr., 116m, 27 III 2010, D.S. Chandler, sift leaf litter by stream ; 1♂, Auckland, 40m, Auckland Domain, 25 III 2001, S.E. Thorpe, on underside of logs in bush ; 1♂, Waikowai, Captains Bush, 10 II 1992, G. Hall ; 1♂, Bethells, Matuku Res., 23 VI 1987, R. C. Craw, sifted rotten wood 87/9 ; 1♂, Symond St. Cemetery, 8 II 2004, J. Nunn ; 1♂, The Dome, 270m, nr Warkworth, 7 I 1983, J.C. Watt, wood mould 83/8 ; 1♂, Waitakere Ranges, 260m, Nohoanga Scenic Res., 8 XII 1984 – 25 I 1985, hdwd.-podocarp forest, A. Newton, M. Thayer 679, FIT &amp; Window trap ; 1♂, Symond St. Cemetery, Auckland, 40m, 24–25 III 2010, yellow pan traps, L. Masner; 1♂, Mt. Auckland, Atuanui Scenic Res., 250m, 2.5km NE Glorit, 2 II 2010, D.S. Chandler, sift rotten wood ; 1♂, Mt. Auckland, Atuanui Scenic Res., 250m, 2.5km NE Glorit, 2 II 2010, D.S. Chandler, sift fallen epiphyte mass; Bay of Plenty : 2♀♀, Tapapa, 300m, 25 III 1978, S.B. Peck, litter ; 1♀, Mt. Te Aroha, 2.5km SE Te Aroha, 390m, Tui Rd. at Tui Creek, Mt. Domain tr., 15 II 2010, D.S. Chandler, sift rotten wood ; 1♂, Tapapa Tukorehe Res., 300m, 25 III 1978, S. &amp; J. Peck, berl., litter ; 1♂, Tapapa Tukorehe Res., 25–29 III 1978, S. &amp; J. Peck, 300m, bait (carrion) trap ; 1♂, Lottin Pt Rd, Waenga Bush, 16 IX 1992, G. Hall, Wood mould 92/52 ; 1♂, Lake Okataina, 9 X 1995, M.C. Lariviere, A. Larochelle, litter 95/12 ; 1♂, Papatea, 26 IV 1993, J. S. Dugdale, litter 93/ 94; Coromandel : 1♂ 1♀, Great Barrier I, Little Windy Hill, 13 XII 2002 – 17 I 2003, P. Sutton, forest edge malaise trap ; 1♀, Great Barrier I, Little Windy Hill, 17 I–27 II 2003, K. Parsons, forest edge Malaise trap; Northland : 4♂♂ 4♀♀ (1♂ 1♀, slide-mounted), Waipoua State Forest, 0.9km e Forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.- podocarp forest, A. Newton, M. Thayer 686, FIT &amp; window trap ; 4♂♂ 1♀, Waipoua SF, Toronui Tr., 150m, 13 IV 1980, kauri-podocarp-broadleafnikau palm forest, A. Newton, M. Thayer, pyrethrin fogging fungusy logs ; 1♂ 2♀♀, Waipoua State Forest, 0.8km nw Wairau Summit, 350m, 27 XI– 6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 689, FIT &amp; window trap ; 1♂, Waipoua State Forest, Wairau Summit, 400m, 27 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 687, FIT &amp; window trap ; 1♂ 1♀, Waipoua SF, 0.8km s Waikohatu Stream bridge, 27m, 28 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 692, FIT &amp; window trap ; 1♀, Waipoua SF, Te Matua Ngahere, 400m, 19 II 1978, berl., under bark of fallen tree with mushroom, S. Peck ; 2♂♂, Waipoua SF, 24 XI 1980, G. Kuschel, on Agathis australis log ; 1♂, Waipoua SF, 25 XI 1980, G. Kuschel, decayed wood 8/120 ; 1♂, Waipoua SF, Waipoua Stm, 70m, 16–21 III 1978, S. &amp; J. Peck, malaise trap in forest clearing; Wellington : 2♂♂, Tinakori Hill, Wellington, 1 VIII 1991, J. Nunn, in decayed wood; 1♂, 4km S of Levin, 30m, 8 III 1978, S. &amp; J. Peck, litter ; 1♀, Kaitoke Regional Park, Pakuratahi Forks, 15 IV 2005, R. Leschen, C. McGuiness, leaf litter ; 2♀♀ (1♀, slide-mounted), Wilton’ s Bush, 110m, 41″15.963′S 175″45.159′E, 24 XI 2005, mixed broadleaf-podocarp forest, FMHD#2005-030, berl., leaf &amp; log litter, M. Thayer, A. Newton, ANMT site 1150 ; 1♂, Tinakori Hill, 1 VI 1991, J. Nunn, decaying wood ; 1♂ 1♀, Tinakori Hill, 1 VI 1991, J. Nunn, decaying wood ; 2♂♂ 1♀, Tinakori Hill, 5 VI 1991, J. Nunn, decaying wood with Mesoponera ants ; 1♂, Tinakori Hill, 10 VIII 1991, J. Nunn, in soggy decaying wood ; 2♀♀, Tinakori Hill, 1 IX 1991, J. Nunn, in partly humified log ; 1♀, Manawatu Gorge, 17 XI 1991, J. Nunn ; 1♀, Tinakori Hill, 15 XII 1991, J. Nunn, in cut grass ; 1♂, Tinakori Hill, 21 IV 1986, J. Nunn, with Mesoponera ants ; 1♂, Johnson’ s Hill, Karori, 17 VII 1988, J. Nunn ; 1♀, Tinakori Hill, 14 IV 1992, J. Nunn, in rotten wood ; 4♂♂ 2♀♀, Eastbourne, 22 VIII 1993, J. Nunn, in decayed wood ; 1♀, Tinakori Hill, 11 IX 1993, J. Nunn, in decaying wood ; 1♀, Klein Tr., Manawatu Gorge, 16 VII 1994, J. Nunn, in decaying wood ; 1♂ 1♀, Karori Res., 25 II 1995, J. Nunn, in twiggy streamside debris ; 1♀, Tinakori Hill, 8 VII 1996, J. Nunn, under plank beneath pines ; 1♂, Keith George Res., Silverstream, 27 XII 1996, J. Nunn, in pulpy decayed wood with Ganoderma fungus .</p><p>Diagnosis. This species can be distinguished from S. denticollis by the bluntly triangular head, larger eye, enlarged male antennomeres 5–6 lacking distinctive tubercles, and unique shape of the genitalia.</p><p>Redescription. Length 2.3–3.8 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 21a). Head: Bluntly triangular with prominent frontal rostrum (Fig. 21f). Male antennomeres 2–10 subquadrate, 5–6 enlarged, 5 largest. Female antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye from frontal fovea. Anterior frontal fovea oval, open anteriorly, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 21a). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe with minor lobe bearing from base (Fig. 21c). Phallobase of median lobe asymmetrical and rounded (Fig. 21c). Parameres asymmetrical, left paramere longer than right with 2 slender branches apically (Fig. 21c).</p><p>Type Locality. New Zealand (unknown specific locality) .</p><p>Distribution. A u c k l a n d, B a y o f P l e n t y, Coromandel, Northland, Taranaki, Wellington (Fig. 22: black circles).</p><p>Habitat. Most specimens of this species were collected using yellow pan and Malaise traps or were taken by sifting leaf litter or rotten wood in hardwood or podocarp forests. Some specimens were collected with Mesoponera ants, and one specimen was collected from a carrion bait trap.</p><p>Comments. Specimens of S. parva are easy to distinguish from other species externally by the enlarged male antennomeres 5–6; large eye that is approximately one-half length of temple; prominent frontal rostrum; shallow frontal sulcus; oval and anteriorly open anterior frontal fovea. The type specimens of S. brevicornis, S. concolorata, S. fovealis, S. insolens, S. punctata, S. rectipes, and S. rotundiceps share these diagnostic characters. For these reasons, we have placed these species in synonymy with S. parva .</p></div>	https://treatment.plazi.org/id/03D82A3EBD4111603F77FA0EFE4DFF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4511603D41FED1FB4FFB95.text	03D82A3EBD4511603D41FED1FB4FFB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola denticollis Broun 1880	<div><p>Sagola denticollis Broun, 1880</p><p>(Figs. 21b, 21d, 22)</p><p>Sagola denticollis Broun, 1880: 138 . Schaufuss 1888: 84; Raffray 1893: 33, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola elevata Broun, 1886: 886 . Raffray 1893: 38, 1904: 498. 1911: 6, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material Examined. Holotype of Sagola denticollis: NEW ZEALAND: Coromandel: 1♀ (BMNH), glued on rectangular card, “253” [green label, printed]; “Type” [red label, printed]; “Tairua” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola denticollis .” [white label, handwritten] . Syntypes of Sagola elevata: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1578.” [white label, handwritten]; “Woodhill” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola elevata .” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “1578.” [white label, handwritten]; “Woodhill” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] .</p><p>Additional Material (n = 18: 9 males; 9 females). NEW ZEALAND: Auckland: 1♂ 1♀, Bethells Matuku FS, 36′53S 174′28E, under bark #157, 29 I 1998, C. Carlton, R. Leschen; 1♀, Waitakere R., Cascade-Kauri Park, up. kauri tr., 170m, 8 XII 1984 – 25 I 1985, kauri-podo-hdwd, A. Newton, M. Thayer 680, FIT &amp; window trap ; 1♀, Matakana nr Warkworth, 27 XI 1995, J. Nunn, in decayed wood from bush remnant ; Coromandel: 5♂♂, Little Barrier Island, 7 I 1952, rotten wood ; 2♀♀, Little Barrier Island, 6 I 1952, rotten wood ; 1♀, Great Barrier Island, Little Windy Hill, 13 XII 2002 – 17 I 2003, P. Sutton, forest edge malaise trap ; 1♀, Little Barrier Island, 17 XII 1983, J. Playfair ; Northland: 3♂♂ 2♀♀ (1♂, slide-mounted), Parua, 4 XI 1981, G. Kuschel, litter and decayed wood 81/125 ;</p><p>Diagnosis. This species can be distinguished from S. parva by the transverse head, small eyes, enlarged antennomeres 4–6 of male with distinctive tubercles, and genitalia.</p><p>Redescription. Length 2.5–3.5 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 21b). Head: Transverse, widest across eyes, frontal rostrum prominent (Fig. 21b). Male antennomeres 2–3 subquadrate, 4–6 enlarged, 6 largest, 7–10 subquadrate, 3–10 with distinctive tubercles. Female antennomeres 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching hind point of eye from frontal fovea. Anterior frontal fovea oval, posterior frontal fovea surrounded by setae. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 21b). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe with triangular lobe at midpoint (Fig. 21d). Phallobase of median lobe symmetrical and rectangular (Fig. 21d). Parameres asymmetrical, left paramere shorter than right, right divided anteriorly (Fig. 21d).</p><p>Type Locality. Tairua, Coromandel, New Zealand .</p><p>Distribution. Auckland, Coromandel, Northland (Fig. 22: triangles).</p><p>Habitat. Specimens of this species were collected using window and flight intercept traps or by sifting leaf litter or rotten wood in kauri, podocarp, and hardwood forests.</p><p>Comments. Specimens of S. denticollis are easily distinguished from other species externally by the transverse head with a prominent frontal rostrum. The type specimens of S. elevata share these diagnostic characters. For these reasons, we have placed S. elevata in synonymy with S. denticollis .</p></div>	https://treatment.plazi.org/id/03D82A3EBD4511603D41FED1FB4FFB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4511603F04FBCAFCB5F99D.text	03D82A3EBD4511603F04FBCAFCB5F99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola valida Broun Desig. Park and Carlton 2013	<div><p>7. valida species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the valida species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 3.0– 3.5 mm; antennomere 1 approximately 2.5 times longer than wide, with dull surface; anterior frontal fovea absent, posterior frontal fovea elongate (Fig. 23c); gular region of male head with small, transverse depression (Fig. 23d); eye distinctly prominent due to emarginate gena (Fig. 23c–d); hind wings well-developed in male, reduced to small pads in female; fore femur with semicircular depression; abdominal tergites IV–VI with discal carinae; present on South Island, not known from North Island (Fig. 24).</p></div>	https://treatment.plazi.org/id/03D82A3EBD4511603F04FBCAFCB5F99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4511623F6AF9C1FB50FD2F.text	03D82A3EBD4511623F6AF9C1FB50FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola valida Broun Desig. Park and Carlton 2013	<div><p>Sagola valida Broun, 1921</p><p>(Figs. 23–24)</p><p>Sagola valida Broun, 1921a: 490 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Type Material Examined. L e c t o t y p e o f Sagola valida: NEW ZEALAND: Nelson: 1♂ (BMNH), glued on rectangular card, “3998. ♂ ” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “mt. owen. 27.12.1914 ” [white label, handwritten]; “ Sagola valida . ♂.” [white label, handwritten]; “ LECTOTYPE Sagola valida Broun Desig. Park and Carlton 2013 ” [red label, printed]. Paralectotypes (2 males) . NEW ZEALAND: Nelson: 2♂♂ (BMNH), Woodhen near Glenhope, T. Hall. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that three specimens were examined (Broun 1921a: 490). This designation will fix the identity of the species and facilitate its recognition by future workers .</p><p>Additional Material (n = 59: 28 males; 31 females). NEW ZEALAND: Buller: 2♂♂ 2♀♀, Nelson Lakes NP, Mt. Robert Speargrass Tr., 875m, 41″49.469′S, 172″48.311′E, 30 XI–17 XII 2005, Nothofagus forest, FMHD#2005-059, FIT, A. Newton, M. Thayer, ANMT site 1161 ; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass Tr., 875m, 41″49.469′S, 172″48.311′E, 17 XII 2005, Nothofagus forest, FMHD#2005-110, berl., leaf &amp; log litter, A. Solodovnikov, D. Clarke, ANMT site 1161 ; 2♂♂ 1♀, Lewis Pass N. R., 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, berl., leaf &amp; log litter, forest floor ; 2♀♀, Lewis Pass N. R., 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest , A. Newton, M. Thayer 715, window trap; 1♂ 1♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &amp;window trap; 1♂ 1♀, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, FIT &amp;window trap; 1♂, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &amp;window trap; 1♂ 1♀, Mt. Robert, carpark, 9 IV 2004, R. Leschen, H. Haman, T. Buckley, litter RL864, 41.49′S, 172.48′E; 1♂, Speargrass Tr., Mt. Robert, 30 XI 2005, J. Nunn; 1♂, Lewis Pass, Asa Pass, 914m, 25 IV 1996, G.W. Ramsay, moss 66/141; 1♂, Rahu Saddle, 670m, 16 X 1984, J.W. Early, moss in Nothofagus forest, LCNZ 84/7; 1♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr., 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp;window trap; 1♀, Nelson Lakes NP, n slope Mt. Robert, Pinchgut tr., 1290m, 18–26 XII 1984, Noth. Sol. elfin forest, A. Newton, M. Thayer 716, FIT &amp;window trap; 2♀♀, Barrytown Croesus Tr., 10 XI 2005, R. Leschen, S. Nomura, litter RL1022, 42°14′S, 171°19′E; 1♀, Inland Pack tr., 1km n of Punakaiki R, 29 XII 2010, J. Nunn, sifted ground litter; Fiordland: 1♂, Hollyford tr., #110, 44′42S 168′08E, Nothofagus coastal forest leaf litter berlese, 22 I 1998, C. Carlton, R. Leschen; Marlborough: 1♂, Fell Pk, Hut Area, 1300m, Richmond Ra., 13 III 1969, A.C. Eyles, litter 69/109; North Canterbury: 1♂, Arthurs Pass, Mt. Aicken, 950m, 9 II 1984, J.C. Watt, sifted litter 84/24; 1♀, 2.9km N Arthur’ s Pass, Bealey Valley tr., 840–950m, 42′55S 171′33E, #040, under bark, 11 I 1998, C. Carlton, R. Leschen; Nelson: 2♂♂ 2♀♀ (1♂, slide-mounted), Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S, 172″44.456′E, 28 XI–19 XII 2005, Nothofagus forest, FMHD #2005-044, FIT, A. Newton, M. Thayer, ANMT site 1156; 1♂ 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus cliffortioides forest, FMHD #2005- 051, FIT, A. Newton, M. Thayer, S. Solodovnikov, ANMT site 1159; 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-055, berl., leaf &amp; log litter, M. Thayer, ANMT site 1159; 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, under bark small Nothofagus logs, M. Thayer, ANMT site 1159; 1♂ 1♀, Kahurangi NP, Cobb Dam Rd., Asbestos Tr., 450m, 41″06.333′S, 172″43,174′E, 29 XI–18 XII 2005, mixed broadleaf-podocarp forest, FMHD #2005-111, berl., leaf &amp; log litter, A. Solodovnikov, D. Clarke, et al., ANMT site 1160; 2♀♀, 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717 FIT &amp;window trap, carrion (squid) trap; 1♂ 1♀, Dun Mt., 31 VI 1966, A.K. Walker, litter 66/274; 1♂, Dun Mt., 762m, 13 I 1961, G. Kuschel, litter 61/1; 1♂, Karamea Bluff, 1420ft, J. Nunn; 1♂, Denniston Saddle, 10 II 1999, R. Leschen, R. Hoare, Berl., Nothofagus forest RL288, 41°44′S 171°48′E; 1♂, Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, wet leaves &amp; flood debris forest stream; 1♂, Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD #2005-049, berl., leaf litter, A. Solodovnikov, D. Clarke, ANMT site 1157; 1♂, 14 km se Reefton, 250m, 29 V 1982, FMHD #82-606, beech forest litter, S. Peck; 1♀, 30km nw Motueka, Tasman NP, 22 V 1982, FMHD #82-596, beech forest stump litter, S. Peck; 1♂, Dun Mt., 2000′ 14 II 1942, E.S. Gourlay; 1♀, Punakaiki, Pororari Tr., 27 I 1996, J. Nunn, in forest litter; 1♀, Kohaihai River, Karamea, West Coast, 30 V 1963, G. Kuschel, litter; 1♀, Oparara Basin, Box Canyon Cave, 200m, 27 II–1 III 2007, J.W. Early, R.F. Gilbert, Nothofagus podocarp forest, yellow pan trap; Westland: 1♀, Jackson Bay tr., 16 I 2005, R. Leschen, T. Buckley, podocarp, RL906, 43′58′S, 168.37′E; 1♂, 3.2km ne Haast, 14m, Haast River Walk, 43′52S 169′03E, #072, Nothofagus -podocarp forest leaf litter berl., 17 I 1998, C. Carlton, R. Leschen; 1♂, Franz Joseph, 24 II 1989, J. Nunn; 1♀, Hokitika R. Gorge, S Kowhitirangi, 100m, 17 III 1980, podocarp-broadleaf, A. Newton, M. Thayer, berl. leaf &amp; log litter, forest floor; 1♀, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd.- podo-nikau forest, A. Newton, M. Thayer 719, berl., leaf &amp; log litter, forest floor.</p><p>Diagnosis. This species can be distinguished from other Sagola species by the gular region of the male head with small round depression; both sexes with eyes distinctly prominent due to emarginate gena.</p><p>Redescription. Length 3.0– 3.5 mm. Body brown, antennae, elytra, legs, and maxillary palpi pale (Fig. 23a). Head: Bluntly triangular, widest across eyes (Fig. 23c). Male antennomeres 2–3 subquadrate, 4 longer than wide, 5–10 subquadrate. Female antennomeres 2–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching behind eye from end of frontal rostrum (Fig. 23c). Anterior frontal fovea absent (Fig. 23c), posterior frontal fovea elongate (Fig. 23c). Eye large and prominent, one-half length of temple (Fig. 23c). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular in male (Fig. 23a), subquadrate in female. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle in male, patches absent in female. Aedeagus: Median lobe deeply divided, minor lobe longer and narrower (Fig. 23b). Phallobase of median lobe symmetrical and rounded (Fig. 23b). Parameres symmetrical, ca. 8 setae apically (Fig. 23b).</p><p>Type Locality. Mount Owen and Woodhen Bend, near Glenhope, Nelson, New Zealand .</p><p>Distribution. Buller, Fiordland, Marlborough, North Canterbury, Nelson, Westland (Fig. 24: black circles).</p><p>Habitat. Most specimens of this species were collected using window and flight intercept traps or by sifting leaf litter or rotten wood in Nothofagus forests. Some specimens were collected from carrion bait or yellow pan traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD4511623F6AF9C1FB50FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD4711633F04FD2BFB42FE73.text	03D82A3EBD4711633F04FD2BFB42FE73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola incisa	<div><p>8. incisa species-group</p><p>(3 species)</p><p>Diagnosis. The members of the incisa speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.2–2.9 mm; antennomere 1 approximately 2.5 times longer than wide with dull surface; anterior frontal fovea oval, posterior frontal fovea elongate (Fig. 25g); frontal sulcus reaching length of vertexal fovea (Fig. 25g); gular region of male head with round depression with small process in middle (Fig. 25h: arrow); hind wings well-developed; fore and mid-femur with semicircular depression; mid-tibia bent; male tergite IV with microtrichial patch covering most of the surface; abdominal tergites IV–VI with discal carinae; known from North Island and Three Kings Islands (Fig. 26).</p><p>KEY TO SPECIES OF THE INCISA SPECIES- GROUP</p><p>The key is based on genitalia because most specimens are indistinguishable based on external morphology.</p><p>1. Median lobe of genitalia much shorter than parameres, never reaching midpoint of paramere; parameres asymmetrical (Fig. 25e) ................ S. deformipes Broun</p><p>1′. Median lobe of genitalia shorter than parameres but exceeding midpoint of paramere; parameres symmetrical..........2</p><p>2(1′). Median lobe of genitalia reaching twothirds level of parameres, phallobase of genitalia rectangular (Fig. 25d); only known from Three Kings Islands (Fig. 26: black circles)................................................ .................... S. incisa Théry and Leschen</p><p>2.′ Median lobe of genitalia reaching almost apices of parameres, phallobase of genitalia round (Fig. 25f); only known from Northland (Fig. 26: black square)........................ .................... S. unuwhaoensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD4711633F04FD2BFB42FE73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD46117C3E93FDD0FE8FF95B.text	03D82A3EBD46117C3E93FDD0FE8FF95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola incisa Thery and Leschen 2013	<div><p>Sagola incisa Théry and Leschen, 2013</p><p>(Figs. 25a, 25d, 26)</p><p>Sagola incisa Théry and Leschen, 2013: 42 .</p><p>Type Material. Paratypes. NEW ZEALAND: Three Kings Islands: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, East Sector, E.G. Turbott, 6 X 1948, A.E. Brookes collection ; 1♀ (NZAC), Great Island, Castaway Tr., 9 XI 2008, sifting rotten wood and leaf litter, T. Buckley, R. Leschen, TH 043, 34°09.369, 172°08.496 .</p><p>Additional Material (n = 43; 21 males; 22 females). NEW ZEALAND: Three Kings Islands: 3♂♂ 3♀♀, Summit, 275m, 23 XI 1970, G. Kuschel, litter 70/202 ; 1♂, Castaway Camp, 16 XI 1970, G. Kuschel, litter 70/195 ; 5♂♂ 1♀, Tasman Valley, 24 XI 1970, G. Kuschel, litter 70/211 ; 2♂♂, Tasman Valley, 26 XI 1970, J.C. Watt, litter 70/223 ; 1♂, W Summit, 240m, 26 XI 1970, J.C. Watt, moss 70/219 ; 1♂, Tasman Valley, 25 XI 1970, G. Kuschel, litter 70/215 ; 1♀, Tasman Valley, 24 XI 1970, G. Kuschel, litter 70/210l ; 1♀, Castaway Camp, 16 XI 1970, G. Kuschel, litter 70/195 ; 2♀♀, Castaway Camp, 29 XI 1970, G.W. Ramsay, litter 70/233 ; 4♂♂ 6♀♀, Great Island, 45m, South East Bay, 1 XII 1983, J.C. Watt, litter ; 2♂♂ 3♀♀, Tasman Valley, 2 XII 1983, J.C. Watt, litter ; 1♂, Great Island, 45m, 28–30 XI 1983, C.F. Butcher, pan trap near coast ; 1♀, Great Island, Tasman Bay, XII 1983, C.F. Butcher, moss ; 1♀, Great Island, area below trig litter 96/21, 8 XII 1996, G.L.F. Carlin ; 1♀, Great Island, Baylis Stm, sifted litter 99/24, 13 IV 1999, T. K. Crosby ; 2♀♀, Eastern Sector, E.G. Turbott, 6 X 1948, leaf mould, A.E. Brookes collection .</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia. This species is only known from Three Kings Islands.</p><p>Redescription. Length 2.2–2.6 mm. Body brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 25a). Head: Transverse, widest across temples (Fig. 25a). Antennomere 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate. Eye small and prominent, approximately one-third length of temple. Thorax: Prosternum weakly transverse, widest at midpoint. Elytra rectangular (Fig. 25a). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe slender, shorter than parameres (Fig. 25d). Phallobase of median lobe asymmetrical and rectangular (Fig. 25d). Parameres symmetrical with dense setae along midline (Fig. 25d).</p><p>Type Locality. Great Island, Three Kings Islands, New Zealand .</p><p>Distribution. Three Kings Islands (Fig. 26: black circle).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p><p>Comments. This species was described by Théry and Leschen (2013) with illustrations of habitus and diagnostic characters based on specimens collected only from the Three Kings Islands, an isolated archipelago ca. 60 km northwest of North Island. We based our study of this species on two paratypes.</p></div>	https://treatment.plazi.org/id/03D82A3EBD46117C3E93FDD0FE8FF95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD59117D3F53FBFAFEBFFC6E.text	03D82A3EBD59117D3F53FBFAFEBFFC6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola deformipes Broun 1880	<div><p>Sagola deformipes Broun, 1880</p><p>(Figs. 25b, 25e, 25g –h, 26)</p><p>Sagola deformipes Broun, 1880: 138 . Schaufuss 1888: 84; Raffray 1893: 33, 1904: 497, 1911: 5; Hudson 1923: 365, 1934: 183; Raffray 1924: 231; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “252” [green label, printed]; “Tairua” [white label, printed]; “252. ♂.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola deformipes ” [white label, handwritten].</p><p>Additional Material (n = 7; 4 males, 3 females). NEW ZEALAND: Auckland: 1♀, Waitakere Rgs, 1 VI 1942, M.W. Carter ; Bay of Plenty: 1♂, Tapapa Tukorehe Res., 300m, 25 III 1978, S. &amp; J. Peck, berl., litter ; Northland: 1♂, Waipoua SF, 20 IX 1977, SH12, D.W. Helmore, rotten stumps/ trees 77/102 ; 1♀, Waipoua SF, Taraite Stump, 15 VI 1966, J.C. Watt ; 1♂, Waipoua SF, Wairau Summit, 387m, 11–14 IV 1980, podocarp-mixed broadleaf, A. Newton, M. Thayer, pyrethrin fogging Ganoderma sp .; 1♀, Waipoua SF, Toatoa tr., 270m, 12–15 IV 1980, toatoa-kauri-podocarpbroadlf, A. Newton, M. Thayer, berl., leaf and log litter, forest floor; Taupo: 1♂ (slide-mounted), Pureora Forest, Rimu Walk, 30 I 2008, K. Marske, R. Leschen, T. Buckley, Sifted wood and leaf litter, S 38.33.987′, E 175.43.068′, 479 m.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Redescription. Length 2.6–2.9 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 25b). Head: Transverse, widest across temples (Fig. 25g). Antennomere 2 longer than wide, 3–10 subquadrate, 5–6 weakly enlarged. Eye small and prominent, one-third length of temple. Thorax: Prosternum weakly transverse, widest at midpoint. Elytra rectangular (Fig. 25b). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe slender, much shorter than parameres (Fig. 25e). Phallobase of median lobe asymmetrical and rectangular (Fig. 25e). Parameres asymmetrical, left longer than right with dense setae along mid line except apex (Fig. 25e). Right paramere with dense setae along midline (Fig. 25e).</p><p>Type Locality. Tairua, Coromandel, New Zealand .</p><p>Distribution. Auckland, Coromandel, Northland, Taupo (Fig. 26: triangles).</p><p>Habitat. Specimens of this species were collected by sifting leaf and log litter in podocarp and broadleaf forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD59117D3F53FBFAFEBFFC6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD58117D3F61FE15FB42F9B1.text	03D82A3EBD58117D3F61FE15FB42F9B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola flavipes Broun, 1893 b: 1422	<div><p>9. flavipes species-group</p><p>(2 species)</p><p>Diagnosis. The members of the flavipes species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.4–2.8 mm; head transverse, widest across eyes (Fig. 27e); antennomere 1 approximately 2 times longer than wide with sparse punctures; anterior frontal fovea small round, but not externally observable because it is covered by the connected frontal rostrum (Fig. 27e); posterior frontal fovea elongate (Fig. 27e); frontal sulcus reaching length of vertexal fovea (Fig. 27e); ventral surface of male head with setose horn medially (Fig. 27f: arrow); hind wings reduced to small pads; fore femur with semicircular depression; mid-tibia bent; Male tergite IV without microtrichial patch; abdominal tergites IV–VI with discal carinae; abdominal ventrites IV–VI with basolateral foveae; present on North Island, not known from South Island (Fig. 28).</p><p>KEY TO SPECIES OF THE FLAVIPES SPECIES- GROUP</p><p>The key is mainly based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Horn of ventral surface of head shorter than wide; median lobe of genitalia branched into 3 lobes, major lobe triangular (Fig. 27c); parameres asymmetrical and as long as median lobe (Fig. 27c).................... S. flavipes Broun</p><p>1′. Horn of ventral surface of head as long as wide; median lobe of genitalia branched into 2 lobes, major lobe rectangular (Fig. 27d); parameres symmetrical and reaching midpoint of median lobe (Fig. 27d) .................... .......................................... S. sulcator Broun</p></div>	https://treatment.plazi.org/id/03D82A3EBD58117D3F61FE15FB42F9B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD58117D3CFEFBECFC48FE5C.text	03D82A3EBD58117D3CFEFBECFC48FE5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola unuwhaoensis Park and Carlton	<div><p>Sagola unuwhaoensis Park and Carlton, new species</p><p>(Figs. 25c, 25f, 26)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, ND Unuwhao, 270m, 25 Nov 1982 G. Kuschel ” “sifted litter and decayed wood 82/125”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola unuwhaoensis Park and Carlton 2013 ”.</p><p>Etymology. This species is named after the type locality, Unuwhao, Northland.</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the shape of the antennae and genitalia.</p><p>Description of Male. Length 2.8 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 25c). Head: Transverse, widest across temples (Fig. 25c). Antennomeres 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Eye small and prominent, one-third length of temple. Thorax: Prosternum weakly transverse, widest at midpoint. Elytra rectangular (Fig. 25c). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe slender, slightly shorter than parameres (Fig. 25f). Phallobase of median lobe asymmetrical and rounded (Fig. 25f). Parameres symmetrical with dense setae along midline and thick setae apically (Fig. 25f).</p><p>Distribution. Northland (Fig. 26: black square).</p><p>Habitat. The holotype and only known specimen of this species was collected by sifting leaf litter and decayed wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD58117D3CFEFBECFC48FE5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD58117F3F57F9BEFDDEFB99.text	03D82A3EBD58117F3F57F9BEFDDEFB99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola flavipes Broun, 1893 b: 1422	<div><p>Sagola flavipes Broun, 1893</p><p>(Figs. 27a, 27c, 27e–f, 28)</p><p>Sagola flavipes Broun, 1893b: 1422 . Raffray 1904: 498, 1911: 6; Hudson 1923: 365, 1934:</p><p>183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2471. ♂ ” [white, handwritten]; “Hunua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola flavipes ” [white label, handwritten].</p><p>Additional Material (n = 15: 11 males; 4 females). NEW Z EAL AND: A u c k l a n d: 2♂♂, Clevedon Scenic Reser., 05km n Clevedon, 20m, 19 III 2010, D.S. Chandler, sift forest litter by streams ; 1♂, Waitakere Ranges Reg. Park, 1.3km n Hunua, Parau Tr., 133m, 20 III 2010, D.S. Chandler, sift kauri, Cyathea &amp; etc. leaf litter ; 1♀, Waitakere Ranges Reser. Pk., Upper Huia Reser . tr., 31 I 2010, 340m, 4km sw Waiatarua, D.S. Chandler, sift mixed leaf litter; 2♀♀, Le Roys Bush, Birkenhead, 21 IX 1957 ; 1♂, Kohukohunui, Hunua Ra., 600m, 30 III 1974, G. Kuschel, litter 74/20 ; 1♂, Waitakere Ra., IV 1955, A.G. Smith, litter ; 1♂ (slide-mounted), Goldie Bush Scenic Res., Mokoroa Falls Tr., 6km w Waitakere, 28 III 2010, 106m, D.S. Chandler, rotten wood &amp; branch debris; Bay of Plenty: 1♂, Kaimai-Mamaku Forest Park, Mt. Te Aroha summit rd., 450m, 37”31.429′S 175″44.005′E, 19 XI 2005, mixed broadleaf forest w/many tree ferns, nikau palms, FMHD#2005-016, FIT, A. Newton, M. Thayer, ANMT site 1144; 1♂, Tikitiki Stm., Horohoro SF, Mamaku Plateau, 24 VII 1976, J.S. Dugdale, litter 76/48; Waikato: 3♂♂, Hapuakohe ra., 3km sw of Kaihere, 29 I 1984, J.C. Watt et al., sieved wood mould 84/7; 1♀, Pirongia Forest Park, Mahaukura tr., (above end Grey Rd.), 270m, 37″58.218′S 175″06.523′E, 18 XI 2005, mixed broadleaf forest, FMHD#2005- 011, berl., leaf &amp; log litter, M. Thayer &amp; A. Newton, ANMT site 1142 .</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the horn on the ventral surface of the male head being shorter in length than width and the shape of the antennae and genitalia.</p><p>Redescription. Length 2.5–2.8 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 27a). Head: Transverse (Fig. 27e). Antennomeres 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Eye small and prominent, one-third length of temple (Fig. 27e). Horn on ventral surface of male head triangular and shorter in length than width. Thorax: Prosternum as long as wide, widest at one-third length. Elytra approximately triangular (Fig. 27a). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe branched into 3 lobes, as long as left paramere, major lobe triangular (Fig. 27c). Phallobase of median lobe symmetrical and rounded (Fig. 27c). Parameres asymmetrical with acute bent apices, left paramere longer than right (Fig. 27c).</p><p>Type Locality. Maketu, Auckland, New Zealand .</p><p>Distribution. Auckland, Bay of Plenty, Waikato (Fig. 28: black circles).</p><p>Habitat. Most specimens of this species were collected using flight intercept traps or by sifting leaf and log litter in broadleaf forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD58117F3F57F9BEFDDEFB99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5A11783CBFFBD5FC0DFF68.text	03D82A3EBD5A11783CBFFBD5FC0DFF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sulcator Broun 1886	<div><p>Sagola sulcator Broun, 1886</p><p>(Figs. 27b, 27d, 28)</p><p>Sagola sulcator Broun, 1886: 885 . Raffray 1893: 36, 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Sagola crassulipes Broun, 1915: 283 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1575. ♂.” [green label, printed]; “Woodhill” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola sulcator . ♂.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola crassulipes: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3697. ♂ ” [white, handwritten]; “Epsom. Feby.1912.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂. crassulipes” [white label, handwritten] .</p><p>Additional Material (n = 6: 2 males; 4 females). NEW ZEALAND: Auckland: 1♂, Omahuta SF, 18 III 1978, S.B. Peck, litter ; 1♂, Tauranganui Estate nr Kaiwhetu, 19 XI 1985, B.A. Holloway, litter 85/57a ; 1♀, Waipoua Forest, 7 XII 1961, G. Kuschel, litter 61/13 ; 1♀, Hen &amp; Chickens Is, Lady Alice, 1 I 1982, L. Roberts, litter 82/6 ; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=173.53334&amp;materialsCitation.latitude=-35.616665" title="Search Plazi for locations around (long 173.53334/lat -35.616665)">Waipoua SF</a>, Yakas Tr., 29 III 1999, R. Leschen, R. Hoare, G. Hall, at large, RL358, 35°37′S, 173°32′E ; 1♀, Mt. Huarua, 1200′, Tangihua Range, 25 XII 1936, E. Fairburn, A.E. Brookes Collection .</p><p>Diagnosis. This species is similar in appearance to other members of this species-group but can be distinguished by the horn on the ventral surface of the male head as long as wide and the shape of the antennae and genitalia.</p><p>Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown (Fig. 27b). Head: Transverse (Fig. 27b). Antennomeres 2–3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Eye small and prominent, one-third length of temple. Horn on ventral surface of male head triangular and as long as wide. Thorax: Prosternum as long as wide, widest at one-third length. Elytra approximately triangular (Fig. 27b). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe divided, major lobe rectangular, longer than minor lobe and parameres, minor lobe slender, as long as parameres (Fig. 27d). Phallobase of median lobe symmetrical and rounded (Fig. 27d). Parameres symmetrical, rectangular (Fig. 27d).</p><p>Type Locality. Woodhill, on the Kaipara Railway, near Helensville, Auckland, New Zealand .</p><p>Distribution. Auckland, Northland (Fig. 28: triangles).</p><p>Habitat. Most of specimens were collected by sifting leaf litter.</p><p>Comments. Specimens of S. sulcator can be distinguished externally from those of S. flavipes by the longer horn on the ventral surface of the male head and longer antennomeres. The type specimen of S. crassulipes shares these diagnostic characters with S. sulcator, and so has been placed as a junior synonym.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5A11783CBFFBD5FC0DFF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5D11793F68FEEFFDABFD33.text	03D82A3EBD5D11793F68FEEFFDABFD33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola helmorei	<div><p>10. helmorei species-group</p><p>(2 species)</p><p>Diagnosis. Members of the helmorei speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.3–2.9 mm; head subquadrate with dense setae around frontal sulcus (Fig. 29a–b); antennomere 1 approximately 2.5 times longer than wide with dull surface; anterior and posterior frontal foveae round (Fig. 29e); frontal sulcus reaching midpoint of eye (Fig. 29e); hind wings welldeveloped; legs elongate and slender (Fig. 29a–b); abdominal tergites IV–VI with discal carinae; genitalia small, not exceeding 0.25 mm (Fig. 29c–d); present on North Island, not known from South Island (Fig. 30).</p><p>KEY TO SPECIES OF THE HELMOREI SPECIES- GROUP</p><p>The key is mainly based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Ventral surface of temple normal (Fig. 29f); antennomeres 6–8 longer than wide; left paramere divided for one-third length (Fig. 29c) ...................................... S. helmorei new species</p><p>1′. Ventral surface of temple depressed and flat; antennomeres 6–8 subquadrate; left paramere more deeply divided for two-thirds length (Fig. 29d) ....... S. wellingtonensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD5D11793F68FEEFFDABFD33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5C11793C93FD13FB43FCCF.text	03D82A3EBD5C11793C93FD13FB43FCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola helmorei Park and Carlton	<div><p>Sagola helmorei Park and Carlton, new species</p><p>(Figs. 29a, 29c, 29e–f, 30)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND SH 12, Waipoua SF 20 Sep 1977 D.W. Helmore ”, “Rotten stumps/trees 77/102”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola helmorei Park and Carlton 2013 ” . Paratypes (11 males). NEW ZEALAND: Auckland: 2♂♂ (FMNH), Waitakere Ra., Cascadekauri Park, Upper Kauri tr., 170m, 8 XII 1984 – 25 I 1985, kauri-podo-hdwd, A. Newton, M. Thayer 680, FIT &amp; window trap ; 1♂ (FMNH), Waitakere Range, 260m, Nohoanga Scenic Res., 8 XII 1984 – 25 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 679, FIT &amp; window trap ; Northland: 2♂♂ (slide-mounted; FMNH), Mangamuka Summit, 12km nw Mangamuka, 400m, 25 XI–5 XII 1984, hdwd-podocarp forest, A. Newton, M. Thayer 683, FIT &amp; window trap ; 1♂ (DSC), Waipoua SF, 0.9km e forest hqtrs, 120m, 26 XI– 4 XII 1984, hdwd-podocarp forest, A. Newton, M. Thayer 686, berl., leaf &amp; log litter, forest floor ; 1♂ (FMNH), Waipoua SF, Wairau Summit, 400m, 27 XI–6 XII 1984, hdwd-podocarp forest, A. Newton, M. Thayer 687, FIT &amp; window trap ; 1♂ (FMNH), Waipoua SF, 0.8km nw Wairau Summit, 350m, 27 XI–6 XII 1984, hdwd-podocarp forest, A. Newton, M. Thayer 689, FIT&amp; window trap ; 1♂ (NZAC), Puketona, 20 I 1972, G.W. Ramsay, litter 72/49 ; 1♂ (NZAC), Pahia, 20 I 1972, G.W. Ramsay, litter 72/44 ; 1♂ (FMNH), Waipoua SF, Te Matua Ngahere., 16–17 III 1978, 400m, S. &amp; J. Peck, Kauri forest .</p><p>Etymology. This species is named for the collector of the holotype, Desmond W. Helmore.</p><p>Diagnosis. This species can be distinguished by the normal temple of the male, longer male antennomeres 6–8, and the left paramere divided for one-third its length.</p><p>Description of Male. L e n g t h 2.3– 2.8 m m. Body brown, antennae, legs, and maxillary palpi paler (Fig.). Head: Antennomere 2 longer than wide, 3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Eye small and prominent, one-third length of temple (Fig. 29e). Thorax: Prosternum slightly longer than wide, widest at one-third length. Elytra rectangular (Fig. 29a). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe broad and blunt, shorter than parameres (Fig. 29c). Phallobase of median lobe asymmetrical and rectangular (Fig. 29c). Parameres asymmetrical and divided, with one seta apically, left paramere broader than right (Fig. 29c).</p><p>Distribution. Auckland, Northland (Fig. 30: black circles).</p><p>Habitat. Most specimens of this species were collected using flight intercept traps or by sifting leaf and log litter in hardwood and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5C11793C93FD13FB43FCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5F117A3F7DFE74FB7FFAF2.text	03D82A3EBD5F117A3F7DFE74FB7FFAF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola rustica Broun 1915	<div><p>11. rustica species-group</p><p>(4 species)</p><p>Diagnosis. Members of rustica species-group can be distinguished from other Sagola speciesgroups by the following combination of characters: body length 2.3–2.9 mm; antennomere 1 approximately 1.5–2.0 times longer than wide (Fig. 31i); ventral surface of male head convex with setose gular region; anterior frontal fovea oval (Fig. 31j), posterior frontal fovea elongate (Fig. 31j); hind wings well-developed; abdominal tergites IV–VI with discal carina; parameres long, triangular, and broader than the median lobe (Fig. 31e–h).</p><p>KEY TO SPECIES OF THE RUSTICA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Antennomeres 4–7 elongate; eye as long as temple (Fig. 31b).... S. duplicata Broun</p><p>1′. Antennomeres 4–7 slightly longer than wide or subquadrate; eye one-half length of temple..................................................2</p><p>2(1′). Apical major lobe of genitalia bluntly triangular (Fig. 31g) ..................................... ........... S. marlboroughensis new species</p><p>2′. Apical major lobe of genitalia round.....3</p><p>3(2′). Larger genitalia, ca. 0.35 mm; minor lobe of genitalia longer than major lobe (Fig. 31e) ........................................... S. rustica Broun</p><p>3′. Smaller genitalia, ca. 0.25 mm; minor lobe of genitalia as long as major lobe (Fig. 31h) ............................ S. walkerae new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD5F117A3F7DFE74FB7FFAF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5F117A3C8EFF31FCBAFEC6.text	03D82A3EBD5F117A3C8EFF31FCBAFEC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola wellingtonensis Park and Carlton	<div><p>Sagola wellingtonensis Park and Carlton, new species</p><p>(Figs. 29b, 29d, 30)</p><p>Type Material. Holotype. NEW ZEALAND: Wellington: 1♂ (NZAC), “ NEW ZEALAND, WN Orongorongo V,W.N. 1 Jan 1993 ”, “Hard beech forest litter Tullgren #13”, “ Sagola sp. 2 det. Klimaszewski 1996 ”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola wellingtonensis Park and Carlton 2013 ” . Paratypes (n = 13: 7 males; 6 females). NEW ZEALAND: Auckland: 1♂ (NZAC), Wellsford, 29 IX 1958, R.A. Cumber; Bay of Plenty: 1♀ (FMNH), Tapapa Tukorehe Res., 300m, 25 III 1978, S. &amp; J. Peck, berl., litter; Coromandel : 3♂♂ 2♀♀ (NZAC), Kaitohe, 1334, 1 II 1952, R. Hornabrook ; 1♂ (AMNZ), Great Barrier I, Little Windy Hill, 100m, 19 XII 2002 – 23 I 2003, K. Parsons, coastal forest, malaise trap L11899 ; 1♂ (AMNZ), Great Barrier I, Mt. Hobson, 500m, pitfall trap, 9 XI–17 XII 2003, J.W. Early ; Waikato: 1♀ (FMNH), Pirongia Forest Park, Tirohanga tr. (above end Corcoran rd.), 500m, 37″58.409′S 175″07.173′E, 20 XI 2005, broadleaf forest, FMHD#2005-015, berl., forest litter, D. Clarke, A. Solodovnikov, ANMT site 1146 ; 1♀ (NZAC), Waitomo, 24 V 1983, G. Kuschel, decayed wood and litter 83/56 ; Wellington: 1♂ (slide-mounted; FMNH), Taraua Forest Park, Waitewaewae Tr., 220m, 40″51.98′S 175″15.319′E, 26 XI–21 XII 2005, broadleaf (much Knightia exelsa)-podocarp forest, FMHD#2005-034, FIT &amp; window trap, ANMT site 1152 ; 1♀ (NZAC), Rimutaka Range, 11 VIII 1954, R. Hornabrook .</p><p>Etymology. The species name is based on the biotic region of some of the type material, Wellington.</p><p>Diagnosis. This species can be distinguished by the depressed and flattened temple of the ventral surface of the male head, subquadrate male antennomeres 4–8, and left paramere divided for two-thirds its length.</p><p>Description. Length 2.4–2.9 mm. Body brown, antennae, legs, and maxillary palpi paler (Fig. 29b). Head: Male antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Female antennomeres 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Eye small and prominent, one-third length of temple. Temple of male head ventrally depressed and flat. Thorax: Prosternum slightly longer than wide, widest at one-third length. Elytra rectangular (Fig. 29b). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe broad and blunt, shorter than parameres (Fig. 29d). Phallobase of median lobe asymmetrical and rectangular (Fig. 29d). Parameres asymmetrical and divided, with 1 seta apically, left paramere deeply divided (Fig. 29d).</p><p>Distribution. A u c k l a n d, B a y o f P l e n t y, Coromandel, Waikato, Wellington (Fig.30: triangles).</p><p>Habitat. Most specimens were collected using pitfall, Malaise, window, and flight intercept traps or by sifting leaf litter in broadleaf and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5F117A3C8EFF31FCBAFEC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5F11753F6EFA60FEA8FD8C.text	03D82A3EBD5F11753F6EFA60FEA8FD8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola rustica Broun 1915	<div><p>Sagola rustica Broun, 1915</p><p>(Figs. 31a, 31e, 32)</p><p>Sagola rustica Broun, 1915: 285 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola subcuneata Broun, 1921a: 488 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.</p><p>Sagola brevifossa Broun, 1921a: 501 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3700.♀.” [white label, handwritten]; “Rakaia. 18_5_1912.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♀. rustica.” [white label, handwritten]. Holotype of Sagola brevifossa: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “4014.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Routeburn. 16.2.1914.” [white label, handwritten]; “ Sagola ♂. brevifossa.” [white label, handwritten]. Holotype of Sagola subcuneata: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “3995.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Sec x?” [white label, handwritten]; “Moa Basin . 20.10.1913.” [white label, handwritten]; “ Sagola ♂. subcuneata.” [white label, handwritten] .</p><p>Additional Material (n = 22: 5 males; 17 females). NEW ZEALAND: Fiordland: 1♂ 2♀♀, Secretary I, Gut Bay, 23 XI 1981, C.F. Butcher, sifted litter around hut 81/177; 1♂, Breaksea Sound, Gilbert I No 6, 12 III 1983, C.F. Butcher, sifted rotten wood 83/43; 1♂, South Arm, L. Manapouri, 15 XII 1998, J. Nunn, underside log with white rot; 1♀, South Arm, L. Manapouri, 13 XII 1998, J. Nunn, in moss; 1♂ 1♀, Fiordland NP, Kepler tr. below Mt. Luxmore Hut, 900m, 45°24.865′S, 167°38.675′E, 11 XII 2005, Nothofagus forest, pyt. fogging mossy log, A. Solodovnikov, D. Clarke, ANMT site 1176; 1♂, Murchison Mts, lower east McKenzie Burn, 945m, 8 XII 1983, C.A. Muir, litter in N. menziesii forest; 2♀♀, Fiordland NP, Milford Sound rd., Smithy Creek Campground area, 400m, 44°57.065′S, 168°01.156′E, 9–11 XII 2005, Nothofagus fusca &amp; N. menziesii open forest, pyt. fogging old mossy &amp; fungusy logs, A. Newton, ANMT site 1170; 1♀, Fiordland NP, Milford Sound rd., Gertrude Saddle car park area, 800m, 44″46.13′S, 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD #2005-087, berl., leaf &amp; log litter, A. Newton, M. Thayer, ANMT site 1169; 1♀, Fiordland NP, Borland rd., Mt. Burns Tops tr. above Borland Saddle, 1075m, 45″44.883′S, 167″22.938′E, 10 XII 2005, subalpine tussockland, FMHD #2005-096, berl., unsifted at bases of shrubs ( Dracophyllum, possibly Fillfolium), M. Thayer, ANMT site 1173; 2♀♀, Eglinton Valley Divide, 400m, 13 II 1982, C.F. Butcher, moss/liverworts 82/29; 2♀♀, Mt. Annetta Sdle, Little Red Hill, Barrier R., 1433m, 4 II 1975, litter alpine swards 75/43; 1♀, Simonin Pass, W. Olivine Ra., 1067m, 23 I 1975, G.W. Ramsay, litter in Nothofagus forest 75/37; Westland: 4♀♀, Mt. Aspiring NP, Arawata Biv, 840m, 5 II 1989, LCNZ 89/2, J.W. Early, R.M. Emberson, scrub litter.</p><p>Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: larger body, length 2.6–2.8 mm; ventral surface of male head convex with setose gular region; eye large and prominent, approximately one-half length of temple; frontal sulcus deep and reverse keyholeshaped; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.6–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 31a). Head: Head round, widest across eyes (Fig. 31a). Ventral surface of male head convex with setose gular region. Male antennomere 1 approximately 2 times longer than wide, 2 subquadrate, 3–8 longer than wide, 9–10 subquadrate. Female antennomere longer than male, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 31a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, less developed in female. Aedeagus: Median lobe divided, minor lobe longer with tubercles (Fig. 31e). Phallobase of median lobe symmetrical and rounded (Fig. 31e). Parameres symmetrical with sparse setae from apex to midpoint, broader than median lobe (Fig. 31e).</p><p>Type Locality. Rakaia Gorge, Mid Canterbury, New Zealand .</p><p>Distribution. Fiordland, Mid Canterbury, Otago Lakes, Westland (Fig. 32: black circles).</p><p>Habitat. Most specimens of this species were collected by fogging or sifting mossy leaf or log litter in Nothofagus forests.</p><p>Comments. Specimens of S. rustica can be distinguished from other species by the shape and size of the antennomeres, frontal fovea, frontal sulcus, size of eye, and unique genitalia. The type specimens of S. brevifossa and S. subcuneata share these diagnostic characters, and these species have been collected at or near the type locality of S. rustica . For these reasons, we have placed S. brevifossa and S. subcuneata in synonymy with S. rustica .</p></div>	https://treatment.plazi.org/id/03D82A3EBD5F11753F6EFA60FEA8FD8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5011753CB1FDB5FC67FC85.text	03D82A3EBD5011753CB1FDB5FC67FC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola duplicata Broun 1886	<div><p>Sagola duplicata Broun, 1886</p><p>(Figs. 31b, 31f, 32)</p><p>Sagola duplicata Broun, 1886: 888 . Raffray 1893: 40, 1904: 498, 1911: 5, 1923: 233; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1581. ♀.” [white label, handwritten]; “Parua” [white label, Printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola duplicata ” [white label, handwritten].</p><p>Additional Material (n = 7: 3 males; 4 females). NEW ZEALAND: Bay of Plenty: 2♀♀ (1♀, slide-mounted), Waiaroho, 17 IX–21 X 1992, G. Hall, Malaise trap; Coromandel : 1♀, Great Barrier I, Little Windy Hill, 220m, 22 V –7 VII 2002, P. Sutton, forest edge malaise trap ; Gisborne: 1♂, Taikawakawa, 2 II–18 III 1993, J.S. Dugdale, Malaise trap ; Marlborough Sounds: 1♂, Tennyson Inlet, east side Ducan Bay, 30m, 15 XII 1984 – 5 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, FIT &amp; window trap ; Southland: 1♂, Blue Mountains north end, 9 VIII 2003, J. Nunn, in compact moss cushion on beech bole at treeline ; Waikato: 1♀, Kaimai Range, Matamata, 1520′, 1 III 1948, A.E. Brookes .</p><p>Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: body length 2.7–2.9 mm; ventral surface of male head convex; eye large and prominent, as long as temple; longer antennomeres 4–7; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.7–2.9 mm. Body reddish brown, antenna, elytra, legs, and maxillary palpi paler (Fig. 31b). Head: Male head round, widest across eyes (Fig. 31b), convex ventrally. Male antennomere 1 approximately 1.5 times longer than wide, 2 subquadrate, 3 longer than wide, 4–7 elongate, 8–9 longer than wide, 10 subquadrate. Female antennomeres 2–3 subquadrate, 4 elongate, 5–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, as long as temple, female slightly smaller. Thorax: Prosternum as long as wide, widest at one-third. Elytra rectangular (Fig. 31b). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of oval or transverse patches of microtrichia. Aedeagus: Median lobe elongate triangular, as long as parameres, apical lobe with weak tubercles (Fig. 31f). Phallobase of median lobe symmetrical and rounded (Fig. 31f). Parameres symmetrical, broad with setae apically (Fig. 31f).</p><p>Type Locality. Parua, near Whangarei Harbour, Northland, New Zealand .</p><p>Distribution. Bay of Plenty, Coromandel, Gisborne, Marlborough Sounds, Northland, Southland, Waikato (Fig. 32: triangles).</p><p>Habitat. Specimens of this species were collected using Malaise traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5011753CB1FDB5FC67FC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5011763EFBFCB9FDD0FE7B.text	03D82A3EBD5011763EFBFCB9FDD0FE7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola marlboroughensis Park and Carlton	<div><p>Sagola marlboroughensis Park and Carlton, new species</p><p>(Figs. 31c, 31g, 32)</p><p>Type Material. Holotype. NEW ZEALAND: Marlborough: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: MB: Richmond Ra, 1600m Johnson Pk., 13 III 1969 J.S. Dugdale, swards 69/96”, “ HOLOTYPE Sagola marlboroughensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the holotype, Marlborough.</p><p>Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: body length 2.4 mm; ventral surface of male head transversely convex with setose gular region; eye large and prominent, approximately one-half length of temple; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.4 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 31c). Head: As long as wide, widest across eyes (Fig. 31c), ventrally transversely convex with setose gular region. Antennomere 1 approximately 2 times longer than wide with dull surface, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching hind point of eye. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large, approximately one-half length of temple. Thorax: Prosternum longer than wide, widest at one-third length. Elytra rectangular (Fig. 31c). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia. Aedeagus: Median lobe divided, major apical lobe broader (Fig. 31g). Phallobase of median lobe symmetrical and rounded (Fig. 31g). Parameres symmetrical, broad with setae from apex to midpoint (Fig. 31g).</p><p>Distribution. Marlborough (Fig. 32: black square).</p><p>Habitat. The holotype of this species was collected in a sward (grassy meadow).</p></div>	https://treatment.plazi.org/id/03D82A3EBD5011763EFBFCB9FDD0FE7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5311773F75F9C1FEE8FEED.text	03D82A3EBD5311773F75F9C1FEE8FEED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola lescheni	<div><p>12. lescheni species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the lescheni species-group can be distinguished from other Sagola species-groups by the following combination of characters: small body, length 1.7–1.9 mm; antennomere 1 approximately 2 times longer than wide; frontal rostrum prominent (Fig. 33a); hind wings well-developed; abdominal tergites IV–VI with discal carinae; broad and flattened genitalia (Fig. 33b); only known from Northland of North Island (Fig. 36).</p></div>	https://treatment.plazi.org/id/03D82A3EBD5311773F75F9C1FEE8FEED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5311763CE8FDF8FBB5F99D.text	03D82A3EBD5311763CE8FDF8FBB5F99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola walkerae Park and Carlton	<div><p>Sagola walkerae Park and Carlton, new species</p><p>(Figs. 31d, 31h–j, 32)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND NN Beebys Knob 1220m 5 Feb 1978 A.K.Walker ”, “Moss and rotten wood 78/76”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola walkerae Park and Carlton 2013 ” . Paratypes (n = 56: 35 males; 21 females). NEW ZEALAND: Nelson: 3♂♂ 7♀♀ (FMNH), Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD#2005-048, berl., dead wood, D. Clarke, ANMT site 1157 ; 1♂ 2♀♀ (FMNH), Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD#2005-049, berl., leaf litter, A. Solodovnikov, D. Clarke, ANMT site 1157 ; 6♂♂ 3♀♀ (FMNH), Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S, 172″44.456′E, 28 XI 2005, Nothofagus - dominant forest, pyr.-fogging mossy old logs, A. Newton, M. Thayer, ANMT site 1156 ; 4♂♂ 1♀ (FMNH), Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S, 172″44.456′E, 28 XI 2005, Nothofagus -dominant forest, berl., leaf &amp; log litter, A. Newton, M. Thayer, ANMT site 1156 ; 4♂♂ 2♀♀ (FMNH), Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, pyrethrin fogging fungusy Nothofagus log; Buller: 2♂♂ 1♀ (1♂, slide-mounted; FMNH), Nelson Lks. NP, n slope Mt. Robert, Speargrass tr., 880m, 21 XII 1984, Nothofagus forest, A. Newton, M. Thayer 704, Pyrethrin fogging fungusy Nothofagus logs ; 6♂♂ 4♀♀ (FMNH &amp; DSC), Nelson Lks . NP, n slope Mt. Robert, Pinchgut tr., 1290m, 18–26 XII 1984, Notho. sol. elfin forest, A. Newton, M. Thayer 716, Pyrethrin fogging fungusy Nothofagus logs ; 5♂♂ (FMNH &amp; DSC), Nelson Lks. NP, n slope Mt. Robert, Pinchgut tr., 950m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, window trap ; 1♂ (FMNH), Nelson Lks. NP, Lake Rotoiti, St. Arnaud tr., 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp; window trap ; 1♂ (DSC), Lewis Pass Nat. Res., 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, berl., leaf &amp; log litter, forest floor (DSC) ; 1♂, Nelson Lks. NP, Robert Ridge, nr. Julius Summit, ca. 1700m, 41″52′S 172″46.5′E, 1 XII 2005, FMHD#2005-063, berl., debris at Bases of tussocks, A. Solodovnikov ; 1♂ (NZAC), Paparoa Ra., 1036, Mt. Dewar, 12 XII 1969, J.C. Watt, moss 69/260 ; 1♀ (NZAC), Reefton, 12 IV 1977, J.A. Wightman, pit trap cutover pine .</p><p>Etymology. This species is named for the collector of the holotype, Annette K. Walker.</p><p>Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: body length 2.3–2.5 mm; ventral surface of male head transversely convex with setose gular region; eye large and prominent, approximately one-half length of temple; antennomeres 3–10 subquadrate; shape of genitalia unique to species.</p><p>Description. Length 2.3–2.5 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 31d). Head: Bluntly triangular, widest across eyes (Fig. 31j), ventral surface of male head transversely convex with setose gular region. Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3–10 subquadrate (Fig. 31i). Frontal rostrum prominent, covering anterior frontal fovea. Frontal sulcus deep, reaching hind point of eye from end of frontal rostrum. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum longer than wide, widest at one-third length. Elytra rectangular (Fig. 31d). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, less developed in female. Abdominal ventrites III–V with very small basolateral fovea. Aedeagus: Median lobe divided, major lobe as wide as minor (Fig. 31h). Phallobase of median lobe symmetrical and rounded (Fig. 31h). Parameres symmetrical, broad with setae apically (Fig. 31h).</p><p>Distribution. Buller, Nelson (Fig. 32: stars).</p><p>Habitat. Most specimens of this species were collected at high elevations (&gt; 670 m) by fogging, with window and flight intercept traps, or by sifting leaf litter in Nothofagus mossy forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5311763CE8FDF8FBB5F99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5211773F5FFC96FBD7FB0E.text	03D82A3EBD5211773F5FFC96FBD7FB0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>13. gourlayi species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the gourlayi species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.3 mm (Fig. 34a); antennomere 1 approximately 2 times longer than wide; anterior and posterior frontal fovea oval, anterior frontal fovea open anteriorly (Fig. 34c); hind wings well-developed; only known from Nelson on South Island (Fig. 36).</p></div>	https://treatment.plazi.org/id/03D82A3EBD5211773F5FFC96FBD7FB0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5211773CDFFE6AFC8CFCA0.text	03D82A3EBD5211773CDFFE6AFC8CFCA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola lescheni Park and Carlton	<div><p>Sagola lescheni Park and Carlton, new species</p><p>(Figs. 33, 36)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), “ NEW ZEALAND: ND Waipoua State Forest, 0.9km e Forest Hqtrs. 120m, 26.xi-4.xii.1984 hdwd.-podocarp forest A. Newton / M. Thayer 686”, “flight intercept (window) trap”, “ HOLOTYPE Sagola lescheni Park and Carlton 2013 ” . Paratypes (2 males). NEW ZEALAND: Northland: 1♂ (FMNH), same data as holotype; 1♂ (slide-mounted; FMNH), Omahuta SF, Kauri Sanct., 18 III 1978, S. &amp; J. Peck, 300m, Rangiahua, berl., forest litter .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study and world renowned beetle specialist and musician, Richard “Blind Beetle” Leschen.</p><p>Diagnosis. This species can be distinguished from other species by the following combination of characters: small body 1.7–1.9 mm; frontal rostrum prominent; ventral surface of head weakly convex; eye large and prominent, approximately one-half length of temple; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. L e n g t h 1.7– 1.9 m m. Body brown, antennae and maxillary palpi paler, elytra and legs paler (Fig. 33a). Head: As long as wide, widest across eyes (Fig. 33a), ventral surface weakly convex with setose gular region. Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum prominent. Frontal sulcus reaching midpoint of eye. Anterior frontal fovea oval, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 33a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe broader and longer than paramere (Fig. 33b). Phallobase of median lobe symmetrical and rounded (Fig. 33b). Parameres symmetrical, elongate-triangular with setae apically (Fig. 33b).</p><p>Distribution. Northland (Fig. 36: black circles).</p><p>Habitat. Specimens of this species were collected using window and flight intercept traps or by sifting forest litter in hardwood and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5211773CDFFE6AFC8CFCA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5211703EEAFB4DFE62F9B0.text	03D82A3EBD5211703EEAFB4DFE62F9B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola gourlayi Park and Carlton	<div><p>Sagola gourlayi Park and Carlton, new species</p><p>(Figs. 34, 36)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “Paradise Peak 4500′ 6- 11-38, E. S. Gourlay ”, “E.S. Gourlay Acc. 1970 Ent. Div.”, “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “HOLOTYPE Sagola gourlayi Park and Carlton 2013 ”.</p><p>Etymology. This species is named for the collector of the holotype, Edward S. Gourlay.</p><p>Diagnosis. This species can be distinguished from other species by the following combination of characters: body length 2.3 mm; head transverse; eye large and prominent, one-half length of temple; neck bearing dense setae anteroventrally; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.3 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 34a). Head: Transverse, widest across eyes (Fig. 34c). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye. Anterior and posterior frontal foveae oval, anterior frontal fovea open anteriorly. Eye large and prominent, one-half length of temple. Anterior ventral neck bearing dense setae. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 34a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided, apical major lobe elongate and umbrella-shaped (Fig. 34b). Phallobase of median lobe symmetrical and rounded (Fig. 34b). Parameres symmetrical, elongately triangular with setae at apex (Fig. 34b).</p><p>Distribution. Nelson (Fig. 36: triangles).</p><p>Habitat. Unknown.</p><p>Comments. The holotype suffered crushing damage to the frons and mouthparts, so the described shape of the frontal sulcus and fovea may differ from the original condition.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5211703EEAFB4DFE62F9B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5511703D66F9BEFCAFFBFC.text	03D82A3EBD5511703D66F9BEFCAFFBFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sharpi Raffray 1893	<div><p>14. sharpi species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the sharpi species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.3–2.6 mm (Fig. 35a); antennomere 1 approximately 1.5 times longer than wide; eye large and prominent, approximately two-thirds length of temple (Fig. 35c); ventral surface of male head distinctly convex with heart-shaped depression containing a pair of diagonal processes (Fig. 35d); hind wings well-developed; abdominal tergites IV–VI with discal carinae.</p></div>	https://treatment.plazi.org/id/03D82A3EBD5511703D66F9BEFCAFFBFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD5511733F62FB7DFC6FFEEC.text	03D82A3EBD5511733F62FB7DFC6FFEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sharpi Raffray 1893	<div><p>Sagola sharpi Raffray, 1893</p><p>(Figs. 35–36)</p><p>Sagola sharpi Raffray, 1893: 26 . Raffray 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola grata Broun, 1912b: 628 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola clavatella Broun, 1912b: 631 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola posticalis Broun, 1915: 291 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Syntypes. NEW ZEALAND: 1♀ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola sharpi Type Raffray.” [white label, handwritten]. 2♂♂ (MNHN), glued on rectangular card, " N. Zeal MP Type Sharpi AR ” Holotype of Sagola clavatella: NEW ZEALAND: Buller: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “19.–” [white label, handwritten]; “Greymouth, New Zealand Helms.” [white label, printed]; “Sharp Coll. 1905-313” [white label, printed]; “ Sagola ♂. clavatella.” [white label, handwritten]. Holotype of Sagola grata: NEW ZEALAND: Marlborough Sounds: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “18_” [white label, handwritten]; “Sharp Coll. 1905-313” [white label, printed]; “Picton, New Zealand Helms.” [white label, printed]; “ Sagola grata . ♂.” [white label, handwritten]. Syntypes of Sagola posticalis: NEW ZEALAND: Kaikoura: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “3707. ♂ ” [white label, handwritten]; “Wairiri. Kaikoura” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂. posticalis” [white label, handwritten]. ♀ (BMNH), glued on rectangular card, “3707. ♀ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Wairiri. Kaikoura” [white label, handwritten]; “ Sagola . posticalis” [white label, handwritten] .</p><p>Additional Material (n = 70: 47 males; 23 females). NEW ZEALAND: Buller: 1♂, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, FIT &amp;window trap; 1♀, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, pyrethrin fogging fungusy log ; 1♂, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &amp;window trap; 1♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, pyrethrin fogging fungusy log ; 1♀, Nelson Lks NP, n slope Mt. Robert, Speargrass tr., 880m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 704, window trap ; 1♀, Nelson Lks NP, n slope Mt. Robert, Pinchgut Tr., 1290m, 18–26 XII 1984, Noth. sol. elfin forest, A. Newton, M. Thayer 716, FIT &amp;window trap; 1♀, Nelson Lks NP, n slope Mt. Robert, 860m, 23–26 III 1980, Nothofagus forest, A. Newton, M. Thayer, pyrethrin fogging fungusy log ; 1♂, Shenandoah Saddle, 700m, 10 I 1982, R . M. Emberson, moss by stream LCZN 82 /5; 1♂, Punakaiki, Pororari R Valley tr., 80m, 2–4 V 2007, J.W. Early, R . F. Gilbert, Nothofagus -podocarp forest, yellow pan trap L15869 ; 1♂, Nelson Lks NP, Lake Rotoiti, St. Arnaud tr., 645m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 705, window trap ; Coromandel: 1♂, Kaitohe, 12 I 1952, R . Hornabrook; Gisborne : 1♂, Urewera NP, Waikaremoana Rd., s end Matanunui Ridge, 720m, 38″44.404′S 177″05.806′E, 22 XI–23 XII 2005, mixed broadlf (incl. Nothofagus fusca)- podocarp, FMHD #2005-028, FIT , M. Thayer, A. Solodovnikov, ANMT site 1149; Dunedin: 1♂, Government tr., Waipori Valley, 13 XII 2003 – 6 II 2004, J. Nunn, FIT ; Marlborough: 2♂♂, Pelorus Bridge Scenic Reserve, 35m, 41″18.3′S 173″34′E, 27 XI 2005, mixed broadlf (incl. Nothofagus)-podocarp forest, pyr-fogging old fungusy logs, A. Newton, D. Clarke, ANMT site 1155; North Canterbury: 1♀, 2.9km N Arthur’ s Pass Bealey Valley tr., 840–950m, 42′55S 171′33E, #40, under bark, 11 I 1998, C. Carlton, R . Leschen; Nelson : 1♂, Kahurangi NP, Cobb Dam rd., Asbestos tr., 450m, 41″06.333′S 172″43.174′E, 29 XI 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest, pyr-fogging mossy/fungusy logs, A. Newton, M. Thayer, ANMT site 1160; 1♂, Kahurangi NP, Cobb Dam rd., Asbestos tr., 450m, 41″06.333′S 172″43.174′E, 29 XI 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest, FMHD #2005-056, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1160 ; 1♂, Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′ S 172″44.456′E, 28 XI 2005, Nothofagus dominant forest, under bark of small logs, A. Newton, M. Thayer, ANMT site 1156; 1♂, Slaters Rd., 0.7km S Whangamoa Saddle, 410m, 13 XII 1984 – 4 I 1985, Nothofagus forest, A. Newton, M. Thayer 703, window trap ; 1♂, Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S 172″44.456′E, 28 XI 2005, Nothofagus dominant forest, FMHD #2005-044, FIT , A. Newton, M. Thayer, ANMT site 1156; 1♂, Hope Saddle, 25 I 1957, E.S. Gourlay (NZAC) ; 1♂, Cobb rd., summit, 29 XI 2005, J. Nunn, under tight bark of beech log ; 1♀, Slaters rd., 0.7km s Whangamoa Saddle, 410m, 13 XII 1984 – 4 I 1985, Nothofagus forest, A. Newton, M. Thayer 703, FIT &amp;window trap; 1♀, 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &amp; window trap; Otago Lakes: 1♂ 1♀, Paradise, 2 II 1984, J.C. Watt, sifted wood mould 84/9 ; 1♂, 14km nw Glenorchy, beyond Paradise, Mt. Aspiring NP, 44′41S 168′20E, #103, Nothofagus leaf litter berlese, 21 I 1998, C. Carlton, R . Leschen; 1♂, Mt. Aspiring NP, 5.5km nne Makarora, 330m, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 741, under bark rotting log; Marlborough Sounds: 3♂♂, Tennyson Inlet, west side Duncan Bay, 30m, 15 XII 1984 – 5 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, FIT &amp;window; 1♀, Tennyson Inlet, west side Duncan Bay, 30m, 15 XII 1984 – 5 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, pyrethrin fogging fungusy log ; 1♂, Tennyson Inlet, west side Te Mako Bay, 125m, 15 XII 1984 – 5 I 1985, Nothofagus -podohdwd, A. Newton, M. Thayer 710, berl., leaf &amp; log litter, forest floor ; 1♂, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 - 5 I 1985, Nothofagus forest, A. Newton, M. Thayer 708, window trap ; 1♂, Opouri Saddle, 28 I 1979, L.A. Mound, wet moss on roadside bank ; 1♂, Picton, IX 1969, G. Kuschel ; Stewart Island: 1♂, Golden Bay, 10 I 1996, J. Nunn, in dead Rimu wood; Southland : 1♂, Catlins Forest Park, Table Hill Scenic Res., nne Papatowai, 180m, 46″30.05′S 169″30.06′E, 13 XII 2005, Nothofagus menziesii, FMHD #2005-106, berl., leaf &amp; log litter, A. Newton, M. Thayer, et al., ANMT site 1179; Wairarapa: 1♂, upper Manawatu Gorge, 24 X 1993, J. Nunn ; 3♂♂ 3♀♀, Klein Tr., Manawatu Gorge, 16 VII 1994, J. Nunn, in much decayed wood ; 3♀♀, Hikurangi, 13 IX 1982, J.C. Watt, sifted woodmould 82/76 ; Westland: 2♂♂, Lake Kaniere rd., 2.8km nw Lake Kaniere, 120m, 8–19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 732, FIT &amp;window trap; 2♂♂ (1♂, slidemounted), Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 731, FIT &amp;window trap; 2♂♂ 1♀, 1.8km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, FIT &amp;window trap; 1♀, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap ; 1♀, Hokitika Lake Mahinapua Res., 28 I 1978, S. &amp; J. Peck, bracket fungi berl; Wellington : 1♂ 1♀, Wilton’ s <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=175.75266&amp;materialsCitation.latitude=-41.26605" title="Search Plazi for locations around (long 175.75266/lat -41.26605)">Bush</a>, 110m, 41°15.963′S 175°45.159′E, 24 XI 2005, mixed broadleaf-podocarp forest, FMHD #2005- 030 , berl., leaf &amp; log litter, M. Thayer, A. Newton, ANMT site 1150; 1♂, Tararua Ra., Dundas Hut Ridge, 750m, 11 II 1985, C.F. Butcher sifted rotten wood 85/31 ; 1♂, Nikau Preserve, Paraparaumu, 23 VII 1995, in kohekohe with white rot, on forest floor ; 1♂, Mana Island, 5–7 XI 1993, J. Nunn, in decayed wood ; 1♂, Nikau Reserve, Paraparaumu, 9 I 1996, in humfied log on forest floor ; 1♂, Kaitoke Regional Park, Pakuratahi Forks, 15 IV 2005, R. Leschen, C. McGuiness, leaf litter, RL975, 41.03′S 175.11′E ; 2♂♂ 1♀, Pakurarai Forks, Kaitoke, 24 VII 1993, J. Nunn, in decayed wood ; 1♀, Nikau Res., Paraparaumu, 10 X 1993, J. Nunn ; 1♀, Wilton Bush, Wellington City, 24 XI 2005, in decayed wood .</p><p>Diagnosis. This species can be distinguished from other species by the following combination of characters: body length 2.3–2.6 mm; ventral surface of male head distinctly convex with heart-shaped depression containing pair of diagonal processes; eye large and prominent, approximately two-thirds length of temple; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.3–2.6 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 35a). Head: Round, widest across eyes (Fig. 35c). Ventral surface of male head distinctly convex with heart-shaped depression containing pair of diagonal processes (Fig. 35d). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep, reaching midpoint of eye (Fig. 35c). Anterior frontal fovea round (Fig. 35c), posterior frontal fovea oval (Fig. 35c). Eye large and prominent, approximately two-thirds length of temple (Fig. 35c). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 35a). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe simple and small, &lt;0.25 mm (Fig. 35b). Phallobase of median lobe symmetrical and rounded (Fig. 35b). Parameres symmetrical with setae apically (Fig. 35b).</p><p>Type Locality. New Zealand (specific locality unknown) .</p><p>Distribution. Buller, Coromandel, Dunedin, Gisborne, Fiordland, Marlborough, North Canterbury, Nelson, Otago Lakes, Marlborough Sounds, Stewart Island, Southland, Westland, Wairarapa, Wellington (Fig. 36: black squares).</p><p>Habitat. Most specimens of this species were collected using window and flight intercept traps, by fogging, or by sifting mossy leaf or decayed log litter in Nothofagus, podocarp, hardwood, or broadleaf forests.</p><p>Comments. The original description cited two male and one female specimens, but we were only able to find and examine one female. Nevertheless, specimens of S. sharpi can be distinguished from those of other species by the shape and size of the antennomeres, frontal foveae, frontal sulcus, and size of the eye. The type specimens of S. clavatella, S. grata, and S. posticalis share these diagnostic characters. For these reasons, we have placed S. clavatella, S. grata, and S. posticalis in synonymy with S. sharpi .</p></div>	https://treatment.plazi.org/id/03D82A3EBD5511733F62FB7DFC6FFEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD56110C3F59FE5BFD45FD81.text	03D82A3EBD56110C3F59FE5BFD45FD81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola auripila Broun 1911	<div><p>15. auripila species-group</p><p>(8 species)</p><p>Diagnosis. Members of the auripila speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body small, length 1.8–2.4 mm; antennomere 1 approximately 1.5–2.0 times longer than wide; frontal sulcus deep and long extending between posterior edge of eye and end of head (Fig. 38j–k); hind wings reduced to small pads; abdominal tergites IV–VI with discal carinae.</p><p>KEY TO SPECIES OF THE AURIPILA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Antennomere 1 approximately 2 times longer than wide, with dull surface; frontal sulcus reaching posterior margin of head...............................................................2</p><p>1′. Antennomere 1 approximately 1.5 times longer than wide; frontal sulcus not extending posterior to vertexal fovea..................3</p><p>2(1). Left paramere longer than wide and bent (Fig. 38g)............ S. gimmeli new species</p><p>2′. Right paramere longer than wide with round depression at base (Fig. 38i) .......... ........................... S. sokolovi new species</p><p>3(1′). Frontal sulcus distinctly wide (Fig. 38j: arrow) ................... S. spiniventris Broun</p><p>3′. Frontal sulcus narrow (Fig. 38k: arrow) ............................................................ 4</p><p>4(3′). Antennomeres 5–10 weakly enlarged; hind femur with triangular process; only known from North Island........................5</p><p>4′. Antennomeres 5–10 simple; hind femur simple; known from Chatham Islands or Three Kings Islands .................................. 6</p><p>5(4). Lateral median lobe of genitalia rectangular with emarginate apical lobe; parameres C-shaped (Fig. 38a)..... S. auripila Broun</p><p>5′. Lateral median lobe of genitalia long curved with wave-shaped ventral margin; parameres weakly curved (Fig. 38e)......... ......................... S. chandleri new species</p><p>6(4′). Parameres asymmetrical and setae only apically (Fig. 38f); only known from Three Kings Islands (Fig. 40: black square) ........ .................. S. threekingsensis new species</p><p>6′. Parameres symmetrical and setae on middle; only known from Chatham Islands ........... 7</p><p>7(6′). Median lobe of genitalia S-shaped (Fig. 38c) ...................... S. chathamensis new species</p><p>7′. Median lobe of genitalia rectangular (Fig. 38d) ....... S. pittensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD56110C3F59FE5BFD45FD81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD29110E3D54FDB5FD9FFC28.text	03D82A3EBD29110E3D54FDB5FD9FFC28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola auripila Broun 1911	<div><p>Sagola auripila Broun, 1911</p><p>(Figs. 37a, 38a, 39)</p><p>Sagola auripila Broun, 1911: 500 . Hudson 1923: 365, 1934: 184; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. Holotype. NEW ZEALAND: Taupo: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3371.” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Erua. Feby.1911.” [white label, handwritten]; “ Sagola ♂ auripila.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female.</p><p>Additional Material (n = 5: 4 males; 1 female). NEW ZEALAND: Gisborne: 1♂, Taikawakawa, 300m, 1 V 1993, C. T. Duval, litter 93/106 ; Taupo: 1♂, Urewera NP, Ngamoko tr., nr. jct Tawa loop, berlese, 8 III 2000, C. Carlton, A. Weir, #027 ; 1♂, National Park, Taurewa Lookout, 13 XII 1961, G. Kuschel, litter 61/14 ; 1♂, Minginui SF, 27 VII 1977, J.S. Dugdale, rotten Podocarpus; Waikato: 1♀, Pirongia Forest Park, Mahaukura tr (above end Grey rd.), 270m, 37”58.218′S 175”06.523′E, 18 XI–27 XII 2005, mixed broadleaf forest, FMHD#2005-009, FIT, A. Newton, M. Thayer et al., ANMT site 1142 .</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: larger body, length 2.1–2.3 mm; weakly enlarged male antennomeres 5–10; frontal sulcus linear and deep, extending slightly posterior to eye; male hind femur with triangular process at middle of c) S. chathamensis, d) S. pittensis, e) S. chandleri, f) S. threekingsensis, g) S. gimmeli, h) S. sokolovi . Scale bars = 1 mm. mesal margin; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.1–2.3 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37a). Head: Round, widest across eyes (Fig. 37a). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 5–10 weakly enlarged. Female antennomere 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus linear and deep, slightly exceeding eye. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37a). Meso- and metathorax trapezoidal, as long as wide. Male hind femur with triangular process at middle of mesal margin. Aedeagus: Lateral median lobe rectangular and apical lobe weakly emarginate (Fig. 38a). Phallobase of median lobe symmetrical and rectangular (Fig. 38a). Parameres symmetrical and curved to C-shaped with setae apically and 1 seta at middle (Fig. 38a).</p><p>Type Locality. Erua, Taupo, New Zealand .</p><p>Distribution. Gisborne, Taupo, Waikato (Fig. 39: black circles).</p><p>Habitat. Most specimens of this species were collected using flight intercept traps or by sifting leaf litter in broadleaf or podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD29110E3D54FDB5FD9FFC28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2B110F3CBBFC27FE8FFC85.text	03D82A3EBD2B110F3CBBFC27FE8FFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola spiniventris Broun 1912	<div><p>Sagola spiniventris Broun, 1912</p><p>(Figs. 37b, 38b, 38j, 39)</p><p>Sagola spiniventris Broun, 1912b: 627 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Sagola occipitalis Broun, 1912b: 624 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Marlborough Sounds: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “17.” [white label, handwritten]; “Picton, New Zealand Helms.” [white label, printed]; “Sharp Coll. 1905-313” [white label, printed]; “ Sagola ♂ spiniventris.” [white label, handwritten] . Holotype of Sagola occipitalis: NEW ZEALAND: Buller: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “11.” [white label, handwritten]; “Greymouth, New Zealand. Helms.” [white label, printed]; “Sharp Coll. 1905-313” [white label, printed]; “ Sagola occipitalis .” [white label, handwritten] .</p><p>Additional Material (n = 15: 8 males; 7 females). NEW ZEALAND: Buller: 1♂, Nelson Lks NP, Lake Rotoiti, St. Arnaud tr , 645m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 705, window trap; 1♂, Mt. Robert, carpark, 9 IV 2004, R. Leschen, H. Harman, T. Buckley, litter RL864, 41.49′S 172.48′E; Fiordland: 1♀, Fiordland NP, Kepler tr, below Mt. Luxmore Hut, 900m, 45”24.86′S, 167”38.675′E, 11 XII 2005, Nothofagus forest, FMHD #2005-101, berl, leaf &amp; log litter, A. Solodovnikov, D. Clarke, ANMT site 1176; Mid Canterbury: 1♂, Banks Peninsula Ahuriri Scenic Res., 480m, 1 IV 1982, G. Kuschel, litter and rotten wood 82/48; Nelson: 1♂, 30km sw Collingwood 15mi Ck, 100m, 23 V 1982, FMHD #82-597, log litter, S. Peck; 2♀♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41”06.351′S, 172”41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1159; 1♀, Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41” 11.351′S, 172”44.456′E, 28 XI 2005, Nothofagus dominant forest, under bark of small logs, A. Newton, M. Thayer, ANMT site 1156; 1♀, Dun Mt., 2500′, 13 II 1961, G. Kuschel; Marlborough Sounds: 1♂, Tennyson Inlet, e side Duncan Bay, 30m, 15 XII 1984 – 5 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, pyrethrin fogging fungusy log; Westland: 1♀, Deception R, 20 XI 1978, S.P. Worner, mixed podocarp litter; 1♀, Okuku Scenic Reserve, 46.7km w Otira, 75m, 42′43S 171′14E, #050, Laurelia novae-zelandiae leaf litter berlese, 12 I 1998, C. Carlton, R. Leschen; Wellington: 1♂, Manukau Rd, north road end, 20 X 1996, in moderately wet forest litter, J. Nunn; 1♂, Atatapia Stm, Tararua FP, 19 IX 1993, J. Nunn, in wood mould.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: body length 2.1–2.3 mm; male antennomeres 5–10 enlarged; frontal sulcus deep and wide, reaching vertexal fovea; posterior frontal fovea large and oval; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.1–2.3 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37b). Head: Round, widest across eyes (Fig. 37j). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–10 subquadrate, male 5–10 enlarged. Frontal sulcus deep and wide, reaching vertexal fovea. Anterior frontal fovea round, posterior frontal fovea large oval. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37b). Meso-and metathorax trapezoidal as long as wide. Aedeagus: Median lobe slender (Fig. 38b). Phallobase of median lobe symmetrical and rectangular (Fig. 38b). Parameres symmetrical and curved to C-shaped with setae apically and 1 seta at middle (Fig. 38b).</p><p>Type Locality. Picton, Marlborough Sounds, New Zealand .</p><p>Distribution. Buller, Fiordland, Nelson, Mid Canterbury, Marlborough Sounds, Westland, Wellington (Fig. 39: triangles).</p><p>Habitat. Specimens of this species were collected by fogging or sifting leaf or decayed log litter in podocarp or Nothofagus forests.</p><p>Comments. Specimens of S. spiniventris can be distinguished from other species by the enlarged male antennomeres 5–10; deep and wide frontal sulcus reaching a point even with the vertexal fovea; posterior frontal fovea large and oval; and the shape of the antennomeres and genitalia. The type specimens of S. occipitalis share these diagnostic characters. For these reasons, we have placed S. occipitalis in synonymy with S. spiniventris .</p></div>	https://treatment.plazi.org/id/03D82A3EBD2B110F3CBBFC27FE8FFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2A110F3CFBFCB9FC84FB95.text	03D82A3EBD2A110F3CFBFCB9FC84FB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola chathamensis Park and Carlton	<div><p>Sagola chathamensis Park and Carlton, new species</p><p>(Figs. 37c, 38c, 38k, 39)</p><p>Type Material. Holotype. NEW ZEALAND: Chatham Islands: 1♂ (LUNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, CH Mangere I Robin Bush 28.x.1993 R. M. Emberson ”, “litter ex Olearia, Melicytus, Muehlenbeckia forest and feathers”, “ HOLOTYPE Sagola chathamensis Park and Carlton 2013 ” . Paratypes (n = 23: 14 males; 9 females). NEW ZEALAND: Chatham Islands: 3♂♂ 5♀♀ (LUNZ), same data as holotype; 2♂♂ 1♀ (1♀, slide-mounted; LUNZ), Star Keys, 23 I 1998, R. M. Emberson, under rocks in coastal sward ; 4♂♂ (LUNZ), Mangere, Robin Bush, 30 XI 2004, R. M. Emberson, P. Syrett, from leaf litter ; 1♂ (LUNZ), Star Keys, 23 I 1998, J.W.M. Marris, under rock in low coastal vegetation ; 1♂ (LUNZ), Mangere I, Robin Bush, 30 XI–3 XII 1992, J.W.M. Marris, J.W. Early, pitfall trap ; 2♂♂ 2♀ (1♂, slide-mounted, NZAC), Taiko Camp, 10 XI 1991, J.S. Dugdale, sifted rotten wood ; 1♀ (LUNZ), Mangere I, 28 X 1993, R. M. Emberson, litter ex coastal scrub LCNZ 93 /10 ; 1♂ (NZAC), Rungaika, 240–260m, 6 III 1991, R. C. Craw, litter 91/14 .</p><p>Etymology. This species is named after the type locality, Chatham Islands.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: body smaller, length 1.8–2.0 mm; ventral surface of head weakly convex; eye small, approximately one-third length of temple; frontal sulcus deep and linear, almost reaching vertexal fovea; anterior frontal fovea round and covered by frontal rostrum; posterior frontal fovea elongate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 1.8–2.0 mm. Body brown, antennae, maxillary palpi, elytra, and legs paler (Fig. 37c). Head: Male head as long as wide, widest across eyes (Fig. 38k), ventral surface of head weakly convex. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–7 subquadrate, 8–10 weakly transverse. Frontal sulcus deep and linear, reaching almost vertexal fovea. Anterior frontal fovea round and covered by frontal rostrum, posterior frontal fovea elongate. Eye small, approximately onethird length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra approximately triangular (Fig. 37c). Meso- and metathorax trapezoidal, longer than wide. Aedeagus: Median lobe bent S-shaped (Fig. 38c). Phallobase of median lobe symmetrical and rectangular (Fig. 38c). Parameres symmetrical and weakly curved with 1 seta apically and 3 setae on middle (Fig. 38c).</p><p>Distribution. Chatham Islands (Fig. 39: black square).</p><p>Habitat. Specimens of this species were collected using pitfall traps or by sifting forest litter in a coastal area.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2A110F3CFBFCB9FC84FB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2A11083EAAFBCAFC01FF68.text	03D82A3EBD2A11083EAAFBCAFC01FF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola pittensis Park and Carlton	<div><p>Sagola pittensis Park and Carlton, new species</p><p>(Figs. 37d, 38d, 40)</p><p>Type Material. Holotype. NEW ZEALAND: Chatham Islands: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ Pitt Island, 16–26-I-44, E.S.Gourlay. ”, “ E.S.Gourlay Acc. 1970 Ent. Div.”, “ N.Z. Arthropod Collection, NZAC Private Bag 92710 AUCKLAND New Zealand”, “ HOLOTYPE Sagola pittensis Park and Carlton 2013 ” . Paratypes (n = 10: 4 males; 6 females). NEW ZEALAND: Chatham Islands: 1♂ (NZAC), Taiko Camp, 10 XI 1991, J.S. Dugdale, sifted rotten wood ; 1♂ 3♀♀ (NZAC), Awatotara, 182m, 21 II 1967, G.W. Ramsay, litter 67/139 ; 1♂ (NZAC), North Coast, 2 III 1967, G.W. Ramsay ; 3♀♀ (NZAC), Pitt Island, 16–26 I 1944, E.S. Gourlay .</p><p>Etymology. This species is named after the type locality, Pitt Island, Chatham Islands.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: body smaller, length 1.8–2.0 mm; ventral surface of male head transversely depressed with rectangular setose process behind labium; eye small, approximately one-third length of temple; frontal sulcus deep and linear, almost reaching vertexal fovea; anterior frontal fovea round and covered by frontal rostrum; posterior frontal fovea elongate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 1.8–2.0 mm. Body brown, antennae, maxillary palpi, elytra, and legs paler (Fig. 37d). Head: Male head as long as wide, widest across eyes (Fig. 37d). Ventral surface of male head transversely depressed with rectangular setose process behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–7 subquadrate, 8–10 weakly transverse. Frontal sulcus deep and linear, almost reaching vertexal fovea. Anterior frontal fovea round and covered by frontal rostrum, posterior frontal fovea elongate. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra approximately triangular (Fig. 37d). Meso-and metathorax trapezoidal, longer than wide. Aedeagus: Median lobe rectangular (Fig. 38d). Phallobase of median lobe symmetrical and rectangular (Fig. 38d). Parameres symmetrical and weakly curved with 1 seta apically and 3 setae at middle (Fig. 38d).</p><p>Distribution. Chatham Islands (Fig. 40: black circle).</p><p>Habitat. Specimens of this species were collected by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2A11083EAAFBCAFC01FF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2D11083EBBFEEFFBEFF95B.text	03D82A3EBD2D11083EBBFEEFFBEFF95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola chandleri Park and Carlton	<div><p>Sagola chandleri Park and Carlton, new species</p><p>(Figs. 37e, 38e, 40)</p><p>Type Material. Holotype. NEW ZEALAND: Wellington: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND WN Wilton Bush 4/1/93 Wellington ”, “in fallen Astelia ”, “ HOLOTYPE Sagola chandleri Park and Carlton 2013 ”. The original label does not mention who collected this specimen, but the collector was J. T. Nunn . Paratypes (n = 5: 4 males; 1 female). NEW ZEALAND: Rangitikei: 1♂ (JTN), Ruahine FP, entrance Wharite, 22 X 1994, J. Nunn ; Wairarapa: 1♂ (JTN), West L Wairarapa Res., 3 X 1995, J. Nunn; Wanganui: 1♂ (JTN), Ashhurst Domain, 25 X 1993, J. Nunn; Wellington : 1♂ 1♀ (JTN), same data as holotype .</p><p>Etymology. This species is named for worldrenown beetle specialist and one of the most influential specialists of Pselaphinae during modern time, Donald S. Chandler.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: body larger, length 2.1–2.3 mm; male antennomeres 5–10 weakly enlarged; frontal sulcus linear and deep, slightly exceeding posterior margin of eye; male hind femur with triangular process at middle of mesal margin; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.1–2.3 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37e). Head: Round, widest across eyes (Fig. 37e). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate, male 5–10 weakly enlarged. Frontal sulcus deep, reaching just behind eye. Anterior frontal fovea round, posterior frontal fovea oval. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37e). Meso-and metathorax trapezoidal, as long as wide. Male hind femur with triangular process on middle of inside edge. Aedeagus: Median lobe simple (Fig. 38e). Phallobase of median lobe symmetrical and rectangular (Fig. 38e). Parameres symmetrical and weakly curved with 4–5 setae apically and 1 seta preapically (Fig. 38e).</p><p>Distribution. Rangitikei, Wairarapa, Wanganui, Wellington (Fig. 40: triangles).</p><p>Habitat. Specimens of this species were collected by sifting forest litter or fallen Astelia Banks &amp; Sol. ex R. Br. (Asteliaceae) .</p></div>	https://treatment.plazi.org/id/03D82A3EBD2D11083EBBFEEFFBEFF95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2C11093CDEFAAEFC6DFBB3.text	03D82A3EBD2C11093CDEFAAEFC6DFBB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola gimmeli Park and Carlton	<div><p>Sagola gimmeli Park and Carlton, new species</p><p>(Figs. 37g, 38g –h, 40)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: ND Waipoua State Forest, vic. Wairau Summit, 460m, 1.xii.1984 hdwd.- podocarp forest A.Newton / M.Thayer 683”, “pyrethrin-fogging fungusy logs”, “ HOLOTYPE Sagola gimmeli Park and Carlton 2013 ”.</p><p>Etymology. This species is named for LSAM alumnus, lab mate, and enthusiastic supporter of this study, Matthew L. Gimmel.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: larger body, length 2.4 mm; antennomere 1 approximately 2 times longer than wide, with dull surface; gular region of male head depressed and densely setose; frontal sulcus deep, reaching vertexal fovea; shape of genitalia unique to species.</p><p>Description of Male. Length 2.4 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37g). Head: Round, widest across eyes (Fig. 37g). Gular region depressed and densely setose. Antennomere 1 approximately 2 times longer than wide, with dull surface, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching end of head from frontal rostrum. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea elongate. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37g). Meso-and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe slender and simple (Fig. 38g). Phallobase of median lobe symmetrical and rounded (Fig. 38g –h). Parameres asymmetrical, left long, bent with divided apex (Fig. 38g –h).</p><p>Distribution. Northland (Fig. 40: star).</p><p>Habitat. The holotype of this species was collected by pyrethrin-fogging fungusy logs in hardwood and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2C11093CDEFAAEFC6DFBB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2C11093CF3FF31FE4FFAB2.text	03D82A3EBD2C11093CF3FF31FE4FFAB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola threekingsensis Park and Carlton	<div><p>Sagola threekingsensis Park and Carlton, new species</p><p>(Figs. 37f, 38f, 40)</p><p>Type Material. Holotype. NEW ZEALAND: Three Kings Islands: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: TH: Tasman Valley 24 XI 1970, G. Kuschel Litter 70/210”, “ HOLOTYPE Sagola threekingsensis Park and Carlton 2013 ”.</p><p>Etymology. This species is named after the type locality, Three Kings Islands.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: larger body, length 1.8 mm; antennomere 1 with dull surface, 2–8 subquadrate; frontal sulcus deep, reaching vertexal fovea; gular region of male head depressed transversely; shape of genitalia unique to species; only known from Three Kings Islands.</p><p>Description of Male. Length 1.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37f). Head: Round, widest across eyes (Fig. 37f). Gular region of ventral surface of head depressed transversely. Antennomere 1 approximately 1.5 times longer than wide, with dull surface, 2–8 subquadrate, 9–10 weakly transverse. Frontal sulcus deep, reaching vertexal fovea from frontal rostrum. Anterior frontal fovea round and covered by frontal rostrum, posterior frontal fovea elongate. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37f). Meso- and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe as long as paramere, with triangular apical lobe (Fig. 38f). Phallobase of median lobe symmetrical and rounded (Fig. 38f). Parameres asymmetrical, right weakly curved and divided, left rectangular (Fig. 38f).</p><p>Distribution. Three Kings Islands (Fig. 40: black square).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2C11093CF3FF31FE4FFAB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2C110A3EAAFBA7FBBAFEDB.text	03D82A3EBD2C110A3EAAFBA7FBBAFEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sokolovi Park and Carlton	<div><p>Sagola sokolovi Park and Carlton, new species</p><p>(Figs. 37h, 38i, 40)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND ND Pakohu 3 Dec 1977 A.K.Walker Litter 77/178”, “ N.Z. Arthropod Collection, NZAC Private Bag 92710 AUCKLAND New Zealand”, “ HOLOTYPE Sagola sokolovi Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Northland: 1♂ (slide-mounted; AMNZ), North Cape area, Taputaputa Bay, 7 XII 1967, K. Wise, in rotten wood .</p><p>Etymology. This species is named for our lab mate, carabid specialist, and enthusiastic supporter of this study, Igor M. Sokolov.</p><p>Diagnosis. This species can be distinguished from the other species of the auripila speciesgroup by the following combination of characters: larger body, length 2.3 mm; antennomere 1 approximately 2 times longer than wide, with dull surface; gular region of male head depressed roundly and densely setose; frontal sulcus deep reaching vertexal fovea; shape of genitalia unique to species.</p><p>Description of Male. Length 2.3 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 37h). Head: Round, widest across eyes (Fig. 37h). Gular region roundly depressed and densely setose. Antennomere 1 approximately 2 times longer than wide, with dull surface, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching end of head from frontal rostrum. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea elongate. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately triangular (Fig. 37h). Meso-and metathorax trapezoidal, as long as wide. Aedeagus: Median lobe weakly curved as S-shaped (Fig. 38i). Phallobase of median lobe symmetrical and rounded (Fig. 38i). Parameres asymmetrical with round depression basally (Fig. 38i).</p><p>Distribution. Northland (Fig. 40: white circles).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter and rotten wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2C110A3EAAFBA7FBBAFEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2F110B3F60FE9FFDABFD74.text	03D82A3EBD2F110B3F60FE9FFDABFD74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola arboricola Broun Desig. Park and Carlton 2013	<div><p>16. arboricola species-group</p><p>(2 species)</p><p>Diagnosis. Members of the arboricola speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.3–3.0 mm; antennomere 1 approximately 1.5 times longer than wide; frontal sulcus deep reaching one-third length of eye; anterior frontal fovea oval, posterior frontal fovea round (Fig. 41e); eye large and prominent, approximately as long as temple (Fig. 41e); ventral surface of male head with process bearing dense setae behind mouthparts (Fig. 41f); hind wings well-developed; abdominal tergites IV–VI with discal carinae.</p><p>KEY TO SPECIES OF THE ARBORICOLA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Body length 2.3–2.7 mm; antennomeres 2–10 with distinct tubercles; ventral surface of head with inverted triangular process bearing dense setae behind mouthparts (Fig. 41f); abdominal ventrite VII with round depression medially ................................... S. arboricola Broun</p><p>1′. Larger body, length 2.5–3.0 mm; ventral surface of head with blunt process bearing dense setae behind mouthparts; antennomeres 2–10 subquadrate and weakly enlarged; abdominal ventrite VII simple ........... S. tenebrica Broun</p></div>	https://treatment.plazi.org/id/03D82A3EBD2F110B3F60FE9FFDABFD74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2E11053CABFCE9FDAEFE96.text	03D82A3EBD2E11053CABFCE9FDAEFE96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola arboricola Broun Desig. Park and Carlton 2013	<div><p>Sagola arboricola Broun, 1921</p><p>(Figs. 41a, 41c, 41e–f, 42)</p><p>Sagola arboricola Broun, 1921a: 502 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. Lectotype. NEW ZEALAND: Otago Lake: 1♀ (BMNH), glued on rectangular card, “4015. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hollyford. 19.2.1914.” [white label, handwritten]; “ Sagola ♀ arboricola.” “LECTOTYPE Sagola arboricola Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotypes (n = 4: 2 males; 2 females). NEW ZEALAND: Otago Lakes: 2♂♂ 2♀♀ (BMNH), same as holotype. The lectotype designation is required because Broun did not explicitly designate a type specimen, and his comments suggest that multiple specimens were examined (Broun 1921a: 502). This designation will fix the identity of the species and facilitate its recognition by future workers .</p><p>Additional Material (n = 75: 42 males; 33 females). NEW ZEALAND: Bay of Plenty: 1♀, Kaimai-Mamaku Forest Park, Mt. Te Aroha, up. end Tui Mine tr, nr. summit rd, 775m, 37″31.685′S 175″44.684′E, 19 XI–26 XII 2005, low Nothofagus menziesii forest w/ Astelia ground layer, FMHD #2005-019, FIT, A. Newton, M. Thayer, ANMT site 1145; Buller: 2♀♀, Nelson Lks NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp;window trap; 2♂♂ 1♀, Nelson Lks NP, n slope Mt. Robert, Pinchgut tr, 1290m, 18–26 XII 1984, Noth. sol. elfin forest, A. Newton, M. Thayer 716, FIT &amp;window trap; 2♀♀, Nelson Lks NP, n slope Mt. Robert, Pinchgut tr, 950m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, FIT &amp;window trap; 1♂, Nelson Lks NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E, 30 XI 2005, Nothofagus forest, pyr.-fogging &amp; hand-collecting litter in crotch of Nothofagus menziesii ca. 1m above ground, A. Newton, ANMT site 1161; 1♂, Nelson Lks NP, n slope Mt. Robert, 860m, 23–26 III 1980, Nothofagus forest, A. Newton, M. Thayer, fogging; Dunedin: 1♂ 1♀, Woodside Glen, Outram, 13 XI 2005, J. Nunn, in moss on tree trunk; 1♂ 1♀, Mt. Maungatua nr bushline, 1 VII 2000, J. Nunn, on sooty mould on bole of Melicytus; 1♂, Woodside Glen, Outram, 30 X 2005, J. Nunn, in moss and lichen on tree trunk; 1♂, Leith Saddle, 31 VIII 2003, J. Nunn, in dead tree branch; 1♂, Mt. Cargill, Dunedin, 28 IX 1998, J. Nunn, Beaten from Totara; 1♀, Mt. Cargill, Dunedin, 27 VII 2002, J. Nunn, in dead Pseudopanax twig; 1♀, Swampy Spur, 29 XI 1997, J. Nunn; 1♀, Cloud Forest of Leith Tr, 7 XII 2003, J. Nunn, general beating; 1♀, Woodside Glen, Outram, 29 XII 2003, J. Nunn, in moss and sooty mould on tree trunk; Fiordland: 3♂♂ 5♀♀, Gertrude Vly, Milford rd, 23 I 2008, J. Nunn; in moss on beech tree; 1♀, Gertrude Vly, Homer, 19 II 2007, Amongst moss on boulder in riverbed, J. Nunn; 1♀, Kepler tr. start, Te Anau, 22 I 2008, J. Nunn, in moss sample; 1♀, Borland Saddle, 24 I 2008, J. Nunn, sifted moss; Mid Canterbury: 2♂♂ 4♀♀, Banks Peninsula, Mt. Sinclair Scenic Res., 775m, 43″42.977′S, 172″51.098′E, 3–16 XII 2005, ridge top mixed broadleaf w/emergent Podocarpus totara, FMHD #2005-070, FIT, A. Newton, M. Thayer, ANMT site 1163; 1♂, Banks Peninsula, Ahuriri Scen. Res., 450m, 43″39.971′S 172″37.427′E, 16 XII 2005, mixed broadleaf w/emergent podocarp, fogging old log &amp; mossy trunk, A. Newton, ANMT site 1162; 1♂ 1♀, McLennans Bush, Mt. Hutt, 25 XII 1999, J. Nunn, from lichen on Nothofagus bole; 1♂, upper Kaituna V, Banks Penin., 12 I 2008, J. Nunn, in moss from forest; Nelson: 1♂ (slide-mounted), 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &amp;window trap; 1♂, Kahurangi NP, Cobb Ridge, above Cobb Resevoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1159; 1♂, Mt. Arthur, 1200m, 20 XI 1969, J.I. Townsend, mats 69/230; 1♂, Mt. Domett, 30 XI 1971, G. Kuschel, moss 71/169; 1♀, Cobb Ridge, 1040m, 29 XI 2005, J. Nunn, Amongst moss on tree trunk; Otago Lakes: 1♂, Mt. Shrimpton tr, at Pipson Ck, Makarora West, 22 X 2006, in moss sample 400m, J. Nunn; Rangitikei: 1♂, Ruahine Ra, 1036m, Shuteye Shack, 7 II 1980, C.F. Butcher, Malaise trap; South Canterbury: 1♀, Otaio Gorge nr Pareora, 23 III 2008, J. Nunn, in moss on rock face; Marlborough Sounds: 2♂♂, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton, M. Thayer 708, window trap; 1♂, Tennyson Inlet, east side Duncan Bay, 30m, 15 XII 19845 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, FIT &amp;window trap; 1♂, Tennyson Inlet, west side Te Mako Bay, 125m, 15 XII 1984 – 5 I 1985, Nothofagus -podohdwd, A. Newton, M. Thayer 710, FIT &amp;window trap; Southland: 2♂♂, Catlins For. Park, Chloris Pass, wnw Owaka, 285m, 46°23.47′S 169°27.482′E, 7 XII 2005, Wienmannia-Metrosideros-Nothofaguspodocarpus forest, fogging, A. Newton, D. Clarke, ANMT site 1167; 1♂, Tutuku, sw of Owaka, 19 I 1978, G. Kuschel, sifted 78/44; 3♂♂ 1♀, Whiskey Gulley, Blue Mtns, 3 VIII 2002, J. Nunn, Amongst lichen from fallen beech branch; 1♂ 3♀♀, Blue Mountains, north end, 9 VIII 2003, in compact moss cushion on beech bole at treeline; 1♂, Dollamore Park, Hokonui Hills, 28 X 2002, J. Nunn, in moss on bole of Rimu; 1♂, Princhester Base Hut, Takitimu Fst, 21 II 2003, J. Nunn, in moss sample; 1♀, Blue Mountains, mw relay stn, 13 II 2005; Taranaki: 1♂, Pouakai Ra., 10–13 I 1978, J.S. Dugdale, malaise trap; Waikato: 2♂♂ 1♀, Mt. Pirongia, Wharauroa, 854m, 9 VI 1977, B.A. Holloway, moss/lichen 77/60; 1♂, Mt. Pirongia, Wharauroa, 854m, 9 VI 1977, B.M. May, moss 77/58; 1♂, Mt. Pirongia, 853m, 24 XI 1964, G. Kuschel, moss 64/105; Westland: 1♂, Otira E., 6.8km ne Otira, 280m, 18-21 III 1980, Nothofagus fusca -podocarp, A. Newton, M. Thayer, fogging.</p><p>Diagnosis. This species can be distinguished from the other species of the arboricola speciesgroup by the following combination of characters: body length 2.3–2.7 mm; male antennomeres 2–10 with tubercles; ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts; male abdominal ventrite VII with round depression medially; shape of genitalia unique to species.</p><p>Redescription. Length 2.3–2.7 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 41a). Head: Bluntly triangular, widest across eyes (Fig. 41e). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts (Fig. 41f). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate, male 2–10 with tubercles. Frontal sulcus deep, reaching onethird length of eye (Fig. 41e). Anterior frontal fovea oval (Fig. 41e), posterior frontal fovea round (Fig. 41e). Eye large and prominent, approximately as long as temple (Fig. 41e). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 41a). Mesoand metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, less developed in female. Male abdominal ventrite VII with round depression medially. Basolateral foveae of abdominal ventrites V–VI very small. Aedeagus: Median lobe triangular, dorsal process large, V-shaped (Fig. 41c). Phallobase of median lobe symmetrical and rectangular (Fig. 41c). Parameres symmetrical and rectangular with sparse setae anteriorly (Fig. 41c).</p><p>Type Locality. Hollyford, Otago Lakes, New Zealand .</p><p>Distribution. Bay of Plenty, Buller, Dunedin, Fiordland, Nelson, Mid Canterbury, Otago Lakes, Rangitikei, South Canterbury, Marlborough Sounds, Southland, Taranaki, Westland, Waikato (Fig. 42: black circles).</p><p>Habitat. Most specimens of this species were collected by fogging, using Malaise, flight intercept, or window traps, or by sifting mossy leaf or log litter in Nothofagus, podocarp, or hardwood forests. Many specimens were collected at high elevations.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2E11053CABFCE9FDAEFE96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2011053F67FCC1FB43FB2B.text	03D82A3EBD2011053F67FCC1FB43FB2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola strialis Broun 1921	<div><p>17. strialis species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the strialis species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.2–2.8 mm; frontal rostrum prominent (Fig. 43c); antennomere 1 approximately 2 times longer than wide; anterior and posterior frontal foveae deep and largely oval (Fig. 43c); ventral surface of male head with distinct triangular process bearing dense setae behind mouthparts (Fig. 43d); hind wings well-developed; abdominal tergites IV–VI with discal carinae; only known from South Island.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2011053F67FCC1FB43FB2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2011053CB1FECFFC76FC9F.text	03D82A3EBD2011053CB1FECFFC76FC9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola tenebrica Broun 1921	<div><p>Sagola tenebrica Broun, 1921</p><p>(Figs. 41b, 41d, 42)</p><p>Sagola tenebrica Broun, 1921a: 487 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♀ (BMNH), glued on rectangular card, “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Moa Basin. 20.10.1913.” [white label, handwritten]; “ Sagola ♀ tenebrica.” [white label, handwritten].</p><p>Additional Material (n = 19; 11 males, 8 females). NEW ZEALAND: Fiordland: 4♂♂ 6♀♀, Borland Saddle, 24 I 2008, sifted moss, J, Nunn ; 1♀, Borland Saddle, 1050m, 16 XII 1998, in moss on Nothofagus bole, J. Nunn ; 1♂, Breaksea So, 12 III 1983, C.F. Butcher ; 1♂, Secretary I, Mt. Grono, 853m, 27 XI 1981, C.F. Butcher, mats, moss, tussock 91/186 ; 1♂, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S, 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD2005-088, recent flood debris floating in stream, J. Nunn, A. Newton, ANMT site 1169; Mid Canterbury: 1♂, Arthur Pass, Dobson Nature Walk, 8 II 1982, C.F. Butcher, sifted litter 82/26 ; 1♂, Moa Basin, A.E. Brookes Collection, T. Broun Collection ; Mackenzie: 1♂, Mt. Burns, 1700m, 13 I 1970, J.I. Townsend, Mats 70/101 ; Nelson: 1♂, Mt. St. Arnaud, 1900m, 1 VII 1973, J.C. Watt, litter 73/109 ; Westland: 1♂, Fox G 1200m, Chancellor Hut, 15 XII 1984, J.W. Early, P Syrett, Subalpine herb field tussock litter LCNZ 84 /8 .</p><p>Diagnosis. This species can be distinguished from the other species of the arboricola speciesgroup by the following combination of characters: larger body length, 2.5–3.0 mm; ventral surface of male head with blunt process bearing dense setae behind mouthparts; male antennomeres 2–10 subquadrate and weakly enlarged; shape of genitalia unique to species.</p><p>Redescription. Length 2.5–3.0 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 41b). Head: Bluntly triangular, widest across eyes (Fig. 41b). Ventral surface of male head with blunt process bearing dense setae behind mouthparts. Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate and weakly enlarged. Female antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching one-third length of eye. Anterior frontal fovea oval, posterior frontal fovea round. Eye large and prominent, approximately as long as temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 41b). Mesoand metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, less developed in female. Aedeagus: Median lobe triangular, dorsal process crescent-shaped (Fig. 41d). Phallobase of median lobe symmetrical and rectangular (Fig. 41d). Parameres symmetrical and rectangular with sparse setae anteriorly and posteriorly (Fig. 41d).</p><p>Type Locality. Moa Basin, Mid Canterbury, New Zealand .</p><p>Distribution. Fiordland, Mid Canterbury, Mackenzie, Nelson, Westland (Fig. 42: triangles).</p><p>Habitat. Most specimens of this species were collected by sifting mossy leaf or log litter in Nothofagus forests. Many specimens were collected at high elevations.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2011053CB1FECFFC76FC9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2011013F53FB2DFDEAFC9F.text	03D82A3EBD2011013F53FB2DFDEAFC9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola strialis Broun 1921	<div><p>Sagola strialis Broun, 1921</p><p>(Figs. 43–44)</p><p>Sagola strialis Broun, 1921a: 489 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Sagola fasciculata Broun, 1921a: 497 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “3996. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Moa Hill. 20.11.1913.” [white label, handwritten]; “ Sagola . ♂ strialis.” [white label, handwritten] . Syntypes. NEW ZEALAND: Nelson: 1♂ (BMNH), glued on rectangular card, “4007.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.12.1914 ” [white label, handwritten]; “ Sagola fasciculata ♂.”[white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “4007. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.2.1915 ” [white label, handwritten]; “ Sagola ♂ fasciculata.”[white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4007. ♀ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.10.1914 ” [white label, ♂♂ handwritten]; “ Sagola ♀ fasciculata.” [white label, handwritten] .</p><p>Additional Material (n = 246: 160 males; 86 females). NEW ZEALAND: Buller: 13♂♂ 3♀♀, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 17 XII 2005, Nothofagus forest, FMHD #2005-110, berl leaf &amp; log litter, A. Solodovnikov, D. Clarke, ANMT site 1161; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 30 XI–17 XII 2005, Nothofagus forest, FMHD #2005-061, pitfall trap (10), A. Solodovnikov, D. Clarke, ANMT site 1161; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 30 XI 2005, Nothofagus forest, FMHD #2005-110, fogging, A. Newton, ANMT site 1161; 7♂♂ 3♀♀, Lewis Pass Nat Res, 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, litter; 3♂♂, 18km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, litter; 3♂♂, Nelson Lks NP, Lake Rotoroa, Braeburn tr, 470m, 16 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 712, window trap; 4♂♂ 2♀♀, Nelson Lks NP, n slope Mt. Robert, Pinchgut tr, 950m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, litter; 1♂, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713; 3♂♂ 4♀♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, litter; 2♂♂ 3♀♀, Nelson Lks NP, n slope Mt. Robert, 860m, 23–26 III 1980, Nothofagus forest, A. Newton, M. Thayer, litter; 2♂♂, 0.8km n Bullock Ck nr Punakaiki, 50m, 23 III 1980, broadleaf-nikau palm-podocp, A. Newton, M. Thayer, litter; 1♂, Nelson Lks NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp;window trap; 1♂, Greymouth, Boddytown, 2 II 1984, J.C. Watt, sifted litter 84/23; 1♀, Paparoa Ra, 610m, Buckland Peaks Tk, 25 XI 1984, B.A. Holloway, C.F. Butcher, moss &amp; lichen 84/77; 3♂♂, Flagstaff Bio. Res., 13 VII 1972, J.S. Dugdale, litter; 1♂, Mawhera SF, 15km se of Ngahere, 305m, 10 XI 1971, J.S. Dugdale, litter 71/143; 1♂, Mawhera SF, 19km se of Ngahere, 305m, 10 XI 1971, J. McBurney, litter 71/144; 1♂, Mawhera SF, 16km se of Ngahere, 305m, 11 XI 1971, J.S. Dugdale, litter 71/145; 2♂♂ 1♀, Mawhera SF, Kangaroo Ck, 27 I 1972, J.S. Dugdale, litter 72/86; 2♂♂, Paparoa Ra, Croesus tr, 730m, 21 I 1982, J.W. Early, moss &amp; litter in podocarp –broadleaf forest; 1♂ 1♀, Fletchers Ck, Biological Reserve, XI 1971, McBurney, litter 71/127; 1♂, W. Inangahua, St. 127 Coll Ck, 19 IX 1972, J.S. Dugdale, moss 72/184; 1♂, Tawhai SF, Big R. Rd, 3km S of Reefton, 28 I 1972, J. McBurney, litter 72/76; 1♂, Fletcher Ck, MacMahan Ck, 25 I 1972, J.S. Dugdale, XPB9, litter 72/79; 1♂, Fletchers Ck, 6km sw Rotokohu, 25 I 1972, J.S. Dugdale, PB1 litter 72/103; 1♂, Fletchers Ck, 6km sw Rotokohu, 25 I 1972, J. McBurney, PB15 litter 72/93; 1♂, Capleston, Redmans Ck, 21 IX 1972, J.C. Watt, litter 72/186; 1♂, Reefton, 12 IV 1977, J.A. Wightman, pit trap burnt pine; 1♂, Lewis Pass, 19 X 2007, J. Nunn, washed soil sample; 1♂, Inland Pack tr, 1km n of Runakaiki River, sifted moss, J. Nunn, 29 XII 2010; 1♀, Coal Ck, Runanga, 11 II 2007, J. Nunn, moss; 1♀, W Inangahua SF, 127m, Stoney Ck, 18 IV 1972, J. McBurney, litter PB1; 1♀, Fletcher Ck, 7 III 1972, J.S. Dugdale, litter 72/106; 1♀, Reefton, 5 XI 1958, J.M. Hoy; 1♂, Nelson Lks NP, Mt. Robert car park, 28 IX 1973, R.M. Emberson, beech litter; Fiordland: 1♂ 1♀, W. Olivine Ra, Simonin Pass, 1 II 1975, G.W. Ramsay, under decayed branches lower bush edge; 2♂♂ 1♀, Fiordland NP, Borland rd, Borland Saddle vic, 980m, 45″44.81′S 167″22.88′E, 10 XII 2005, Nothofagus menziesii forest in ravine, FMHD #2005-090, litter, A. Newton, ANMT site 1171; 1♂ 4♀♀, Fiordland NP, Milford Sound rd, Tukoko tr, 70m, 44″40.517′S 167″57.895′E, 9 XII 2005, very mossy Nothofagus forest, emergent podocarp, FMHD #2005-086, litter, A. Newton, M. Thayer, ANMT site 1168; 1♂ 1♀, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD #2005-087, litter, A. Newton, M. Thayer, ANMT site 1169; 1♂ 6♀♀, Fiordland NP, Milford Sound rd, Smithy Creek Campground area, 400m, 44″57.065′S 168″01.155′E, 9 XII 2005, Nothofagus fusca &amp; N. menziesii open forest, FMHD #2005-089, litter, M. Thayer, A. Newton, ANMT site 1170; 1♂, Borland rd, Limestone Cave tr, 45′46S 167′28E, #140, Nothofagus log/bark litter berlese, 24 I 1998, C. Carlton, R. Leschen; 1♂, Borland rd, Borland Saddle tr, 45′46S 167′23E, #138, Nothofagus leaf litter berlese, 24 I 1998, C. Carlton, R. Leschen; 1♂, Bauza Is, Doubtfull sd, III 1986, pitfall trap, G.M. Bremner; 1♂, Lower Hollyford rd, 44′44S 168′08E, #111, Nothofagus /coastal forest leaf litter berlese, 22 I 1998, C. Carlton, R. Leschen; 1♂, Hump Ridge, 1220m, 6 II 1976, G.W. Ramsay, litter 76/38; 1♂, Milford rd, 60′, 12 XII 1966, A.K. Walker, K. Wilson, cordyline litter 66/505; 1♂, Eglington Valley Divide, 400m, 13 II 1982, C.F. Butcher, moss/liverworts 82/29; 1♂, Lake Hauroko, 2 II 1966, J.I. Townsend, mixed moss 66/363; 1♀, Beside Tutoko R, 60m, 13 II 1980, J.W. Early, litter, N. menziesii forest; 1♀, Lake Hauroko Road end, 19 II–22 III 2003, FIT, R. Leschen, J. Nunn; 1♂ 1♀ (1♂, slide-mounted), Te Anau Control Gates, #104, 45′26S 167′41E, Nothofagus leaf litter berlese, 22 I 1998, R. Leschen, D. Gleeson; 1♀, Fiordland NP, Kepler tr, above Brod Bay, 680m, 45″24.37′S 167″39.105′E, 11 XII 2005, Nothofagus forest, FMHD #2005- 102, litter, A. Solodovnikov, D. Clarke, ANMT site 1177; 1♀, Fiordland NP, Kepler tr, below Mt. Luxmore Hut, 900m, 45″24.865′S 167″38.675′E, 11 XII 2005, Nothofagus forest, fogging, A. Solodovnikov, D. Clarke, ANMT site 1176; 1♀, Tempest Spur, W. Olivine Ra, 1220–1463m, 25 I 1975, G.W. Ramsay, litter 75/33; Mid Canterbury: 3♂♂, Arthur’ s Pass, Dobson Nature Walk, 8 II 1982, C.F. Butcher, sifted litter 82/26; 3♂♂, Arthur’ s Pass, Bealy tr, 8 II 1982, C.F. Butcher, litter 82/19; 1♂, Craigieburn SF, Dracophyllum Flat tr, 43′10S 171′42E, FIT, #150, 11–27 I 1998, C. Carlton, R. Leschen; North Canterbury: 1♂ 1♀, 2.9km N Arthur’ s Pass, 840m, Bealey Valley Trailhead, 42′55S 171′33E #036, Nothofagus solandri leaf litter berlese, 11 I 1998, C. Carlton, R. Leschen; 1♂, 3km n Arthur’ s Pass, 900m, 42′55S 171′33E, #042, Nothofagus / Dracophyllum leaf litter berlese, 11 I 1998, C. Carlton, R. Leschen; 3♂♂, Arthur’ s Pass NP, Bealey Vy tr, 840m, 18–21 III 1980, subalpine Nothofagus, A. Newton, M. Thayer, litter; 1♂, Arthur’ s Pass NP, Klondyke Cnn, 700m, 26 X 1981, J.W. Early, moss from forest floor; 1♂, Mt. Arthur Pass, Avalanche Peak, 3 I 1950, E. Dawson, leaf litter; 1♂, Arthur’ s Pass NP, Halpine Creek, 17 V 1970, D.S. Homink litter &amp; rotten wood; 1♂, Arthur’ s Pass Mt. Aicken, 950m, 9 II 1984, J.C. Watt, sifted litter 84/24; 1♀, Arthur’ s Pass, Mt. Aicken, 305m, 11 II 1984, J. C. Watt, W. Pond flood debris; Nelson: 5♂♂, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, AMNT site 1159; 23♂♂ 2♀♀, Kahurangi NP, Cobb Dam rd, Asbestos tr, 450m, 41″06.333′S 172″43.174′E, 29 XI–18 XII 2005, mixed braodleaf (incl. Nothofagus fusca)-podocarp forest, FMHD #2005- 057, carrion(octopus) trap, A. Solodovnikov, D. Clarke, ANMT site 1160; 1♂ 2♀♀, Kahurangi NP, Mt. Arthur tr, below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD #2005-049, litter, A. Solodovnikov, D. Clarke, ANMT site 1157; 1♀, Cobb Ridge east of Cobb Reservoir, 990m, 2 I 1985, Nothofagus forest, A. Newton, M. Thayer 728, litter; 2♂♂, Kahurnagi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S 172″44.456′E, 28 XI–19 XII 2005, Nothofagus dominant forest, FMHD #2005-044, FIT, A. Newton, M. Thayer, ANMT site 1156; 3♂♂, 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &amp;window trap; 3♂♂ 4♀♀, Third House, Dun Mt, 14 IX 1971, G.W. Ramsay, litter 7/114; 1♂ (slide-mounted), 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap; 2♂♂, Karamea sdle, 13 X 1970, J.I. Townsend, litter 70/157; 1♂ 1♀, Karamea Bluff, 1420ft, 29 IX 1966, J. Nunn; 1♂, Cobb Valley, Mytton Hut, 1220m, 13 XII 1967, S. Edridge, litter 67/287; 1♂, Dun Mt, 610m, 4 IV 1966, J.I. Townsend, moss 66/127; 1♂, Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, wet litter; 1♂, Punakaiki, Pororari tr, 27 I 1996, forest litter, J. Nunn; 1♂, Oparara Basin, Box Canyon Cave, 200m, 27 II–1 III 2007, J.W. Early, R.F. Gilbert, Nothofagus -podocarp forest, yellow pan trap; 1♂, Glenhope, T. Hall, 30 I 1915; 1♀, Dun Mt, 2000′, 14 II 1942, E.S. Gourlay; 1♀, Dun Mt., 580m, 12 VII 1966, J.I. Townsend, moss 66/204; 1♀, Upper Maitai, 3 V 1950, E.S. Gourlay; 1♀, Wooded Peak, 609m, 31 VIII 1966, J.C. Watt, rotten Nothofagus stump; Otago Lakes: 4♂♂, Mt. Aspiring NP, 12.5km nne Makarora, 370m, 11–17 I 1985, Noth. menz. - hdwd-podocarp forest, A. Newton, M. Thayer 740, FIT &amp;window trap; 1♂ 1♀, Davis Flat, Makarora vly, 19 I 2008, moss, J. Nunn; 1♂, Mt. Aspiring NP, se of Haast Pass, 600m, 11–17 I 1985, Noth. menz. forest, A. Newton, M. Thayer 739, FIT &amp;window trap; 1♂, Mt. Aspiring NP, 5.5km nne Makarora, 330m, 11–17 I 1985, Noth. menz. forest, A. Newton, M. Thayer 741, FIT &amp;window trap; 1♂, Makarora, 23–25 I 1978, 500m, S. &amp; J. Peck, litter; 1♂, McKerrow Range, Makarora, 1150m, 23 I 1978, G. Kuschel, litter 78/51; 1♂, Dart Hut, 13–15 II 1980, J.S. Dugdale, malaise trap; 1♀, 44.5km nw Wanaka, 350m, Matukituki Valley, 44′29S 168′47E, #079, Nothofagus menziesii leaf litter berlese, 18 I 1998, C. Carlton, R. Leschen; 1♀, Cameron Flat, Makarora V, 22 I 1996, J. Nunn; Southland: 1♂, N. Te Anau, 50m, 19 II 1965, A.K. Walker, litter 65/52; 1♂, Owaka Glenomaru Res, 18 I 1978, S. &amp; J. Peck, litter; 1♀, MacLennan, 100’, 14 II 1968, J.I. Townsend, moss 68/20; Westland: 1♂♂ 3♀♀, Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1986, podocarp-hdwd forest, A. Newton, M. Thayer 731, litter; 1♂, Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1986, podocarp-hdwd forest, A. Newton, M. Thayer 731, fogging; 2♂♂ 1♀, 1.8km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, FIT &amp;window trap; 5♂♂, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap; 1♂, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, litter; 2♂♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, FIT &amp;window trap; 1♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, litter; 2♂♂ 1♀, Lake Ianthe, 7 II 1984, J.C. Watt, sifted litter &amp; wood mould 84/22; 4♂♂ 4♀♀, Hokitika Gorge, 29 I 1978, S.B. Peck, litter; 2♂♂, Okuku State Forest, 10km e of Kumara, 28 VIII 1977, R.M. Emberson, podocarp-broadleaf forest, moss&amp;litter; 1♂, Hokitika R.Gorge, S Kowhitirangi, 100m, 17 III 1980, podocarpbroadleaf, A. Newton, M. Thayer, litter; 1♂ 1♀, Okuku Ck, 11.3km sse Kumara, 60m, 18–22 III 1980, podocarp-broadlf, A. Newton, M. Thayer, litter; 1♂, L. Mahinapua Scen Res, 30m, 16–22 III 1980, podocarp mixed broadleaf, A. Newton, M. Thayer, litter; 1♂ 1♀, Lake Wahapo, 27 XI 1966, J.I. Townsend, litter 66/345; 1♂, Jacksons Bay, 23 IX 1979, A.K. Walker, moss &amp; dead wood litter; 1♂ 1♀, Hokitika Gorge Res, Hokitika, 29 I 1978, S. &amp; J. Peck, litter, 100m; 1♂, Westland NP, Castle Rocks Hut, 1220m, 14–16 I 1996, yellow pan trap, subalpine tussock herb field; 1♂, Haast Pass, road summit, 6 II 2009, sifted ground moss, J. Nunn; 1♂, Haast Pass, 31 I 2005, J. Nunn, forest floor moss; 1♂, Otira, Barretts Creek, 9 II 1982, C.F. Butcher, litter 82/27; 1♂ 1♀, Fantail Falls, 3km ne Haast Pass, 4 II 1984, J.C. Watt, wood mould &amp; litter; 1♂, Summit Haast Pass rd, 1850’, 7 XII 1966, A.K. Walker, litter 66/471; 1♂, Pakihi Walk, 5.2km se Okarito, #062, 43′15S 170′12E, 14 I 1998, litter, C. Carlton, R. Leschen; 1♂, 3.2km ne Haast, 14m, Haast River Walk, 43′52S 169′03E, #072, Nothofagus -podocarp forest litter, 17 I 1998, C. Carlton, R. Leschen; 1♂, 56.6km se Haast, Fantail Falls tr, 475m, 44′05S 169′23E, #073, Nothofagus menziesii litter, 17 I 1998, C. Carlton, R. Leschen; 1♂, Victoria Range, 300’, 26 X 1940, E.S. Gourlay; 3♀♀, Jacksons, Taramakau R, 150m, 26 I 1978, G. Kuschel, sifted litter &amp; rotten wood; 1♀, Haast Pass w of summit, 550m, 24 I 1978, G. Kuschel, sifted litter &amp; rotten wood 78/52; 1♀, Haast Pass, 20 III 1968, R.A. Cumber, litter; 1♀, Otira Valley, Dobsons tr, 1005m, 11 XI 1966, A.K. Walker, moss 66/382; 1♀, Nile River Valley, moss, 1 IX 1971, R.M. Emberson; 1♀, Haast Pass, 518m, 23 X 1966, J.I. Townsend, moss 66/349; 1♀, Lake Kaniere rd, 2.8km nw Lake Kaniere, 120m, 8-19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 732, litter; 1♀, 3km n of Bruce Bay, 6 II 2009, sifted moss from podocarp swamp forest, J. Nunn; 1♀, Kelly’ s Ck, Otira, 25 II 1989, J. Nunn.</p><p>Diagnosis. This species can be distinguished from other species by the following combination of characters: body larger, length 2.2–2.8 mm; frontal rostrum prominent; anterior and posterior frontal foveae deep and largely oval; ventral surface of male head with distinct triangular process bearing dense setae behind mouthparts; shape of antennomeres and genitalia unique to this species.</p><p>Redescription. Length 2.2–2.8 mm. Body reddish to blackish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 43a). Head: Round, widest across eyes (Fig. 43c). Ventral surface of male head with distinct triangular process bearing dense setae behind mouthparts (Fig. 43d). Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum prominent. Frontal sulcus deep, slightly exceeding eye (Fig. 43c). Anterior and posterior frontal foveae oval (Fig. 43c). Eye prominent, one-third length of temple (Fig. 43c). Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 43a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe elongate and divided vertically, dorsal process small and triangular (Fig. 43b). Phallobase of median lobe symmetrical and rounded (Fig. 43b). Parameres symmetrical and wide with setae anteriorly (Fig. 43b).</p><p>Type Locality. Moa Hill, Mid Canterbury, New Zealand .</p><p>Distribution. Buller, Mid Canterbury, North Canterbury, Nelson, Otago Lakes, Southland, Westland (Fig. 44: black circles).</p><p>Habitat. Most specimens of this species were collected using flight intercept and window traps or by sifting mossy leaf or log litter in hardwood, podocarp, broadleaf, or Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD2011013F53FB2DFDEAFC9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2411013CBFFCDBFB45FA8D.text	03D82A3EBD2411013CBFFCDBFB45FA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola anisarthra Broun, 1893 b: 1053	<div><p>18. anisarthra species-group</p><p>(5 species)</p><p>Diagnosis. Members of the anisarthra speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.3–3.2 mm; antennomere 1 approximately 2 times longer than wide; ventral surface of head weakly convex, but not modified (Fig. 45m); anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea oval (Fig. 45l); hind wings well-developed; abdominal tergites IV–VI with discal carinae.</p><p>KEY TO SPECIES OF THE ANISARTHRA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Abdominal ventrite VII deeply emarginate, surrounded by dense setae........................ 2</p><p>1′. Abdominal ventrite VII normal..............3</p><p>2(1). Antennomeres 9–10 transverse; elytra triangular, as long as wide; hind wings reduced to small pads ............................ .............. S. wairarapaensis new species</p><p>2′. Antennomeres 9–10 subquadrate; elytra rectangular, longer than wide; hind wings fully developed ..... S. clarkei new species</p><p>3(1′). Antennomeres 4–6 enlarged; frontal sulcus deep (Fig. 45l); eye larger, one-half length of temple (Fig. 45l) ................................... ................................. S. anisarthra Broun</p><p>3′. Antennomeres 4–6 normal, with tubercles; frontal sulcus shallow; eye smaller, onethird length of temple.............................. 4</p><p>4(3′). Median lobe of genitalia slender, as wide as paramere (Fig. 45g –h) .......................... ............................... S. planipennis Broun</p><p>4′. Median lobe of genitalia at least 3 times broader than paramere (Fig. 45i) .............. ........................ S. townsendi new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD2411013CBFFCDBFB45FA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD2411033F41FAB0FCE4F99D.text	03D82A3EBD2411033F41FAB0FCE4F99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola anisarthra Broun, 1893 b: 1053	<div><p>Sagola anisarthra Broun, 1893</p><p>(Figs. 45a, 45f, 45 l-m, 46)</p><p>Sagola anisarthra Broun, 1893b: 1053 . Raffray 1904: 498, 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Sagola suturalis Broun, 1914b: 158 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.</p><p>Sagola dickensis Broun, 1917: 377 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Dunedin: 1♂ (BMNH), glued on rectangular card, “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola anisarthra fide Sandager Moeraki” [white label, handwritten] . Syntypes of Sagola dickensis: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed], “3827. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Mt. Dick. 10.3.1914 ” [white label, handwritten], “ Sagola . ♂ dickensis.” [white label, handwritten]; 1♂ (BMNH), glued on rectangular card, “3827. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Dick. 10.3.1914 ” [white label, handwritten]; “ Sagola . ♂ dickensis.” [white label, handwritten]; 1♂ (NZAC), glued on rectangular card, “Coll. T.Hall. 10-3-1914.” [white label, handwritten], “Mt. Dick, W. of L. Wakatipu, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Paratype.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “ T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]; 1♂ (NZAC), glued on rectangular card and aedeagus dissected and mounted in balsam on a clear plastic card, “ Sagola dickensis Broun. ” [white label, handwritten], “3827.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “Coll. T.Hall, march, 1914.” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed] ; 1♂ (NZAC), glued on rectangular card, “Coll. T.Hall, march, 1914.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed] ; 2♂♂ (NZAC), glued on rectangular card, “Coll. T.Hall, march, 1914.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed]. *Original description examined eight specimens, but we could not find one of them. Syntypes of Sagola suturalis: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “type” [red label, printed], “3522. ♂ ” [white label, handwritten], “ Rakaia. 5.6.1912 ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “ Sagola . ♂ suturalis.” [white label, handwritten] ; ♂, glued on rectangular card, “3522. ♂ ” [white label, handwritten], “ Rakaia Gorge. 5.6.1912 ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “ Sagola . ♂. suturalis.” [white label, handwritten] .</p><p>Additional Material (n = 37: 30 males; 7 females). N E W Z E A L A N D: D u n e d i n: 10♂♂ (1♂, slide-mounted), Boulder Beach, Otago Penin., 19 XI 2004, washed up live with incoming tide, J. Nunn ; 1♀, Mt. Watkin nr Waikouaiti, XII 2007, pit trap, Kelvin Lloyd ; 1♂, Boulder Beach, Otago Penin., 7 XII 2002, washed up live with incoming tide, J. Nunn ; 1♂, Boulder Beach, Otago Penin., 14 I 2005, washed up live with incoming tide, J. Nunn ; 2♂♂, Mt. Cargill, Grahams Bush, 260m, 45″48.299′S 170″34.905′E, 14 XII 2005, mixed broadleaf forest on steep slope w/emergent podocarp, FMHD#2005-107, litter, A. Newton, M. Thayer, ANMT site 1164; Fiordland: 4♂♂ 1♀, S of Lake Hauroko, 12 XI 1966, F. Alack, litter 66/420; Mid Canterbury : 4♂♂ 2♀♀, Goat Hill, nr. Metheven, T. Hall, 20 X 1913 ; 1♂, Mt Hutt, 11 V 2003, A.C. Eyles, mountain beech litter; Southland : 1♀, Clifden, Limestone Bluffs, 11 II 1968, J.I. Townsend, moss ; 5♂♂ 1♀, MW relay stn, Blue Mountains, 19 III 2006, J. Nunn ; 1♂, Lookout Point Bluff, 7 VII 2001, amongst tussock grasses, J. Nunn ; 1♀, Pounawea Catlins, 25 X 2003, litter at beach streamline, J. Nunn .</p><p>Diagnosis. This species can be distinguished from the other species of the anisarthra speciesgroup by the following combination of characters: larger body, length 2.4–2.8 mm; male antennomeres 4–6 enlarged and 4–11 bearing tubercles; frontal sulcus deep reaching midpoint of eye; anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, approximately twothirds length of temple; shape of genitalia unique to species.</p><p>Redescription. Length 2.4–2.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 45a). Head: Bluntly triangular, widest across eyes (Fig. 45l). Ventral surface of head weakly convex. Male antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate, 4–6 enlarged, 7–10 bearing tubercles. Female antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum (Fig. 45l), posterior frontal fovea oval (Fig. 45l). Eye large and prominent, approximately one-half length of temple (Fig. 45l). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 45a). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe as long and wide as parameres (Fig. 45f). Phallobase of median lobe symmetrical and rounded (Fig. 45f). Parameres symmetrical with setae apically (Fig. 45f).</p><p>Type Locality. Moeraki, Dunedin, New Zealand .</p><p>Distribution. Dunedin, Fiordland, Mid Canterbury, Otago Lakes, Southland (Fig. 46: black circles).</p><p>Habitat. Most specimens of this species were collected by sifting mossy leaf or log litter in forests or the intertidal zone of beach areas.</p><p>Comments. Specimens of S. anisarthra can be distinguished from other species by the enlarged male antennomeres 4–6 and dorsal shape of head. The type specimens of S. dickensis and S. suturalis share these diagnostic characters, and additionally those species have been collected at or near the type localities. For these reasons, we have placed S. dickensis and S. suturalis in synonymy with S. anisarthra .</p></div>	https://treatment.plazi.org/id/03D82A3EBD2411033F41FAB0FCE4F99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD26111C3EBBF9C1FB9AFA82.text	03D82A3EBD26111C3EBBF9C1FB9AFA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola planipennis Broun 1921	<div><p>Sagola planipennis Broun, 1921</p><p>(Figs. 45b, 45g –h, 46)</p><p>Sagola planipennis Broun, 1921a: 500 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Additional Material (n = 15: 13 males; 2 females). NEW ZEALAND: Buller: 1♀, Lake Rotoiti, 15 II 1916, T. Broun collection, A.E. Brookes collection ; 2♂♂, Rotoiti, 3 IV 1916, T. Broun collection, A.E. Brookes collection ; Fiordland: 1♀, Barrier River, 1067m, little Red Hill, 2 II 1975, G.W. Ramsay, litter 75/50; Marlborough : 2♂♂, nr Wards Pass, 1218m, Molesworth, 19 III 1968, J.I. Townsend, moss 68/80 ; 1♂, Wairau V, Schrodes Ck, 7 IX 1966, J.I. Townsend, litter 66/295 ; Mid Canterbury: 1♂ 1♀ (1♀, slide-mounted), Arthurs Pass, Temple Basin, 1493m, 8 II 1982, J.S. Dugdale, sedges, moss and mats ; Mackenzie: 1♂, Ohau Skifield, Ohau Rng, 3 III 2007, J. Nunn, under stone, 1800m; Nelson : 4♂♂, Saint Arnaud, 15 VI 1916, T. Broun collection, A.E. Brookes collection ; 1♂, Iron Hill Summit ridge, Lake Sylvester, 1600m, 18 XI 1972, J.S. Dugdale, 72/193; Otago Lakes: 1♂, Haast Pass, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=169.1&amp;materialsCitation.latitude=-44.433334" title="Search Plazi for locations around (long 169.1/lat -44.433334)">Mt. Albert</a>, 5 III 2003, R. Leschen, under rocks, RL779, 44°26′S 169°06′E .</p><p>Diagnosis. This species can be distinguished from the other species of the anisarthra speciesgroup by the following combination of characters: body length 2.3–2.7 mm; antennomeres 4–11 bearing tubercles; frontal sulcus shallow, reaching midpoint of eye; anterior frontal fovea round partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, one-half length of temple; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.3–2.7 mm. Body reddish brown, antennae, maxillary palpi, elytra, and legs paler (Fig. 45b). Head: Male head round, widest across eyes (Fig. 45b). Male antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate, 4–11 bearing tubercles. Female antennomeres 2–3 subquadrate, 4 longer than wide, 5–7 subquadrate, 8–10 weakly transverse, 4–11 bearing tubercles. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea round partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, one-half of temple. Thorax: Prosternum as long as wide, widest at one-third. Elytra rectangular (Fig. 45b), female slightly shorter. Meso-and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of oval or transverse weak patches of microtrichia, patches absent in female. Aedeagus: Median lobe elongate and bent to right (Fig. 45g –h). Phallobase of median lobe symmetrical and rounded (Fig. 45g –h). Parameres symmetrical and long with setae apically (Fig. 45g –h).</p><p>Type Locality. Mount Oakden, Mid Canterbury, New Zealand .</p><p>Distribution. Buller, Fiordland, Marlborough, Mid Canterbury, Mackenzie, Nelson, Otago Lakes (Fig. 46: triangles).</p><p>Habitat. Most specimens of this species were collected by sifting leaf and moss litter.</p><p>Comments. We could not examine any of the syntypes of S. planipennis because they are apparently missing from the collection at the Natural History Museum, London. We based our concept of this species on two specimens from the type locality that were identified by T. Broun, who described this species, as S. planipennis .</p></div>	https://treatment.plazi.org/id/03D82A3EBD26111C3EBBF9C1FB9AFA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD39111D3E86FAB0FDA0FC3E.text	03D82A3EBD39111D3E86FAB0FDA0FC3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola townsendi Park and Carlton	<div><p>Sagola townsendi Park and Carlton, new species</p><p>(Figs. 45c, 45i, 46)</p><p>Type Material. Holotype. NEW ZEALAND: South Canterbury: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: SC: Mt. Dalgety, 1737m 19 I 1966, G.W. Ramsay J.I. Townsend, moss 66/36”, “ HOLOTYPE Sagola townsendi Park and Carlton 2013 ”.</p><p>Etymology. This species is named for one of the collectors of the holotype, James I. Townsend.</p><p>Diagnosis. This species can be distinguished from the other species of the anisarthra species-group by the following combination of characters: body length 2.8 mm; antennomeres 4–11 bearing tubercles; frontal sulcus shallow, reaching midpoint of eye; anterior frontal fovea round partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, one-half length of temple; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.8 mm. Body reddish brown and antenna, elytra, legs, and maxillary palpi paler (Fig. 45c). Head: Round, widest across eyes (Fig. 45c). Antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third. Elytra rectangular (Fig. 45c). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of oval or transverse weak patches of microtrichia. Aedeagus: Median lobe broad with triangular apical lobe (Fig. 45i). Phallobase of median lobe symmetrical and rounded (Fig. 45i). Parameres symmetrical and long with setae apically (Fig. 45i).</p><p>Distribution. South Canterbury (Fig. 46: black square).</p><p>Habitat. The holotype was collected in moss.</p></div>	https://treatment.plazi.org/id/03D82A3EBD39111D3E86FAB0FDA0FC3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD38111D3CF2FC3CFCDDFD2F.text	03D82A3EBD38111D3CF2FC3CFCDDFD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola wairarapaensis Park and Carlton	<div><p>Sagola wairarapaensis Park and Carlton, new species</p><p>(Figs. 45d, 45j, 46)</p><p>Type Material. Holotype. NEW ZEALAND: Wairarapa: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand WA Putangirua Stm Palliser Bay 23-Oct- 95”, “ In forest leaf litter”, “ HOLOTYPE Sagola wairarapaensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the type material, Wairarapa.</p><p>Diagnosis. This species can be distinguished from the other species of the anisarthra speciesgroup by the following combination of characters: body length 2.8 mm; antennomeres 4–11 bearing tubercles; frontal sulcus shallow, reaching midpoint of eye; anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, one-half length of temple; male abdominal ventrite VII emarginate and surrounded by dense setae; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.7 mm. Body yellowish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 45d). Head: Head round, widest across eyes (Fig. 45d). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–7 subquadrate, 9–10 transverse, 4–11 bearing tubercles. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third. Elytra approximately triangular (Fig. 45d). Hind wings reduced to small pads. Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV without patches of microtrichia. Abdominal ventrite VII deeply emarginate and surrounded dense setae. Aedeagus: Median lobe elongate triangular and broader than parameres (Fig. 45j). Phallobase of median lobe symmetrical and rounded (Fig. 45j). Parameres symmetrical and long with setae apically (Fig. 45j).</p><p>Distribution. Wairarapa (Fig. 46: star).</p><p>Habitat. The holotype was collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD38111D3CF2FC3CFCDDFD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD38111E3EF2FD13FE8FFC9F.text	03D82A3EBD38111E3EF2FD13FE8FFC9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola clarkei Park and Carlton	<div><p>Sagola clarkei Park and Carlton, new species</p><p>(Figs. 45e, 45k, 46)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: NN: Kahurangi N.P., Cobb Dam Rd., Asbestos Track, 450m, 41″06.333′S, 172″43.174′E, 29 XI-18 XII 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest; FMHD#2005- 111, berl., leaf &amp; log litter, A. Solodovnikov, D. Clarke, et al.; ANMT site 1160”, “ HOLOTYPE Sagola clarkei Park and Carlton 2013 ” . Paratypes (n = 9: 2 males; 7 females). NEW ZEALAND: Nelson: 2♂♂ 3♀♀ (1♂, slide-mounted; FMNH), same data as holotype; 4♀♀ (FMNH), 20km ne Takaka, Tasman NP, 21 V 1982, FMHD#82-591, mixed forest litter, S. Peck .</p><p>Etymology. This species is named for one of the collectors of the holotype and staphylinid specialist, Dave J. Clarke.</p><p>Diagnosis. This species can be distinguished from the other species of the anisarthra speciesgroup by the following combination of characters: body length 2.9–3.2 mm; antennomeres 4–11 bearing tubercles; frontal sulcus shallow, reaching midpoint of eye; anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, one-third length of temple; male abdominal ventrite VII deeply emarginate and surrounded by dense setae; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.9–3.2 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 45e). Head: Male head bluntly triangular, widest across eyes (Fig. 45e). Antennomere 1 approximately 2 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third. Male elytra rectangular (Fig. 45e), female elytra triangular. Male hind wings well-developed, female hind wings reduced to small pads. Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV without patches of microtrichia. Male abdominal ventrite VII deeply emarginate and surrounded by dense setae. Aedeagus: Median lobe elongate as wide as parameres (Fig. 45k). Phallobase of median lobe symmetrical and rounded (Fig. 45k). Parameres symmetrical and C-shaped with setae apically (Fig. 45k).</p><p>Distribution. Nelson (Fig. 46: white circles).</p><p>Habitat. Specimens of this species were collected by sifting forest litter in broadleaf or podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD38111E3EF2FD13FE8FFC9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3B111E3D6CFCDEFB7EFE1C.text	03D82A3EBD3B111E3D6CFCDEFB7EFE1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola socia Broun 1915	<div><p>19. socia species-group</p><p>(5 species)</p><p>Diagnosis. Members of the socia species-group can be distinguished from other Sagola speciesgroups by the following combination of characters: body length 2.2–2.8 mm; antennomere 1 approximately 1.5–2.0 times longer than wide; anterior frontal fovea oval and mostly covered by frontal rostrum, posterior frontal fovea oval (Fig. 47k); hind wings well-developed; abdominal tergites IV–VI with discal carinae; median lobe of genitalia divided vertically, major lobe partially covering minor lobe (Fig. 47f–j).</p><p>KEY TO SPECIES OF THE SOCIA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Antennomere 7 enlarged, at least twice as long as 8..................................................2</p><p>1′. Antennomere 7 normal or larger but not exceeding twice length of 8....................3</p><p>2(1). Elytra rectangular, hind wings fully developed ..................... S. socia Broun</p><p>2′. Elytra subquadrate, hind wings reduced to small pads.................................................. ................ S. kahurangiensis new species</p><p>3(1′). Antennomeres 8–10 enlarged, bearing distinctive tubercles; ventral head simple ......4</p><p>3′. Antennomeres 8–10 normal; ventral surface of head with inverted triangular process bearing dense setae behind mouthparts ......... S. canterburyensis new species</p><p>4(3). Eye one-third length of temple; antennomeres 5–10 normal.................................... .................. S. solodovnikovi new species</p><p>4′. Eye as long as temple; antennomeres 5–10 enlarged ......... S. bullerensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD3B111E3D6CFCDEFB7EFE1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3B11183F76FE5FFE61FB01.text	03D82A3EBD3B11183F76FE5FFE61FB01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola socia Broun 1915	<div><p>Sagola socia Broun, 1915</p><p>(Figs. 47a, 47f, 47k, 48)</p><p>Sagola socia Broun, 1915: 281 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.</p><p>Sagola unicalis Broun, 1917: 376 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “type” [red label, printed], “3695. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Pudding Hill. 10.12.1912.” [white label, handwritten]; “ Sagola socia . ♂.” [white label, handwritten]. One of two syntypes of Sagola unicalis . NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “3826. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Moa Basin 20.10.1913.” [white label, handwritten]; “ Sagola unicalis ” [white label, handwritten].</p><p>Additional Material (n = 9: 8 males; 1 female). NEW ZEALAND: Fiordland: 1♂, Barrier River, 1067m, little Red Hill, 2 II 1975, G.W. Ramsay, litter 75/50 ; 1♂, W. Olivine Ra., Tempest Spur Summit, 1463m, 30 I 1975, G.W. Ramsay, litter 75/46 ; 1♂, Middle basin, Tutoko bench, Darran mts, 1524m, 14 I 1977, J.S. Dugdale, swards 77/9; Mid Canterbury: 2♂♂ (1♂, slide-mounted), Arthurs Pass, Temple Basin, 1493m, 8 II 1982, J.S. Dugdale, sedges and mats 82/24 ; Westland: 1♂ 1♀, Mt Aspiring NP, Arawata Biv, 840m, 5 II 1989, LCNZ 89 /2, J.W. Early, R. M. Emberson, scrub litter ; 1♂, Chancellor Hut, Fox Glacier, 29 I 2005, J. Nunn, in tussock litter at 1140m ; 1♂, Alex Knob, 1280m, nr Franz Josef, 6 II 1984, J.C. Watt, turf 84/15 .</p><p>Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: larger body, length 2.5–2.8 mm; male antennomere 7 enlarged and longer than wide; eye prominent, one-third length of temple; male hind wings welldeveloped, female hind wings reduced to small pads; shape of genitalia unique to species.</p><p>Redescription. Length 2.5–2.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 47a). Head: As long as wide, widest across eyes (Fig. 47k). Ventral surface of head weakly convex. Male antennomere 1 approximately 2 times longer than wide, 2–6 subquadrate, 4–7 enlarged, 7 distinctly enlarged and longer than wide, 8–10 subquadrate, 4–11 bearing tubercles. Female antennomeres 2–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus deep, reaching posterior end of eye (Fig. 47k). Anterior frontal fovea oval and partially covered by frontal rostrum (Fig. 47k), posterior frontal fovea oval (Fig. 47k). Eye large and prominent, approximately one-third length of temple (Fig. 47k). Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 47a), female elytra subquadrate. Male hind wings well-developed, female hind wings reduced to small pads. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided vertically, minor lobe broader and triangular apically (Fig. 47f). Phallobase of median lobe symmetrical and rounded (Fig. 47f). Parameres symmetrical with setae apically (Fig. 47f).</p><p>Type Locality. Pudding Hill, near Methven, Mid Canterbury, New Zealand .</p><p>Distribution. F i o r d l a n d, M i d C a nt e r b ur y, Westland (Fig. 48: black circles).</p><p>Habitat. Specimens of this species were collected by sifting forest litter at high elevations.</p><p>Comments. We could not examine one of the two syntypes of S. unicalis, but they share a unique enlarged male antennomere 7 that is longer than wide, a diagnostic species character for S. socia . One specimen was collected at the same locality as S. socia and was identified as S. unicalis by T. Broun, the author of the species, and additional specimens have been collected near the type locality. For these reasons, we have placed S. unicalis in synonymy with S. socia .</p></div>	https://treatment.plazi.org/id/03D82A3EBD3B11183F76FE5FFE61FB01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3D11183C96FB4DFBB2FAA9.text	03D82A3EBD3D11183C96FB4DFBB2FAA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola solodovnikovi Park and Carlton	<div><p>Sagola solodovnikovi Park and Carlton, new species</p><p>(Figs. 47b, 47g, 48)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: NN: Kahurangi N.P., Cobb Dam Rd., Asbestos Track, 450m, 41″06.333′S, 172″43.174′E, 29 XI–18 XII 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest; FMHD#2005-056, flight intercept trap, A. Newton, M. Thayer &amp; A. Solodovnikov; ANMT site 1160”, “HOLOTYPE Sagola solodovnikovi Park and Carlton 2013 ” . Paratypes (n = 7: 3 males; 4 females). NEW ZEALAND: Nelson: 1♂ 1♀ (NZAC), Upper Maitai, 19 V 1941, E.S. Gourlay ; 1♂ 2♀♀ (NZAC), Upper Maitai, 13 II 1957, E.S. Gourlay ; 1♀ (NZAC), Upper Maitai, 7 III 1949, E.S. Gourlay ; 1♂ (slide-mounted; FMNH), Kahurangi NP, Cobb Dam Rd, Asbestos tr, 450m, 41″06.333′S 172″43.174′E, 29 XI–18 XII 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp forest, FMHD #2005-058, pitfall trap, A. Solodovnikov, D. Clarke, ANMT site 1160.</p><p>Etymology. This species is named for the collector of the holotype and staphylinid specialist, Alexey Y. Solodovnikov.</p><p>Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: body larger, length 2.4–2.6 mm; male antennomeres 8–10 enlarged with tubercles; eye prominent, one-third length of temple; male hind wings well-developed, female hind wings reduced to small pads; shape of genitalia unique to species.</p><p>Description. Length 2.4–2.6 mm. Body brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 47b). Head: Male head round, widest across eyes (Fig. 47b). Ventral surface of head weakly convex. Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–7 subquadrate, 8–10 transverse, male 8–10 enlarged with tubercles. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 47b), female elytra subquadrate. Male hind wings well-developed, female hind wings reduced to small pads. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided vertically, minor lobe broader and wave-shaped apically (Fig. 47g). Phallobase of median lobe symmetrical and rounded (Fig. 47g). Parameres symmetrical with setae apcially (Fig. 47g).</p><p>Distribution. Nelson (Fig. 48: triangles).</p><p>Habitat. Specimens of this species were collected using flight intercept or pitfall traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3D11183C96FB4DFBB2FAA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3D11193EA4FAAEFE56FBC4.text	03D82A3EBD3D11193EA4FAAEFE56FBC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola kahurangiensis Park and Carlton	<div><p>Sagola kahurangiensis Park and Carlton, new species</p><p>(Figs. 47c, 47h, 48)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: NN: Kahurangi N.P., track Mt. Arthur Hut to Mt. Arthur Summit, 1550m, 41″13′S, 172″42′E, 19 xii 2005, subalpine area; FMHD#2005-115, berl., sifted tussocks, speargrass &amp; daisy litter, A. Solodovnikov &amp; D. Clarke; ANMT site 1182”, “ HOLOTYPE Sagola kahurangiensis Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Nelson: 1♂ (FMNH), same data as holotype (FMNH) .</p><p>Etymology. This species is named after the type locality, Kahurnagi National Park, Nelson.</p><p>Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: larger body, length 2.5–2.6 mm; male antennomere 7 enlarged, as long as wide; eye prominent, onethird length of temple; male hind wings reduced to small pads; shape of genitalia unique to species.</p><p>Description of Male. L e n g t h 2.5– 2.6 m m. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 47c). Head: Head round, widest across eyes (Fig. 47c). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–10 subquadrate, 7 enlarged, 4–11 bearing tubercles. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 47c). Hind wings reduced to small pads. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe divided vertically, major lobe narrower than minor lobe (Fig. 47h). Phallobase of median lobe symmetrical and rounded (Fig. 47h). Parameres symmetrical with setae apically to middle (Fig. 47h).</p><p>Distribution. Nelson (Fig. 48: black square).</p><p>Habitat. Specimens of this species were collected by sifting tussocks, speargrass, and daisy litter in subalpine areas.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3D11193EA4FAAEFE56FBC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3C11193CD2FB7AFC5BFD2F.text	03D82A3EBD3C11193CD2FB7AFC5BFD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola bullerensis Park and Carlton	<div><p>Sagola bullerensis Park and Carlton, new species</p><p>(Figs. 47d, 47i, 48)</p><p>Type Material. Holotype. NEW ZEALAND: Buller: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “Rotoiti 25-12-15”, “ A.E.Brookes Collection ”, “ T.Broun Collection”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola bullerensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the type material, Buller.</p><p>Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: body larger, length 2.3 mm; male antennomeres 5–10 transverse and enlarged with tubercles; eye large and prominent, as long as temple; male hind wings reduced to small pads; shape of genitalia unique to species.</p><p>Description of Male. Length 2.3 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 47d). Head: Head round, widest across eyes (Fig. 47d). Antennomere 1 approximately 1.5 times longer than wide, 2–4 subquadrate, 5–10 transverse and enlarged with tubercles. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, as long as temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 47d). Hind wings reduced to small pads. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe divided vertically, minor lobe as long as wide (Fig. 47i). Phallobase of median lobe symmetrical and rounded (Fig. 47i). Parameres symmetrical, apical lobe triangular with setae (Fig. 47i).</p><p>Distribution. Buller (Fig. 48: star).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3C11193CD2FB7AFC5BFD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3C111A3E8FFD13FEFDFE36.text	03D82A3EBD3C111A3E8FFD13FEFDFE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola canterburyensis Park and Carlton	<div><p>Sagola canterburyensis Park and Carlton, new species</p><p>(Figs. 47e, 47j, 48)</p><p>Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, MC Prices Valley 5-24. iii. 1981 J.W. Early Malaise trap ”, “ HOLOTYPE Sagola canterburyensis Park and Carlton 2013 ”.</p><p>Etymology. The species name is based on the biotic region of the type material, Mid Canterbury.</p><p>Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: body larger, length 2.2 mm; ventral surface of male head with reversed triangular process bearing dense setae behind mouthparts; eye large and prominent, slightly longer than temple; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.2 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 47e). Head: As long as wide, widest across eyes (Fig. 47e). Ventral surface of head with reversed triangular process bearing dense setae behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–8 subquadrate, 9–10 transverse, 4–11 bearing tubercles. Frontal sulcus reaching one-third length of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, slightly longer than temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 47e). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided vertically, minor lobe as long as wide (Fig. 47j). Phallobase of median lobe symmetrical and rounded (Fig. 47j). Parameres symmetrical, apical lobe rectangular with setae (Fig. 47j).</p><p>Distribution. Mid Canterbury (Fig. 48: white circle).</p><p>Habitat. The holotype was collected using a Malaise trap.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3C111A3E8FFD13FEFDFE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3F111B3D6FFE3AFDABFEDB.text	03D82A3EBD3F111B3D6FFE3AFDABFEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>20. gilae species-group</p><p>(2 species)</p><p>Diagnosis. Members of the gilae species-group can be distinguished from other Sagola speciesgroups by the following combination of characters: body length 1.8–2.3 mm; antennomere 1 approximately 2 times longer than wide with dull surface; anterior frontal fovea oval and partially covered by frontal rostrum, posterior frontal fovea oval (Fig. 49e); hind wings well-developed; abdominal tergites IV–VI with discal carinae; parameres divided (Fig. 49c–d).</p><p>KEY TO SPECIES OF THE GILAE SPECIES- GROUP</p><p>The key is based on male specimens because female specimens are unknown.</p><p>1. Frontal rostrum bluntly rectangular (Fig. 49a); gular depression without process (Fig. 49f); median lobe of genitalia bent to left and parameres deeply divided (Fig. 49c) ........... .................................... S. gilae new species</p><p>1′. Frontal rostrum bluntly triangular; gular depression with small process from anterior middle; median lobe of genitalia straight and right paramere shallowly divided (Fig. 49d) .......... ............................. S. tishechkini new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD3F111B3D6FFE3AFDABFEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3E111B3F5CFB00FB8DF9B8.text	03D82A3EBD3E111B3F5CFB00FB8DF9B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>21. owensae species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the owensae species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 1.9 mm; antennomere 1 approximately 2 times longer than wide; ventral surface of male head with a pair of cone-shaped temporal depressions and small triangular process medially (Fig. 50c: arrows); hind wings well-developed; abdominal tergites IV–VI with discal carinae.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3E111B3F5CFB00FB8DF9B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3E111B3CEEFE9FFDD5F9B8.text	03D82A3EBD3E111B3CEEFE9FFDD5F9B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola gilae Park and Carlton	<div><p>Sagola gilae Park and Carlton, new species</p><p>(Figs. 49a, 49c, 49e–f, 51)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (LUNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “Beech litter, Desert Rd, 10ml. N. Waioru 17. ii. 1973 R. M. Emberson ”, “ HOLOTYPE Sagola gilae Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Waikato: ♂ (slide-mounted; NZAC), Waikato-Haipakihi Junction, 3 km s of Turangi, 915m, 19 II 1979, L. A. Moond, litter 79/22.</p><p>Etymology. This species is named for LSAM alumna, database specialist, and enthusiastic supporter of this study, Stephanie Gil.</p><p>Diagnosis. This species can be distinguished from the other species of the gilae species-group by the following combination of characters: body length 2.3 mm; frontal rostrum bluntly rectangular; gular region transversely depressed; shape of genitalia unique to species.</p><p>Description of Male. Length 2.3 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 49a). Head: Round, widest across eyes. Gular region transversely depressed with dense setae (Fig. 49f). Antennomere 1 approximately 2.5 times longer than wide with dull surface, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum bluntly rectangular (Fig. 49a). Frontal sulcus deep, slightly exceeding eye (Fig. 49e). Anterior frontal fovea oval and partially covered by frontal rostrum (Fig. 49e), posterior frontal fovea oval (Fig. 49e). Eye prominent, one-third length of temple (Fig. 49e). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 49a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe bent left with pointed apical lobe (Fig. 49c). Phallobase of median lobe asymmetrical and triangular (Fig. 49c). Parameres asymmetrical and deeply divided (Fig. 49c).</p><p>Distribution. Waikato (Fig. 51: black circle).</p><p>Habitat. Specimens of this species were collected by sifting beech and leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3E111B3CEEFE9FFDD5F9B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3E111B3CD2F9BEFCDDFB53.text	03D82A3EBD3E111B3CD2F9BEFCDDFB53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola tishechkini Park and Carlton	<div><p>Sagola tishechkini Park and Carlton, new species</p><p>(Figs. 49b, 49d, 51)</p><p>Type Material. Holotype. NEW ZEALAND:</p><p>Gisborne: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: GB: Lake Waikaremoana 17 I 1972, G.W. Ramsay Litter 72/21”, “ HOLOTYPE Sagola tishechkini Park and Carlton 2013 ” .</p><p>Etymology. This species is named for LSAM alumnus, histerid specialist, and enthusiastic supporter of this study, Alexey K. Tishechkin.</p><p>Diagnosis. This species can be distinguished from the other species of the gilae species-group by the following combination of characters: body smaller, length 1.8 mm; frontal rostrum bluntly triangular; gular depression with small process from anterior middle; shape of genitalia unique to species.</p><p>Description of Male. Length 1.8 mm. Body brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 49b). Head: As long as wide, widest across eyes (Fig. 49b). Gular region transversely depressed with small process medially and bearing dense setae. Antennomere 1 approximately 2.5 times longer than wide with dull surface, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum bluntly triangular. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval and partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 49b). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe straight and slender (Fig. 49d). Phallobase of median lobe asymmetrical and rounded (Fig. 49d). Parameres asymmetrical, left divided and apical lobe of major lobe arrow head-shaped, right shallowly divided (Fig. 49d).</p><p>Distribution. Gisborne (Fig. 51: triangle).</p><p>Habitat. The holotype was collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3E111B3CD2F9BEFCDDFB53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3E11143EEAF9BEFC92FC10.text	03D82A3EBD3E11143EEAF9BEFC92FC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola owensae Park and Carlton	<div><p>Sagola owensae Park and Carlton, new species</p><p>(Figs. 50–51)</p><p>Type Material. Holotype. NEW ZEALAND: Wairarapa: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND MB Pongaroa 31 Dec 1980 J.C. Watt. Wood mould 80/158”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola owensae Park and Carlton 2013 ” .</p><p>Etymology. This species is named for LSAM alumna, pselaphine specialist, and an enthusiastic supporter of this study, Brittany Owens.</p><p>Diagnosis. This species can be distinguished from other species by the following combination of characters: body length 1.9 mm; head bluntly transverse; ventral surface of male head with a pair of cone-shaped temporal depressions and small triangular process medially; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 1.9 mm. Body yellowish brown, maxillary palpi, elytra, and legs paler (Fig. 50a). Head: Bluntly transverse, widest across eyes (Fig. 50a). Ventral surface with pair of cone-shaped temporal depressions and small triangular process in center (Fig. 50c: long arrow). Antennomere 1 approximately 2 times longer than wide, 2–5 longer than wide, 6–10 subquadrate. Frontal rostrum prominent and covering anterior frontal fovea. Frontal sulcus reaching behind eye, posterior frontal sulcus deep and oval. Eye large and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 50a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe twisted with pointed apical lobe (Fig. 50b). Phallobase of median lobe symmetrical and rounded (Fig. 50b). Parameres symmetrical, apical lobe broad and round with long and dense setae (Fig. 50b).</p><p>Distribution. Wairarapa (Fig. 51: black square).</p><p>Habitat. The holotype was collected by sifting moldy wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3E11143EEAF9BEFC92FC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3011163D4CFF31FD44FEFE.text	03D82A3EBD3011163D4CFF31FD44FEFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola furcata Broun 1921	<div><p>22. furcata species-group</p><p>(6 species)</p><p>Diagnosis. Members of the furcata speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body small, length 1.8–2.2 mm; frontal sulcus inverted Y- or V-shaped, divided with each posterior end meeting vertexal fovea (Fig. 52o, q); ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts (Fig. 52p, r); hind wings fully developed; abdominal tergites IV–VI with discal carinae; male abdominal ventrites V–VI weakly depressed or flattened medially and bearing dense setae; only known from South Island (Fig. 53).</p><p>KEY TO SPECIES OF THE FURCATA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Frontal sulcus inverted V-shaped (Fig. 52q); eye large, as long as temple (Fig. 52q); antennomeres 4–5 subquadrate..............2</p><p>1′. Frontal sulcus inverted Y-shaped (Fig. 52o); eye small, approximately one-third length of temple (Fig. 52o); antennomeres 4–5 longer than wide ..................................................... 3</p><p>2(1). Apical lobe of genitalia with V-shaped depression (Fig. 52k)...... S. ignota Broun</p><p>2′. Apical lobe of genitalia blunt apically (Fig. 52l–m)..... S. lewisensis new species</p><p>3(1′). Median lobe of genitalia at least 3 times broader than paramere (Fig. 52g) ............. ...................................... S. furcata Broun</p><p>3′. Median lobe of genitalia as wide as paramere................................................4</p><p>4(3′). Median lobe of genitalia twisted (Fig. 52h) .............................. S. punctulata Raffray</p><p>4′. Median lobe of genitalia curved vertically .................................................................5</p><p>5(4′). Only apical part of median lobe of genitalia hook-shaped (Fig. 52i)......................... ................................ S. pecki new species</p><p>5′. Entire lobe of median lobe of genitalia hook-shaped (Fig. 52j) .............................. ............................. S. muirae new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD3011163D4CFF31FD44FEFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3311163D56FE62FB7CFCCF.text	03D82A3EBD3311163D56FE62FB7CFCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola furcata Broun 1921	<div><p>Sagola furcata Broun, 1921</p><p>(Figs. 52a, 52g, 53)</p><p>Sagola furcata Broun, 1921a: 495 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (BMNH), glued on rectangular card, “ 4005. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922– 4 82.” [white l a b e l, p r i n t e d]; “ B e l g r o v e. 24.10.1914.” [white label, handwritten]; “ Sagola ♂ furcata.” [white label, handwritten].</p><p>Additional Material (n = 14: 10 males; 4 females). NEW ZEALAND: Nelson: 1♂ 1♀, 20km nw Motueka, Riwaka River Res, 28 V 1982, FMHD#2005-605, mixed forest litter, S. Peck ; 2♂♂, Slaters Road, 0.7km s Whangamoa Saddle, 410m, 13 XII 1984 – 4 I 1985, Nothofagus forest, A. Newton, M. Thayer, 703, FIT &amp; window trap ; 1♂ 1♀, Dun Track, 14 II 1942, E.S. Gourlay ; 1♂, Dun Mt., 1850’, 4 IV 1966, J.C. Watt, fungus in Nothofagus forest ; 1♀, Dun Mt., 2000′, 11 IV 1943, E.S. Gourlay ; 1♂, Upper Maitai, 19 X 1941, E.S. Gourlay ; Marlborough Sounds: 1♂ 1♀, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton, M. Thayer, litter ; 1♂, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton, M. Thayer, window trap ; 1♂, Tennyson Inlet, e side Duncan Bay, 30m, 15 XII 1984 – 5 I 1985, podo- Nothofagus forest, A. Newton, M. Thayer 709, FIT &amp; window trap ; 1♂, Port Underwood saddle, 17 XII 1968, J.C. Watt .</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters: antennomeres 4–6 longer than wide; frontal sulcus inverted Y-shaped; eye prominent, approximately one-third length of temple; male abdominal ventrites V–VI weakly depressed medially and surrounded by dense setae; shape of genitalia unique to species.</p><p>Redescription. Length 1.8–2.1 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52a). Head: Weakly transverse, widest across eyes (Fig. 52a). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus inverted Y-shaped. Anterior frontal fovea rectangular covered by frontal rostrum, posterior frontal fovea absent. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI weakly depressed on middle with dense setae. Aedeagus: Median lobe of genitalia broader with hook-shaped lobe at apex (Fig. 52g). Phallobase of median lobe symmetrical and rounded (Fig. 52g). Parameres symmetrical with serrate outer edge of apical lobe and setae inside (Fig. 52g).</p><p>Type Locality. Belgrove, near Nelson, New Zealand .</p><p>Distribution. Nelson, Marlborough Sounds (Fig. 53: black circles).</p><p>Habitat. Specimens of this species were collected using flight intercept and window traps or by sifting leaf litter in podocarp or Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3311163D56FE62FB7CFCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3211173CA3FF31FCA3FD2F.text	03D82A3EBD3211173CA3FF31FCA3FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola punctulata Raffray 1893	<div><p>Sagola punctulata Raffray, 1893</p><p>(Figs. 52b, 52h, 52o–p, 53)</p><p>Sagola punctulata Raffray, 1893: 21 . Raffray 1904: 497, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Type Material. Two of three syntypes. NEW ZEALAND: Nelson: 2♂ ♀ (MNHN), glued on rectangular card, “Nov. Zea”, “ MP Type punctulata AR ”.</p><p>Additional Material (n = 20: 16 males; 4 females). NEW ZEALAND: Buller: 1♂, 18km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 718, litter ; 3♂♂, Tawhai SF, 6 III 1972, J. McBurney, litter 72/125 ; 1♂, Tawhai SF, 6 III 1972, J.S. Dugdale, litter 72/121 ; Nelson: 1♂, Kongahu, XI 1980, J. Jones, Malaise trap near swamp ; 1♂ (slide-mounted), 0.6km e Gowanbrige, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &amp; window trap ; Westland: 5 ♂ ♂ 1♀ (1♂, s l i d e -m o u n t e d), 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, FIT &amp; window trap ; 2♂♂ 1♀, 1.8km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwdnikau forest, A. Newton, M. Thayer 718, FIT &amp; window trap ; 1♂ 1♀, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap ; 1♂, Okuku Scen Res, 2 VIII 1981, R. M. Emberson, litter LUNZ 81 /6 ; 1♀, Lake Kaniere, 6 II 1984, J.C. Watt, dead tree fern .</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters: antennomeres 4–7 longer than wide; frontal sulcus inverted Y-shaped; eye prominent, approximately one-third length of temple; male abdominal ventrites V–VI weakly depressed medially and surrounded by dense setae; shape of genitalia unique to species.</p><p>Redescription. Length 1.9–2.2 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52b). Head: Weakly transverse, widest across eyes (Fig. 52o). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts (Fig. 52p). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus inverted Y-shaped (Fig. 52o). Anterior frontal fovea rectangular, covered by frontal rostrum (Fig. 52o), posterior frontal fovea absent (Fig. 52o). Eye prominent, approximately onethird length of temple (Fig. 52p). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52b). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI weakly depressed on middle with dense setae. Aedeagus: Median lobe of genitalia slender and twisted (Fig. 52h). Phallobase of median lobe symmetrical and rounded (Fig. 52h). Parameres symmetrical with serrate outer edge of apical lobe and setae inside (Fig. 52h).</p><p>Type Locality. “ New Zealand ” (specific locality unknown).</p><p>Distribution. Buller, Nelson, Westland (Fig. 53: triangles).</p><p>Habitat. Most specimens of this species were collected using flight intercept and window traps or by sifting leaf litter in hardwood, nikau, podocarp, or Nothofagus forests. One specimen was collected from dead tree ferns.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3211173CA3FF31FCA3FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3211103EFAFD13FDD2FDAA.text	03D82A3EBD3211103EFAFD13FDD2FDAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola pecki Park and Carlton	<div><p>Sagola pecki Park and Carlton, new species</p><p>(Figs. 52c, 52i, 53)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: So. Island, 13 km W Collingwood, Mangarakau, 20 V 1982, FMHD#82-589, mixed forest litter, S. Peck ”, “ HOLOTYPE Sagola pecki Park and Carlton 2013 ” . Paratypes (n = 2: 1 male; 1 female). NEW ZEALAND: Nelson: 2♀♀ (1♀, slide-mounted; FMNH), same data as holotype .</p><p>Etymology. This species is named for the collector of the holotype and world-renown beetle specialist, Stewart B. Peck.</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters: antennomeres 4–5 longer than wide; frontal sulcus inverted Y-shaped; eye prominent, approximately one-third length of temple; male abdominal ventrites V–VI weakly depressed medially and surrounded by dense setae; shape of genitalia unique to species.</p><p>Description. Length 1.9–2.1 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52c). Head: Weakly transverse, widest across eyes (Fig. 52c). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–9 subquadrate, 10 weakly transverse. Frontal sulcus inverted Y-shaped. Anterior frontal fovea rectangular covered by frontal rostrum, posterior frontal fovea absent. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52c). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI weakly depressed on middle with dense setae. Aedeagus: Median lobe of genitalia slender with hook-shaped apical lobe (Fig. 52i). Phallobase of median lobe symmetrical and rounded (Fig. 52i). Parameres symmetrical with serrate outer edge of apical lobe and setae inside (Fig. 52i).</p><p>Distribution. Nelson (Fig. 53: black square).</p><p>Habitat. Specimens of this species were collected by sifting mixed forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3211103EFAFD13FDD2FDAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3511103CF0FDA8FBAAFE36.text	03D82A3EBD3511103CF0FDA8FBAAFE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola muirae Park and Carlton	<div><p>Sagola muirae Park and Carlton, new species</p><p>(Figs. 52d, 52j, 53)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (LUNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, NN Mt Burnett 450m 8.ii.1981 C.A. Muir sweeping ferns”, “ HOLOTYPE Sagola muirae Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Nelson: 1♂ (slidemounted; NZAC) Whanganui Inlet, Kaihoka Lakes Tr, 17 XII 2007, K. Marske, J. Allwood. Hand collected ex fungus-encrusted dead wood, S 40.33.220′, E 172.36.247′, 52m.</p><p>Etymology. This species is named for the collector of the holotype, Carol A. Muir.</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters: antennomeres 4–5 longer than wide; frontal sulcus inverted Y-shaped; eye prominent, approximately one-third length of temple; male abdominal ventrites V–VI weakly depressed medially and surrounded by dense setae; shape of genitalia unique to species.</p><p>Description of Male. Length 1.8–2.0 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52d). Head: Weakly transverse, widest across eyes (Fig. 52d). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus inverted Y-shaped. Anterior frontal fovea rectangular, covered by frontal rostrum, posterior frontal fovea absent. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52d). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI weakly depressed on middle with dense setae. Aedeagus: Median lobe of genitalia slender and curved, C-shaped (Fig. 52j). Phallobase of median lobe symmetrical and rounded (Fig. 52j). Parameres symmetrical with enlarged apical lobe with serrate midline and setae (Fig. 52j).</p><p>Distribution. Nelson (Fig. 53: stars).</p><p>Habitat. Specimens of this species were collected from tree ferns or fungusy dead wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3511103CF0FDA8FBAAFE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3511113F68FE28FDA9FCF5.text	03D82A3EBD3511113F68FE28FDA9FCF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola ignota Broun 1921	<div><p>Sagola ignota Broun, 1921</p><p>(Figs. 52e, 52k, 52n, 52q–r, 53)</p><p>Sagola ignota Broun, 1921a: 495 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Otago Lakes: 1♂ (BMNH), aedeagus dissected and mounted in balsam on a clear plastic card, “4004. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Routeburn. 16.2.1914.” [white label, handwritten]; “ Sagola ignota . ♂ ” [white label, handwritten].</p><p>Additional Material (n = 15: 8 males; 7 females). NEW ZEALAND: Dunedin: 1♂, Woodside Glen Outram, 30 XI 2005, J. Nunn, in moss and lichen on tree trunk ; Fiordland: 2♂♂ (1♂, slidemounted), Borland Saddle, c 1050m, 16 XII 1998, in moss on Nothofagus bole, J. Nunn ; 3♀♀, Borland Saddle, 24 I 2008, J. Nunn, sifted moss ; 2♀♀, W. Olivine Ra, Simonin Pass, 23 I 1975, G.W. Ramsay, litter 75/35 ; 1♀, Outer Gilbert 2, Breaksea Sound, Nov 1985, on Kiekie ; 1♂, Fiordland NP, S Borland V Biv, 730m, 2–6 II 1982, J.W. Early, Malaise trap at Nothofagus forest edge ; 1♂, Gertude V, Milford Rd, 23 I 2008, J. Nunn, in moss on beech tree ; 1♂, Barrier Ra, Little Red Hill, Mt. Allison, 4 II 1972, J.S. Dugdale, little 75/52 ; Stewart Island: 1♂, ne Big S. Cape I, 122m, 18 XI 1968, G. Kuschel, litter 68/191 ; 1♀, Island Hill Homestead, 2–8 II 1991, C.J. Vink, J.W. Early, yellow pan trap in bush ; Southland: 1♂, 1km S Longwood Trig, 700m, Longwood Ra, 1 II 1976, L.L. Deitz, mats 76/12 .</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters:antennomeres 4–7 subquadrate; frontal sulcus inverted V-shaped; eye prominent, as long as temple; male abdominal ventrites V–VI flattened roundly, surrounded by dense setae; shape of genitalia unique to species.</p><p>Redescription. Length 1.9–2.2 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52e). Head: Bluntly triangular, widest across eyes (Fig. 52q). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts (Fig. 52r). Antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate (Fig. 52n). Frontal sulcus inverted V-shaped (Fig. 52q). Anterior frontal fovea rectangular, covered by frontal rostrum (Fig. 52q), posterior frontal fovea absent (Fig. 52q). Eye prominent, approximately as long as temple (Fig. 52q). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52e). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI roundly flattened on middle with dense setae. Aedeagus: Median lobe of genitalia weakly curved and long with V-shaped depression (Fig. 52k). Phallobase of median lobe symmetrical and rounded (Fig. 52k). Parameres symmetrical and as long as median lobe, with setae apically (Fig. 52k).</p><p>Distribution. Dunedin, Fiordland, Stewart Island, Southland (Fig. 53: white circles).</p><p>Habitat. Specimens of this species were collected using Malaise and yellow pan traps or by sifting leaf, moss, and leaf litter in Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3511113F68FE28FDA9FCF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3411113F5DFC69FB42F95B.text	03D82A3EBD3411113F5DFC69FB42F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola excavata Broun 1886	<div><p>23. excavata species-group</p><p>(2 species)</p><p>Diagnosis. Members of excavata species-group can be distinguished from other Sagola speciesgroups by the following combination of characters: body small, length 1.9–2.9 mm; frontal sulcus deep and reaching vertexal fovea (Fig. 54e); anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea keyholeshaped (Fig. 54e); hind wings fully developed; abdominal tergites IV–VI with discal carinae; Male tergite V distinctly depressed, with processes from anterior and posterior edge with long and dense setae between processes (Fig. 54a–b, f); only known from North Island (Fig. 55).</p><p>KEY TO SPECIES OF THE EXCAVATA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Larger size, length 2.5–2.9 mm; abdominal tergite V distinctly depressed with triangular processes from anterior and posterior margins (Fig. 54f)......................... S. excavata Broun</p><p>1′. Smaller size, length 1.9–2.2 mm; abdominal tergite V distinctly depressed with rectangular processes from anterior and posterior margins ............ S. aucklandensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD3411113F5DFC69FB42F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3411113CD2FC69FCBFFCF5.text	03D82A3EBD3411113CD2FC69FCBFFCF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola lewisensis Park and Carlton	<div><p>Sagola lewisensis Park and Carlton, new species</p><p>(Figs. 52f, 52l–m, 53)</p><p>Type Material. Holotype. NEW ZEALAND: Buller: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: BR: Lewis, Pass Nat. Res., 11.9km ese Springs Junction, 540m, 17 XII 84 - 21 I 85, Nothofagus spp. forest, A. Newton/M. Thayer 715, window trap”, “HOLOTYPE Sagola lewisensis Park and Carlton 2013 ”. Paratypes (2 males). NEW ZEALAND: Buller: 1♂ (slide-mounted; NZAC), same data as holotype; Nelson: 1♂ (FMNH), Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1159.</p><p>Etymology. This species is named after the type locality, Lewis Pass, Buller.</p><p>Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters: antennomeres 4–7 subquadrate; frontal sulcus inverted V-shaped; eye prominent, as long as temple; male abdominal ventrites V–VI roundly flattened roundly and surrounded by dense setae; shape of genitalia unique to species.</p><p>Description of Male. Length 1.8–2.1 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 52f). Head: Bluntly triangular, widest across eyes (Fig. 52f). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts. Antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate. Frontal sulcus inverted V-shaped. Anterior frontal fovea rectangular, covered by frontal rostrum, posterior frontal fovea absent. Eye prominent, approximately as long as temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 52f). Meso-and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI roundly flattened on middle, with dense setae. Aedeagus: Median lobe of genitalia weakly curved vertically and long with blunt apical lobe (Fig. 52l–m). Phallobase of median lobe symmetrical and rounded (Fig. 52l–m). Parameres symmetrical and as long as median lobe, with setae apically (Fig. 52l–m).</p><p>Distribution. Buller, Nelson (Fig. 53: white squares).</p><p>Habitat. Specimens of this species were collected using flight intercept or window traps in Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3411113CD2FC69FCBFFCF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD3711123D4EFADBFC7BF9E6.text	03D82A3EBD3711123D4EFADBFC7BF9E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola excavata Broun 1886	<div><p>Sagola excavata Broun, 1886</p><p>(Figs. 54a, 54c, 54e–f, 55)</p><p>Sagola excavata Broun, 1886: 884 . Raffray 1893: 35, 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola brevitarsis Broun, 1886: 887 . Raffray 1893: 38, 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola citima Broun, 1893a: 177 . Hudson 1923: 365, 1934: 47; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola parallela Broun, 1893b: 1053 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola cognata Broun, 1911: 494 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola diversa Broun, 1911: 495 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p></div>	https://treatment.plazi.org/id/03D82A3EBD3711123D4EFADBFC7BF9E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD09112C3C94FA8BFC62FB34.text	03D82A3EBD09112C3C94FA8BFC62FB34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola aucklandensis Park and Carlton	<div><p>Sagola aucklandensis Park and Carlton, new species</p><p>(Figs. 54b, 54d, 55)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND AK Lynfield Wattle Bay 8 Oct 1986 G. Kuschel ”, “ Sifted rotten wood 86/21”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola aucklandensis Park and Carlton 2013 ” . Paratypes (n = 2: 1 male; 1 female). NEW ZEALAND: Auckland: 1♀ (NZAC), same data as holotype; 1♂ (NZAC), Waikowhai Park, 5 X 1051, dead wood, P328 .</p><p>Etymology. The species name is based on the biotic region of the type material, Auckland.</p><p>Diagnosis. This species can be distinguished from the other species of the excavata speciesgroup by the following combination of characters: size smaller, length 1.9–2.2 mm; Male tergite V distinctly depressed with rectangular processes from anterior and posterior edges; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 1.9–2.2 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 54b). Head: Round, as long as wide, widest across eyes (Fig. 54b). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 subquadrate, 8–10 transverse. Frontal sulcus deep and reaching vertexal fovea. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea keyhole-shaped. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 54b). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male tergite V distinctly depressed, with rectangular processes from anterior and posterior edges and with long and dense setae between processes. Aedeagus: Median lobe of genitalia slender, longer than parameres (Fig. 54d). Phallobase of median lobe symmetrical and reversed triangular (Fig. 54d). Parameres symmetrical with 3 setae apically (Fig. 54d).</p><p>Distribution. Auckland (Fig. 55: triangle).</p><p>Habitat. Specimens of this species were collected from rotten wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD09112C3C94FA8BFC62FB34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD09112D3F6BFB2AFDA8FA65.text	03D82A3EBD09112D3F6BFB2AFDA8FA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola notabilis Broun Desig. Park and Carlton 2013	<div><p>24. notabilis species-group</p><p>(3 species)</p><p>Diagnosis. Members of the notabilis speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.4–3.0 mm; antennomere 1 approximately twice as long as wide (Fig. 56g); frontal sulcus deep and reaching posterior margin of eye (Fig. 56h); anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea elongate (Fig. 56h); ventral surface of male head distinctly depressed with a pair of processes (Fig. 56i); hind wings fully developed; hind femur as wide as tibia (Fig. 56j); hind claws enlarged (Fig. 56k); abdominal tergites IV–VI with discal carinae; male abdominal ventrites VI–VII with triangular and rectangular depression, respectively (Fig. 56l).</p><p>KEY TO SPECIES OF THE NOTABILIS SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Antennomere 8 enlarged; fore femur with weak semicircular depression; only known from North Island.....................................2</p><p>1′. Antennomeres 4–6 enlarged (Fig. 56g); fore femur simple; only known from South Island (Fig. 57: triangles)........................... ................................. S. monstrosa Reitter</p><p>2(1). Antennomeres 4–7 longer than wide; known from north to midpoint of North Island (Fig. 57: black circles) .................... ..................................... S. notabilis Broun</p><p>2′. Antennomeres 4–7 subquadrate, 4–6 weakly enlarged; only known from Wellington (Fig. 57: black squares)............................ ................... S. tararuaensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD09112D3F6BFB2AFDA8FA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD08112F3CB2F9C1FE69FE61.text	03D82A3EBD08112F3CB2F9C1FE69FE61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola notabilis Broun Desig. Park and Carlton 2013	<div><p>Sagola notabilis Broun, 1880</p><p>(Figs. 56a, 56d, 57)</p><p>Sagola notabilis Broun, 1880: 137 . Schaufuss 1888: 85; Raffray 1893: 32, 1904: 496, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola macronyx Broun, 1893b: 1418 . Raffray 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola electa Broun, 1914a: 91 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola fulvipennis Broun, 1915: 289 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Lectotype. NEW ZEALAND: Coromandel: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “251” [white label, printed]; “Tairua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola notabilis ” [white label, handwritten]; “ LECTOTYPE Sagola notabilis Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotype (1 male). NEW ZEALAND: Coromandel: 1♂ (BMNH), Tairua, New Zealand, Sharp Coll. 1905-313 (BMNH). The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined (Broun 1880: 137). This designation will fix the identity of the species and facilitate its recognition by future workers. Holotype of Sagola electa: NEW ZEALAND: Taupo: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3400. ♂ ” [white label, handwritten]; “Erua. Jany.1911.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola electa . ♂.” [white label, handwritten] . Holotype of Sagola fulvipennis: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3705. ♂ ” [white label, handwritten]; “Tairua. Novr.1875.” [white label, handwritten]; “ Sagola ♂. fulvipennis” [white label, handwritten]. The original description states that the specimen was collected in December 1874, but the single specimen examined was collected in November 1875. Perhaps the date provided in the original description was incorrect . Holotype of Sagola macronyx: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “ ♂ ” [white label, printed]; “977” [white label, printed]; “Hunua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola macronyx ” [white label, handwritten] .</p><p>Additional Material (n = 72: 27 males; 45 females). NE W Z EAL AND: Au ck la nd: 23♂♂ 34♀♀, Lynfield, G. Kuschel, 23 XI 1974 – 29 III 1981, malaise trap or litter; 2♂♂, Waitakere Range, 260m, Nohoanga Scenic Res, 8 XII 1984 – 25 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 679, FIT &amp;window trap; 1♀, Hunua Ranges, Vining Res, Mangatangi tr, 240m, 37″07,747′S, 175″13.024′E, 17 XI–28 XII 2005, Nothofagus - Agathis -Phyllocladus trichomanoides, etc, FMHD #2005-005, FIT, A. Newton, M. Thayer et al, ANMT site 1141; 2♀♀, Orere Saddle, 2 I 1984, J.C. Watt, bark and subcortical debris 84/1; 1♀, Manurewa, Murphy’ s Bush, 2 III 1981, G. Kuschel, rotten wood 81/59; 1♀, Papatoetoe, Murphy’ s Bush, 17 VIII 1978, D.W. Helmore, woodmould 78/204; Bay of Plenty: 1♂, Mt Te Aroha Summit, 1000m, 3 XI 1977, B.M. May, dead trunk; 1♂, Te Aroha, 19 III 1975, G. Kuschel, sifted rotten wood bottom stream; 1♀, Okere Falls Scenic Res, 21 X 1979, J.S. Dugdale, liverworts 79/99; Gisborne: 2♀♀, Gray’ s Bush, 30 VII 1951, under bark; Northland: 2♀♀, Omahuta Kauri Sanctuary, 4 II 1975, G. Kuschel, rotten wood; Rangitikei: 1♀, Ruahine Range, Armstrong Saddle, 10 II 1980, C.F. Butcher, litter 80/11.</p><p>Diagnosis. This species can be distinguished from the other species of the notabilis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; male antennomere 8 enlarged; male fore femur with weak semicircular depression; shape of ventral surface of male head and genitalia unique to species; only known from North Island.</p><p>Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi,and elytra paler (Fig. 56a). Head: Round, as long as wide, widest across eyes (Fig. 56a). Antennomere 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 5–10 subquadrate, 8 enlarged in male. Eye prominent, approximately two-thirds length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 56a). Meso- and metathorax trapezoidal, longer than wide. Male fore femur with shallow, semicircular depression. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia triangular and shorter than parameres (Fig. 56d). Phallobase of median lobe symmetrical and rectangular (Fig. 56d). Parameres symmetrical, setose from midpoints to apices (Fig. 56d).</p><p>Type Locality. Tairua, Coromandel, New Zealand .</p><p>Distribution. Auckland, Bay of Plenty, Coromandel, Gisborne, Northland, Rangitikei (Fig. 57: black circles).</p><p>Habitat. Specimens of this species were collected using Malaise, flight intercept, and window traps or by sifting dead logs and wood litter in hardwood, podocarp, or Nothofagus forests.</p><p>Comments. Male specimens of S. notabilis can be distinguished from those of other species by the enlarged male antennomere 8 and modification of the ventral surface of the male head. The type specimens of S. electa, S. fulvipennis, and S. macronyx share these diagnostic characters, and these species have been collected at or near the type locality. For these reasons, we have placed S. electa, S. fulvipennis, and S. macronyx in synonymy with S. notabilis .</p></div>	https://treatment.plazi.org/id/03D82A3EBD08112F3CB2F9C1FE69FE61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0A11293CA8FDD8FDFCFE96.text	03D82A3EBD0A11293CA8FDD8FDFCFE96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola monstrosa Reitter 1880	<div><p>Sagola monstrosa Reitter, 1880</p><p>(Figs. 56b, 56e, 56g –l, 57)</p><p>Sagola monstrosa Reitter, 1880: 168 . Schaufuss 1888: 84; Raffray 1893: 17, 1904: 497, 1911: 5, 1924: 231; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola nitida Broun, 1911: 492 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola bifoveiceps Broun, 1912b: 629 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola biimpressa Broun, 1912b: 630 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: 1♂ (MNHN), glued on rectangular card, “New Zealand. Helms Reitter”, “15”, “ MP ”, “ monstrosa AR ” . Holotype of Sagola bifoveiceps: NEW ZEALAND: Buller: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “20.” [white label, handwritten], “Greymouth, New Zealand. Helms.” [white label, printed]; “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola ♂. foviecpes” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola biimpressa: NEW ZEALAND: Buller: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “21.” [white label, handwritten], “Greymouth, New Zealand. Helms.” [white label, printed]; “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola ♂. biimpressa” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola nitida: NEW ZEALAND: Buller: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3364. ♂.” [white label, handwritten], “Greymouth, New Zealand. Helms.” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Greymouth.-Lewis.” [white label, handwritten]; “ Sagola nitida ” [white label, handwritten]. The original label indicates this specimen is a male, but it is female.</p><p>Additional Material (n = 92: 40 males; 52 females). NEW ZEALAND: Buller: 1♂ 6♀♀, Nelson Lakes NP, n slope Mt Robert, Pinchgut tr, 950m, 15 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, window trap; 1♂ 1♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &amp;window trap; 1♂ 4♀♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 645m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 705, window trap; 4♀♀, Lewis Pass NR, 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, window trap; 1♂ 2♀♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp;window trap; 2♀♀, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E, 30 XI–17 XII 2005, Nothofagus forest, FMHD #2005-059, FIT, A. Newton, M. Thayer, ANMT site 1161; 1♂ 1♀, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, FIT &amp;window trap; 1♂, Nelson Lakes NP, n slope Mt. Robert, Speargrass tr, 880m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 704, window trap; 1♂, Nelson Lakes NP, Mt. Robert, 660m, 26 XII 1984 – 6 I 1985, Leptospermum - Nothofagus scrub, A. Newton, M. Thayer 722, FIT &amp;window trap; 1♂, Lake Rotoiti, 600m, 8 II 1978, A.K. Walker, ant nest in Nothofagus forest; 1♀, Nelson Lakes NP, Lake Rotoroa, Braeburn tr, 470m, 16 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 712, window trap; 1♀, Nelson Lakes NP, St. Arnaud tr, Lake Rotoiti, 610–650m, 24–26 III 1980, Nothofagus forest, A. Newton, M. Thayer, fogging; 1♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 645m, 14 XII 1984 – 6 I 1985, A. Newton, M. Thayer 705, fogging; Dunedin: 2♂♂, Mt. Cargill, 12 VII 2003, J. Nunn, dead Pseudopanax bole; 1♀, Leith Saddle, Waitati Rd, 17 I 2008, J. Nunn, decayed wood; 1♂, Leith Saddle, 27 I 2001, under bark, J. Nunn; 1♂, Cloud Forest of Leith Tr, 19 VIII 2010, J. Nunn, wood mould and frass under bark dead tree; Fiordland: 1♀, Keplar track start, Te Anau, 22 I 2008, J. Nunn; moss; Mid Canterbury: 3♂♂, Banks Peninsula, Ahuriri Sce Res, 480m, 1 IV 1982, G. Kuschel, litter and rotten wood 82/48; 1♂, McClennans Bush, 12km nw Methven, 560m, 3 III 2010, R. Emberson, sift leaf litter; 1♂, Kowhai Bush, Springfield, 24 IV 2005, J. Nunn, decayed wood; 1♂, Banks Peninsula, Hay Scen Res, Pigeon Bay, 25m, 11 XII 1984 – 22 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 702, window trap; 1♂, Craigieburn SF, Dracophyllum Flat tr, 43′10S, 171′42E, FIT, 11–27 I 1998, C. Carlton, R. Leschen; 1♂, Banks Peninsula, Mt. Sinclair Scen Res, 775m, 43″42.977′S, 172″51.098′E, 3–16 XII 2005, ridgetop mixed broadleaf w/emergent Podocarpus totara, FMHD #2005-070, FIT, A. Newton, M. Thayer, ANMT site 1163; 1♀, Banks Peninsula, Ahuriri Scen Res, 450m, 43″39.971′S, 172″37.427′E, 3–16 XII 2005, ridgetop mixed broadleaf w/emergent Podocarpus totara, FMHD #2005- 066, FIT, A. Newton, M. Thayer, ANMT site 1162; 1♀, Banks Peninsula, Peraki Saddle Scen Res, 500m, 11 XII 1984 hdwd- Podocarpus elfin forest, A. Newton, M. Thayer 701, litter; 8♂♂ 4♀♀, Cooper’ s Knobs nr Christchurch, 1800’, rotten wood, 18 X 1951; North Canterbury: 1♂, 4.5km se Arthur’ s Pass, 725m, #044, 42′55S, 171′33E, under bark, 11 I 1998, C. Carlton, R. Leschen; Nelson: 2♂♂ 2♀♀ (1♂, slide-mounted), 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &amp;window trap; 1♂, Kahurangi NP, Cobb Ridge, above Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1159; 6♀♀, Slaters Rd, 0.7km s Whangamoa Saddle, 410m, 13 XII 1984 – 4 I 1985, Nothofagus forest, A. Newton, M. Thayer 703, window trap; 1♂, Upper Maitai, 14 II 1943, E.S. Gourlay; Otago Lakes: 1♂ 1♀, 2km se Kinloch, 14 I 2006, R. Leschen, T. Buckley, R. Hoare, under rotten logs, RL170, 44.51S, 168.20E; 1♂, 19km nw Glenorchy, #098, 44′41S 168′27E, under bark, rotten logs, 20 I 1998, C. Carlton, R. Leschen; 1♂, Makarora, 21 I 1978, G. Kuschel, litter and rotten wood 78/47; 1♀, 17km nw Glenorchy (beyond Paradise), 44′41S, 168′20E, #100, under bark, under log, 20 I 1998, C. Carlton, R. Leschen; 1♂, Mt. Aspiring NP, 12.5km nne Makarora, 370m, 11–17 I 1985, Nothofagus menziesii -hdwd-podocarp forest, A. Newton, M. Thayer 740, FIT &amp;window trap; 2♀♀, Lake Hawea, Hunter Valley rd, Sawyer Burn, 400m, 10–17 I 1985, Nothofagus solandri forest, A. Newton, M. Thayer 737, FIT &amp;window trap; 1♀, Mt. Aspiring NP, se of Haast Pass, 600m, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 739, window trap; Marlborough Sounds: 4♀♀, Tennyson Inlet, west side Te Mako Bay, 125m, 15 XII 1984 – 5 I 1985, Nothofagus - podo-hdwd, A. Newton, M. Thayer 710, FIT &amp;window trap; 1♂ 1♀, Tennyson Inlet, west side Duncan Bay, 30m, 15 XII 1984 – 5 I 1985, Nothofagus -podocarp forest, A. Newton, M. Thayer 709, FIT &amp;window trap; 1♀, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton, M. Thayer 708, window trap; Westland: 1♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwdnikau forest, A. Newton, M. Thayer 720, FIT &amp;window trap; 1♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, litter; 1♀, Okuku Scen Res, 9.2km sse Kumara, 120m, 8–19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 731, FIT &amp;window trap; 1♀, Haast Pass w of summit, 550m, 24 I 1978, G. Kuschel, sifted litter and rotten wood 78/52.</p><p>Diagnosis. This species can be distinguished from the other species of the notabilis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; male antennomeres 4–6 enlarged; shape of ventral surface of male head and genitalia unique to species; only known from South Island.</p><p>Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 56b). Head: Round, as long as wide, widest across eyes (Fig. 56h). Antennomere 2 longer than wide, 3–10 subquadrate, 4–6 enlarged in male (Fig. 56g). Eye prominent, approximately two-thirds length of temple (Fig. 56h). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 56b). Meso- and metathorax trapezoidal, longer than wide. Male fore femur with shallow semicircular depression. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia triangular, shorter than parameres (Fig. 56e). Phallobase of median lobe symmetrical and rectangular (Fig. 56e). Parameres symmetrical with process at one-third length, with setae and processes apically (Fig. 56e).</p><p>Type Locality. “ New Zealand ” (specific locality not mentioned) .</p><p>Distribution. Buller, Dunedin, Fiordland, Mid Canterbury, North Canterbury, Nelson, Otago Lakes, Marlborough Sounds, Westland (Fig. 57: triangles).</p><p>Habitat. Specimens of this species were collected using flight intercept or window traps or by sifting dead logs and wood litter in mostly Nothofagus forests. One specimen was found in an ant nest.</p><p>Comments. Male specimens of S. monstrosa can be distinguished from those of other species by antennomeres 3–10 being subquadrate, enlarged male antennomeres 4–6, and modification of the ventral surface of male head. The type specimens of S. bifoveiceps, S. biimpressa, and S. nitida share these diagnostic characters. For these reasons, we have placed S. bifoveiceps, S. biimpressa, and S. nitida in synonymy with S. monstrosa .</p></div>	https://treatment.plazi.org/id/03D82A3EBD0A11293CA8FDD8FDFCFE96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0C11293C81FEC4FC65FE36.text	03D82A3EBD0C11293C81FEC4FC65FE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola tararuaensis Park and Carlton	<div><p>Sagola tararuaensis Park and Carlton, new species</p><p>(Figs. 56c, 56f, 57)</p><p>Type Material. Holotype. NEW ZEALAND: Wellington: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: WN: Tararua Forest Park, above Akatarawa Saddle, 455m, 40″56.936′S, 175″06.529′E, 26 XI–21 XII 2005, broadleafpodocarp forest on slope; FMHD#2005-0112, flight intercept trap, A. Newton &amp; M. Thayer; ANMT site 1151”, “ HOLOTYPE Sagola tararuaensis Park and Carlton 2013 ” . Paratypes (n = 12: 7 males; 5 females). NEW ZEALAND: Wellington: 1♂ (FMNH), same data as holotype; 2♂♂ (JTN), Pakurarahi Forks, Kaitoke, 24 VII 1993, dead kiekie stem, J. Nunn ; 1♂ 1♀ (JTN), Manawatu Gorge, 17 XI 1991, J. Nunn ; 1♂ 1♀ (JTN), Keith George Park, Silverstream, 5 V 1995, J. Nunn, decayed wood ; 1♂ (JTN), Waiotauru Rd, Tararua FP, 23 IV 1995, J. Nunn, decayed wood ; 1♂ (JTN), Graces Stream tr, Rimutaka FP, 15 VIII 1993, J. Nunn, decayed wood ; 1♀ (JTN), Wainui Trig, Wainuiomata, 14 VII 1993, J. Nunn, decayed wood ; 1♀ (JTN), West Lake Wairarapa Res, 20 IV 1997, J. Nunn, subhumified wood ; 1♀ (JTN), Trentham Memorial Park, 7 VIII 1993, J. Nunn, decayed wood .</p><p>Etymology. This species is named after the type locality, Tararua Forest Park, Wellington.</p><p>Diagnosis. This species can be distinguished from the other species of the notabilis speciesgroup by the following combination of characters: size larger, length 2.7–3.0 mm; male antennomeres 4–6 weakly enlarged, 8 enlarged; male fore femur with weak semicircular depression; shape of ventral surface of male head and genitalia unique to species; only known from Wellington of North Island.</p><p>Description. Length 2.7–3.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 56c). Head: Round, as long as wide, widest across eyes (Fig. 56c). Antennomere, 2 longer than wide, 3–10 subquadrate, 4–6 weakly enlarged in male, 8 enlarged in male. Eye prominent, approximately two-thirds length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 56c). Meso- and metathorax trapezoidal, longer than wide. Male fore femur with shallow, semicircular depression. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia divided vertically, major lobe triangular and minor lobe oval (Fig. 56f). Phallobase of median lobe symmetrical and rectangular (Fig. 56f). Parameres symmetrical with triangular membranous process at one-third length, with setae apically (Fig. 56f).</p><p>Distribution. Wellington (Fig. 57: black squares).</p><p>Habitat. Specimens of this species were collected mostly by sifting decayed wood litter or using flight intercept traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0C11293C81FEC4FC65FE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0C112A3F65FE2EFD44FA8C.text	03D82A3EBD0C112A3F65FE2EFD44FA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola misella Sharp Desig. Park and Carlton 2013	<div><p>25. misella species-group</p><p>(13 species)</p><p>Diagnosis. Members of the misella speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.1–3.3 mm; antennomere 1 approximately 1.5–2.0 times longer than wide, male antennomeres enlarged; frontal rostrum rectangular, lobes continuous, reaching posterior frontal fovea (Fig. 59o); frontal sulcus reaching midpoint of eye, posterior frontal fovea present (Fig. 59o); female hind wings reduced to small pads; abdominal tergites IV–VI with discal carinae.</p><p>KEY TO SPECIES OF THE MISELLA SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Eye small, approximately one-third length of temple .................................. 2</p><p>1′. Eye large, approximately one-half length of temple................................................8</p><p>2(1). Apical lobe of genitalia arrowheadshaped (Fig. 59b) .................................. .............................. S. bituberata Broun</p><p>2′. Apical lobe of genitalia not arrowheadshaped ..................................................3</p><p>3(2′). Major process of median lobe of genitalia S-shaped (Fig. 59a) ....................... ................................... S. misella Sharp</p><p>3′. Median lobe of genitalia not curved.... 4</p><p>4(3′). Median lobe of genitalia triangular with membranous tube (Fig. 59c) ................. ..................................... S. prisca Sharp</p><p>4′. Medina lobe of genitalia not triangular.............................................................5</p><p>5(4′). Median lobe of genitalia depressed, left apex and as long as paramere............. 6</p><p>5′. Median lobe of genitalia divided, minor lobe longer than major lobe and paramere .................................................................. 7</p><p>6(5). Paramere depressed apically (Fig. 59d); only known from Nelson (Fig. 61: triangle) ............................................. ....... S. murchisonensis new species</p><p>6′. Paramere blunt; known from Fiordland (Fig. 62: star)......................................... ....................... S. clunieae new species</p><p>7(5′). Abdominal ventrite VII with small and acute process; major lobe of median lobe of genitalia slender and simple (Fig. 59g) .............. S. humpridgensis new species</p><p>7′. Abdominal ventrite VII without process; major lobe of median lobe of genitalia diamond-shaped and broad (Fig. 59f)............................................... ................. S. hunterensis new species</p><p>8(1′). Elytra subquadrate, hind wings reduced to small pads ..... S. huvali new species</p><p>8′. Elytra rectangular, hind wings fully developed.............................................9</p><p>9(8′). Antennomeres 4–8 longer than wide .... .............. S. tennysonensis new species</p><p>9′. Antennomeres 4–8 subquadrate........10</p><p>10(9′). Median lobe of genitalia twisted, with membranous tube (Fig. 59i).................. .................... S. turretensis new species</p><p>10′. Median lobe of genitalia not twisted, without tube....................................... 11</p><p>11(10′). Apical lobe of genitalia rectangular vertically (Fig. 59j–k)............................ .................... S. otagoensis new species</p><p>11′. Apical lobe of genitalia acute...........12</p><p>12(11′). Median lobe of genitalia C-shaped vertically (Fig. 59e) ............................... .............. S. fiordlandensis new species</p><p>12′. Median lobe of genitalia S-shaped vertically (Fig. 59l)....................................... ............ S. southlandensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD0C112A3F65FE2EFD44FA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0F11253D50FABBFC1FFB6A.text	03D82A3EBD0F11253D50FABBFC1FFB6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola misella Sharp Desig. Park and Carlton 2013	<div><p>Sagola misella Sharp, 1874</p><p>(Figs. 58a, 59a, 59o–r, 60)</p><p>Sagola misella Sharp, 1874: 508 . Broun 1880: 136; Schaufuss 1888: 84; Raffray 1893: 26, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola foveiventris Broun, 1921a: 492 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Lectotype. NEW ZEALAND: 1♀ (BMNH), glued on rectangular card, “ Type ” [ red label, printed]; “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola misella . ♂? Type. D.S.” [white label, handwritten]; “Sex ♀ DSC`87” [white label, handwritten]; “ LECTOTYPE Sagola misella Sharp Desig. Park and Carlton 2013 ” [red label, printed]. Paralectotypes (14 females). NEW ZEALAND: 14♀♀, same data as holotype. The lectotype designation is required because Sharp did not explicitly designate a type specimen. His comment suggested that multiple specimens were examined, but the exact number was not clear (Sharp 1874: 508). We examined 15 specimens and could not locate additional specimens from museums where most Sharp specimens are deposited. This designation will fix the identity of the species and facilitate its recognition by future workers . Syntypes of Sagola foveiventris: NEW ZEALAND: Otago Lakes: 2♂♂ (BMNH), glued on rectangular card, “4000. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922– 482.” [white label, printed]; “Hollyford. 19.2.1914.” [white label, written]; “ Sagola ♂. foveiventris.” [white label, handwritten] .</p><p>Additional Material (n = 157: 70 males; 87 females). NEW ZEALAND: Fiordland: 4♂♂ 2♀♀, Fiordland NP, Borland rd, Borland saddle vic, 980m, 45″44.81′S, 167″22.88′E, 10 XII 2005, Nothofagus menziesii forest in ravine, FMHD#2005-090, litter, A. Newton, ANMT site 1171 ; 2♂♂, Fiordland NP, Milford Sound rd, Tutoko tr, 70m, 44″40.517′ S, 167″57.895′E, 9 XII 2005, very mossy Nothofagus forest, emergent podocarp, FMHD#2005-086, litter, A. Newton, M. Thayer, ANMT site 1168 ; 1♂ 2 ♀♀, Fiordland NP, Borland rd, Mt. Burns Tops tr, above Borland Saddle, 1075m, 45″44.883′S, 167″22.938′E, 10 XII 2005, subalpine tussockland, FMHD#2005-096, unsifted litter at bases of shrubs ( Dracophyllum, possibly filifolium), M. Thayer, ANMT site 1173 ; 1♂, Fiordland NP, Kepler tr, below Mt. Luxmore Hut, 900m, 45″24.86′S, 167″38.675′E, 11 XII 2005, Nothofagus forest, FMHD#2005-101, litter, A. Solodovnikov, D. Clarke, ANMT site 1176 ; 3♀♀, Fiordland NP, Kelper tr, above Brod Bay, 680m, 45″24.37′S, 167″39.105′E, 11 XII 2005, Nothofagus forest, FMHD#2005-102, litter, A. Solodovnikov, D. Clarke, ANMT site 1177 ; 1♀, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S, 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD#2005- 087, litter, A. Newton, M. Thayer, ANMT site 1169 ; 1♀, Fiordland NP, Borland rd, Mt. Burns Tops tr, above Borland Saddle, 1035m, 45”44.82′S, 167″22.87′E, 10 XII 2005, short ridgetop Nothofagus menziesii forest near timberline, FMHD#2005-098, litter, M. Thayer, ANMT site 1174 ; 4♂♂ 10♀♀, Doubtful Sound, 1571, 5 IV 1953, R. Hornabrook ; 2♂♂ 5♀♀, W. Olivine Ra, Simonin Pass, 1067m, 27 I 1975, G.W. Ramsay, litter 75/38 ; 1♂ 2♀♀, W. Olivine Ra, Simonin Pass, 23 I 1975, G.W. Ramsay, litter 75/35 ; 1♂, Wilmot Pass, 630m, 26 I 1970, G.W. Ramsay, litter 70/93 ; 1♀, Wilmot Pass, 630m, 24 I 1970, J.S. Dugdale, mats 70/94 ; 1♂, Turret Ra, Wolfe Flat, 900m, 22 I 1970, A.C. Eyles, litter 70/90 ; 1♂, Doubtful Sound, Wilmot Pass, 21 I 1970, A.C. Eyles, litter 70/64 ; 1♂, Hollyford V, Hd L. Marion, 640m, 13 XII 1966, A.K. Walker, litter 66/510; 2♂ 7♀♀, Hump Ridge, Fairburn, 12 III 1939 ; 3♂♂, Secretary I, Grono Bay, 28 XI 1981, C.F. Butcher, litter ; 1♂ 1♀, W. Olivine Ra, 1220–1463m, 25 I 1975, G.W. Ramsay, litter ; 1♀, W. Olivine Ra, 1067m, 23 I 1975, G.W. Ramsay, litter ; 4♂ 7♀♀, Fiordland NP, Murchison Mts, 1 XII 1980 – 9 XII 1983, R. M. Emberson, C.A. Muir ; 1♂ 4♀♀, Lower Hollyford rd, 44′44S 168′08′E, #111, Nothofagus /coastal forest leaf litter berlese, 22 I 1998, C. Carlton, R. Leschen ; 1♂ 1♀, Camp 1020m, Tutoko Bench, Darran Mts, 15 I 1977, T. K. Crosby, litter 77/10 ; 1♂ 1♀, Percy Stm, 460m, 5 II 1982, J.W. Early, Nothofagus fusca forest ; 2♂♂, Secretary I, Gut Bay, 23 XI 1981, C.F. Butcher, litter ; 1♂, Dusky Sound, Passage I, 8 III 1983, C.F. Butcher, litter ; 1♂, W. Olivine Ra, Tempest Spur Summit, 1463m, 30 I 1975, G.W. Ramsay, litter ; 1♂, Wilmot Pass, 670m, 5 II 1983, J.W. Early, litter ; 1♂, Hollyford tr, #110, 44′42S 168′08E, Nothofagus coastal forest litter, 22 I 1998, C. Carlton, R. Leschen ; 3♀♀, Breaksea Sound, Gilbert I, 6 V 1982, C.F. Butcher, litter ; 1♀, Borland saddle, Hunter Mtns, 28 I 2011, litter, 45”44.765′S, 167”22.709′E, 1015m, J. Nunn ; 1♀, Borland rd, Borland Saddle tr, 45′46S 167′23E, #138, Nothofagus leaf litter, 24 I 1998, C. Carlton, R. Leschen ; 1♀, Beside Tutoko R, 60m, 13 II 1980, J.W. Early, litter in Nothofagus menziesii forests ; North Canterbury: 1♂, Lake Marion, Holly rd, 2000′ 13 XII 1966, A.K. Walker, fern litter 66/514; Otago Lakes: 1♂ 6♀♀, McKerrow range. Makarora, 1000m, 23 I 1978, G. Kuschel, litter ; 2♂♂ 1♀, nr Dart Hut, 940m, 12–20 II 1980, J.C. Watt, pit fall ; 1♂ 2♀♀, Haast Pass summit, 560m, 22 I 1978, G. Kuschel, litter ; 1♂, Makarora, 21–24 I 1978, S. &amp; J. Peck, malaise trap ; 1♂, Dart Hut, 13–14 II 1980, J.S. Dugdale, pan trap in bush ; 1♂, Makarora, 20 I 1996, J. Nunn; in beech forest ; 1♀, Makarora, 23–24 I 1978, 500m, S. &amp; J. Peck, litter ; 1♀, Makarora, 350m – 400m, 22 I 1978, S. &amp; J. Peck, litter ; 1♀, Haast River, 1000m, Sunny Flat, 25 I 1978, G. Kuschel, litter ; 1♂, Paradise, 10 I 1945, E.S. Gourlay ; Stewart Island: 10♂♂ 4♀♀, Codfish I, N.W. Bay, 15 XII 1966, A.H. Whittaker, litter ; 1♂ 2♀♀, Pegasus Creek, 24 II 1968, G. Kuschel, litter ; 1♂, Pegasus Creek, 24 XI 1968, G. Kuschel, litter ; 2♂♂, Christmas Village, J.I. Townsend, 28 I 1962, litter ; 2♂♂, N.E. Big S. Cape I, 121m, 18 XI 1968, G. Kuschel, litter; 1♂ (slide-mounted), Codfish I, Valley tr, 25 XI 1981, B.A. Holloway, litter; Southland : 2♀♀, Manapouri, 21 I 1970, G.W. Ramsay, litter ; 2♀♀, Spence Burn Basin, 1150m, Takitimu mtns, 17–18 I 2000, J. Nunn, tussock litter ; 1♀, Bluff Hill, Bluff, 7 VII 2002, J. Nunn, litter in kamahi forest ; 1♀, Orepuki, 450’ 7 II 1968, litter, J.I. Townsend ; Westland: 2♂♂, Mt. Aspiring NP, Duglas Ck at hwy. 6, 65m, 11–18 I 1985, Nothofagus menziesii -hdwd-podocarp forest, A. Newton, M. Thayer 738, FIT &amp; window trap ; 1♂, Lower Haast, 16 III 1966, J. Nunn ; 2♀♀, 2.7km s Franz Josef Glacier, #063, Trailhead of Alex Knob, 43′25S 173′10E, 150m, coastal forest leaf litter, 15 I 1998, C. Carlton, R. Leschen ; 1♀, 2km se Fox Glacier, 225m, #066, 43′29S 170′01E, litter, 16 I 1998, C. Carlton, R. Leschen ; 2♀♀, 11.2km ne Franz Josef Glacier, nr Lake Mapourika, 150m, Otto’ s Corner picnic area, 43′18S 170′14E, #064, coastal forest leaf litter, 15 I 1998, C. Carlton, R. Leschen ; 2♀♀, 3.2km ne Haast, 14m, Haast River Walk, 43′52S 169′03E, #072, Nothofagus /podocarp forest litter, 17 I 1998, C. Carlton, R. Leschen ; 1♀, Doughboy Creek, 6km sw Mahitahi, 5 II 1984, J.C. Watt, sifted wood mould ; 3♂♂, Mt Aspiring NP, Arawata Blv, 840m, 5 II 1989, J.W. Early, R. M. Emberson, scrub litter .</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body length 2.9–3.7 mm; male antennomeres 5–10 weakly enlarged bearing tubercles; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.9–3.7 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58a). Head: Round, as long as wide, widest across eyes (Fig. 59o). Male antennomere 1 approximately 1.5 times longer than wide, 2–4 longer than wide, 5–10 subquadrate and weakly enlarged, 5–11 with tubercles. Female antennomeres 2–8 longer than wide, 9–10 subquadrate, 7–11 with tubercles. Frontal rostrum rectangular, lobes continuous (Fig. 59o). Frontal sulcus reaching midpoint of eye (Fig. 59o). Posterior frontal fovea teardrop-shaped (Fig. 59o). Eye prominent, approximately one-third length of temple (Fig. 59o). Thorax: Prosternum as long as wide, widest at midpoint (Fig. 59p). Male elytra rectangular (Fig. 58a), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide (Fig. 59q), female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia divided, major lobe curved as S-shape, minor lobe longer and membranous (Fig. 59a). Phallobase of median lobe symmetrical and rounded (Fig. 59a). Parameres symmetrical longer than median lobe and with setae apically (Fig. 59a).</p><p>Type Locality. “ New Zealand ” ( Specific locality not mentioned) .</p><p>Distribution. Fiordland, North Canterbury, Otago Lakes, Stewart Island, Southland, Westland (Fig. 60: black circles).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf litter. A few male specimens were collected using flight intercept or window traps.</p><p>Comments. Specimens of S. misella can be distinguished from those of other species by the shape of the antennomeres. The type specimens of S. foveiventris share these diagnostic characters. For these reasons, we have placed S. foveiventris in synonymy with S. misella .</p></div>	https://treatment.plazi.org/id/03D82A3EBD0F11253D50FABBFC1FFB6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0011263F40FAE0FC9DFC85.text	03D82A3EBD0011263F40FAE0FC9DFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola bituberata Broun 1914	<div><p>Sagola bituberata Broun, 1914</p><p>(Figs. 58b, 59b, 60)</p><p>Sagola bituberata Broun, 1914b: 160 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. One of three syntypes. NEW ZEALAND: Buller: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “ 3525. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Greymouth. J.H. Lewis ” [white label, handwritten]; “ Sagola ♂. bituberata” [white label, handwritten] .</p><p>Additional Material (n = 52: 24 males; 28 females). NEW ZEALAND: Buller: 3♂♂, Lake nr Mt. Priestly, Paparoa Ra, 1060m, 10 XII 1969, J. McBurney, litter; 1♂, Fletchers Ck, 6km sw of Rotokohu, 25 I 1972, J.M. McBurney, litter ; 1♀, Relax Shelter, Mt Robert Ridge, 1 X 2005, J. Nunn, moist tussock litter at 1410m ; 1♂, Arthur Pass, Kellys Ck, 8 XI 2005, R. Leschen, S. Nomura, litter, 42′47S, 171′24E; Nelson : 2♂♂ 3♀♀, Canaan, 18 IV 1966, J.I. Townsend, litter ; 1♂, Mt. Domett, 1250m, 30 XI 1971, G. Kuschel, litter ; 2♀♀, Canaan, Harwoods tr, 4 II 1965, L.P. Marchant, litter; Otago Lakes : 1♂ 1♀, Haast River, 100m, Sunny Flat, 25 I 1978, G. Kuschel, litter ; 2♀♀, Mt. Aspiring NP, 5.5km, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 741, window trap; Westland : 4♂♂ 8♀♀, Kellys Creek, Otira, 22 III 2009, J. Nunn, mossy forest floor litter ; 1♂ 2♀♀, Kellys Creek, Otira, 25 II 1989, J. Nunn ; 1♀, Haast Pass n of summit, 550m, 24 I 1978, G. Kuschel, sifted litter and rotten wood ; 1♂ 2♀♀, Okuku Ck, 11.3km sse Kumara, 60m, 18–22 III 1980, podocarpbroadlf, A. Newton, M. Thayer, litter ; 3♂♂, Franz Joseph. 24 II 1989, J. Nunn ; 2♀♀, Lake Wombat tr, Franz Joseph, 7 II 2009, J. Nunn, washed soil sample, tree fern/broadleaf forest ; 2♂♂, Jackson Bay, Smoothwater tr, 14 II 1977, T. K. Crosby, litter ; 1♂, Kellys Cr, Otira Vly, 9 IV 2010, J. Nunn, litter in kamahi forest ; 1♂, 2.7km s Franz Josef Glacier, #063, Trailhead of Alex Knob, 43′28S 1173′10E, 150m, coastal forest litter, 15 I 1998, C. Carlton, R. Leschen ; 1♂, Mt. Aspiring NP, 5.5km nne Makarora, 330m, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 741, FIT &amp; window trap ; 1♂, Mt. Aspiring NP, Douglas Creek at Hwy. 6, 65m, 11–18 I 1985, Nothofagus menziesii -hdwd-podocarp forest, A. Newton, M. Thayer 738, FIT &amp; window trap ; 1♀, Open Bay I, Taumaka I, 27 IX 1976, D.S. Horning, litter ; 1♀, Mt Tuhua, 1067m, e side of L. Kaniere, 20 XI 1984, C.F. Butcher, litter ; 1♀, Lake Ianthe, 7 II 1984, J.C. Watt, sifted litter and wood mould ; 1♀, Doughboy Creek, 6km sw Mahitahi, 5 II 1984, J.C. Watt, sifted wood mould .</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.5–3.1 mm; male antennomeres 4–10 enlarged; male abdominal ventrite V with weak process medially; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.5–3.1 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58b). Head: Round, as long as wide, widest across eyes (Fig. 58b). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 5–11 with tubercles. Female antennomeres 2–8 longer than wide, 9–10 subquadrate, 5–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardrop-shaped. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 58b), female subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, patches absent in female. Male abdominal ventrite V with weak process medially. Aedeagus: Apical lobe of genitalia arrowhead-shaped (Fig. 59b). Phallobase of median lobe symmetrical and rounded (Fig. 59b). Parameres asymmetrical, right paramere longer than left with setae apically (Fig. 59b).</p><p>Type Locality. Greymouth, Buller, New Zealand .</p><p>Distribution. Buller, Nelson, Otago Lakes, Westland (Fig. 60: triangles).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf litter, and a few male specimens were from flight intercept or window traps.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0011263F40FAE0FC9DFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0311213F75FCBCFDD6FB1B.text	03D82A3EBD0311213F75FCBCFDD6FB1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola prisca Sharp Desig. Park and Carlton 2013	<div><p>Sagola prisca Sharp, 1874</p><p>(Figs. 58c, 59c, 61)</p><p>Sagola prisca Sharp, 1874: 507 . Broun 1880: 136; Reitter 1880: 167; Schaufuss 1888: 85; Raffray 1893: 24, 1904: 498, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Jeannel 1950: 45; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola fuscipalpis Broun, 1914b: 159 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola disparata Broun, 1914b: 160 . Hudson 1923: 365; Hudson 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola striatifrons Broun, 1921a: 492 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.</p><p>Sagola acuminata Broun, 1921a: 498 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola rufescens Broun, 1921a: 499 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Lectotype. NEW ZEALAND: 1♂ (BMNH), left specimen glued on rectangular card, “Type” [red label, printed]; “ N. Zeal. ” [white label, printed]; “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola prisca ♂ Type. D.S.” [white label, handwritten]; “ LECTOTYPE Sagola prisca Sharp Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotypes (n = 6: 4 males; 2 females). NEW ZEALAND: 4♂♂ 2♀♀ (BMNH), same data as holotype . Holotype of Sagola acuminata: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “4009. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Routerburn. 16.2.1914.” [white label, written]; “ Sagola ♂. acuminata.” [white label, handwritten]. One of two syntypes of Sagola disparata: NEW ZEALAND: Central Otago: 1♂ (BMNH), glued on rectangular card, “3524. ♂ ” [white label, handwritten]; “Old Man Range.” [white label, written]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂. disparata” [white label, handwritten] . Syntypes of Sagola fuscipalpis: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “type” [red label, printed]; “3523. ♂ ” [white label, handwritten]; “Mt. Hutt. 12.4.1912 ” [white label, written]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂. fuscipalis” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “3523. ♂ ” [white label, handwritten]; “Mt. Hutt. 12.4.1912 ” [white label, written]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂. fuscipalpis” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “var. 3523. ♀.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Hutt. 12.4.1912 ” [white label, written] . Holotype of Sagola rufescens: NEW ZEALAND: Central Otago: 1♀ (BMNH), glued on rectangular card, “4010.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Staircase. 13.3.1914.” [white label, written]; “ Sagola rufescens .” [white label, handwritten] . Syntypes of Sagola striatifrons: NEW ZEALAND: Otago Lakes: 1♀ (BMNH), glued on rectangular card, “4001. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Harris Saddle. 13.2.1914.” [white label, written]; “ Sagola ♀ striatifrons.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4001. ♀ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Harris Saddle. 13.2.1914.” [white label, written]; “ Sagola striatifrons .” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4001.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hollyford. 19.2.1914.” [white label, written]; “ Sagola striatifrons .” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “4001.” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hollyford. 19.2.1914.” [white label, written]; “ Sagola ♂ striatifrons.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4001. var.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hollyford. 19.2.1914.” [white label, written]. The lectotype designation is required because Sharp did not explicitly designate a type specimen, and he mentioned that seven specimens were examined (Sharp, 1874: 508). This designation will fix the identity of the species and facilitate its recognition by future workers .</p><p>Additional Material (n = 157: 81 males; 76 females). NEW ZEALAND: Buller: 2♂♂, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, FIT &amp;window trap; 1♂, Nelson Lakes NP, Lake Rotoiti St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp;window trap; 1♀, Bell Hill Scenic Res, 11 V 1996, K.W. Drew &amp; E.B. Spurr, bait trap; Central Otago: 1♂, Dunstan Ra, 1646m, Summit, 13 I 1971, J.S. Dugdale, moss; 2♂♂, Lake Alta, The Remarkables, 21 I 2008, 1♂, Carrick Ra, Watts rk, 1300m, 5–8 II 1986, J.W. Early, yellow pan trap in tussock grassland; Dunedin: 2♂♂ 3♀♀, Leith Saddle, Leith Trig tr, 420m, 26 IX 2011, washed soil, J. Nunn; 1♂ 6♀♀, Mt. Cargill, 400-450m, 21 IX 1981, J.C. Watt, litter; 2♂♂ 2♀♀, Mt. Cargill Sec Res, Organ Pipes tr, nr. Mt. Cargill summit, 600m, 45”48.505′S, 170″33.999′E, 5 XII 2005, podocarp-broadleaf forest, FMHD #2005-076, litter, A. Solodovnikov, D. Clarke, ANMT site 1165; 1♂ 4♀♀, Leith Saddle, 17 V 2010, J. Nunn, litter; 1♂ 1♀, Mt. Cargill, 30 VII 2000, litter, J. Nunn; 1♂, Allison Res nr Akatore, 1 IV 2001, J. Nunn; 1♂, Leith Saddle, 20 V 2006, J. Nunn, washing soil; 1♂, Leith Saddle, J. Nunn, litter; 1♂, Leith Saddle, 11 XI 2000, J. Nunn, litter; 1♂, Leith Saddle, Leith Trig Tr, 24 IX 2010, J. Nunn, washed soil; 1♂ 1♀, Leith Trig, 430m, 17 X 2010, J. Nunn, litter; 1♂, Allison Res, nr Akatore, 1 IV 2001, J. Nunn, litter; Gisborne: 2♂♂ 16♀♀, Urewera NP, Maungapohatu rd, 3.2km e Taupeupe Saddle, 1110 m, 38 ″ 36.975 ′ S 177″02.753′E, 22 XI 2005, mossy Nothofagus menziesii forest, FMHD #2005-027, litter, A. Newton, M. Thayer, ANMT site 1148; 3♀♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S 172″41.658′E, 29 XI 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-055, litter, M. Thayer, ANMT site 1159; Fiordland: 3♂♂ 5♀♀, Secretary I, Grono Bay, 24 III 1984, C.F. Butcher, litter; 1♂ 6♀♀, Lake Hauroko, 2 II 1966, F.D. Alack, litter; 1♂, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S 168″00.16′E, 9 XII 2005, mossy Nothofagus menziesii forest, FMHD #2005-088, recent flood debris floating in stream, J. Nunn, A. Newton, ANMT site 1169; 1♂, Cameron Mts, Lake Mike, 488m, 13 II 1963, J.I. Townsend, litter; 3♂♂ 1♀ (2♂♂ 1♀, slide-mounted), Lower Hollyford rd, 44′44S 168′08E, #111, Nothofagus / coastal forest leaf litter, 22 I 1998, C. Carlton, R. Leschen; Mid Canterbury: 2♂♂, Mt. Hutt, McClennan’ s Bush, 11 XII 1973, G. Kuschel, litter; Nelson: 9♂♂ 1♀, Kahurangi NP, Cobb Ridge, Above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, ANMT site 1159; 6♂♂ 10♀♀, Kahurangi NP, Cobb Ridge, Above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI– 18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-055, litter, M. Thayer, ANMT site 1159; 1♂ 1♂, Kahurangi NP, Cobb Ridge, Above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii &amp; N. solandri cliffortioides forest, FMHD #2005-053, pitfall trap (5), A. Solodovnikov, D. Clarke, ANMT site 1159; 3♂♂ 2♀♀, Kahurangi NP, Mt. Arthur tr, ca. 1400m, 28 XI 2005, above treeline, FMHD #2005- 050, alpine tussock litter, D. Clarke, ANMT site 1158; 1♂, Cobb Ridge, east of Cobb Reservoir, 990m, 2 I 1985, Nothofagus forest, A. Newton, M. Thayer 728, litter; 1♀, Kahurangi NP, Mt. Arthur tr, below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD #2005-049, litter, A. Solodovnikov, D. Clarke, ANMT site 1157; 4♂♂, Mt. Arthur, 1220m, 20 III 1964, J.I. Townsend, litter; Otago Lakes: 1♂ 2♀♀, Rees Saddle, 1600m, 14 II 1980, J.S. Dugdale, swards; 2♂♂ 1♀, Headlong Pk, 1680m – 1830m, 13–19 II 1980, J.C. Watt, pit fall; Rangitikei: 1♀, Ruahine Ra, Triplex, 10 II 1980, C.F. Butcher, litter; Stewart Island: 1♂, Kundy I, 17 XII 1981, S.L. Lobb, litter; Taranaki: 1♂, Dawson Falls, 914m, 23 I 1972, G.W. Ramsay, litter; 1♂, Pouakai Ra, 1067m, Ahukawakawa tr, 11 I 1978, litter &amp; moss, J.C. Watt; 1♂, Pouakai Trig, Pouakai ra, 3 XII 1975, J.S. Dugdale, litter; Westland: 3♂♂ 1♀, At. Aspiring NP, Arawata biv, 840m, 31 I–5 II 1989, J.W. Early, yellow pan trap; 7♂♂ 4♀♀, Open Bay I, Taumaka I, 1 VIII 1975 – 27 IX 1976, R.H. Mattlin &amp;, D.S. Horning, litter; 2♂♂, Doughboy Creek, 6km sw Mahitahi, 5 II 1984, J.C. Watt, wood mould; 1♂ 1♀, Mt. Tuhua, 1067m, e side of L. Kaniere, 20 XI 1984, C.F. Butcher, litter; 1♂, 2.7km se Lake Moeraki, 30m, 10–18 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 736, FIT &amp;window; 1♂, Lake Ianthe, 7 II 1984, J.C. Watt, litter &amp; wood mould; 1♀, Okuku Ck, 11.3km sse Kumara, 60m, 18-22 III 1980, podocarp-broadlf, A. Newton, M. Thayer, litter; Wellington: 1♂ 1♀, Tararua Ra, Dundas Hut Ridge, 950m, 3 XII 1984, R.C. Craw, litter; 1♀, Tararua Ra, Dundas Hut Ridge, 800m, 13 II 1985, G.W. Ramsay, litter.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.4–3.8 mm; male antennomeres 4–10 enlarged; male abdominal ventrite V with weak process medially; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.4–3.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58c). Head: Round, as long as wide, widest across eyes (Fig. 58c). Male antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4 longer than wide, 5–10 subquadrate, 4–10 enlarged, 5–11 with tubercles. Female antennomeres 2–6 longer than wide, 7–10 subquadrate, 6–11 with tubercles. Frontal rostrum rectangular, meeting each other. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardrop-shaped. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 58c), elytra subquadrate in females and some males. Male meso- and metathorax trapezoidal, longer than wide, females and some males as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, patches absent in females and some males. Male abdominal ventrite V with weak process medially. Aedeagus: Apical lobe of genitalia triangular with membranous tube (Fig. 59c). Phallobase of median lobe symmetrical and rounded (Fig. 59c). Parameres symmetrical, as long as median lobe, and with setae apically (Fig. 59c).</p><p>Type Locality. New Zealand (specific locality not mentioned) .</p><p>Distribution. Buller, Central Otago, Dunedin, Gisborne, Fiordland, Mid Canterbury, Nelson, Otago Lakes, Rangitikei, Stewart Island, Taranaki, Westland, Wellington (Fig. 61: black circles).</p><p>Habitat. Specimens of this species were collected using pitfall, yellow pan, flight intercept, and window traps or by sifting leaf and wood litter mostly in Nothofagus forests.</p><p>Comments. Male specimens of S. prisca can be distinguished from those of other species by the shape of the antennomeres. The type specimens of S. acuminata, S. disparata, S. fuscipalpis, S. rufescens, and S. striatifrons share these diagnostic characters. For these reasons, we have placed S. acuminata, S. disparata, S. fuscipalpis, S. rufescens, and S. striatifrons in synonymy with S. prisca . This species exhibits polymorphism in the shape of the antennomeres, length of male elytra, and size of genitalia. However, these characters do not vary sufficiently to warrant separating the complex into additional species.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0311213F75FCBCFDD6FB1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0411213CF1FB50FC05FCA0.text	03D82A3EBD0411213CF1FB50FC05FCA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola murchisonensis Park and Carlton	<div><p>Sagola murchisonensis Park and Carlton, new species</p><p>(Figs. 58d, 59d, 61)</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND NN Mt Murchison 914m 26 mar 1975 K.W. Walker ”, “Tussock Litter 75/123”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola murchisonensis Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Nelson: 1♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named after the type locality, Mt. Murchison, Nelson.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.4–2.6 mm; male antennomeres not enlarged; male elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Nelson.</p><p>Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58d). Head: Round, as long as wide, widest across eyes (Fig. 58d). Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardrop-shaped. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 58d). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patches of microtrichia. Aedeagus: Median lobe of genitalia divided, minor lobe membranous (Fig. 59d). Phallobase of median lobe symmetrical and rounded (Fig. 59d). Parameres symmetrical, as wide as median lobe, with setae apically (Fig. 59d).</p><p>Distribution. Nelson (Fig. 61: triangle).</p><p>Habitat. Specimens of this species were collected by sifting tussock litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0411213CF1FB50FC05FCA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0411223E97FC96FDDFFBC4.text	03D82A3EBD0411223E97FC96FDDFFBC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola fiordlandensis Park and Carlton	<div><p>Sagola fiordlandensis Park and Carlton, new species</p><p>(Figs. 58e, 59e, 62)</p><p>Type Material. Holotype. NEW ZEALAND: Fiordland: 1♂ (NZAC), “ NEW ZEALAND FD Secretary I ridge towards Mt Grono 853m 30 Nov 1981 ”, “ C.F. Butcher moss &amp; mats 81/189”, “Duplicate specimens in alcohol”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola fiordlandensis Park and Carlton 2013 ” . Paratypes (n = 20: 10 males; 10 females). NEW ZEALAND: Fiordland: 2♀♀ (NZAC), same data as holotype; 3♂♂ 5♀♀ (NZAC), Secretary I, Mt. Grono, 853m, 27 XI 1981, C.F. Butcher, mats, moss &amp; tussock ; 1♂ (NZAC), Secretary I, Grono Bay, 24 III 1984, C.F. Butcher, litter ; 1♂ (NZAC), Secretary I, Grono Bay, 29 III 1984, T. Haslam, mats plants (NZAC) ; 1♀, Secretary I, Grono Bay, 28 XI 1981, C.F. Butcher, litter ; 1♂ (LUNZ), Fiordland NP, Borland Saddle, 940m, 2 II 1982, J.W. Early, sweeping tussock ; 1♂ 1♀ (NZAC), summit, Tempest Spur, w Olivine Ra, 1463m, 25 I 1975, G.W. Ramsay, litter; Marlborough : 1♂ (NZAC), Richmond Ra, Fell pk, 1311m, 18 III 1969, A.C. Eyles, litter ; Westland: 2♂♂ 1♀ (LUNZ), Mt. Cook NP, 1500m, 23 I 1977, W.J. Sweney .</p><p>Etymology. The species name is based on the biotic region of some of the type material, Fiordland.</p><p>Diagnosis. This species can be distinguished from the other species of the misella species-group by the following combination of characters: size larger, 2.6–3.0 mm; male antennomeres 4–10 enlarged; eye larger, approximately one-half length of temple; male abdominal ventrites V–VI medially flattened; shape of genitalia unique to species.</p><p>Description. Length 2.6–3.0 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58e). Head: Round, as long as wide, widest across eyes (Fig. 58e). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 6–11 tubercles. Female antennomere 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate, 6–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardrop-shaped. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 58e), elytra subquadrate in females and some males. Male meso- and metathorax trapezoidal, longer than wide, females and some males as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in females and some males. Male abdominal ventrites V–VI medially flatten. Aedeagus: Apical lobe of genitalia acute (Fig. 59e). Phallobase of median lobe symmetrical and rounded (Fig. 59e). Parameres asymmetrical, right longer than left, with setae apically (Fig. 59e).</p><p>Distribution. Fiordland, Marlborough, Westland (Fig. 62: black circles).</p><p>Habitat. Specimens of this species were collected mostly by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0411223E97FC96FDDFFBC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0711233C98FB7AFD88FE36.text	03D82A3EBD0711233C98FB7AFD88FE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hunterensis Park and Carlton	<div><p>Sagola hunterensis Park and Carlton, new species</p><p>(Figs. 58f, 59f, 62)</p><p>Type Material. Holotype. NEW ZEALAND: Fiordland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: FD: Hunter Mts., N. Mt. Burns 1300m, 10 I 1970 J.I. Townsend, mats 70/6”, “ HOLOTYPE Sagola hunterensis Park and Carlton 2013 ” . Paratypes (n = 5: 2 males; 3 females). NEW ZEALAND: Fiordland: 3♀♀ (NZAC), Hunter Mts, S. Borland, 760m, 14 I 1970, J.I. Townsend, mats 70/106 ; 1♂ (JTN), Mt. Burns, Hunter Mtns, 10 XII 2005, J. Nunn, in tussock litter ; 1♂ (FMNH), Fiordland NP, Borland rd, Borland Saddle vic, 980m, 45″44.81′S 167″22.88′E, 10 XII 2005, Nothofagus menziesii forest in ravine, FMHD#2005-090, litter, A. Newton, ANMT site 1171 .</p><p>Etymology. This species is named after the type locality, Hunter Mts., Fiordland.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: larger body, length 2.1–2.5 mm; male antennomeres 5–10 weakly enlarged; posterior frontal fovea elongate; male elytra subquadrate; male abdominal ventrite V medially flattened, VII small with acute process medially; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.1–2.5 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58f). Head: Round, as long as wide, widest across eyes (Fig. 58f). Male antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–10 subquadrate, 5–10 weakly enlarged, 6–11 with tubercles. Female antennomere 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate, 6–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea elongate. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 58f). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patches of microtrichia. Male abdominal ventrite V medially flattened, VII small with acute process medially. Aedeagus: Apical lobe of genitalia divided, major lobe diamond-shaped (Fig. 59f). Phallobase of median lobe symmetrical and rounded (Fig. 59f). Parameres symmetrical, blunt with setae apically (Fig. 59f).</p><p>Distribution. Fiordland (Fig. 62: triangles).</p><p>Habitat. Specimens of this species were collected by sifting tussock litter and material from mats.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0711233C98FB7AFD88FE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD0611233CFCFE25FBC4FE0B.text	03D82A3EBD0611233CFCFE25FBC4FE0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola humpridgensis Park and Carlton	<div><p>Sagola humpridgensis Park and Carlton, new species</p><p>(Figs. 58g, 59g, 62)</p><p>Type Material. Holotype. NEW ZEALAND: Fiordland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: FD: Hump Ridge, 1220m 6 II 1976, G.W. Ramsay Litter 76/38”, “ HOLOTYPE Sagola humpridgensis Park and Carlton 2013 ” . Paratypes (n = 17: 7 males; 10 females). NEW ZEALAND: Fiordland: 4♂♂ 2♀♀ (NZAC), same data as holotype; 1♂ (NZAC), Wilmot Pass, 630m, 26 I 1970, G.W. Ramsay, litter 70/93 ; 3♀♀ (NZAC), Turret Ra, Wolfe Flat, 900m, 22 I 1970, A.C. Eyles, litter 70/90 ; 3♀♀ (NZAC), Hump Ridge, 12 III 1939, Fairburn . Southland: 2♂♂ 1♀ (NZAC), MacLennan, 31m, 13 II 1968, J.I. Townsend, moss 68/30 ; 1♀ (NZAC), Manpouri, 21 I 1970, G.W. Ramsay, litter 70/96 .</p><p>Etymology. This species is named after the type locality, Hump Ridge, Fiordland.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.6–2.9 mm; male antennomeres 6–10 enlarged; male elytra subquadrate; male abdominal ventrite V medially flattened, VII small with acute process medially; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.6–2.9 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58g). Head: Round, as long as wide, widest across eyes (Fig. 58g). Male antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3–10 subquadrate, 6–10 enlarged, 6–11 with tubercles. Female antennomeres 2–5 longer than wide, 6–10 subquadrate, 7–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea elongate. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 58g). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patches of microtrichia. Male abdominal ventrite V medially flattened, VII small with acute process medially. Aedeagus: Apical lobe of genitalia divided, major lobe shorter and more slender (Fig. 59g). Phallobase of median lobe symmetrical and rounded (Fig. 59g). Parameres symmetrical, blunt with setae apically (Fig. 59g).</p><p>Distribution. Fiordland, Southland (Fig. 62: black squares).</p><p>Habitat. Specimens of this species were collected by sifting moss and leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD0611233CFCFE25FBC4FE0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD06113C3EEAFE4FFE17FF68.text	03D82A3EBD06113C3EEAFE4FFE17FF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola clunieae Park and Carlton	<div><p>Sagola clunieae Park and Carlton, new species</p><p>(Figs. 58h, 59h, 62)</p><p>Type Material. Holotype. NEW ZEALAND: Fiordland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND FD Secretary I Top of Ridge 29 May 1982 S. Brasch ”, “Litter 82/65”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola clunieae Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Fiordland: 1♀ (NZAC), Secretary I, ridge towards, Mt Grono, 853m, 27 XI 1981, C.F. Butcher, Alpine mats, moss and tussock .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Leonie Clunie. She assisted with specimen access through loans and during the first author’ s visit to the New Zealand Arthropod Collection.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.2–2.3 mm; male antennomeres 6–10 enlarged; male abdominal ventrite V medially flattened; shape of antennomeres and genitalia unique to species; only known from Secretary Island in Fiordland.</p><p>Description. Length 2.2–2.3 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58h). Head: Round, as long as wide, widest across eyes (Fig. 58h). Male antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 weakly enlarged, 6–11 with tubercles. Female antennomeres 2–7 longer than wide, 8–10 subquadrate, 8–11 with tubercles. Frontal rostrum rectangular, meeting each other. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea oval. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 58h). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patches of microtrichia. Male abdominal ventrite V medially flattened. Aedeagus: Apical lobe of genitalia triangular (Fig. 59h). Phallobase of median lobe symmetrical and rounded (Fig. 59h). Parameres symmetrical blunt with setae apically (Fig. 59h).</p><p>Distribution. Fiordland (Fig. 62: star).</p><p>Habitat. Specimens of this species were collected by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD06113C3EEAFE4FFE17FF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD19113C3CA6FEEFFB46FEEC.text	03D82A3EBD19113C3CA6FEEFFB46FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola turretensis Park and Carlton	<div><p>Sagola turretensis Park and Carlton, new species</p><p>(Figs. 58i, 59i, 63)</p><p>Type Material. Holotype. NEW ZEALAND: Fiordland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: FD: Turret Ra., N of Percy Sdle 1250m, 16 I 1970, G.W. Ramsay, Grass 70/52”, “ HOLOTYPE Sagola turretensis Park and Carlton 2013 ” . Paratypes (n = 10: 3 males; 7 females). NEW ZEALAND: Fiordland: 1♀ (NZAC), same data as holotype; 1♂ (NZAC), Hunter Mts, W. Borland Sdle, 760m, 12 I 1970, J.I. Townsend, moss 70/10; Marlborough : 1♀ (NZAC), Richmond Ra, Johnson Pk, 1500m, 13 III 1969, J.C. Watt, litter 69/102; Marlborough Sounds: 1♂ 1♀ (NZAC), Ship Cove, 30 XI 1972, J.S. Dugdale, litter 72/273; Southland : 2♂♂ 1♀ (JTN), Spence Burn Basin, 1150m, Takitimu Mtns, 18 I 2000, tussock litter, J. Nunn ; 1♀ (JTN), Spence Burn Basin, 1150m, Takitimu Mtn, 17 I 2000, J. Nunn, tussock litter ; 1♀ (JTN), Spence Basin, Takitimu Rng, 17 I 2000, J. Nunn, tussock litter .</p><p>Etymology. This species is named after the type locality, Turret Range, Fiordland.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.5–2.7 mm; male antennomeres 4–10 enlarged; eye larger, approximately one-half length of temple; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.5–2.7 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58i). Head: Round, as long as wide, widest across eyes (Fig. 58i). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 6–11 with tubercles. Female antennomere 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate, 6–11 with tubercles. Frontal rostrum rectangular, lobe continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea elongate. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular (Fig. 58i), elytra subquadrate in female and some males. Male meso- and metathorax trapezoidal, longer than wide, female and some males as long as wide. Abdomen: Male tergite IV with a pair of transverse patches of microtrichia, patches absent in females and some males. Aedeagus: Median lobe of genitalia twisted with membranous tube (Fig. 59i). Phallobase of median lobe symmetrical and rounded (Fig. 59i). Parameres asymmetrical, right longer than left, with setae apically (Fig. 59i).</p><p>Distribution. Fiordland, Marlborough, Marlborough Sounds, Southland (Fig. 63: black circles).</p><p>Habitat. Specimens of this species were collected by sifting grass, moss, and tussock litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD19113C3CA6FEEFFB46FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD19113D3EBCFE6AFE6BFBDE.text	03D82A3EBD19113D3EBCFE6AFE6BFBDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola otagoensis Park and Carlton	<div><p>Sagola otagoensis Park and Carlton, new species</p><p>(Figs. 58j, 59j–k, 63)</p><p>Type Material. Holotype. NEW ZEALAND: Central Otago: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand CO Ailsa Craig Lammermoor Rng 7-Dec- 08”, “Sifted moss and tussock litter”, “ HOLOTYPE Sagola otagoensis Park and Carlton 2013 ”. The original label does not mention who collected the type specimen, but it was collected by J. Nunn. Paratypes (n = 11: 4 males; 7 females). NEW ZEALAND: Central Otago: 2♂♂ 7♀♀ (JTN), same data as holotype; 2♂♂ (JTN), Ailsa Craig, Lammerlaw Rng, 14 XII 2008, J. Nunn .</p><p>Etymology. The species name is based on the biotic region of the type material, Central Otago.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.6–2.8 mm; male antennomeres 4–10 enlarged; eye larger, approximately one-half length of temple; male abdominal ventrite V with round process medially; shape of antennomeres and genitalia unique to species; known from Central Otago of South Island.</p><p>Description. Length 2.6–2.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58j). Head: Round, as long as wide, widest across eyes (Fig. 58j). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 6–11 with tubercles. Female antennomere 2 longer than wide, 3–10 subquadrate, 6–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea long. Eye prominent, approximately onehalf length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 58j). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV without patches of microtrichia. Male abdominal ventrite V with round process medially. Aedeagus: Median lobe of genitalia triangular, apical lobe rectangular vertically (Fig. 59j–k). Phallobase of median lobe symmetrical and rounded (Fig. 59j). Parameres asymmetrical, right longer than left, with setae apically (Fig. 59j).</p><p>Distribution. Central Otago (Fig. 63: triangle).</p><p>Habitat. Specimens of this species were collected by sifting tussock litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD19113D3EBCFE6AFE6BFBDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD18113D3CFDFB9DFC17FCA0.text	03D82A3EBD18113D3CFDFB9DFC17FCA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola southlandensis Park and Carlton	<div><p>Sagola southlandensis Park and Carlton, new species</p><p>(Figs. 58k, 59l, 63)</p><p>Type Material. Holotype. NEW ZEALAND: Southland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand SL Croyden Bush Gore 5-Sep-04”, “Forest floor litter from area subject to flooding”, “76.03 Southland Hills Hokonui”, “ HOLOTYPE Sagola southlandensis Park and Carlton 2013 ”. Original label does not mention who collected the type specimen, but it was collected by J. Nunn. Paratype (1 male). NEW ZEALAND: Southland: 1♀ (JTN), same data as holotype .</p><p>Etymology. The species name is based on the biotic region of the type material, Southland.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: larger body, length 2.7–2.8 mm; male antennomeres 4–10 enlarged; eye larger, approximately one-half length of temple; shape of antennomeres and genitalia unique to species; only known from Southland.</p><p>Description of Male. Length 2.7–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58k). Head: Round, as long as wide, widest across eyes (Fig. 58k). Antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 6–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea elongate. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 58k). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV without pair of patches of microtrichia. Aedeagus: Median lobe of genitalia triangular, curved as S-shaped vertically (Fig. 59l). Phallobase of median lobe symmetrical and rounded (Fig. 59l). Parameres asymmetrical, right longer than left, with setae apically (Fig. 59l).</p><p>Distribution. Southland (Fig. 63: black square).</p><p>Habitat. Specimens of this species were collected by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD18113D3CFDFB9DFC17FCA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD18113E3EFCFC96FE21FD2F.text	03D82A3EBD18113E3EFCFC96FE21FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola huvali Park and Carlton	<div><p>Sagola huvali Park and Carlton, new species</p><p>(Figs. 58l, 59m, 63)</p><p>Type Material. Holotype. NEW ZEALAND: Southland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND Longwood Range 8 Feb 1976 G.W. Ramsay litter 76/36”, “Duplicate specimens in alcohol”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola huvali Park and Carlton 2013 ” . Paratypes (n = 2: 1 male; 1 female). NEW ZEALAND: Southland: 1♂ 1♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named for one of our lab mates and an enthusiastic supporter of this study, Forest Huval.</p><p>Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: size larger, 2.6–2.8 mm; male antennomeres 6–10 enlarged; eye larger, approximately one-half length of temple; male abdominal ventrites V–VI with round medial process, VII with a pair of acute processes; shape of antennomeres and genitalia unique to species; known from Southland.</p><p>Description. Length 2.6–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58l). Head: Round, as long as wide, widest across eyes (Fig. 58l). Male antennomere 1 approximately 1.5 times longer than wide, 2–4 longer than wide, 5–10 subquadrate, 6–10 enlarged, 6–11 with tubercles. Female antennomeres 2–7 longer than wide, 8–10 subquadrate, 5–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea elongate. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate (Fig. 58l). Male meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patches of microtrichia. Male abdominal ventrites V–VI with round medial process, VII with a pair of acute processes. Aedeagus: Apical lobe of genitalia triangular, with membranous tube (Fig. 59m). Phallobase of median lobe symmetrical and rounded (Fig. 59m). Parameres asymmetrical, right wider than left, with setae apically (Fig. 59m).</p><p>Distribution. Southland (Fig. 63: star).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD18113E3EFCFC96FE21FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1B113E3F5DFD86FB7FFA74.text	03D82A3EBD1B113E3F5DFD86FB7FFA74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>26. moehauensis species-group</p><p>(2 species)</p><p>Diagnosis. Members of the moehauensis species - group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.5–2.8 mm; antennomere 1 approximately 2 times longer than wide with dull surface; male head triangular with projecting temporal lobe (Fig. 64e–f) and female head longer than wide; anterior frontal fovea oval, partially covered by frontal rostrum, and posterior frontal fovea oval (Fig. 64f); hind wings fully developed; abdominal tergites IV–VI with discal carinae; parameres divided (Fig. 64c–d).</p><p>KEY TO SPECIES OF THE MOEHAUENSIS SPECIES - GROUP</p><p>The key is based on male specimens because female specimens are indistinguishable based on external morphology.</p><p>1. Temporal lobes of ventral surface of head flattened (Fig. 64e: arrow); median lobe of genitalia subquadrate and phallobase of genitalia subquadrate (Fig. 64c); parameres narrowly divided (Fig. 64c) ................................ ......................... S. moehauensis new species</p><p>1′. Temporal lobes of ventral surface of head concave; median lobe of genitalia rectangular and phallobase of genitalia triangular (Fig. 64d); paramere broadly divided (Fig. 64d) .......................... S. horokakaensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD1B113E3F5DFD86FB7FFA74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1B113E3CE8FD13FC01FDD1.text	03D82A3EBD1B113E3CE8FD13FC01FDD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola tennysonensis Park and Carlton	<div><p>Sagola tennysonensis Park and Carlton, new species</p><p>(Figs. 58m, 59n, 63)</p><p>Type Material. Holotype. NEW ZEALAND: Marlborough Sounds: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: SD: Tennyson Inlet Road, 115-135m, 30 XII 84-5 I 85, forest streams. hdwd.-podo.-nikau for., A. Newton / M. Thayer 723, berl., leaf &amp; log litter, forest floor”, HOLOTYPE Sagola tennysonensis Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Westland: 1♂ (LSAM), 11.2km ne Franz Josef Glacier, nr Lake Mapourika, 150m, Otto’ s Corner picnic area, 43′18S 170′14E, #064, Coastal forest leaf litter berlese, 15 I 1998, C. Carlton, R. Leschen .</p><p>Etymology. This species is named after the type locality, Tennyson Inlet, Marlborough Sounds.</p><p>Diagnosis. This species can be distinguished from the other species of the misella species-group by the following combination of characters: body larger, length 2.7–2.9 mm; male antennomeres 4–10 longer than wide and enlarged; eye larger, approximately one-half length of temple; male abdominal ventrites V–VI with round medial process, VII with rounded concave process medially; shape of antennomeres and genitalia unique to species.</p><p>Description of Male. Length 2.7–2.9 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 58m). Head: Round, as long as wide, widest across eyes (Fig. 58m). Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate, 4–10 enlarged, 4–11 bearing tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardropshaped. Eye prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 58m). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia. Male abdominal ventrites V–VI with round medial process, VII with rounded concave process medially. Aedeagus: Apical lobe of genitalia rectangular, vertically with membranous tube (Fig. 59n). Phallobase of median lobe symmetrical and rounded (Fig. 59n). Parameres asymmetrical, right longer than left, with setae apically (Fig. 59n).</p><p>Distribution. Marlborough Sounds, Westland (Fig. 63: white circle).</p><p>Habitat. Specimens of this species were collected by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1B113E3CE8FD13FC01FDD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1B11383E86F9E4FD8EFD70.text	03D82A3EBD1B11383E86F9E4FD8EFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola moehauensis Park and Carlton	<div><p>Sagola moehauensis Park and Carlton, new species</p><p>(Figs. 64a, 64c, 64e–f, 65)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND CL Mt Moehau 600m 16 Oct 1980 J.C. Watt Litter 80/89”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola moehauensis Park and Carlton 2013 ”. Paratypes (n = 6: 5 males; 1 female). NEW ZEALAND: Bay of Plenty: 1♂ (AMNZ), Orete Forest, Te Puia Hut, 240m, 25 I 1993, J.W. Early, podocarp-broadleaf forest, yellow pan trap; 1♂ (slide-mounted; LUNZ), Papatea, NZMS 260 Y14 386806, 22 X 1992 – 25 I 1993, J.W. Marris, R. M. Emberson, pit trap, podocarp-broadleaf forest; 1♂ (NZAC), Upper Kaimai, 26 XII 1943, leaf mould, A.E. Brookes collection; Coromandel: 1♂ (AMNZ), Moehau Ra, Mist Trust main ridge, 200m, 27 VI–18 VII 1999, Y. Forbes, Kanuka-broadleaf-rata forest, malaise trap L7169; Taupo: 1♀ (FMNH), Whirinaki Forest Park , Arohaki Lagoon tr, 500m, 38″40.82′S 176″40.032′E, 21 XI 2005, mixed broadleaf forest incl. Belischmidia tawa, FMHD#2005-024, litter, A. Newton, M. Thayer et al., ANMT site 1147; Waikato: 1♂ (JTN), Mahaukura tr, Mt. Pirongia, 18 XI 2005, J. Nunn .</p><p>Etymology. This species is named after the type locality, Mt. Moehau, Coromandel.</p><p>Diagnosis. This species can be distinguished from the other species of the moehauensis speciesgroup by the flat, ventrally projecting temporal lobe of the head and the genitalia being small, &lt;0.2 mm.</p><p>Description. Length 2.5–2.7 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler (Fig. 64a). Head: Male head triangular, widest across temples (Fig. 64f). Ventral aspect of temple of male head projecting and flattened (Fig. 64e: arrow). Antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus reaching posterior margin of eyes. Eye small, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 64a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of patches of microtrichia. Aedeagus: Male genitalia small, &lt;0.2 mm (Fig. 64c). Median lobe subquadrate (Fig. 64c). Phallobase of median lobe symmetrical and subquadrate (Fig. 64c). Parameres symmetrical and shallowly divided (Fig. 64c).</p><p>Distribution. Bay of Plenty, Coromandel, Taupo, Waikato (Fig. 65: black circles).</p><p>Habitat. Specimens of this species were collected using Malaise or pitfall traps or by sifting leaf litter in broadleaf and podocarp forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1B11383E86F9E4FD8EFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1D11383C96FCFEFDD7FCA7.text	03D82A3EBD1D11383C96FCFEFDD7FCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola horokakaensis Park and Carlton	<div><p>Sagola horokakaensis Park and Carlton</p><p>(Figs. 64b, 64d, 65)</p></div>	https://treatment.plazi.org/id/03D82A3EBD1D11383C96FCFEFDD7FCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1D113A3F7EFBD7FD45FEDB.text	03D82A3EBD1D113A3F7EFBD7FD45FEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola major Sharp 1874	<div><p>27. major species-group</p><p>(6 species)</p><p>Diagnosis. Members of the major species-group can be distinguished from other Sagola speciesgroups by the following combination of characters: body small, length 1.5–2.8 mm; male head triangular, widest across temples (Fig. 66n), female head bluntly triangular, widest across eyes; temple of male head depressed and projecting ventrally (Fig. 66n); male neck with ventral patch of dense setae anteriorly (Fig. 66o–t); male hind wings fully developed; abdominal tergites IV–VI with discal carinae; genitalia robust and convex (Fig. 66g –l).</p><p>KEY TO SPECIES OF THE MAJOR SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Frontal sulcus exceeding eyes; posterior frontal fovea elongate................................ ............... S. triregia Théry and Leschen</p><p>1′. Frontal sulcus reaching midpoint of eye; posterior frontal fovea oval.....................2</p><p>2(1′). Fore trochanter with acute triangular process (Fig. 66m: arrow); abdominal ventrite VIII with pair of small processes ............. ......................................... S. major Sharp</p><p>2′. Fore trochanter round; abdominal ventrite VIII simple..............................................3</p><p>3(2′). Temporal lobe of ventral surface of head with acute process; only known from Three Kings Islands (Fig. 67: black square)......... ............................... S. marrisi new species</p><p>3′. Temporal lobe of ventral surface of head concave; known from North Island or South Island.............................................4</p><p>4(3′). Smaller size, length 1.5–1.7 mm (Fig. 66e); antennomere 4 subquadrate; paramere narrower than median lobe of genitalia (Fig. 66k)........ S. prowellae new species</p><p>4′. Larger size, length&gt; 1.8 mm; antennomere 4 longer than wide; paramere broader than median lobe of genitalia..........................5</p><p>5(4′). Median lobe of genitalia with rectangular process dorsally (Fig. 66j); paramere widest at middle, with setae from apex to midpoint (Fig. 66j); only known from Auckland of North Island (Fig. 67: star).......................... ................................. S. boothi new species</p><p>5′. Median lobe of genitalia without process but divided vertically (Fig. 66l); paramere widest at base, with setae apically (Fig. 66l); known from South Island............................. ..................... S. mackenziensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD1D113A3F7EFBD7FD45FEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1F113B3D5FFE82FDC8FDFC.text	03D82A3EBD1F113B3D5FFE82FDC8FDFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola major Sharp 1874	<div><p>Sagola major Sharp, 1874</p><p>(Figs. 66a, 66g, 66m –o, 67)</p><p>Sagola major Sharp, 1874: 507 . Broun 1880: 135; Schaufuss 1888: 84; Raffray 1893: 18, 1904: 497, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola ruficeps Broun, 1893b: 1053 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola mimica Broun, 1893b: 1419 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola elongata Broun, 1893b: 1423 . Raffray 1904: 498, 1911: 6; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola longipes Broun, 1915: 287 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “ N. Zeal. ” [white label, printed]; “Sharp. Coll. 1905-313.” [white label, printed]; “ Sagola major ♂ Type. D.S.” [white label, handwritten] . Holotype of Sagola elongata . NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2472.” [white label, handwritten]; “Karaka.” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola . elongata” [white label, handwritten] . Holotype of Sagola longipes . NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3702. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Woodhill. Waitakerei.” [white label, handwritten]; “ Sagola longipes ♀.” [white label, handwritten] . Holotype of Sagola mimica . NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2467. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hunua. Clevedon.” [white label, handwritten]; “ Sagola mimica ” [white label, handwritten] . Syntypes of Sagola ruficeps . NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “1882.” [white label, handwritten]; “ ♀ ” [white label, handwritten]; “ Hunua. Clevedon. ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ruficeps ” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . 1♂ (BMNH), glued on rectangular card, “1882.” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “ Hunua. Clevedon. ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] . 1♂ (BMNH), glued on rectangular card, “1882.” [white label, handwritten]; “N. Zealand. 93–56.” [white label, printed]; “ Sagola ruficeps Broun ” [white label, handwritten] .</p><p>Additional Material (n = 28: 14 males; 14 females). NEW ZEALAND: Northland: 1♂, Pekarau, Mangonui, A.E. Brookes, 24 III 1918; Wellington: 13♂♂ 14♀♀ (1♂, slidemounted), Waikawa Beach, 18 III 1990 .</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body length 2.5–2.8 mm; eye large and prominent, two-thirds length of temple; ventral temple of male head with depressed and projecting temporal lobe and surface flattened; male fore trochanter with acute triangular process; male abdominal ventrite VIII with a pair of small processes; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.5–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66a). Head: Male head triangular, widest across temples (Fig. 66n). Female head bluntly triangular, widest across eyes. Ventral temple of male head with depressed and projecting temporal lobe and surface flattened (Fig. 66o). Antennomere 1 approximately 2.5 times longer than wide, 2–7 longer than wide, 8–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, two-thirds length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Male fore trochanter with acute triangular process. Elytra rectangular (Fig. 66a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Male abdominal ventrite VIII with pair of small processes. Aedeagus: Median lobe of genitalia broader apically with tube extending from apex (Fig. 66g). Phallobase of median lobe symmetrical and rounded (Fig. 66g). Parameres symmetrical, broader than median lobe, setose from midpoints to apices (Fig. 66g).</p><p>Type Locality. New Zealand (specific locality not mentioned) .</p><p>Distribution. Auckland, Northland, Wellington (Fig. 67: black circles).</p><p>Habitat. Unknown.</p><p>Comments. Specimens of S. major can be distinguished from those of other species by the shape of the antennomeres, large eye, ventral temple of male head depressed and projecting, surface flat; male fore trochanter with acute triangular process; male abdominal ventrite VIII with a pair of small processes. The type specimens of S. elongata, S. longipes, S. mimica, and S. ruficeps share these diagnostic characters. For these reasons, we have placed S. elongata, S. longipes, S. mimica, and S. ruficeps in synonymy with S. major .</p></div>	https://treatment.plazi.org/id/03D82A3EBD1F113B3D5FFE82FDC8FDFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1E11343CFDFD7BFE78FE61.text	03D82A3EBD1E11343CFDFD7BFE78FE61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola triregia Thery and Leschen 2013	<div><p>Sagola triregia Théry and Leschen, 2013</p><p>(Figs. 66b, 66h, 66p, 67)</p><p>Sagola triregia Théry and Leschen, 2013: 45 .</p><p>Type Material. Paratype. New Zealand: Three Kings Islands: 1♀ (NZAC), glued on rectangular card, “NEW ZEALAND, TH Great Is, Tasman Valley along Tasman Stm 10 Nov 2008” [white label, printed]; “sifting leaf litter and rotten wood” [white label, printed]; “ T. Buckley, R. Leschen TH053 34°09.733/172°08.610″ [white label, printed]; “Paratype Sagola triregia Théory &amp; Leschen 2012 ” [blue label, printed] .</p><p>Additional Material (n = 97: 30 males; 67 females). NEW ZEALAND: Three Kings Islands: 1♂ 1♀, Great I, Bald Hill, sifted litter 99/27, 13 IV 1999, T. K. Crosby ; 1♂ 3♀♀, Great I, sifted litter 99/25, 13 IV 1999, T. K. Crosby ; 2♂♂ 8♀♀, South West I, summit, 25 XI 1983, J.C. Watt, sifted litter 83/126 ; 1♂ 1♀, South West I, 26 XI 1983, J.C. Watt, litter 83/128 ; 2♂♂ 1♀, North East I, summit, 1 XII 1983, J.C. Watt, litter and humus 83/114 ; 1♀, South West I, 200m, 21–21 XI 1985, M. Potter, litter 85/81 ; 11♂♂ 21♀♀, Castaway Camp, 29 XI 1970, G.W. Ramsay, litter ; 4♂♂ 10♀♀, South West Island, 1 XII 1970, G. Kuschel, litter 70/237 ; 2♂♂ 3♀♀, Tasman Valley, 26 XI 1970, J.C. Watt, litter 70/223 ; 1♂, Castaway Camp, 80m, 16 XI 1970, G. Kuschel, litter 70/188 ; 1♂, South West Is, 1 XII 1970, G.W. Ramsay, litter 70/240 ; 5♀♀, Castaway Camp, 85m, 16 XI 1970, G. Kuschel, litter 70/190 ; 2♀♀, Castaway Camp, 85m, 16 XI 1970, G. Kuschel, litter 70/191 ; 1♀, Castaway Camp, 85m, 12 XI 1970, G. Kuschel, litter 70/183 ; 1♀, Tasman Valley, 25 XI 1970, G. Kuschel, litter 70/215 ; 4♂♂ 9♀♀, South West I, summit ridge, 13 I 1951, E.G. Turbott, leaf litter under puka forest .</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body length 2.4–2.6 mm; eye one-half length of temple; ventral temple of male head with depressed and projecting temporal lobe and surface flattened; shape of antennomeres and genitalia unique to species; only known from Three Kings Islands.</p><p>Redescription. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66b). Head: Male head triangular, widest across temples (Fig. 66b). Female head bluntly triangular, widest across eyes. Ventral temple of male head with depressed and projecting temporal lobe, surface flattened (Fig. 66p). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus shallow, extended beyond eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea elongate. Eye prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 66b), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia funnel-shaped with round process at middle (Fig. 66h). Phallobase of median lobe symmetrical and rounded (Fig. 66h). Parameres symmetrical, setose from midpoints to apices (Fig. 66h).</p><p>Type Locality. Three Kings Islands, New Zealand .</p><p>Distribution. Three Kings Islands (Fig. 67: triangle).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1E11343CFDFD7BFE78FE61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1111343CF1FDD5FC12FEEC.text	03D82A3EBD1111343CF1FDD5FC12FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola marrisi Park and Carlton	<div><p>Sagola marrisi Park and Carlton, new species</p><p>(Figs. 66c, 66i, 66q, 67)</p><p>Type Material. Holotype. NEW ZEALAND: Three Kings Islands: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: TH: Castaway Camp 29 XI 1970, G.W. Ramsay Litter 70/234”, “ HOLOTYPE Sagola marrisi Park and Carlton 2013 ” . Paratypes (7 males). NEW ZEALAND: Three Kings Islands: 7♂♂ (NZAC), same data as holotype .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, John Marris. He provided many valuable specimens from South Island.</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body length 2.2–2.4 mm; eye one-half length of temple; ventral temple of male head with depressed and weakly projecting temporal lobe and surface bearing small process; pair of thick, dense setae on gular depression; shape of antennomeres and genitalia unique to species; only known from Three Kings Islands.</p><p>Description of Male. Length 2.2–2.4 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66c). Head: Bluntly triangular, widest across eyes (Fig. 66c). Ventral temple of head with depressed and weakly projecting temporal lobe and surface bearing small process (Fig. 66q). Pair of thick, dense setae on gular depression. Antennomere 1 approximately 2 times longer than wide with dull surface, 2 longer than wide, 3–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 66c). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia rectangular with tube extending from apex (Fig. 66i). Phallobase of median lobe symmetrical and rounded (Fig. 66i). Parameres symmetrical, broader than median lobe, setose from midpoints to apices (Fig. 66i).</p><p>Distribution. Three Kings Islands (Fig. 67: black square).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1111343CF1FDD5FC12FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1111353E8BFE52FE81FE7B.text	03D82A3EBD1111353E8BFE52FE81FE7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola boothi Park and Carlton	<div><p>Sagola boothi Park and Carlton, new species</p><p>(Figs. 66d, 66j, 66r, 67)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND AK Bethells Matuku Reserve 23 Jun 1987 R. C. Craw ”, “Sifted rotten wood 87/9”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola boothi Park and Carlton 2013 ” . Paratypes (n = 4: 3 males; 1 female). NEW ZEALAND: Auckland: 1♂ (NZAC), same data as holotype; 1♂ 1♀ (DSC), Matuku Reserve, 45m, 6.5km w Waitakere end of Jonker’ s Road, 2–5 IV 2010, L. Masner, yellow pan traps ; 1♂ (AMNZ), Waitakere Ra, Karekare, 14 II 2000, Wasp Survey, malaise trap .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Roger G. Booth. He assisted with type examinations and access to specimens during the first author’ s visit to the Natural History Museum, London.</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body length 1.8–2.0 mm; eye large, two-thirds length of temple; ventral temple of male head with depressed and projecting temporal lobe, surface concave; shape of antennomeres and genitalia unique to species; only known from Auckland.</p><p>Description. Length 1.8–2.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66d). Head: Male head triangular, widest across temples (Fig. 66d). Female head bluntly triangular, widest across eyes. Ventral temple of male head with depressed and projecting temporal lobe and surface concave (Fig. 66r). Thick, dense setae on male gular depression. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, two-thirds length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 66d). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broader apically with long process dorsally and membranous tube extending from apex (Fig. 66j). Phallobase of median lobe symmetrical and rounded (Fig. 66j). Parameres symmetrical, broader than median lobe, setose from midpoints to apices (Fig. 66j).</p><p>Distribution. Auckland (Fig. 67: star).</p><p>Habitat. Specimens of this species were collected using yellow pan and Malaise traps or by sifting rotten wood.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1111353E8BFE52FE81FE7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1011353CD2FDF8FC5BFE5C.text	03D82A3EBD1011353CD2FDF8FC5BFE5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola prowellae Park and Carlton	<div><p>Sagola prowellae Park and Carlton, new species</p><p>(Figs. 66e, 66k, 66s, 67)</p><p>Type Material. Holotype. NEW ZEALAND: Wellington: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand WN Johnson’ s Hill Karori 22 IV 1991 ”, “ HOLOTYPE Sagola prowellae Park and Carlton 2013 ”. The original label does not mention who collected the specimen, but it was collected by J. Nunn. Paratypes (n = 2: 1 male; 1 female). NEW ZEALAND: Wellington: 1♂ (JTN), Johnson’ s Hill, Karori, 17 VII 1988, J. Nunn; 1♀ (JTN), Johnson’ s Hill, Karori, 28 II 1987, J. Nunn .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Dorothy Prowell. She was a member of the first author’ s PhD advisory committee and provided valuable advice during this study, a major component of his dissertation.</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body small, length 1.5–1.7 mm; eye large and prominent, two-thirds length of temple; ventral temple of male head with depressed and distinctly projecting temporal lobe, surface concave; shape of antennomeres and genitalia unique to species; only known from Wellington.</p><p>Description. Length 1.5–1.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66e). Head: Male head triangular, widest across temples (Fig. 66e). Female head bluntly triangular, widest across eyes. Ventral temple of male head with depressed and distinctly projecting temporal lobe, surface concave (Fig. 66s). Thick, dense setae on male gular depression. Antennomere 1 approximately 2 times longer than wide with dull surface, 2 longer than wide, 3–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, two-thirds length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular (Fig. 66e). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia broader apically with tube extending from apex (Fig. 66k). Phallobase of median lobe symmetrical and rounded (Fig. 66k). Parameres symmetrical and narrow, with process at midpoint, setose from midpoints to apices (Fig. 66k).</p><p>Distribution. Wellington (Fig. 67: white circle).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1011353CD2FDF8FC5BFE5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1011363E94FE02FE78FDAA.text	03D82A3EBD1011363E94FE02FE78FDAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola mackenziensis Park and Carlton	<div><p>Sagola mackenziensis Park and Carlton, new species</p><p>(Figs. 66f, 66l, 66t, 67)</p><p>Type Material. Holotype. NEW ZEALAND: Mackenzie: ♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand MK Hooker Valley Mt Cook NP 14-Apr-06”, “ HOLOTYPE Sagola mackenziensis Park and Carlton 2013 ” . Paratypes (n = 5: 3 males; 2 females). NEW ZEALAND: Central Otago: 1♂ (NZAC), Mt. Ida, 3000′, Naseby, 8 XI 1968, J.I. Townsend, moss 68/161 ; Mid Canterbury: 1♀ (LUNZ), Mt Hutt, 1000m, Scotts Saddle, 27 XI 1981, R. M. Emberson, litter ; 1♀ (NZAC), Mt. Hutt, 11 V 2003, A.C. Eyles 2003, mountain beech litter ; Mackenzie: 1♂ (JTN), same data as holotype; 1♂ (JTN), Tasman Vly, Mt. Cook NP, 3 II 2006 .</p><p>Etymology. The species name is based on the biotic region of some of the type material, Mackenzie.</p><p>Diagnosis. This species can be distinguished from the other species of the major species-group by the following combination of characters: body length 1.9–2.1 mm; eye one-half length of temple; ventral temple of male head with depressed and projecting temporal lobe and surface weakly concave; shape of antennomeres and genitalia unique to species; found from South Island.</p><p>Description. Length 1.9–2.1 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 66f). Head: Male head triangular, widest across temples (Fig. 66f). Female head bluntly triangular, widest across eyes. Ventral temple of male head with depressed and projecting temporal lobe and surface weakly concave (Fig. 66t). Thick, dense setae on male gular depression. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus shallow, reaching midpoint of eye. Anterior frontal fovea oval, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, one-half length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 66f), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia divided vertically, upper lobe narrow apically (Fig. 66l). Phallobase of median lobe symmetrical and rounded (Fig. 66l). Parameres symmetrical broader than median lobe, with setae apically (Fig. 66l).</p><p>Distribution. Central Otago, Mid Canterbury, Mackenzie (Fig. 67: white squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBD1011363E94FE02FE78FDAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1311373D59FDADFDA8F988.text	03D82A3EBD1311373D59FDADFDA8F988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola insignis Broun, 1893 b: 1049	<div><p>28. insignis species-group</p><p>(24 species)</p><p>Diagnosis. Members of the insignis speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.2–3.2 mm; antennomere 1 approximately 1.5 times longer than wide; male head triangular, widest across temples (Fig. 70y–z) and female head bluntly triangular, widest across eyes; anterior and posterior frontal fovea present (Fig. 70y–z); temples of male head depressed and projecting ventrally (Fig. 69); male neck ventrally with patch of long and dense setae at anteriorly (Fig. 69); female hind wings reduced to small pads; abdominal tergites IV–VI with discal carinae; genitalia broad and flattened (Fig. 70a–x).</p><p>KEY TO SPECIES OF THE INSIGNIS SPECIES- GROUP</p><p>The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</p><p>1. Anterior margin of temporal lobe depressed and L-shaped (Fig. 69b, s) ..... ............................................................... 2</p><p>1′. Anterior margin of temporal lobe straight or weakly curved...................................3</p><p>2(1). Apical lobe of genitalia triangular (Fig. 70b); known from South Island (Fig. 71: triangles) ................................ ................................. S. hectorii Broun</p><p>2′. Apical lobe of genitalia round (Fig. 70s); known from North Island (Fig. 74: black circles) ...... S. hanae new species</p><p>3(1′). Temporal lobe of head weakly projecting, as long as eye or barely exceeding eye (Fig. 69c, f–g, k)..........................4</p><p>3′. Temporal lobe of head distinctly projecting, obviously exceeding eye (Fig. 69a– b, d–e, h–j, l–x)..................................... 7</p><p>4(3). Median lobe of genitalia with a pair of processes on lateral margins...............5</p><p>4′. Median lobe of genitalia without processes on lateral margins.....................6</p><p>5(4). Apical lobe of genitalia longer than wide (Fig. 70c)..... S. bipunctata Broun</p><p>5′. Apical lobe of genitalia subquadrate (Fig. 70f)............... S. robustula Broun</p><p>6(4′). Elytra rectangular; median lobe of genitalia widest at midpoint (Fig. 70g)....... ................................ S. pertinax Broun</p><p>6′. Elytra subquadrate; median lobe of genitalia widest at one-third length (Fig. 70k) ..... S. sheldoni new species</p><p>7(3′). Elytra rectangular................................8</p><p>7′. Elytra subquadrate.............................18</p><p>8(7). Median lobe of genitalia with pair of acute processes laterally......................9</p><p>8′. Median lobe of genitalia without process ............................................................. 14</p><p>9(8). Apical lobe of genitalia with round depression at apex.............................10</p><p>9′. Apical lobe of genitalia with round or straight apex ...................................... 11</p><p>10(9). Apical lobe of genitalia semicircular (Fig. 70a) .................. S. insignis Broun</p><p>10′. Apical lobe of genitalia subquadrate (Fig. 70d)............. S. angulifera Broun</p><p>11(9′). Pair of acute processes on lateral margin of genitalia reaching apex of apical lobe (Fig. 70m) ......................... ................. S. cuvierensis new species</p><p>11′. Pair of acute processes on lateral margin of genitalia shorter than apex of apical lobe....................................................12</p><p>12(11′). Pair of acute processes on lateral margin of genitalia larger, reaching one-third to one- fourth length of apical lobe ....... 13</p><p>12′. Pair of acute processes on lateral margin of genitalia minute, not exceeding onefifth length of apical lobe (Fig. 70q) .... ............... S. boudreauxae new species</p><p>13(12). Median lobe at least three times broader than parameres (Fig. 70e); known from North Island (Fig. 71: white circles) .... ................................. S. eminens Broun</p><p>13′. Median lobe approximately twice broader than parameres (Fig. 70i); known from South Island (Fig. 72: black squares) .... .................................... S. laticeps Broun</p><p>14(8′). Parameres with more than 10 setae apically (Fig. 70j)........ S. castanea Broun</p><p>14′. Parameres with less than 5 setae apically...................................................... 15</p><p>15(14′). Apical lobe of genitalia with semicircular depression apically (Fig. 70r) .......... ......................... S. taegyui new species</p><p>15′. Apical lobe of genitalia without depression apically.......................................16</p><p>16(15′). Apical lobe of genitalia rectangular (Fig. 70l)................................................ ............... S. mercuryensis new species</p><p>16′. Apical lobe of genitalia triangular....17</p><p>17(16′). Median lobe of genitalia widest at midpoint (Fig. 70h) .... S. monticola Broun</p><p>17′. Median lobe of genitalia widest at base (Fig. 70u)............................................... ................. S. egmontensis new species</p><p>18(7′). Apical lobe of genitalia with acute apex................................................................19</p><p>18′. Apical lobe of genitalia with blunt apex...........................................................20</p><p>19(18). Median lobe of genitalia with pair of acute processes laterally (Fig. 70n) ...... ....................... S. buckleyi new species</p><p>19′. Median lobe of genitalia without processes laterally (Fig. 70p) ..................... ............................ S. otteai new species</p><p>20(18′). Parameres triangular (Fig. 70t) ............. ...................... S. gyudongi new species</p><p>20′. Parameres slender..............................21</p><p>21(20′). Apical lobe of genitalia triangular....22</p><p>21′. Apical lobe of genitalia rectangular .... 23</p><p>22(21). Median lobe of genitalia widest at onethird length (Fig. 70x); known from South Island (Fig. 74: white squares) ... ................. S. jeongnamae new species</p><p>22′. Median lobe of genitalia widest at midpoint (Fig. 70o); known from North Island (Fig. 73: black squares).............. .................... S. brumfieldi new species</p><p>23(21′) Median lobe of genitalia widest at onethird length (Fig. 70w); known from Stewart Island (Fig. 74: white circles) .............. S. stewartensis new species</p><p>23′. Median lobe of genitalia widest at base (Fig. 70v); known from North Island (Fig. 74: stars) ..... S. boonei new species</p></div>	https://treatment.plazi.org/id/03D82A3EBD1311373D59FDADFDA8F988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1211333CBEF9BEFDB9FBEC.text	03D82A3EBD1211333CBEF9BEFDB9FBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola insignis Broun, 1893 b: 1049	<div><p>Sagola insignis Broun, 1893</p><p>(Figs. 68a, 69a, 70a, 71)</p><p>Sagola insignis Broun, 1893b: 1049 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola sobrina Broun, 1893b: 1050 . Hudson 1923: 365, 1934: 183; Raffray 1924: 233; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New Synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1875.” [white label, handwritten]; “Mokohinau” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola insignis ” [white label, handwritten] . Syntypes of Sagola sobrina: NEW ZEALAND: Northland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1877.” [white label, handwritten]; “Mokohinau” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola sobrina ” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “1877.” [white label, handwritten]; “Mokohinau I” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♀ (BMNH), glued on rectangular card, “1877. ♀ ” [white label, handwritten]; “Mokohinau I” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] .</p><p>Additional Material (n = 35: 12 males; 23 females). NEW ZEALAND: Auckland: 2♂♂, Lynfield WB, 13 VI 1980, G. Kuschel; 1♀, Lynfield, 28 VIII 1976, G. Kuschel, litter; 1♂, Manurewa, Murphy’ s Bush, 2 V 1981, G. Kuschel, rotten wood; 1♂, 20–40m, Grafton Gully, 4 III 2001 , S.E. Thorpe, bush litter; Coromandel: 5♂♂ 9♀♀, The Alderman I, Ruamahuaiti I, 40m, 6–11 XII 1994, J.W. Early, R.F. Gilbert, yellow pan trap ; 1♂ 1♀, Little Barrier I, 3 I 1952; 1♀, Little Barrier I, 2 I 1952; Northland: 4♂♂ 1♀, Mokohinau I, Trig I, 27 II 1978, G. Kuschel, litter; 2♀♀, Poor Knights I, Aorangi, ridge to Oneho Hill, 17 XI 1981 , J.C. Watt, litter; 2♀♀, Poor Knights I, Tawhiti Rahi, 4 XII 1980 , G. Kuschel, litter; 1♀, Poor Knights I, Aorangi, Puweto Valley, 14 XI 1981 , J.C. Watt, litter; 1♀, Hen &amp; Chickens I, Lady Alice I, I 1982 , L. Roberts, litter; 1♀, Poor Knights I, Aorangi, Puweto Valley, 9–17 XI 1981 , J.C. Watt, pit trap; 1♀, Poor Knights I, Aorangi, Crater Bay, 9–17 XI 1981 , J.C. Watt, pit trap.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–2.9 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts with three patches of dense setae medially; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.6–2.9 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68a). Head: Male head triangular, widest across temples (Fig. 69a). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye with concave ventral surface (Fig. 69a). Ventral surface of male head with transverse process behind mouthparts with 3 patches of dense setae medially (Fig. 69a). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68a), female elytra subquadrate. Male meso-and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad with 3 round depressions apically (Fig. 70a). Phallobase of median lobe symmetrical and rounded (Fig. 70a). Parameres symmetrical and slender, setose from midpoints to apices (Fig. 70a).</p><p>Type Locality. Mokohinou Island, New Zealand .</p><p>Distribution. Auckland, Coromandel, Northland (Fig. 71: black circles).</p><p>Habitat. Specimens of this species were collected using pitfall and yellow pan traps or by sifting leaf litter.</p><p>Comments. Specimens of S. insignis can be distinguished from those of other species by the shape of the ventral surface of the male head and the antennomeres. The type specimens of S. sobrina share these diagnostic characters, and additional specimens have been collected at or near the type locality. For these reasons, we have placed S. sobrina in synonymy with S. insignis .</p></div>	https://treatment.plazi.org/id/03D82A3EBD1211333CBEF9BEFDB9FBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBD1611333CB9FB5AFCC1F95B.text	03D82A3EBD1611333CB9FB5AFCC1F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hectorii Broun 1917	<div><p>Sagola hectorii Broun, 1917</p><p>(Figs. 68b, 69b, 70b, 71)</p><p>Sagola hectorii Broun, 1917: 378 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola distorta Broun, 1921b: 602 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b. Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3829.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Belgrove. 7.11.1914.” [white label, written]; “ Sagola . ♂ hectori” [white label, handwritten] . Holotype of Sagola distorta . NEW ZEALAND: Wairarapa: 1♂ (BMNH), glued on rectangular card, “4166. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] ; “Featherston. 6.10.1916.” [white label, written]; “ Sagola distorta . ♂ ” [white label, handwritten].</p><p>Additional Material (n = 8: 3 males; 5 females). NEW ZEALAND: Nelson: 1♂ 2♀♀, Upper Maitai, 7 IV 1963, E.S. Gourlay ; 1♂, Upper Maitai, 7 III 1949, E.S. Gourlay ; 3♀♀, Upper Maitai, 3 V 1950, E.S. Gourlay ; 1♂, Dovedale, 11 X 1963, J.I. Townsend, litter .</p><p>Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.4–2.8 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; male temporal lobe with L-shaped depression anteriorly; ventral surface of male head with transverse process behind mouthparts with depression medially and setae along edge; elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.4–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68b). Head: Male head triangular, widest across temples (Fig. 69b). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Male eye prominent, approximately one-fourth length of temple, female slightly smaller. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface (Fig. 69b). Male temporal lobe with L-shaped depression anteriorly (Fig. 69b). Ventral surface of male head with transverse process behind mouthparts with depression medially, with setae along edge (Fig. 69b). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68b). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe of genitalia broad with triangular head (Fig. 70b). Phallobase of median lobe symmetrical and rounded (Fig. 70b). Parameres symmetrical and triangular, setose from midpoints to apices (Fig. 70b).</p><p>Type Locality. Belgrove, near Nelson, New Zealand .</p><p>Distribution. Nelson (Fig. 71: triangles).</p><p>Habitat. One specimen was collected by sifting leaf litter.</p><p>Comments. Specimens of S. hectorii can be distinguished from those of other species by the shape of the ventral surface of the male head and the antennomeres. The type specimen of S. distorta shares these diagnostic characters. For these reasons, we have placed S. distorta in synonymy with S. hectorii .</p></div>	https://treatment.plazi.org/id/03D82A3EBD1611333CB9FB5AFCC1F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE911CE3D47FF31FC29FEDB.text	03D82A3EBDE911CE3D47FF31FC29FEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola bipunctata Broun 1886	<div><p>Sagola bipunctata Broun, 1886</p><p>(Figs. 68c, 69c, 70c, 70y, 71)</p><p>Sagola bipunctata Broun, 1886: 887 . Raffray 1893: 39, 1904: 498, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Sagola osculans Broun, 1886: 885 . Raffray 1893: 36, 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola frontalis Raffray, 1893: 23 . Raffray 1904: 498, 1911: 6; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola fulva Broun, 1893b: 1052 . Raffray 1904: 498, 1911: 6; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola laminata Broun, 1893b: 1421 . Raffray 1904: 498, 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Watt 1983: 44; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola immota Broun, 1893b: 1422 . Raffray 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola lawsoni Broun, 1912b: 632 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola latula Broun, 1912b: 633 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola halli Broun, 1914b: 155 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola brevisternis Broun, 1915: 284 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola confusa Broun, 1915: 286 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola laetula Broun, 1915: 291 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1580. ♀ ” [white label, handwritten]; “Whangarei” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola bipunctata ” [white label, handwritten] . Holotype of Sagola brevisternis: NEW ZEALAND: Mid Canterbury: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3699. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Pudding Hill. 4.5.1912.” [white label, handwritten]; “ Sagola - ♀. brevisternis” [white label, handwritten] . Holotype of Sagola confusa: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3701. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Hunua. Clevedon.” [white label, handwritten]; “ Sagola ♂. confusa.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola frontalis: NEW ZEALAND: 1♀ (BMNH), glued on rectangular card, “Bealey, New Zealand. Helms.” [white label, printed]; “Sharp. Coll. 1905-313.” [white label, printed]; “7.-” [white label, handwritten]; “ Sagola . frontalis ♀ ” [white label, handwritten] . Holotype of Sagola fulva: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1881” [white label, handwritten]; “Hunua. Clevedon” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola fulva .” [white label, handwritten] . Syntypes of Sagola halli: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3518. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Pudding Hill. 4.5.1912.” [white label, handwritten]; “ Sagola halli . ♂.” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “3518. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Pudding Hill. 4.5.1912.” [white label, handwritten]; “ Sagola halli . ♂.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “3518. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Pudding Hill. 4.5.1912.” [white label, handwritten]; “ Sagola halli . ♀.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “3518. ♀ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Pudding Hill. 4.5.1912.” [white label, handwritten]; “ Sagola halli . ♀.” [white label, handwritten] . Syntypes of Sagola immota: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2470. ♀ ” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola . immota” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “2470. ♀ ” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola . immota” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “2470” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♀ (BMNH), glued on rectangular card, “Immota” [white label, handwritten]; “ N. Zealand. 93–56.” [white label, printed]; “ Sagola immota Broun ” [white label, handwritten] . Holotype of Sagola laetula: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3708. ♂ ” [white label, handwritten]; “Hunua. Clevedon” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ laetula.” [white label, handwritten] . Syntypes of Sagola laminata: NEW ZEALAND: Auckland: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2469” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola . laminata” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “2469. ♂ ” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♂ (BMNH), glued on rectangular card, “2469.” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “Hunua. Maketu.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♂ (BMNH), glued on rectangular card, “ S. laminata ” [white label, handwritten]; “ N. Zealand. 93-56.” [white label, printed]; “ Sagola laminata Broun ” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “2469.” [white label, handwritten]; “Sharp. Coll. 1905-313.” [white label, printed]; “ Sagola laminata 2nd.auk. N.Z. Broun. ” [white label, handwritten] . Holotype of Sagola latula: NEW ZEALAND: Buller: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “12.” [white label, handwritten]; “Greymouth, New Zealand. Helms.” [white label, printed]; “Sharp. Coll. 1905-313.” [white label, printed]; “ Sagola ♂ latula.” [white label, handwritten]. The original label indicates this specimen is a male, but it is female . Holotype of Sagola lawsoni . NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “ Auckland, New Zealand.” [white label, printed]; “Sharp. Coll. 1905-313.” [white label, printed]; “ Sagola lawsoni auckland.” [white label, handwritten] . Holotype of Sagola osculans: NEW ZEALAND: Auckland: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “9576” [white label, handwritten]; “Woodhill” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola osculans ” [white label, handwritten] .</p><p>Additional Material (n = 170: 77 males; 93 females). NEW ZEALAND: Auckland: 15♂♂ 13♀♀, Symonds St. Cemetery, Auckland, 40m, 23 II 2010, D.S. Chandler, sift leaf litter by stream; 17♂♂ 21♀♀, Murphy’ s Bush, 39m, 5km ne Manukau, 19 III 2010 , D.S. Chandler &amp; K.P. Puliafico, sift litter in dry ravine ; 2♂♂, Symonds St. Cemetery, Auckland, 40m, 2–6 IV 2010, L. Masner, yellow pan trap; 22♂♂ 26♀♀, Lynfield, 23 XI 1974 – 13 XI 1981, G. Kuschel, litter; 3♂♂ 2♀♀, Grafton Gully, 28 II 1978 , litter, S. Peck; 1♂, Auckland City, Albert Park, tussock at night, 12 IV 2006 , S.E. Thorpe; 1♂, Lynfield, Wattle Bay, 11 II 1977 , B.A. Holloway, litter; 1♂, Grafton Gully, 5 XI 1985 , R.C. Craw, litter; 1♂ 1♀, Hunua Ranges Reg. Park, 7km w Kaiaua, Workman Tr, 116m, 27 III 2010 , D.S. Chandler, sift stream drift piles; 1♂, 20-40m, Grafton Gully, 4 III 2001 , S.E. Thorpe, litter; 1♂, Auckland City, bush under Grafton bridge, sedges 15 I 2005 , S.E. Thorpe; 3♀♀, Wattle Bay Res, Lynfield, 19 III 1994 , J. Nunn; 1♀, Grafton Gully, 31 V 1994, J. Nunn, underside log; 3♀♀, Little Barrier I, Pohutukawa Flat, foreshore, 3 X 1975 , B.M. May, litter; 2♀♀, little barrier I, upper Pohutukawa Flat, 3 X 1975 , B.M. May, litter; 1♀, Grafton Gully, 26 VI 1996, J. Nunn, underside of log; 1♀, Wattle Bay, 11 X 1981 , G. Kuschel, rotten wood and soil; 1♀, Wattle Bay, 24 I 1977 , C.F. Butcher, pit trap ; 1♀, Kepa Bush Reserve, 9 II 2010 , 20m, D.S. Chandler, sift leaf litter along stream; 1♀, MacLeans Reserve, Howick, 10 II 2010 , 17m, D.S. Chandler, sift rotten willow logs; Buller: 2♂♂ 2♀♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp; window trap ; 3♂♂ (1♂, slide-mounted), Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 645m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 705, window trap ; 1♂, Lake Rotoiti, 600m, 8 II 1978 , S.B. Peck, J. Peck, under bark frass ; 1♀, Nelson Lakes NP, Lake Rotoiti, 600m, 6 II 1978 , S. &amp; J. Peck, Nothofagus forest, streamside litter; Northland: 1♂, Hokianga Heads, 7 XII 1961 , G. Kuschel, Nothofagus litter; 1♂, Kerikeri, 20 I 1972 , G.W. Ramsay, litter; 2♂♂, Waipoua SF, 1km e Headquarters, 11–15 IV 1980 , J.C. Watt, pitfall trap ; 1♂, Above Hihi, 6 I 1969 , K. Wise, juvenile Kahikatea; 10♀♀, Mangonui, 22 XI 1994 , J. Nunn; 1♀, Radar Bush, Te Paki, 4 XII 1995, J. Nunn, litter; 1♀, Waipoua Stm, 100m, 19 III 1978, S.B. Peck, litter; 1♀, Waitohue rd, route to Takahue, 8 XII 2008, R. Leschen, T. Buckley, D. Seldon, RL1387, dead wood, 35°16.049, 173°17.684; Mid Canterbury: 1♂, Pudding Hill nr Methven, T. Hall, 12 VII 1912 ; 1♂, Pudding Hill nr Methven, T. Hall, 4 V 1912.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.5–2.8 mm; temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface; ventral surface of male head with setose triangular process behind mouthparts; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.5–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68c). Head: Head bluntly triangular, widest across eyes (Fig. 69c). Antennomere 1 approximately 1.5 times longer than wide, 2–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye (Fig. 70y). Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round (Fig. 70y). Eye large and prominent, approximately one-half length of temple (Fig. 69c). Temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface (Fig. 69c). Ventral surface of male head with setose triangular process behind mouthparts (Fig. 69c). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68c), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with pair of small, triangular processes at one-third length (Fig. 70c). Phallobase of median lobe symmetrical and rounded (Fig. 70c). Parameres symmetrical and slender with setae apically and 1 seta at middle (Fig. 70c).</p><p>Type Locality. Near Whangarei Harbour, Northland, New Zealand .</p><p>Distribution. Auckland, Buller, Northland, Mid Canterbury (Fig. 71: black squares).</p><p>Habitat. Specimens of this species were collected using pitfall, yellow pan, flight intercept, and window traps or by sifting leaf and wood litter.</p><p>Comments. Specimens of S. bipunctata can be distinguished from those of other species by the shape of the ventral surface of the male head and the antennomeres. The type specimens of S. brevisternis, S. confusa, S. frontalis, S. fulva, S. halli, S. immota, S. laetula, S. laminata, S. latula, S. lawsoni, and S. osculans share these diagnostic characters, and additional specimens have been collected at or near the type locality. For these reasons, we have placed S. brevisternis, S. confusa, S. frontalis, S. fulva, S. halli, S. immota, S. laetula, S. laminata, S. latula, S. lawsoni, and S. osculans in synonymy with S. bipunctata .</p></div>	https://treatment.plazi.org/id/03D82A3EBDE911CE3D47FF31FC29FEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDEB11CF3F5FFE82FE7DFD01.text	03D82A3EBDEB11CF3F5FFE82FE7DFD01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola angulifera Broun 1911	<div><p>Sagola angulifera Broun, 1911</p><p>(Figs. 68d, 69d, 70d, 71)</p><p>Sagola angulifera Broun, 1911: 491 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. Holotype. NEW ZEALAND: Taupo: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3363.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Waimarino. Jany 1909.” [white label, handwritten]; “ Sagola ♂ angulifer” [white label, handwritten].</p><p>Additional Material (n = 19: 10 males; 9 females). NEW ZEALAND: Wellington: 2♂♂ 4♀♀, Nikau Res, Paraparaumu, 11 VI 1988, J. Nunn ; 4♂♂ 2♀♀, Nikau Res, Paraparaumu, 2 VII 1988, J. Nunn ; 2♂♂, Nikau Res, Paraparaumu, 23 VII 1988, J. Nunn ; 1♂, Nikau Res, Paraparaumu, 1 V 1988, J. Nunn, leaf litter ; 1♀, Nikau Res, Paraparaumu, 8 V 1988, J. Nunn ; 2♀♀, Nikau Res, Paraparaumu, 25 IV 1988, J. Nunn ; Taranaki: 1♂, Potaema Walk, Mt. Egmont, 9 XII 1995, J. Nunn, forest litter .</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.3–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts with three patches of long and dense setae medially; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.3–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68d). Head: Male head triangular, widest across temples (Fig. 69d). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface (Fig. 69d). Ventral surface of male head with transverse process behind mouthparts with 3 patches of long, dense setae medially (Fig. 69d). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68d), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with pair of triangular processes apically (Fig. 70d). Apical lobe with semicircular depression apically (Fig. 70d). Phallobase of median lobe symmetrical and rounded. Parameres symmetrical and slender, setose from midpoints to apices (Fig. 70d).</p><p>Type locality. Waimarino, Taupo, New Zealand .</p><p>Distribution. Taranaki, Taupo, Wellington (Fig. 71: stars).</p><p>Habitat. Specimens of this species were collected by sifting forest litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDEB11CF3F5FFE82FE7DFD01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDEA11C83CBDFD38FDBAF99D.text	03D82A3EBDEA11C83CBDFD38FDBAF99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola eminens Broun Desig. Park and Carlton 2013	<div><p>Sagola eminens Broun, 1895</p><p>(Figs. 68e, 69e, 70e, 70z, 71)</p><p>Sagola eminens Broun, 1895: 75 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Lectotype. NEW ZEALAND: Waikato: 1♂ (BMNH), glued on rectangular card, “2724. ♂.” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “ Tarukenga. ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ LECTOTYPE Sagola eminens Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotypes (2 females). NEW ZEALAND: Bay of Plenty: 2♀♀ (BMNH), glued on rectangular card, “2724. ♀.” [white label, handwritten]; “ ♂ ” [white label, handwritten]; “ Tarukenga. ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola eminens ” [white label, handwritten]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that three specimens were examined (Broun, 1895: 75). This designation will fix the identity of the species and facilitate its recognition by future workers.</p><p>Additional Material (n = 76: 39 males; 37 females). NEW ZEALAND: Bay of Plenty: 1♂ 5♀♀, Motuotau I, off Mt Maunganui, 10 XI 1972, L. R. Moran, litter ; 2♂♂, Waiaroho, 26 IV 1993, G. Hall, litter ; 1♂, Waiaroho, 10 III 1993, J.S. Dugdale, litter ; 2♂♂, Te Koau, 243m, 24 XI 1992 – 31 I 1993, R. M. Emberson, pitfall trap, mixed broadleaf-podocarp forest ; 2♂♂, Hicks Bay Bush, Loop tr, 6 XII, Nothofagus forest ; 1♂, Mamaku / Kaimai Ra, Otanewainuku, 640m, 21 IX 1981, B.M. May, litter; 1♀, Te Koau, 140m, 14 III 1993, R. F. Gilbert, beneath bark of Puriri tree; 1♀, 8km from Opatiki, 20 VIII 1976, A. R. Ferguson, litter ; 1♀, Rereauira Swamp, 9 III–26 IV 1993, J.S. Dugdale, pit tap ; 1♀, Waiaroho, 26–29 IV 1993, G. Hall, pan traps edge of forest nr stream ; 1♂, Mayor I, 14 XI 1955, J.C. Watt; Gisborne: 4♂♂ 2♀♀, Urewa NP, Waikaremoana rd, s end Maranunui Ridge, 720m, 38″44.404′S 177″05.806′E, 22 XI–23 XII 2005, mixed broadleaf (incl. Nothofagus fusca)-podocarp, FMHD#2005- 028, FIT, M. Thayer, A. Solodovnikov, ANMT site 1149 ; 1♂ 1♀, Urewa NP, Ngamoko tr, pit trap fish and banana, 22 III 2000, C. Carlton, A. Weir #076 ; 3♂♂ 1♀, Te Koau, track to Hovells Watching dog, 160m, 31 I–4 II 1993, J.W. Early, Puriri /nikau forest, yellow pan trap ; 1♂, Lake Waikaremoana, 17 I 1972, G.W. Ramsay, litter ; 1♂, Te Urewera NP, Ngomoko tr, 11 V 2001, R. Leschen, litter ; 1♀, Tapuaeroa V, Waiorongomai, 12 I 1994, J.S. Dugdale, Agaric on tawa ; 1♀, Waimata V, Kaharoa Stn, 22 XI 1993 – 10 I 1994, G. Hall, pit traps ; 2♀♀, Urewera NP, Lake Waikareiti tr, Nothofagus litter, 19 III 2000, C. Carlton, A. Weir, #059 ; 1♀, Urewera NP, Black Beech tr. at Lake Waikaremoana, rotten wood, 17 III 2000, C. Carlton, A. Weir, #048 ; 2♀♀, Urewera NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=176.16667&amp;materialsCitation.latitude=-38.783333" title="Search Plazi for locations around (long 176.16667/lat -38.783333)">Ngamoko</a> tr, 38°47S 176°10E, slime flux, Nothofagus, 18 III 2000, C. Carlton, A. Weir #057 ; 1♂ (slide mounted), Urewera NP, Waikaremoana rd., s end Matanunui Ridge, 720m, 38″44.404′S 177″05.806′E, 22 XII 2005, mixed broadleaf (incl. Nothofagus fusca)- podocarp, FMNH#2005-118, berl., leaf &amp; log litter, M. Thayer, A. Solodovnikov, ANMT site 1149; Hawkes Bay : 2♂♂, Balls Clearing, 13 III 1980, C.F. Butcher, litter; Taranaki : 5♂♂ 7♀♀, Mt. Egmont, 914m, 23 I 1972, G.W. Ramsay, litter ; 1♂, Mt. Egmont, 946n, Dawson Falls, 14 I 1962, G. Kuschel, litter ; 2♂♂, Dawson Falls, 914m, 23 I 1972, G.W. Ramsay, litter ; 1♀, Mt. Messenger, 182m, 23 I 1972, G.W. Ramsay, litter ; 2♂♂ 1♀, Mt. Egmont NP, Stratford Mtn House, 846m, 23 V 1986, C.L. Lyal, litter; Taupo: 2♂♂ 5♀♀, Minginui SF, 28 VII 1977, J.S. Dugdale, litter ; 1♂, Whirinaki Forest Park, Arohaki Lagoon tr, 500m, 38″40.82′S 176″40.032′E, 21 XI 2005, mixed broadleaf forest incl. Belischmidia tawa, FMHD#2005-024, litter, A. Newton, M. Thayer, ANMT site 1147 ; 1♀, Lake Rotopounamu, sw Turangi, 650m, 4–9 IV 1980, mixed broadleafpodocarp forest, A. Newton, M. Thayer, pitfall tap ; 1♀, Scenic Res Turangi, s side of Pihanga, 13 I 1972, litter, G.W. Ramsay; Wellington : 1♂, Lake Wairarapa, 1 IX 1965, J.I. Townsend, moss ; 1♂, Johnson Hill, Karori, 28 VII 1996, J. Nunn; Waikato : 1♂ 1♀, Maungatautari, Mainland Reserve, 15 XI 2007, S.A. Forgie, litter ; 1♂, Waikato, 20 VII 1965, M. Luxton, Leptospermum scaparium litter.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–2.9 mm; temporal lobe of male head distinctly projecting, exceeding eye with concave ventral surface; ventral surface of male head with transverse process behind mouthparts with three patches of dense setae medially; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.6–2.9 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68e). Head: Male head triangular, widest across temples (Fig. 68z). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching end of eye (Fig. 68z). Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round (Fig. 68z). Eye prominent, approximately one-third length of temple (Fig. 68z). Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface (Fig. 69e). Ventral surface of male head with transverse process behind mouthparts with 3 patches of dense setae medially (Fig. 69e). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68e), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with pair of triangular processes apically (Fig. 70e). Apical lobe with semicircular depression apically (Fig. 70e). Phallobase of median lobe symmetrical and rounded. Parameres symmetrical and slender, setose from midpoints to apices (Fig. 70e).</p><p>Type Locality. Tarukenga, near Rotorua, Bay of Plenty, New Zealand .</p><p>Distribution. Bay of Plenty, Gisborne, Hawkes Bay, Taranaki, Taupo, Wellington, Waikato (Fig. 71: white circles).</p><p>Habitat. Specimens of this species were collected using pitfall, yellow pan, flight intercept, and window traps or by sifting leaf and wood litter in broadleaf, podocarp, or Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBDEA11C83CBDFD38FDBAF99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDED11CA3D4DF9C1FDADFE96.text	03D82A3EBDED11CA3D4DF9C1FDADFE96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola robustula Broun Desig. Park and Carlton 2013	<div><p>Sagola robustula Broun, 1917</p><p>(Figs. 68f, 69f, 70f, 71)</p><p>Sagola robustula Broun, 1917: 377 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Sagola cordiceps Broun, 1921a: 493 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola aemula Broun, 1921a: 496 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola bipuncticeps Broun, 1921a: 499 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola affinis Broun, 1921a: 500 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola remixta Broun, 1921a: 502 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Lectotype of Sagola robustula: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3828. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] “Hollyford. 19.2.1914.” [white label, handwritten]; “ Sagola ♂ robustula.” [white label, handwritten]; “ LECTOTYPE Sagola robustula Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotype (1 male). NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “3828. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] “Hollyford. 19.2.1914.” [white label, handwritten]; “ Sagola ♂ robustula.” [white label, handwritten] . Syntypes of Sagola aemula . NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “4006. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mistake. 9.10.1913.” [white label, handwritten]; “ Sagola aemula . ♂.” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “4006. ♂ ” [white label, handwritten]; “Mistake. 9.10.1913.” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola aemula . ♂.” [white label, handwritten]. 2♀♀ (BMNH), glued on rectangular card, “4006. ♀ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mistake. 9.10.1913.” [white label, handwritten]; “ Sagola aemula . ♀.” [white label, handwritten] . Holotype of Sagola affinis . NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “4012.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Algidus. 25.9.1913.” [white label, handwritten]; “ Sagola affinis .♂.” [white label, handwritten]. The original description indicates that the specimen was collected on 25 October 1913, but we found a single specimen collected on 25 September 1913. One of them may be misspelled . Holotype of Sagola bipuncticeps . NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “4011. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Algidus. 25.9.1913.” [white label, handwritten]; “ Sagola ♂ bipuncticeps.” [white label, handwritten]. The original description indicates that the specimen collected in 25 th October 1913, but I found a single specimen collected in 25 th September 1913. One of them may be misspelled . Holotype of Sagola cordiceps . NEW ZEALAND: Otago Lakes: 1♀ (BMNH), glued on rectangular card, “4002” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Alfred. 9.2.1914.” [white label, handwritten]; “ Sagola cordiceps ” [white label, handwritten] . Holotype of Sagola remixta . NEW ZEALAND: Nelson: 1♂ (BMNH), glued on rectangular card, “4016. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Mt. Owen. 27.12.1914.” [white label, handwritten]; “ Sagola ♂ remixta.” [white label, handwritten]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined (Broun, 1917: 377). This designation will fix the identity of the species and facilitate its recognition by future workers .</p><p>Additional Material (n = 32: 18 males; 14 females). NEW ZEALAND: Fiordland: 2♂♂ 1♀, Fiordland NP, Milford Sound rd, Smithy Creek Campground area, 400m, 44″57.065′S 168″01.155′E, 9 XII 2005, Nothofagus fusca &amp; N. menziesii open forest, FMHD #2005-089, litter, M. Thayer, A. Newton, ANMT site 1170; 1♂, Fiordland NP, Martins Bay Hut, 5 II 1980, R. Scott; 1♂, Thicket Burn, Lake Hauroko, 3 I 1996, J. Nunn; 1♀, Lake Hauroko, Southland, 2 II 1966, J.I. Townsend, mixed moss; 4♂♂, Lake Hauroko, 13 IV 1952, R. Hornabrook; Hawkes Bay: 1♂ 1♀, Waipatiki Res, 23 XII 1983, J.C. Watt, sifted woodmould; Mid Canterbury: 1♂ 1♀, Mt. Algidus nr Methven, 3 XII 1913, T. Hall; Nelson: 1♂ 1♀, Mouth of Karamea River, 22 VI 1967, A.K. Walker, litter; 1♂, Puponga Farm P, 7–10 II 1981, J.W. Early, Malaise trap; 1♂, Karamea, 30 I 1996, J. Nunn; 2♀♀, Oparara River Mouth, 1 I 2011, J. Nunn, under drift wood after flood; 3♀♀, Lake Rotoiti, 27 VII 1965, A.K. Walker, moss; Otago Lakes: 1♂, Makarora, 23–25 I 1978, 500m, S. &amp; J. Peck, litter with carrion; 2♀♀, Makarora, 21 I 1978, G. Kuschel, litter; Taranaki: 1♂, Hawera, High St Swinburns, 27 XII 1976, H.P. McColl, litter; Westland: 2♂♂, Aspiring NP, Douglas Creek at Hwy 6, 65m, 11–18 I 1985, Nothofagus menziesii -hdwd-podocarp forest, A. Newton, M. Thayer 738; 2♀♀, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720; 1♂, Lake Mahinapua, 6 I 1951, R. Jacob, leaf mould.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.7–3.0 mm; temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface; ventral surface of male head with transverse setose process behind mouthparts; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.7–3.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68f). Head: Bluntly triangular, widest across eyes (Fig. 69f). Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head weakly projecting, as long as eye, with flat ventral surface (Fig. 69f). Ventral surface of male head with transverse setose process behind mouthparts (Fig. 69f). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68f), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with pair of triangular processes apically (Fig. 70f). Apical lobe subquadrate. Phallobase of median lobe symmetrical and rounded (Fig. 70f). Parameres symmetrical and slender, sparsely setose from midpoints to apices (Fig. 70f).</p><p>Type Locality. Routeburn, north of Lake Wakatipu, Otago Lakes, New Zealand .</p><p>Distribution. Fiordland, Hawkes Bay, Mid Canterbury, Nelson, Otago Lakes, Taranaki, Westland (Fig. 71: white squares).</p><p>Habitat. Specimens of this species were collected using pitfall, yellow pan, flight intercept, and window traps or by sifting leaf and wood litter in broadleaf, podocarp, or Nothofagus forests.</p><p>Comments. Specimens of S. robustula can be distinguished from those of other species by the shape of the ventral surface of the male head and antennomeres. The type specimens of S. aemula, S. affinis, S. bipuncticeps, S. cordicep, and S. remixta share these diagnostic characters, and additional specimens have been collected at or near the type locality. For these reasons, we have placed S. aemula, S. affinis, S. bipuncticeps, S. cordiceps, and S. remixta in synonymy with S. robustula .</p></div>	https://treatment.plazi.org/id/03D82A3EBDED11CA3D4DF9C1FDADFE96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDEF11CA3D48FECFFC74FCCF.text	03D82A3EBDEF11CA3D48FECFFC74FCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola pertinax Broun 1893	<div><p>Sagola pertinax Broun, 1893</p><p>(Figs. 68g, 69g, 70g, 72)</p><p>Sagola pertinax Broun, 1893a: 176 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Type Material. Lectotype. NEW ZEALAND: Waikato: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “2720. ♂.” [white label, handwritten]; “ Waikato ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola pertinax ” [white label, handwritten]; “ LECTOTYPE Sagola pertinax Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotypes (n = 2: 1 male; 1 female). NEW ZEALAND: Waikato: 1♂ (BMNH), glued on rectangular card, “2720. ♂ ” [white label, handwritten]; “ Waikato ” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola pertinax ” [white label, handwritten] . 1♀ (BMNH), glued on rectangular card, “2720.♀” [white label, handwritten]; “head. elytra” [white label, handwritten]; “Ohaupo. Waikato ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that three specimens were examined (Broun, 1893a: 176). This designation will fix the identity of the species and facilitate its recognition by future workers .</p><p>Additional Material (n = 3 males). NEW ZEALAND: Auckland: 1♂, Hunua, Clevedon, T. Broun collection ; 1♂, Rangitoto I, 8 XII 1951, moss ; 1♂, Eaves Bush, Orewa, 15 II 1945, A.E. Brookes collection .</p><p>Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.2–2.5 mm; temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface; ventral surface of male head with triangular setose process behind mouthparts; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.2–2.5 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68g). Head: Bluntly triangular, widest across eyes (Fig. 69g). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface (Fig. 69g). Ventral surface of male head with triangular setose process behind mouthparts (Fig. 69g). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68g), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, in female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with narrower rectangular apical lobe (Fig. 70g). Phallobase of median lobe symmetrical and rounded (Fig. 70g). Parameres symmetrical and slender, with sparse setae from apex to middle (Fig. 70g).</p><p>Type Locality. Ohaupo, near Mr. Kusab’ s sawmill, Waikato, New Zealand .</p><p>Distribution. Auckland, Waikato (Fig. 72: black circles).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBDEF11CA3D48FECFFC74FCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDEE11CB3CB5FF31FBD5FE36.text	03D82A3EBDEE11CB3CB5FF31FBD5FE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola monticola Broun 1912	<div><p>Sagola monticola Broun, 1912</p><p>(Figs. 68h, 69h, 70h, 72)</p><p>Sagola monticola Broun, 1912a: 402 . Hudson 1923: 365, 1934: 184; Raffray 1924: 233; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Type Material. O n e o f s y n t y p e s. N E W ZEALAND: Taupo: 1♂ (BMNH), glued on rectangular card, “ Type ” [red label, printed]; “3195. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Ngauruhoe. march. 1910.” [white label, handwritten]; “ Sagola ♂. monticola.” [white label, handwritten] .</p><p>Additional Material (n = 3: 2 males; 1 female). NEW ZEALAND: Bay of Plenty: 1♂, Rotorua, Arikikapakapa Reserve, 25 VII 1951, moss ; Taupo: 1♂ 1♀, Desert Rd, Mangatawai, 13 I 1972, G.W. Ramsay, litter 72/1 .</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; temporal lobe of male head projecting forward posterio-laterally with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68h). Head: Male head triangular, widest across temples (Fig. 69h). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-half length of temple. Temporal lobe of male head projecting forward posterior-laterally with weakly concave ventral surface (Fig. 69h). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69h). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68h), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, in female as long as wide. Abdomen: Male tergite IV with pair of small transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia widest at midpoint and narrower apically (Fig. 70h). Phallobase of median lobe symmetrical and rounded (Fig. 70h). Parameres symmetrical and slender, with sparse setae from apex to midpoint (Fig. 70h).</p><p>Type Locality. Mount Ngauruhoe, Taupo, New Zealand .</p><p>Distribution. Bay of Plenty, Taupo (Fig. 72: triangles).</p><p>Habitat. Specimens of this species were collected by sifting moss or leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDEE11CB3CB5FF31FBD5FE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDEE11C43F51FE28FE79FBDE.text	03D82A3EBDEE11C43F51FE28FE79FBDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola laticeps Broun 1911	<div><p>Sagola laticeps Broun, 1911</p><p>(Figs. 68i, 69i, 70i, 72)</p><p>Sagola laticeps Broun, 1911: 490 . Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Type Material. Holotype. NEW ZEALAND: Buller: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3362. ♂.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Greymouth.-Lewis-” [white label, handwritten]; “ Sagola ♂ laticeps.” [white label, handwritten]. The original label indicates the specimen is male, but it is female.</p><p>Additional Material (n = 29: 14 males; 15 females). NEW ZEALAND: Buller: 9♂♂ 5♀♀, Lewis Pass NR, 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, litter; 1♂ 1♀, Greymouth, Grandjeans tk, 10 XI 2005, R. Leschen, S. Nomura ; 1♂, 4km n of Rapahoe, 13 IX 1981, R. M. Emberson, coastal scrub and flax ; 1♀, Barrytown Croesus tk, 10 XI 2005, R. Leschen, S. Nomura, rotten log ; Westland: 1♂ 1♀, Waiata Haast, 16 III 1968, R. A. Cumber, leaf litter ; 2♀♀, Otira, Barretts Creek, 9 II 1982, C.F. Butcher, litter ; 1♂ 2♀♀, Kelly Creek, Otira, 22 III 2009, J. Nunn, moss litter ; 1♂, Otira R, 6.8km ne Otira, 280m, 18–21 III 1980, Nothofagus fusca - podocarp, A. Newton, M. Thayer, litter ; 1♀, Doughboy Creek, 6km sw Mahitahi, 5 II 1984, J. C. Watt, wood mould ; 1♀, Lake Mahinapua, 20m, 23 X 1978, J.W. Early, broadleaf-punga litter ; 1♀, Arthur Pass NO, Otira, 580m, 1 III 1987, R. M. Emberson, rata-kamahi litter .</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.8–3.2 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.8–3.2 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68i). Head: Male head triangular (Fig. 69i), widest across temples. Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with flat ventral surface (Fig. 69i). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69i). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68i), female elytra subquadrate. Male meso-and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with pair of triangular processes apically (Fig. 70i). Phallobase of median lobe symmetrical and rounded (Fig. 70i). Parameres symmetrical and slender, with setae at apex and midpoint (Fig. 70i).</p><p>Type Locality. Greymouth, Buller, New Zealand .</p><p>Distribution. Buller, Westland (Fig. 72: black squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf, wood, or moss litter in broadleaf or Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBDEE11C43F51FE28FE79FBDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE111C53D4EFB87FDD0FB95.text	03D82A3EBDE111C53D4EFB87FDD0FB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola castanea Broun 1886	<div><p>Sagola castanea Broun, 1886</p><p>(Figs. 68j, 69j, 70j, 72)</p><p>Sagola castanea Broun, 1886: 884 . Raffray 1893: 20, 1904: 497, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.</p><p>Type Material. Holotype. NEW ZEALAND: Dunedin: 1♂ (BMNH), glued on rectangular card, “Type” [red label, printed]; “1573. ♂.” [white label, handwritten]; “Taieri” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ castanea.” [white label, handwritten].</p><p>Additional Material (n = 39: 21 males; 18 females). NEW ZEALAND: Central Otago: 1♂, Omarama, Ahuriri Valley, 762m, 23 I 1966, J.I. Townsend, moss; Dunedin : 1♂, Allison Res, Akatore, 13 VIII 1997, J. Nunn, forest litter; Fiordland : 1♂ 2♀♀, Lake Hauroko, 2 XI 1966, J.I. Townsend, litter ; 1♂ 2♀♀, Hollyford Camp, 10 XII 1966, A.K. Walker, litter ; 2♂♂, Cameron Mts, Lake Mike, 488m, 13 II 1963, J.I. Townsend, litter; 1♂ 1♀, Borland Lodge, 26 IX 2009, J. Nunn, moss; 2♂♂, Lake Hauroko rd, 5km e of lake, 26 IX 2009, J. Nunn, moss on trunk of fallen beech tree; 1♂, Keplar Tck start, Te Anau, 21 II 2003, J. Nunn, moss; 1♂, Lake Hauroko rd end, 19 II–22 III 2003, J. Nunn, R. Leschen, FIT; 1♂, First Bay, Lake Hauroko, 22 III–19 IV 2003, R. Leschen, J. Nunn, FIT; 1♂, Fiordland NP, Monowai Lake, 4km sw Monowai, 11 III 2010, L. Masner, yellow pan traps, Nothofagus solandri forest; 1♂, Fiordland NP, Milford Sound rd, Smithy Creek Campground area, 400m, 44″57.065′S 168″01.155′E, 9 XII 2005, Nothofagus fusca &amp; N. menziesii open forest, FMHD #2005-089, litter, M. Thayer, A. Newton, ANMT site 1170; 1♀, Fiordland NP, Kepler tr, above Brod Bay, 680m, 45″24.37′S, 167″39.105′E, 11 XII 2005, Nothofagus forest, FMHD #2005-102, litter, A. Solodovnikov, D. Clarke, ANMT site 1177; 2♀♀, Thicket Burn, Lake Hauroko, 4 I 1996, J. Nunn, litter; 1♀, Gertrude Vly, Milford rd, 22 I 2008, J. Nunn, moss on beech tree; 1♀, Lake Te Anau, control gates, 22 I 2011, J. Nunn, red beech litter; Otago Lakes: 1♂ 1♀, Paradise, 2 II 1984, J.C. Watt, wood mould; Stewart Island: 1♂ 1♀, Stewart Island, 14 II 1953, R. Hornabrook; 1♀, Stewart Island, 12 IV 1953, R. Hornabrook; Southland: 1♂ 2♀♀, Otatara, Otatara South Scen Res, 5m, 46″27.475′S 168″16.858′E, 12 XII 2005, coastal totara ( Podocapus totara) forest, FMHD #2005-103, litter, A. Newton, D. Clarke, A. Solodovnikov, ANMT site 1178; 1♀, Otatara, Otatara South Scen Res, 5m, 46″2 7.4 7 5′ S 168″16.858′E, 12 XII 2005, coastal totara ( Podocapus totara) forest, FMHD #2005-104, totara logs litter, M. Thayer, ANMT site 1178; 2♂♂ 1♀, Papatowai Picnic Point tr, 20m, 46″33.915′S 169″28.446′E, 6 XII 2005, mixed broadleafpodocarp forest, FMHD #2005-077, litter, A. Newton, M. Thayer, et. al, ANMT site 1166; 1♂, Catlins, Papatown SR, 15 II–24 III 2003, R. Leschen, FIT, 46°34′S, 169°29′E; 1♂ 1♀, Catlins, Tahakopa Scenic Reserve, 30 I 1983, C. F. Butcher, moss and rotten wood litter.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.8–3.1 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse setose process behind mouthparts; male elytra rectangular, fe,ale elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.8–3.1 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68j). Head: Male head triangular, widest across temples (Fig. 69j). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head distinctly projecting, exceeding eye with concave ventral surface (Fig. 69j). Ventral surface of male head with transverse setose process behind mouthparts (Fig. 69j). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68j), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia rectangular and broad (Fig. 70j). Phallobase of median lobe symmetrical and rounded (Fig. 70j). Parameres symmetrical and longer than median lobe, with setae at apex and midpoint (Fig. 70j).</p><p>Type Locality. West Taieri bush, Dunedin, New Zealand .</p><p>Distribution. Central Otago, Dunedin, Fiordland, Otago Lakes, Stewart Island, Southland (Fig. 72: stars).</p><p>Habitat. Specimens of this species were collected by sifting leaf, wood, or moss litter in broadleaf, podocarp, or Nothofagus forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE111C53D4EFB87FDD0FB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE011C53CDCFBCAFCB4FBE3.text	03D82A3EBDE011C53CDCFBCAFCB4FBE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola sheldoni Park and Carlton	<div><p>Sagola sheldoni Park and Carlton, new species</p><p>(Figs. 68k, 69k, 70k, 72)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card “ NEW ZEALAND ND Ruakaka 22 Jan 1977 C.F. Butcher ”, “ Pit trap in pasture”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola sheldoni Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Northland: 1♀ (NZAC), Waipoua SF, 24 XI 1980, G. Kuschel, mats on roadside .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Frederick H. Sheldon. a member of the first author’ s PhD advisory committee who provided valuable advice during this study, a major component of his dissertation.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.6 mm; temporal lobe of male head projecting and exceeding eye, with flattened ventral surface; ventral surface of male head with triangular setose process behind mouthparts; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Northland.</p><p>Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68k). Head: Male head triangular, widest across temples (Fig. 69k). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head projecting and exceeding eye, with flattened ventral surface (Fig. 69k). Ventral surface of male head with triangular setose process behind mouthparts (Fig. 69k). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68k). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia widest at one-third length, with long rectangular apical lobe (Fig. 70k). Phallobase of median lobe symmetrical and rounded (Fig. 70k). Parameres symmetrical and slender, with sparse setae from apex to midpoint (Fig. 70k).</p><p>Distribution. Northland (Fig. 72: white circles).</p><p>Habitat. Specimens of this species were collected using pitfall traps in a pasture or by sifting mats on roadsides.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE011C53CDCFBCAFCB4FBE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE011C63E93FB57FCEAFD8C.text	03D82A3EBDE011C63E93FB57FCEAFD8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola mercuryensis Park and Carlton	<div><p>Sagola mercuryensis Park and Carlton, new species</p><p>(Figs. 68l, 69l, 70l, 72)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND CL Mercury Is Green I 11– 23 Dec 1987 G. Hall ”, “ Pit traps under Coprosma ”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola mercuryensis Park and Carlton 2013 ” . Paratypes (n = 151: 70 males; 81 females). NEW ZEALAND: Coromandel: 12♂♂ (NZAC), same data as holotype (NZAC); 2♂♂ 3♀♀ (NZAC), Ohena I, 25 XI 1972, G.W. Ramsay, litter 72/236; 2♂♂ 1♀ (NZAC), Ohena I, Koruenga I, 27 XI 1972, G.W. Ramsay, litter 72/247; 3♂♂ 1♀ (NZAC), Ohena I, G.W. Ramsay, 25 XI 1972, litter 72/233; 13♂♂ 20♀♀ (NZAC), Ohena I, 25 XI 1972, S.W. Ramsay, litter 72/235; 3♂♂ 1♀ (NZAC), Mercury I, Korapuke I, 28 XI 1972, G.W. Ramsay, litter 72/249; 4♂♂ 3♀♀ (NZAC), Mercury I, Stanley I, 24 XI 1972, G. Ramsay, litter 72/225; 4♂♂ 4♀♀ (NZAC), Mercury I, Middle I, 21 XI 1972, G.W. Ramsay, litter 72/214; 3♂♂ 14♀♀ (NZAC), Mercury I, Middle I, 21 XI 1972, G.W. Ramsay, litter 72/212; 1♂ 3♀♀ (NZAC), Mercury I, Stanley I, 23 XI 1972, G.W. Ramsay, 72/220; 2♂♂ 1♀ (NZAC), Mercury I, Korapuke I, 28 XI 1972, G.W. Ramsay, litter 72/254; 1♂ 1♀ (NZAC), Mercury I, Korapuka I, 28 XI 1972, G.W. Ramsay, litter 72/253; 1♂ 1♀ (NZAC), Mercury I, 24 XI 1972, G.W. Ramsay, litter 72/227; 1♂ 1♀ (NZAC), Mercury I, Korapuke I, 28 XI 1972, G.W. Ramsay, litter 72/252; 1♀ (NZAC), Mercury I, Korapuke I, 28 XI 1972, G.W. Ramsay, litter 72/249; 6♂♂ 6♀♀ (NZAC), Mercury I, Korapuki I, 16 XII 1987, G. Hall, rotten wood litter; 6♂♂ 13♀♀ (NZAC), Mercury I, Middle I, 19 II 1984, G. Hall, litter; 3♂♂ 3♀♀ (NZAC), Mercury I, Middle I, 20 X 1983, I. Southey, litter; 1♂ 1♀ (NZAC), Mercury I, Middle I, 16–19 II 1984, G. Hall, pit trap; 1♂ (NZAC), Mercury I, Middle I, 23 XII 1987, G. Hall, litter; 1♂ 3♀♀ (AMNZ), Ohinau I, 5 XII 2000, C. McGuinness, pit trap .</p><p>Etymology. This species is named after the type locality, Mercury Island, Coromandel.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.3–2.6 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Coromandel of North Island.</p><p>Description. Length 2.3–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68l). Head: Male head triangular, widest across temples (Fig. 69l). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface (Fig. 69l). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69l). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68l). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia rectangular (Fig. 70l). Phallobase of median lobe symmetrical and rounded (Fig. 70l). Parameres symmetrical and slender with setae at apex and 1seta at midpoint (Fig. 70l).</p><p>Distribution. Coromandel (Fig. 72: white squares).</p><p>Habitat. Specimens of this species were collected using pitfall traps or by sifting moss, wood, or leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE011C63E93FB57FCEAFD8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE311C73EB3FDB3FEEFFBE3.text	03D82A3EBDE311C73EB3FDB3FEEFFBE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola cuvierensis Park and Carlton	<div><p>Sagola cuvierensis Park and Carlton, new species</p><p>(Figs. 68m, 69m, 70m, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Coromandel: 1♂ (NZAC), “ NEW ZEALAND CL Cuvier I Radar Point 27 Feb 1982 G.Hall ”, “Litter 82/37”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola cuvierensis Park and Carlton 2013 ” . Paratypes (n = 41: 24 males; 17 females). NEW ZEALAND: Coromandel: 1♀ (NZAC), same data as holotype; 2♀♀ (NZAC), Cuvier I, Shooters Ridge, 2 III 1982, G. Hall, litter ; 1♂ (NZAC), Cuvier I, 25 II–2 III 1982, G. Hall, pan trap in ferns and bracken; 14♂♂ 8♀♀ (AMNZ), Cuvier I, Old Radar Station, 200m, 10–18 XI 1999, J.W. Early, S.E. Thorpe, yellow pan trap in grassy clearing : 3♂♂ 1♀ (AMNZ), Cuvier I, Landing Bay, 11–13 XI 1999, J.W. Early, S.E. Thorpe, yellow pan trap, base of Phormium tenex; 4♀♀ (AMNZ), Cuvier I, Wilson Peak, 100m, 13–18 XI 1999, J.W. Early, S.E. Thorpe, coastal Pohutukawa forest, yellow pan trap ; 3♂♂ (AMNZ), Cuvier I, Old Radar Station, 180m, 29 III–5 IV 2000, J.W. Early, R. F. Gilbert, forest edge, malaise trap ; 2♂♂ (AMNZ), Cuvier I, Ridge Track, 120m, 29 III–6 IV 2000, J.W. Early, R. F. Gilbert, forest edge, malaise trap ; 1♂ (AMNZ), Cuvier I, West Ridge, 160m, 29 III–6 IV 2000, J.W. Early, R. F. Gilbert, manuka scrub, yellow pan trap ; 1♀ (AMNZ), Cuvier I, Ridge Track, 100m, 10–18 XI 1999, J.W. Early, S.E. Thorpe, yellow pan trap .</p><p>Etymology. This species is named after the type locality, Cuvier Island, Coromandel.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.4–2.6 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with blunt process behind mouthparts and round setose depression; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species; known only from Coromandel.</p><p>Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68m). Head: Male head triangular, widest across temples (Fig. 69m). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye and with weakly concave ventral surface (Fig. 69m). Ventral surface of male head with blunt process behind mouthparts and round setose depression (Fig. 69m). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68m), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with pair of processes apically, reaching apex of apical lobe (Fig. 70m). Phallobase of median lobe symmetrical and rounded (Fig. 70m). Parameres symmetrical and slender, sparsely setose from midpoints to apices (Fig. 70m).</p><p>Distribution. Coromandel (Fig. 73: black circle).</p><p>Habitat. Specimens of this species were collected using yellow pan and Malaise traps or by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE311C73EB3FDB3FEEFFBE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE211C03C9DFB57FE78FD4A.text	03D82A3EBDE211C03C9DFB57FE78FD4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola buckleyi Park and Carlton	<div><p>Sagola buckleyi Park and Carlton, new species</p><p>(Figs. 68n, 69n, 70m, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND:ND, Poor Knights Islands Tawhiti Rahi, Main Track, XII-182009 TB270, T. R. Buckley, R. Leschen, D. Seldon sift flood debris”, “ HOLOTYPE Sagola buckleyi Park and Carlton 2013 ” . Paratypes (n = 20: 15 males; 5 females). NEW ZEALAND: Northland: 5♂♂ 4♀♀ (DSC), same as holotype; 3♂♂ (NZAC), Poor Knights I, Aorangi, ridge to Oneho Hill, 17 XI 1981, J.C. Watt, litter ; 2♂♂ (NZAC), Poor Knights I, Aorangi, Puweto Valley, 14 XI 1981, J.C. Watt, litter ; 2♂♂ (NZAC), Poor Knights I, Tawhiti Rahi, Shag Bay, 40m, 20 IX 1980, J.C. Watt, litter ; 2♂♂ (NZAC), Poor Knights I, Tawhiti Rahi, 4 XII 1980, G. Kuschel, litter ; 1♂ 1♀ (NZAC), Poor Knights I, Tawhiti Rahi, Shag Bay, 30m, 12 IX 1980, J.C. Watt, litter .</p><p>Etymology. This species is named for the collector of the holotype, Thomas R. Buckley.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round setose depression; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Poor Knights Islands of Northland.</p><p>Description. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68n). Head: Male head triangular, widest across temples (Fig. 69n). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye and with weakly concave ventral surface (Fig. 69n). Ventral surface of male head with transverse process behind mouthparts and round setose depression (Fig. 69n). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68n). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia broad with pair of triangular processes apically, apical lobe triangular (Fig. 70n). Phallobase of median lobe symmetrical and rounded (Fig. 70n). Parameres symmetrical and slender, with sparse setae at apex and midpoint (Fig. 70n).</p><p>Distribution. Only known from Poor Knights Islands, Northland (Fig. 73: triangle).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE211C03C9DFB57FE78FD4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE511C03CA6FD09FC12FC10.text	03D82A3EBDE511C03CA6FD09FC12FC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola brumfieldi Park and Carlton	<div><p>Sagola brumfieldi Park and Carlton, new species</p><p>(Figs. 68o, 69o 70o, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Taupo: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand: Tongariro N.P., Mangawhero R. Rd. near Ohakune, S. Slope of Mt. Ruapehu TO”, “Forest litter berlese at FIT 4, 10 Mar 2000 C Carlton, A Weir, #034”, “ HOLOTYPE Sagola brumfieldi Park and Carlton 2013 ” . Paratypes (5 females). NEW ZEALAND: Waikato: 2♀♀ (NZAC), Mahoenui, Gribbons Rd, 26 VI 1977, G.W. Ramsay, litter ; 1♀ (NZAC), Mahoenui, Gribbons Rd, 26 VI 1977, N.H. Mancer, litter ; 2♀♀ (NZAC), nr Waitomo, Mathers Rd, Fred Catchment, 23 V 1983, litter .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Robb T. Brumfield. He was a member of the first author’ s PhD advisory committee and provided valuable advice during this study, a major component of his dissertation.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.6 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68o). Head: Male head triangular, widest across temples (Fig. 69o). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-fourth length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface (Fig. 69o). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69o). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68o). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia widest at middle, with flat apical margin (Fig. 70o). Phallobase of median lobe symmetrical and rounded (Fig. 70o). Parameres symmetrical and slender, with sparse setae at apex and midpoint (Fig. 70o).</p><p>Distribution. Taupo, Waikato (Fig. 73: black squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE511C03CA6FD09FC12FC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE511C13E97FC47FE4FFB95.text	03D82A3EBDE511C13E97FC47FE4FFB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola otteai Park and Carlton	<div><p>Sagola otteai Park and Carlton, new species</p><p>(Figs. 68p, 60p, 70p, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND WO Mt Karioi near base 11 Oct 1981 C.F.Butcher ”, “Litter 81/96”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola otteai Park and Carlton 2013 ” . Paratypes (n = 6: 4 males; 2 females). NEW ZEALAND: Waikato: 1♂ 1♀ (NZAC), same data as holotype; 1♂ (NZAC), Mt. Karioi, 11 Oct 1981, C.F. Butcher, litter 81/95 ; 2♂♂ 1♀ (NZAC), Piripiri, Caves Reserve, 25 V 1986, C.L. Lyal, leaf litter CL480 .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, James Ottea. He was a member of the first author’ s PhD advisory committee and provided valuable advice during this study, a major component of his dissertation.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.5–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species; known from Waikato of North Island.</p><p>Description. Length 2.5–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68p). Head: Male head triangular, widest across temples (Fig. 69p). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye and with flattened ventral surface (Fig. 69p). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69p). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68p). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia widest at middle and triangular apically (Fig. 70p). Phallobase of median lobe symmetrical and rounded (Fig. 70p). Parameres symmetrical and slender, with sparse setae at apex and midpoint (Fig. 70p).</p><p>Distribution. Waikato (Fig. 73: stars).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE511C13E97FC47FE4FFB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE411C13CD2FBCAFC79FB64.text	03D82A3EBDE411C13CD2FBCAFC79FB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola boudreauxae Park and Carlton	<div><p>Sagola boudreauxae Park and Carlton, new species</p><p>(Figs. 68q, 69q, 70q, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), “ NEW ZEALAND BP Rurima Rocks 23 Nov 1983 I. McFadden Litter (3B)”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola boudreauxae Park and Carlton 2013 ” . Paratype (n = 26: 7 males; 19 females). NEW ZEALAND: Bay of Plenty: 3♂♂ 8♀♀ (NZAC), same data as holotype; 1♂ (NZAC), Rurima Rocks, 21 V 1986, I. McFadden, 86/9 ; 3♂♂ (NZAC), Rurima Rocks, 23 XI 1983, I. McFadden, litter ; 5♀♀ (NZAC), Rurima Rocks, 7 X 1983, I. McFadden, litter ; 5♀♀ (NZAC), Rurima Rocks, 9 VII 1987, I. McFadden, litter 87/16 ; 1♀ (NZAC), Rurima Rocks, 19 IV 1985, I. McFadden, litter 85/38 .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Bonnie Boudreaux. She was a member of the first author’ s PhD advisory committee and provided valuable advice during this study, a major component of his dissertation.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.3–2.6 mm; temporal lobe of male head projecting, slightly exceeding eye and with weakly concave ventral surface; ventral surface of male head with triangular setose process behind mouthparts; elytra subquadrate; shape of antennomeres and genitalia unique to species; known from Bay of Plenty.</p><p>Description. Length 2.3–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68q). Head: Male head triangular, widest across temples (Fig. 69q). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-third length of temple. Temporal lobe of male head projecting, slightly exceeding eye, with weakly concave ventral surface (Fig. 69q). Ventral surface of male head with triangular setose process behind mouthparts (Fig. 69q). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68q). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad, with pair of small triangular processes apically, apical lobe with flat margin (Fig. 70q). Phallobase of median lobe symmetrical and rounded (Fig. 70q). Parameres symmetrical and slender, sparsely setose from midpoints to apices (Fig. 70q).</p><p>Distribution. Bay of Plenty (Fig. 73: white circle).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE411C13CD2FBCAFC79FB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE411C23EB1FADBFE21FA07.text	03D82A3EBDE411C23EB1FADBFE21FA07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola taegyui Park and Carlton	<div><p>Sagola taegyui Park and Carlton, new species</p><p>(Figs. 68r, 69r, 70r, 73)</p><p>Type Material. Holotype. NEW ZEALAND: Bay of Plenty: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “NEW ZEALAND, BP Orete Forest Te Puia Hut Bush 15 Sep 1992 G. Hall &amp;”, “ R.C. Henderson Sifted litter 92/50”, “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “HOLOTYPE Sagola taegyui Park and Carlton 2013 ” . Paratypes (n = 3: 2 males; 1 female). NEW ZEALAND: Taupo: 2♂♂ 1♀ (NZAC), Ahimanawa Ra, 609m, 14 I 1972, G.W. Ramsay, litter 72/10 .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Tae-Gyu Kim. He is one of the first author’ s colleagues and assisted with molecular components of the phylogenetic analysis.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.5–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round setose depression; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.5–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68r). Head: Male head triangular, widest across temples (Fig. 69r). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface (Fig. 69r). Ventral surface of male head with transverse process behind mouthparts and round setose depression (Fig. 69r). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68r), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with semicircular depression apically (Fig. 70r). Phallobase of median lobe symmetrical and rounded (Fig. 70r). Parameres symmetrical and slender, sparsely setose from midpoints to apices (Fig. 70r).</p><p>Distribution. Bay of Plenty, Taupo (Fig. 73: white squares).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE411C23EB1FADBFE21FA07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE711C33CFCFA3BFDACFBD6.text	03D82A3EBDE711C33CFCFA3BFDACFBD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola hanae Park and Carlton	<div><p>Sagola hanae Park and Carlton, new species</p><p>(Figs. 68s, 69s, 70s, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Wairarapa: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand WA Castlehill / Haunui area. SE Puketoi Ra March 2010 ”, “ HOLOTYPE Sagola hanae Park and Carlton 2013 ” The Original label does not indicate who collected the holotype, but it was collected by J. Nunn. Paratypes (n = 6: 3 males; 3 females). NEW ZEALAND: Wairarapa: 1♂ 2♀♀ (JTN), same data as holotype; Wellington: 1♂ (FMNH), Wilton’ s Bush, 110m, 41”15.963′S 175”45.159′E, 24 XI 2005, mixed broadleafpodocarp forest, FMHD#2005-030, litter, M. Thayer, A. Newton, ANMT site 1150 ; Rangitikei: 1♂ 1♀ (NZAC), Utiku, Main Trunk Line, J. Ford, 27 VII 1917 .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Hana Kim. She is one of the first author’ s colleagues and assisted with molecular components of the phylogenetic analysis.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.2–2.4 mm; temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface; male temporal lobe with L-shaped depression anteriorly; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.2–2.4 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68s). Head: Male head triangular, widest across temples (Fig. 69s). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 subquadrate, 3–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface (Fig. 69s). Male temporal lobe with L-shaped depression anteriorly (Fig. 69s). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69s). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68s), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia widest at middle, with round apical lobe (Fig. 70s). Phallobase of median lobe symmetrical and rounded (Fig. 70s). Parameres symmetrical and triangular, setose from midpoints to apices (Fig. 70s).</p><p>Distribution. Rangitikei, Wairarapa, Wellington (Fig. 74: black circles).</p><p>Habitat. One specimen was collected by sifting forest litter from a broadleaf and podocarp forest.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE711C33CFCFA3BFDACFBD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE611C33CD2FB9DFB43FA19.text	03D82A3EBDE611C33CD2FB9DFB43FA19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola gyudongi Park and Carlton	<div><p>Sagola gyudongi Park and Carlton, new species</p><p>(Figs. 68t, 69t, 70t, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Marlborough: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: MB: Pelorus Bridge, 25 VII 1967 G. Kuschel, litter 67/220”, “ HOLOTYPE Sagola gyudongi Park and Carlton 2013 ” . Paratypes (n = 10: 8 males; 2 females). NEW ZEALAND: Buller: 2♂♂ (DSC &amp; FMNH), Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &amp; window trap ; 2♂♂ 2♀♀ (FMNH), Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr, 645m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 705, window trap; Nelson : 1♂ (FMNH), 20km ne Takaka, Tasman NP, 21 V 1982, FMHD#2005-591, mixed forest litter, S. Peck; Marlborough Sounds: 3♂♂ (LUNZ), Outer Chetwode I, ( Te Kakaho), 11–16 II 1988, M.H. Bowie, yellow pan trap in shoreline vegetation .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, Gyu-Dong Han. He is one of the first author’ s colleagues and assisted with molecular components of the phylogenetic analysis.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–3.0 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.6–3.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68t). Head: Male head triangular, widest across temples (Fig. 69t). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface (Fig. 69t). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69t). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68t). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with or without pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with triangular apical lobe (Fig. 70t). Phallobase of median lobe symmetrical and rounded (Fig. 70t). Parameres symmetrical and triangular, setose from midpoints to apices (Fig. 70t).</p><p>Distribution. Buller, Marlborough, Nelson, Marlborough Sounds (Fig. 74: triangles).</p><p>Habitat. Specimens of this species were collected using window and yellow pan traps or by sifting leaf litter in Nothofagus forests or on shorelines.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE611C33CD2FB9DFB43FA19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDE611DC3EE8FA5EFE78FA22.text	03D82A3EBDE611DC3EE8FA5EFE78FA22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola egmontensis Park and Carlton	<div><p>Sagola egmontensis Park and Carlton, new species</p><p>(Figs. 68u, 69u, 70u, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Taranaki: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand TK Patea Walk Mt Egmont 8.12.95”, “ In forest litter”, “ HOLOTYPE Sagola egmontensis Park and Carlton 2013 ”. The original label does not indicate who collected the holotype, but it was collected by J. Nunn. Paratypes (n = 4: 1 male; 3 females). NEW ZEALAND: Taranaki: 1♂ (NZAC), Mt. Egmont, 914m, 23 I 1972, G.W. Ramsay, litter 72/67 ; 3♀♀ (JTN), White Cliffs Tck, Mt. Messenger, 20 III 1994, J. Nunn .</p><p>Etymology. This species is named after the type locality, Mt. Egmont, Taranaki.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.6–3.0 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Mt. Egmont in Taranaki of North Island.</p><p>Description. Length 2.6–3.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68u). Head: Male head triangular, widest across temples (Fig. 69u). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface (Fig. 69u). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69u). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular (Fig. 68u), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia gourd-shaped (Fig. 70u). Phallobase of median lobe symmetrical and rounded (Fig. 70u). Parameres symmetrical and slender, setose from midpoints to apices (Fig. 70u).</p><p>Distribution. Taranaki (Fig. 74: black squares).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDE611DC3EE8FA5EFE78FA22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDF911DC3CF3FA11FC12F9B8.text	03D82A3EBDF911DC3CF3FA11FC12F9B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola boonei Park and Carlton	<div><p>Sagola boonei Park and Carlton, new species</p><p>(Figs. 68v, 69v, 70v, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Waikato: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND WO Mah oe nui Grib bon s R oa d 26 Jun 1977 G.W. Ramsay ”, “Litter 77/81”, “ N. Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ”, “ HOLOTYPE Sagola boonei Park and Carlton 2013 ”. Paratypes (n = 4: 2 males; 2 females). NEW ZEALAND : Taranaki: 1♂ (NZAC), Mt. Messenger, III 1933, A.E. Brookes; Taupo : 1♂ (JTN), Waituhi Saddle, 1 IX 1993, J. Nunn, leaf litter ; Waikato: 2♀♀ (NZAC), same data as holotype .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, James H. Boone. He assisted with specimen access through loans and during the first author’ s visit to the Field Museum of Natural History, Chicago, IL.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.7–3.0 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species.</p><p>Description. Length 2.7–3.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68v). Head: Male head triangular, widest across temples (Fig. 69v). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface (Fig. 69v). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69v). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68v). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad and rectangular (Fig. 70v). Phallobase of median lobe symmetrical and rounded (Fig. 70v). Parameres symmetrical and slender, sparsely setose from apical two-thirds to apex (Fig. 70v).</p><p>Distribution. Taranaki, Taupo, Waikato (Fig. 74: stars).</p><p>Habitat. Specimens of this species were collected by sifting leaf litter.</p></div>	https://treatment.plazi.org/id/03D82A3EBDF911DC3CF3FA11FC12F9B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDF911DD3E86F9BEFC53FEDB.text	03D82A3EBDF911DD3E86F9BEFC53FEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola stewartensis Park and Carlton	<div><p>Sagola stewartensis Park and Carlton, new species</p><p>(Figs. 68w, 69w, 70w, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Stewart Island: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: SI: Codfish IS nr summit, 244m, 14XII1966 J.I.Townsend, litter 66/443”, “HOLOTYPE Sagola stewartensis Park and Carlton 2013 ”. Paratypes (n = 24: 10 males; 14 females). NEW ZEALAND: Stewart Island: 1♂ 2♀♀ (NZAC), Codfish I, North Hut tr, 6 XII 1981, S.A. Holloway, litter 81/207; 2♂♂ 3♀♀ (NZAC), Pegasus Creek, 24 II 1968, G. Kuschel, litter 68/64; 5♂♂ 1♀ (NZAC), Pegasus Creek, 24 II 1968, G. Kuschel, litter 68/63; 1♂ 1♀ (NZAC), ne Big S. Cape I, 91m, 22 II 1969, J.S. Townsend, moss 69/83; 1♀ (NZAC), Codfish I, Upper Valley tr, 27 XI 1981, B.A. Holloway, litter 81/197; 1♀ (NZAC), Codfish I, Summit tk, 250m, 30 XI 1981, B.A. Holloway, litter 81/199; 1♀ (NZAC), Codfish I, Upper Miro tr, 1 XII 1981, B.A. Holloway, litter 81/200; 1♀ (NZAC), Garden Mound tr, 113m, 21 I 2007, R. Leschen, T. Buckley, K. Marske, log fogging; 1♂ 2♀♀ (NZAC), Stewart Island, 14 II 1953, R. Hornabrook.</p><p>Etymology. This species is named after the type locality, Stewart Island.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–2.9 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse, setose process behind mouthparts; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Stewart Island.</p><p>Description. Length 2.6–2.9 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68w). Head: Male head triangular, widest across temples (Fig. 69w). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface (Fig. 69w). Ventral surface of male head with transverse, setose process behind mouthparts (Fig. 69w). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68w). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia broad with transverse apical lobe (Fig. 70w). Phallobase of median lobe symmetrical and rounded (Fig. 70w). Parameres symmetrical and slender, sparsely setose from midpoints to apices (Fig. 70w).</p><p>Distribution. Stewart Island (Fig. 74: white circles).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf and moss litter and one from log fogging.</p></div>	https://treatment.plazi.org/id/03D82A3EBDF911DD3E86F9BEFC53FEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDF811DE3E9EFE9FFD58FDD1.text	03D82A3EBDF811DE3E9EFE9FFD58FDD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola jeongnamae Park and Carlton	<div><p>Sagola jeongnamae Park and Carlton, new species</p><p>(Figs. 68x, 69x, 70x, 74)</p><p>Type Material. Holotype. NEW ZEALAND: Marlborough Sounds: 1♂ (LUNZ), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND, SD Stephens Island Keepers Bush 18-20.ii.1994 J.W.M. Marris ”, “ pitfall trap in Melicytus, Myoporum, Rhopalostylis forest”, “ HOLOTYPE Sagola jeongnamae Park and Carlton 2013 ” . Paratypes (n = 10: 5 males; 5 females). NEW ZEALAND: Marlborough Sounds: 5♂♂ 4♀♀ (LUNZ), same data as holotype; 1♀ (LUNZ), Middle Trio Island, 15 II 1995, J.W.M. Marris, LUNZ 95 /6 litter ex Melicytus / broadleaf forest .</p><p>Etymology. This species is named for one of the enthusiastic supporters of this study, JeongNam Lee. She is one of the first author’ s colleagues and assisted with molecular components of the phylogenetic analysis.</p><p>Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Marlborough Sounds.</p><p>Description. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 68x). Head: Male head triangular, widest across temples (Fig. 69x). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate. 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, approximately onethird length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface (Fig. 69x). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (Fig. 69x). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate (Fig. 68x). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia broad, with bluntly triangular apical lobe (Fig. 70x). Phallobase of median lobe symmetrical and rounded (Fig. 70x). Parameres symmetrical and slender, with setae at apex and midpoint (Fig. 70x).</p><p>Distribution. Marlborough Sounds (Fig. 74: white squares).</p><p>Habitat. Specimens of this species were collected using pitfall traps in forsts of Melicytus J. R. Forst. &amp; G. Forst. (Violaceae), Myoporum Sol. ex G. Forst. (Scrophulariaceae), and Rhopalostylis Klotzsch ex Baill. (Euphorbiaceae); one was taken by sifting leaf litter in Melicytus /broadleaf forests.</p></div>	https://treatment.plazi.org/id/03D82A3EBDF811DE3E9EFE9FFD58FDD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFB11DE3D5AFD85FE5DFBDE.text	03D82A3EBDFB11DE3D5AFD85FE5DFBDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola genalis Broun 1881	<div><p>29. genalis species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the genalis species-group can be distinguished from other Sagola species by the following combination of characters: body length 2.1–2.3 mm; antennomere 1 approximately 1.5 times longer than wide; male head triangular, widest across temples (Fig. 75d) and female head bluntly triangular, widest across eyes; posterior frontal fovea large round, as large as eye (Fig. 75d); temple of male head depressed and projecting ventrally, exceeding eye (Fig. 75d); male neck ventrally with round depression with dense setae anteriorly (Fig. 75e: arrow); abdominal tergites IV–VI with discal carinae; median lobe of genitalia elongate, as wide as paramere (Fig. 75b–c).</p></div>	https://treatment.plazi.org/id/03D82A3EBDFB11DE3D5AFD85FE5DFBDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFB11D93D56FB9DFCC5FD70.text	03D82A3EBDFB11D93D56FB9DFCC5FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola genalis Broun 1881	<div><p>Sagola genalis Broun, 1881</p><p>(Figs. 75, 78)</p><p>Sagola genale Broun, 1881: 663 . Schaufuss 1888: 84; Raffray 1893: 19, 1904: 497, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.</p><p>Sagola pallidula Broun, 1912b: 626 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Sagola bilobata Broun, 1921a: 486 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.</p><p>Sagola fagicola Broun, 1921a: 494 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.</p><p>Sagola minuscula Broun, 1921a: 497 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.</p><p>Type Material. Holotype. NEW ZEALAND: Wellington: 1♂ (BMNH), glued on clear plastic card, “Type” [red label, printed]; “1157.” [green label, printed], “New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Wellington – Sandager-” [white label, handwritten]; “ Sagola genalis .” [white label, handwritten] . Syntypes of Sagola bilobata: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “3993. ♂ ” [white label, handwritten]; “Type.” [white label, handwritten]; “Mt. Dick 10.3.1914.” [white label, handwritten]; “New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ bilobata.” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “3993. ♂ ” [white label, handwritten]; “Mt. Dick. 10.3.1914.” [white label, handwritten]; “ New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♂ (BMNH), glued on rectangular card, “3993. ♂ ” [white label, handwritten]; “New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Dick. 13.3.1914.” [white label, handwritten]. 1♂ (BMNH), glued on rectangular card, “Coll. T.Hall. 10.3.1914.” [white label, handwritten]; “Mt. Dick. W. of L. Wakatipu, Southland.” [white label, handwritten]; “3993.” [white label, handwritten]; “ Sagola bilobata Broun ” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed]. 1♂ (BMNH), glued on rectangular card, “Coll. T.Hall. 10.3.1914.” [white label, handwritten]; “Mt. Dick, Wakatipu, Southland.” [white label, handwritten]; “3993.” [white label, handwritten]; “ Sagola bilobata Broun ” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]. 1♂ (BMNH), glued on rectangular card, “Mt. Dick, 10.3.1914.” [white label, handwritten]; “ Sagola ♂ bilobata” [white label, handwritten]; “3993.” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed]. 3♂♂ (BMNH), glued on rectangular card, “Mt. Dick, 13.3.1914.” [white label, handwritten]; “ Sagola ♂ bilobata” [white label, handwritten]; “3993.” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]. 1♂ (BMNH), glued on rectangular card, “Mt. Alfred, 4.2.1914.” [white label, handwritten]; “ Sagola ♂ bilobata.” [white label, handwritten]; “3993.” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed]. 1♂ (BMNH), glued on rectangular card, “Earnslaw. 5.2.1914.” [white label, handwritten]; “3993.” [white label, handwritten]; “ Sagola ♂ bilobata.” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]. Syntypes of Sagola fagicola: NEW ZEALAND: Otago Lakes: 3♀♀ (BMNH), glued on rectangular card, “4003. ♂ ” [white label, handwritten]; “Mt. Dick. 10.3.1914.” [white label, handwritten]; “New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ fagicola.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4003. ♂ ” [white label, handwritten]; “Mt. Dick. 10.3.1914.” [white label, handwritten]; “ New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 2♀♀ (BMNH), glued on rectangular card, “Coll. T.Hall 10.3.1914.” [white label, handwritten]; “Mt. Dick, W. of L. Wakatipu, Southland.” [white label, handwritten]; “4003.” [white label, handwritten]; “ Sagola fagicola Broun. ” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]. The original label indicates the types are male, but all syntypes are female. Syntypes of Sagola minuscula: NEW ZEALAND: Otago Lakes: 1♀ (BMNH), glued on rectangular card, “4008. ♂ ” [white label, handwritten]; “Heavens. 5.2.1914.” [white label, handwritten]; “New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♂ minuscula.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “4008. ♂ ” [white label, handwritten]; “ New Zealand. Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Heavens. 5.2.1914.” [white label, handwritten]. 1♀ (BMNH), glued on rectangular card, “ Coll. T.Hall 5.2.1914.” [white label, handwritten]; “Heaven’ s Gate, N. of L. Wakatipu, Southland.” [white label, handwritten]; “4008.” [white label, handwritten]; “ Sagola minusculus Broun.” [white label, handwritten]; “ T. Broun Collection” [white label, printed]; “ A.E. Brookes Collection ” [white label, printed]; “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]. The original label indicates the types are male, but all syntypes are female. Holotype of Sagola pallidula: NEW ZEALAND: Buller: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “13.” [white label, printed], “Greymouth, New Zealand. Helms.” [white label, printed]; “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola pallidula ” [white label, handwritten].</p><p>Additional Material (n = 74: 40 males; 34 females). NEW ZEALAND: Auckland: 1♂, Waitakere Range, 260m, Nohoanga Scenic Res, 8 XII 1984 – 25 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 679, FIT &amp; window trap; 1♂ , Waitakere Ra, Kauri Knoll, 22 I 1979, J.S. Dugdale, litter; 1♂ , Riverhead Forest Reserve, 25 IV 1982, P.A. Maddison, litter; 1♂ , Wattle Bay, 11 X 1981, G. Kuschel, rotten wood and soil; 1♂, Waitakere Ra, Cascade-Kauri Park, Up. Kauri tr, 170m, 23 XI–8 XII 1984, kauripodo-hdwd, A. Newton, M. Thayer 680, litter; Buller: 1♀, Greymouth, Boddytown, 8 II 1984, J.C. Watt, litter; Fiordland: 1♂ 1♀, Keplar Tck start, Te Anau, 22 I 2008, J. Nunn, moss; Mid Canterbury: 1♀, Arthurs Pass, Dobson Nature Walk, 8 II 1982, C.F. Butcher, litter; Mackenzie: 1♀ , Ohau skifield, 1475m, 10 III 2007, J. Nunn, moss and tussock near stream; North Canterbury: 2♀♀, Arthur’ s Pass, 900m, 26 I 1978, G. Kuschel, litter ; Nelson: 1♂, Kahurangi NP, Arthur Range, a bo ve Fl ora S ad dl e, 10 00m, 41″11.35 1′ S 172″44.456′E, 28 XI–19 XII 2005, Nothofagus forest, FMHD#2005-044, FIT, A. Newton, M. Thayer, ANMT site 1156; 1♂ 1♀ , Karamea River, estuary, 30 XII 2010, J. Nunn, under driftwood after flood; 1♂ , Slaters Rd, 0.7km s Whangamoa Saddle, 410m, 13 XII 1984 – 4 I 1985, Nothofagus forest, A. Newton, M. Thayer 703, FIT &amp; window trap ; Otago Lakes: 1♂ , Mt. Algidus nr Methven, T. Hall, 3 II 1914; 1♂ , Lake Wakatipu, T. Hall, III 1914; 1♂ , Staircase, T. Hall, 14 III 1914; 2♀♀ , Mt. Alfred near Paradise, 3 II 1914, T. Hall; 1♀ , Heaven’ s Gate, n of L. Wakatipu, 3 II 1914, T. Hall; 6♀♀ , Staircase, Remarkables, 15 III 1914, T. Hall; 2♀♀ , Mt. Dick, w of L. Wakatipu, 10 III 1914, T. Hall ; Marlborough Sounds: 1♂ , Queen Charlotte sd, Bay of Many Coves, 24 X–26 XII 1993, J.W.M. Marris, Malaise trap; 1♂ , Tennyson Inlet, west side Te Mako Bay, 125m, 15 XII 1984 – 5 I 1985, Nothofagus-podo-hdwd, A. Newton, M. Thayer 710, FIT &amp; window trap; 1♂ , Okiwi Bay, IX 1984, T. Jones, Malaise trap; 1♀ , 70km ne Nelson, Tennyson Inlet, 480m, 27 V 1982, FMHD#82-604, Beech forest, litter, S. Peck; Westland: 2♂♂ , 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap; 1♂ , 1.8km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, FIT &amp; window trap; 3♂♂ , Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 731, FIT &amp; window trap; 1♂ , 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwdnikau forest, A. Newton, M. Thayer 720, window trap; 2♂♂ 2♀♀ (1♂, slide-mounted), Westland NP, saddle, 10.2km ne Fox Glacier, 405m, 9–18 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 735, litter; 2♀♀, Lake Kaniere rd, 2.8km nw Lake Kaniere, 120m, 8–19 I 1985, podocarphdwd forest, A. Newton, M. Thayer 732, litter; 2♀♀ , L. Mahinapua Scen Res, 30m, 16–22 III 1980, podocarp mixed broadleaf, A. Newton, M. Thayer, litter; 1♀, Hokitika Lake Mahinapua Res, 28 I 1978, S. &amp; J. Peck, bracket fungi berl; 1♀ , Okuku Scenic Reserve, 46.7km W Otira, 75m, 42′43S 171′14E, #050, litter, 12 I 1998, C. Carlton, R. Leschen; 1♀ , 11.2km ne Franz Josef Glacier nr Lake Mapourika, 150m, Otto’ s Corner picnic area, 43′18S 170′14E, #064, coastal forest leaf litter, 15 I 1998, C. Carlton, R. Leschen; 1♀ , Gillespies Beach, 12 XI 1969, J.I. Townsend, moss ; Wellington: 5♂♂ 2♀♀, Mana Island, 4–6 II 1994, J. Nunn, litter; 4♂♂ , Karori Reservoir, 15 II 1997, beaten from Phormium colensoi, J. Nunn; 2♂♂ , Karori Reservoir, 10 IV 1997, J. Nunn, Phormium litter; 1♂ , Lookout Rock, Karori Reservoir, 29 IX 1994, J. Nunn, decayed gorse wood; 1♂ , Wainui Trig, Wainuiomata, 14 VIII 1993, J. Nunn, decayed wood; 1♂ , Nikau Res, Paraparaumu, 8 VI 1988, J. Nunn; 1♂ , Porirua SR, Elsdon, 18 IX 1993, J. Nunn, decayed wood; 1♂ , Tinakori Hill, 6 IX 1991, J. Nunn, litter; 1♀ , Days Bay, 22 VIII 1993, J. Nunn, litter under sedge; 1♀ , Mt. Holdsworth, Donelly Flat, 30 I 1985, H.P. McColl, under nr giant rata; 1♀ , Mt. Holdsworth, 10 XI 1952, R. Hornabrook .</p><p>Diagnosis. This species can be distinguished from other Sagola species by the following combination of characters: body small, length 2.1–2.3 mm; posterior frontal fovea large round, as large as eye; temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface; ventral surface of male head with triangular setose process behind mouthparts; male neck with round ventral depression anteriorly; shape of antennomeres and genitalia unique to species.</p><p>Redescription. Length 2.1–2.3 mm. Body brown, antennae, legs, maxillary palpi,and elytra paler (Fig. 75a). Head: Male head triangular, widest across temples (Fig. 75d). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye (Fig. 75d). Anterior frontal fovea oval, partially covered by frontal rostrum (Fig. 75d). Eye large and prominent, approximately one-half length of temple (Fig. 75d). Temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface (Fig. 75d). Ventral surface of male head with triangular setose process behind mouthparts (Fig. 75e: arrow). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 75a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia long and curved, C-shaped vertically, as long as paramere (Fig. 75b–c). Phallobase of median lobe symmetrical and rounded (Fig. 75b–c). Parameres symmetrical and elongate with setae apically (Fig. 75b–c).</p><p>Type Locality. Wellington, New Zealand .</p><p>Distribution. Auckland, Buller, Fiordland, Mid Canterbury, Mackenzie, Marlborough Sounds, North Canterbury, Nelson, Otago Lakes, Westland, Wellington (Fig.78: black circles).</p><p>Habitat. Specimens of this species were collected using Malaise, flight intercept, and window traps or by sifting leaf litter in podocarp, broadleaf, nikau, hardwood, or Nothofagus forests.</p><p>Comments. Specimens of S. genalis can be distinguished from those of other Sagola species by their small size, large posterior frontal fovea, the shape of the ventral surface of the male head, and the antennomeres. The type specimens of S. bilobata, S. fagicola, S. minuscula, and S. pallidula share these diagnostic characters, and additional specimens have been collected at or near the type locality. For these reasons, we have placed S. bilobata, S. fagicola, S. minuscula, and S. pallidula in synonymy with S. genalis .</p></div>	https://treatment.plazi.org/id/03D82A3EBDFB11D93D56FB9DFCC5FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFC11DA3F43FCE7FD44FA07.text	03D82A3EBDFC11DA3F43FCE7FD44FA07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola	<div><p>30. flemingensis species - group</p><p>(2 species)</p><p>Diagnosis. Members of the flemingensis species-group can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.1–2.5 mm; antennomere 1 approximately 1.5 times longer than wide; eye small, one-third to one-fourth length of temple (Fig. 76e–f); elytra as long as wide (Fig. 76a–b), hind wings reduced to small pads; abdominal tergites IV–VI with discal carinae; known from Auckland Islands and Snares Islands (Fig. 78).</p><p>KEY TO SPECIES OF THE FLEMINGENSIS SPECIES- GROUP</p><p>The key is based on male specimens because female specimens are indistinguishable based on external morphology.</p><p>1. Antennomeres 9–10 weakly transverse and clubbed (Fig. 76a); anterior frontal fovea keyhole-shaped, posterior frontal fovea absent (Fig. 76e); eye small, one-fourth length of temple (Fig. 76e); abdominal ventrite VII simple; only known from Auckland Islands (Fig. 78: triangle).............................................. .......................... S. flemingensis new species</p><p>1′. Antennomeres 9–10 subquadrate (Fig. 76b); anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round (Fig. 76f); eye small, onethird length of temple (Fig. 76f); abdominal ventrite VII with round depression; only</p><p>known from Snares Islands (Fig. 78: black square) .............. S. snaresensis new species</p></div>	https://treatment.plazi.org/id/03D82A3EBDFC11DA3F43FCE7FD44FA07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFF11DB3C9CFA3BFDBEFBC4.text	03D82A3EBDFF11DB3C9CFA3BFDBEFBC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola flemingensis Park and Carlton	<div><p>Sagola flemingensis Park and Carlton, new species</p><p>(Figs. 76a, 76c, 76e, 78)</p><p>Type Material. Holotype. NEW ZEALAND: Auckland Islands: 1♂ (NZAC), “New Zealand: AU: Auckland I., Fleming Plateau, 563m, swards 20 Feb 1973, JS Dugdale 73/58”, “ HOLOTYPE Sagola flemingensis Park and Carlton 2013 ” . Paratypes (n = 48: 29 males; 19 females).</p><p>NEW ZEALAND: Auckland Islands: 13♂♂ 11♀♀ (1♂, slide-mounted; NZAC), same data as holotype . 3♂♂ 2♀♀ (NZAC), Fleming Plateau, 426m, swards, 9 II 1973, JS Dugdale ; 2♂♂ 1♀ (1♂, slide-mounted; NZAC), Fleming Plateau, 518m, Carnley Harbor, swards, 9 II 1973, JS Dugdale ; 3♂♂ 2♀♀ (NZAC), Fleming Plateau, 487m, swards, 12 II 1973, JS Dugdale ; 4♂♂ 1♀ (NZAC), e end of Tagua Bay, Carnley Harbor, litter, 4 II 1973, JS Dugdale ; 1♂ 1♀ (NZAC), Mt. D’ Urville, 639m, moss &amp; mats, 4 I 1973, J. Rarrell ; 2♂♂ (NZAC), Fleming Plateau, 578m, Carnley Harbor, moss, 7 II 1973, JS Dugdale ; 1♂ (NZAC), Adams I, Magnetic Cove, East Ridge, moss, 22 I 1966, G. Kuschel .</p><p>Etymology. This species is named after the type locality, Fleming Plateau, Auckland Islands.</p><p>Diagnosis. This species can be distinguished from other Sagola species by the following combination of characters: body small, length 2.1–2.4 mm; antennomeres 9–11 weakly clubbed; anterior frontal fovea keyhole-shaped; eye small, one-fourth length of temple; shape of antennomeres and genitalia unique to species; only known from Auckland Islands.</p><p>Description. Length 2.1–2.4 mm. Body brown, antennae, legs, maxillary palpi,and elytra paler (Fig. 76a). Head: Male head round, widest at midpoint of head (Fig. 76e). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–8 subquadrate, 9–10 transverse, 9–11 weakly clubbed. Frontal sulcus shallow, reaching posterior margin of eye. Anterior frontal fovea keyhole-shaped, partially covered by frontal rostrum (Fig. 76e). Eye small, one-fourth length of temple (Fig. 76e). Thorax: Prosternum as long as wide, widest at midpoint. Elytra as long as wide (Fig. 76a). Meso-and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia longer than parameres (Fig. 76c). Phallobase of median lobe symmetrical and rounded (Fig. 76c). Parameres symmetrical, but right slightly longer than left and with setae apically (Fig. 76c).</p><p>Distribution. Auckland Islands (Fig. 78: triangle).</p><p>Habitat. Specimens of this species were collected by sifting litter in swards, moss, or mats.</p></div>	https://treatment.plazi.org/id/03D82A3EBDFF11DB3C9CFA3BFDBEFBC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFE11D43F49FAADFE06FC28.text	03D82A3EBDFE11D43F49FAADFE06FC28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola opercularis Broun 1915	<div><p>31. opercularis species-group</p><p>(1 species)</p><p>Diagnosis. The single member of the opercularis species-group can be distinguished from other Sagola species by the following combination of characters: body length 2.1 mm; antennomere 1 approximately 1.5 times longer than wide; anterior frontal fovea small, round, posterior frontal fovea elongate; frontal sulcus reaching to hind point of eye (Fig. 77a); eye large and prominent, approximately as long as temple (Fig. 77a–b); ventral surface of male head with W-shaped process bearing dense setae behind process (Fig. 77b: arrow); elytra longer than wide (Fig. 77a), hind wings well-developed; abdominal tergites IV–VI with discal carinae; median lobe of male genitalia curved laterally, C-shaped (Fig. 77c–d); known from South Island (Fig. 78).</p></div>	https://treatment.plazi.org/id/03D82A3EBDFE11D43F49FAADFE06FC28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDFE11DB3CD2FB7AFBF7FAA9.text	03D82A3EBDFE11DB3CD2FB7AFBF7FAA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola snaresensis Park and Carlton	<div><p>Sagola snaresensis Park and Carlton, new species</p><p>(Figs. 76b, 76d, 76f, 78)</p><p>Type Material. Holotype. NEW ZEALAND: Snares Islands: 1♂ (NZAC), “New Zealand: SN: Penguin Ck. In wet Olearia lyallii 26 Oct 1972, DS Horning”, “ HOLOTYPE Sagola snaresensis Park and Carlton 2013 ” . Paratypes (n = 22: 10 males; 12 females). NEW ZEALAND: Snares Islands: 1♀ (NZAC), same data as holotype; 2♂♂ 4♀♀ (NZAC), Penguin Ck, rotten wood of Olearia lyallii stump, 11 III 1971, D.S. Horning ; 2♂♂ 1♀ (NZAC), Signpost Hill, Stilbocarpa robusta litter, 26 VIII 1972, D.S. Horning ; 2♂♂ (NZAC), Broughton I, Stilbocarpa robusta, 4 XI 1972, D.S. Horning ; 1♂ (NZAC), sw Promontory, Olearia lyallii litter, 31 VII 1972, D.S. Horning ; 2♂♂ (NZAC), Penguin Creek, 14 XII 1974, D.S. Horning, under branches of Olearia lyallii; 2♀♀ (NZAC), nr Penguin Colony, 19 I 1972, C.J. Horning, rotten Olearia lyallii; 1♀ (NZAC), Opp Mollymauk Islet, 24 I 1967, P.M. Johns, beating Poa astonii; 1♀ (NZAC), Penguin Creek, 26 X 1972, D.S. Horning, wet Olearia lyallii litter ; 1♀ (AMNZ), nr Razorbark, 15 XII 1976, J.W. Early, under loose rock on peat ; 1♀ (NZAC), Rocky Islet, 28 II 1971, D.S. Horning, Tillaea moscha sample from rotten granite .</p><p>Etymology. This species is named after the type locality, Snares Islands.</p><p>Diagnosis. This species can be distinguished from other Sagola species by the following combination of characters: body small, length 2.2–2.5 mm; anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round; eye small, one-third length of temple; shape of antennomeres and genitalia unique to species; only known from Snares Islands.</p><p>Description. Length 2.2–2.5 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 76b). Head: Male head round, widest across eyes (Fig. 76f). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus shallow reaching midpoint of eye (Fig. 76f). Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round (Fig. 76f). Eye small, one-third length of temple (Fig. 76f). Thorax: Prosternum as long as wide, widest at midpoint. Elytra approximately as long as wide (Fig. 76b). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Male ventrite VII with round depression. Aedeagus: Median lobe of genitalia bluntly triangular (Fig. 76d). Phallobase of median lobe symmetrical and rectangular (Fig. 76d). Parameres symmetrical, with thick setae from apex to middle (Fig. 76d).</p><p>Distribution. Snares Islands (Fig. 78: black square).</p><p>Habitat. Specimens of this species were collected mostly by sifting leaf and wood litter of Olearia lyallii Hook. f ( Asteraceae) or Stilbocarpa robusta (Kirk) Cockayne (Araliaceae) .</p></div>	https://treatment.plazi.org/id/03D82A3EBDFE11DB3CD2FB7AFBF7FAA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
03D82A3EBDF111D53CBAFC27FEA1FDD1.text	03D82A3EBDF111D53CBAFC27FEA1FDD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sagola opercularis Broun 1915	<div><p>Sagola opercularis Broun, 1915</p><p>(Figs. 77–78)</p><p>Sagola opercularis Broun, 1915: 284 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.</p><p>Type Material Examined. Holotype: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, aedeagus dissected and mounted in balsam on a clear plastic card, “Type” [red label, printed]; “3698. ♂ ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Rakaia 27.12.1912.” [white label, handwritten]; “ Sagola opercularis . ♂.” [white label, handwritten].</p><p>Diagnosis. This species can be distinguished from other Sagola species by the following combination of characters: posterior frontal fovea elongate; ventral surface of male head with W-shaped process bearing dense setae behind process; median lobe of male genitalia curved laterally, C-shaped.</p><p>Redescription of Male. Length 2.1 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 77a). Head: Head round, widest across eyes (Fig. 77b). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching hind point of eye (Fig. 77a). Anterior frontal fovea small round and partially covered by frontal rostrum, posterior frontal fovea elongate (Fig. 77a). Thorax: Prosternum as long as wide, widest at midpoint. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of microtrichial patches. Aedeagus: Genitalia with claw-shaped apical lobe (Fig. 77d). Phallobase of median lobe symmetrical and round (Fig. 77c–d). Parameres symmetrical, with short setae from base to middle and long setae from middle to apex (Fig. 77c).</p><p>Type Locality. Rakaia Gorge, near Methven, Mid Canterbury, New Zealand .</p><p>Distribution. Mid Canterbury (Fig. 78: star).</p><p>Habitat. Unknown.</p></div>	https://treatment.plazi.org/id/03D82A3EBDF111D53CBAFC27FEA1FDD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, J. - S.;Carlton, C. E.	Park, J. - S., Carlton, C. E. (2014): A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae). The Coleopterists Bulletin (mo 13) 68: 1-156, DOI: 10.1649/072.068.0mo4.1
