taxonID	type	description	language	source
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	description	(FIGS 2 – 4) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. o r g: act: 58 C 958 C 5 - 6656 - 4749 - 94 C 1 - 69 E 7666 B 58 EC	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	type_taxon	Type species: † Baltosidis damzeni.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	diagnosis	Diagnosis: Baltosidis is most similar to the genus Parasidis, having ten antennomere antenna with three-antennomere club, ventrite 1 with complete or almost complete v-shape abdominal postcoxal lines and four tarsomeres. However, it can be distinguished by a presence of distinct sutural carinae placed in the apical half of elytra. From similar-looking Coccidophilus Brèthes, 1905 and Stictospilus Brèthes, 1924, it can be separated by four-segmented tarsi (three segmented in both genera) and an antennal club consisting of three antennomeres (in Coccidophilus it consists of two antennomeres).	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	description	Description: Length 1.05 – 1.28 mm. Body elongate, oval and convex (Figs 2 D, 3 A, 4 A), dorsum smooth (Fig. 3 E) or sparsely setose (Fig. 2 B); setae, if present, associated with micropunctures. Metathoracic wings present (Fig. 4 A) in at least in two species (in. B. damgaardi not observed). Head transverse, eyes large (Fig. 2 G), without interfacetal setae, dorsally not margined (Fig. 2 I). Frons (Fig. 2 G) about as long as length of an eye, emarginate around antennal insertions, lateral sides margined. Subantennal grooves shallow, extending behind eyes. Antenna consisting of ten antennomeres (Fig. 2 I) with three apical forming distinct club (Fig. 3 B). Terminal maxillary palpomere knife-like (cultriform) (Fig. 3 F). Pronotum transverse (Fig. 3 C), disc with complete line separating anterior corners (Figs 3 E, 4 E), posterior and lateral margins bordered (Fig. 4 E), anterior edge not margined. Prosternum with prominent chin-piece (Figs 2 G, 3 B), prosternal process about 0.6 × as wide as procoxa. Elytral surfaces with fine punctures, with sutural line present along posterior half length (Figs 2 E, 3 D), epipleura reaching posterior border of ventrite 3, with bordering line reaching posterior margin of ventrite 2 (Figs 2 A, 3 A). Scutellar shield heart-like (Figs 3 C, 4 E). Mesoventral process transverse (Figs 2 G, 4 B), raised anteriorly. Metaventrite (Figs 2 F, 3 A) transverse, 1.5 × longer than abdominal ventrite 1, mesometaventral junction arcuate anteriorly, discrimen visible in posterior half, metanpisternum with short, transverse sulcus near apical margin, postcoxal lines complete or incomplete laterally, straight or descending. Abdomen with six ventrites, ventrite 6 partially visible (Fig. 3 A), ventrite 1 as long as ventrites 2 and 3 combined, postcoxal lines arcuate and incomplete laterally, accessory line long, often joining postcoxal line at apex forming v-shape (Fig. 4 C). Legs slender with femora slightly swollen (Fig. 3 A), tarsi with four tarsomeres (Fig. 2 C), pretarsal claws barely appendiculate.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	etymology	Etymology: First part of the name is derived from the Baltic Sea, the second refers to the genus Parasidis to which it is related.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E4AFFD6FEE1F9F1A7AC3A46.taxon	description	KEY TO THE SPECIES OF BALTOSIDIS	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	description	(FIG. 2 A – J) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org: act: 96 CACFFB- 9847 - 4 DED- 8212 - 8818 CDDD 25 A 3	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	materials_examined	Material examined: Holotype. GPIH no. 5030, Carsten Gröhn collection no. C 8111, FT-IR 9615 (Fig. 5), sex unknown, no syn-inclusions. Paratypes. GPIH no. 5031, Carsten Gröhn collection no. C 4527, FT-IR 9614 (Fig. 5), female, no syn-inclusions; ALD Co 80, FT-IR 9616 (Fig. 5), sex unknown, no syn-inclusions.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	diagnosis	Diagnosis: Baltosidis damgaardi can be distinguished from congeners by pronotum and elytra covered with two types of setae: long, stiff bristles intermixed with short hairs. Moreover, its abdominal postcoxal lines are widely separated from accessory lines. Ventral surface setose.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	description	Description: Total length = 1.05 – 1.18 mm, total width = 0.73 – 0.79 mm. Body elongate oval (Fig. 2 A, B). Head, pronotum and elytra covered with two types of setae: distinct long setae intermixed with fine, short hairs (Fig. 2 D). Pronotum covered with medium-sized, irregular punctures about same size as those on elytra, punctures associated with setae (Fig. 2 B). Metaventrite covered with moderately large punctures separated by distance of about its double diameter (Fig. 2 F), punctures associated with short setae, additional rows of large punctures present anteriorly to hind coxae, metaventral postcoxal lines complete laterally, slightly descending, discrimen visible in posterior half. Elytra covered with dense punctures of similar size as those on pronotum, irregularly distributed, punctures associated with setae, sutural line present in apical third (Fig. 2 E), elytral margin narrow. Elytral epipleuron punctate, glabrous, incomplete, reaching to posterior border of ventrite 3 (Fig. 2 A). Wings not observed. Ventrite 1 as long as 2 and 3 combined, surface of ventrites covered with punctures of same size and density as those on metaventrite, abdominal postcoxal lines reaching one-seventh of length of ventrite 1 at level of hind coxa, incomplete, accessory line distinctly curved, short, broadly separated from postcoxal line, not forming v-shape pattern (Fig. 2 H). Ventrite 2 longer than ventrite 3, ventrites 3 and 4 subequal in length, ventrite 5 large, rounded, about as long as ventrite 2. Ventrites 2 – 5 covered with long setae. Femora swollen, tibiae extended apically, tarsi tetramerous (Fig. 2 C). A part of female genitalia can be observed in paratype C 4527 (GPIH 5031) (Fig. 2 J): coxites narrow, elongate, with two to three long setae attached to stylus.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	etymology	Etymology: The specific epithet is dedicated to Anders L. Damgaard (Copenhagen, Denmark), collector of one of the paratypes, who allowed me to examine his collection of coccinellid beetles.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFDAFEECFB87A69339C7.taxon	discussion	Locality and horizon: Eocene, Baltic amber, Gulf of Gdańsk area (secondary deposit).	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E48FFD5FC44FA9BA02D3CE7.taxon	description	(FIG. 3 A – G) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org: act: A 2 F 18452 - 6 E 52 - 41 FB- 86 BB- 3 FB 99 A 29 D 35 B Material examined: Holotype. MAIG no. 5988, donated by Jonas Damzen, FT-IR 9694 (Fig. 5), female, no syn-inclusions. Paratypes. MAIG no. 5967, donated by Jonas Damzen, sex unknown, no syn-inclusions; MAIG no. 3687, donated by Tomasz Dębowski & Maciej Szulimowicz, 09.09.2000, sex unknown, syninclusions: Cantharidae 1, Hymenoptera 1; AWI- 154, FT-IR 14954 (Fig. 5), sex unknown, no syn-inclusions. Other material. MAIG no. 5985, sex unknown, no syn-inclusions. Diagnosis: Baltosidis damzeni can be distinguished from other Baltosidis species by glabrous pronotum and elytra, abdominal postcoxal lines almost fused with accessory lines, but not forming distinctly closed v-shape. Ventral surface glabrous. Description: Total length = 1.23 – 1.28 mm, total width = 0.80 – 0.86 mm. Body elongate oval (Fig. 3 A). Head, pronotum and elytra glabrous (Fig. 3 C, G). Pronotum covered with medium-sized, irregular punctures about same size as those on elytra. Metaventrite covered with small punctures separated by a distance of about 3 – 4 × its diameter (Fig. 3 A), punctures not associated with setae, without additional rows of large punctures anteriorly to hind coxae, metaventral postcoxal lines complete laterally, straight, discrimen visible in posterior half. Elytra covered with dense punctures of similar size as those on pronotum, irregularly distributed, punctures not associated with setae, sutural line present in apical third (Fig. 3 D), elytral margin narrow. Elytral epipleuron punctate, glabrous, incomplete, reaching to posterior border of ventrite 3 (Fig. 3 A). Wings well developed. Ventrite 1 as long as 2 and 3 combined, surface covered with punctures of smaller size than those on metaventrite, abdominal postcoxal lines reaching one-seventh of length of ventrite 1 at level of hind coxa, incomplete, accessory line distinctly curved, long, separated from postcoxal line only in apical part, not forming v-shape pattern (Fig. 3 G). Ventrite 2 slightly longer than ventrite 3, ventrites 3 and 4 subequal in length, ventrite 5 large, rounded, about as long as ventrite 2. Ventrites 2 – 5 glabrous. Femora swollen, tibiae extended apically, tarsi tetramerous. Coxites of female genitalia can be observed in holotype (MAIG 5988) specimen: they are narrow, elongate, with one to two long setae attached to stylus (Fig. 3 A). Etymology: The specific epithet is dedicated to my friend Jonas Damzen (Vilnius, Lithuania) collector and donator of the type specimens to the MAIG. Locality and horizon: Eocene, Baltic amber, Gulf of Gdańsk area (secondary deposit).	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	description	(FIG. 4 A – E) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org: act: 0 DE 65 C 57 - 3418 - 455 A-A 64 C- 79 C 12 CBBBD 89	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	materials_examined	Material examined: Holotype. GPIH no. 5032, Carsten Gröhn collection no. C 4673, FT-IR 9613 (Fig. 5), sex unknown, no syn-inclusions. Paratype. CCHH 891, specimen preserved in epoxy resin, sex unknown, no syn-inclusions.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	diagnosis	Diagnosis: Baltosidis szadziewskii is characterized by fine, short setae on pronotum and elytra, abdominal postcoxal lines fused with accessory lines apically forming v-shape. Ventral surface setose.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	description	Description: Total length = 1.13 mm, total width = 0.78 mm. Body elongate oval (Fig. 4 A, B). Head, pronotum and elytra shortly setose. Pronotum covered with small, irregular punctures (Fig. 4 E), punctures associated with setae (Fig. 4 D), setae on pronotum longer than those on elytra. Metaventrite covered with moderately large punctures separated by distance of about its double diameter, punctures associated with short setae, without distinct additional rows of large punctures anteriorly to hind coxae, metaventral postcoxal lines complete laterally, straight, discrimen visible in posterior half. Elytra covered with dense punctures of similar size as those on pronotum, irregularly distributed, punctures associated with short setae, sutural line present in apical half, elytral margin narrow (Fig. 4 A). Elytral epipleuron punctate, punctures associated with short seta, incomplete, reaching to posterior border of ventrite 3. Wings well developed (Fig. 4 A, B). Ventrite 1 distinctly shorter than 2 and 3 combined, surface covered with punctures of same size as on metaventrite but distributed more sparsely, abdominal postcoxal lines reaching oneseventh of length of ventrite 1 at level of hind coxa, incomplete, accessory line distinctly curved and roundly merged to postcoxal line, forming narrow v-shape pattern (Fig. 4 C). Ventrite 2 longer than ventrite 3, ventrites 3 and 4 subequal in length, ventrite 5 large, rounded, slightly shorter than ventrite 2. Ventrites 2 – 5 covered with moderately long setae. Femora swollen, tibiae extended apically, tarsi tetramerous.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	etymology	Etymology: The specific epithet is dedicated to Prof. Ryszard Szadziewski (University of Gdańsk, Poland), entomologist and expert on recent and fossil Ceratopogonidae (Diptera), who encouraged me to investigate Baltic amber coleopterans.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	distribution	Locality and horizon: Eocene, Baltic amber, Gulf of Gdańsk area (secondary deposit).	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
03EDD1467E47FFD6FC75FF51A72C39E7.taxon	discussion	Note: I examined one additional specimen from the Bitterfeld deposit (CCHH 743 - 1), which also belongs to the genus Baltosidis. However, its state of preservation (large bubble on ventral side) prevents correct identification to the species level.	en	Szawaryn, Karol (2021): The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193: 1294-1309, DOI: 10.1093/zoolinnean/zlaa180, URL: https://doi.org/10.1093/zoolinnean/zlaa180
