identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F2440A25FF8D4B39DBD5BB6EFCE1F930.text	F2440A25FF8D4B39DBD5BB6EFCE1F930.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proleonhardella (Proleonhardella) hirtella Jeannel 1934	<div><p>Proleonhardella (Proleonhardella) hirtella Jeannel, 1934</p> <p>SERBIA • 1 ♂; southwestern Serbia, municipality of Priboj, village of Krnjača, Tmuša Gorge, Goveđa Pećina Cave; 17 Jul. 2013; Dragan Antić leg.; IZFB.</p> </div>	http://treatment.plazi.org/id/F2440A25FF8D4B39DBD5BB6EFCE1F930	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF8D4B39DBD5BBE6FD43F861.text	F2440A25FF8D4B39DBD5BBE6FD43F861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proleonhardella (Proleonhardella) remyi Jeannel 1934	<div><p>Proleonhardella (Proleonhardella) remyi Jeannel, 1934</p> <p>SERBIA • 6 ♂♂, 8 ♀♀; southwestern Serbia, municipality of Priboj, village of Krnjača, Tmuša Gorge, Goveđa Pećina Cave; 17 Jul. 2013; Dragan Antić leg.; IZFB • 8 ♂♂, 18 ♀♀; southwestern Serbia, municipality of Prijepolje, Kamena Gora, village of Kamena Gora, Bezdan Pit; 20 May–5 Nov. 2017; Miloš Kuraica leg.; pitfall traps; IZFB.</p> </div>	http://treatment.plazi.org/id/F2440A25FF8D4B39DBD5BBE6FD43F861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF8A4B3FD939BF5DFC86FA35.text	F2440A25FF8A4B3FD939BF5DFC86FA35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bozidaria Curcic & Pavicevic 2021	<div><p>Genus Bozidaria Ćurčić &amp; Pavićević gen. nov.</p> <p>urn:lsid:zoobank.org:act: 8CF69930-741E-4484-9F09-5E8C1FBC86AA</p> <p>Type species</p> <p>Bozidaria serbooccidentalis Ćurčić &amp; Pavićević gen. et sp. nov., by monotypy.</p> <p>Diagnosis</p> <p>Bozidaria gen. nov. is most closely related to the following Dinaric genera of the group Théléomorphes belonging to the phyletic series of “ Leonhardella ” (Jeannel 1924): Proleonhardella, Blattochaeta Reitter, 1910, Augustia Zariquiey, 1927 and Pholeuodromus Breit, 1913. These genera share a similar body form, the presence of tetramerous tarsi in males, the absence of a comb on anterior tibiae, the first antennomere clearly shorter than the second antennomere, apically widened distal antennomeres, the absence of sutural striae, and the presence of a similar type of aedeagus.</p> <p>The new genus differs from its closest relatives in the body shape (elliptical, elongate vs bathyscioid, oval/ovoid, mostly wide in Proleonhardella), TL (R 2.51–2.80 mm vs R 4.0– 5.5 mm in Blattochaeta and R 3.8–4.6 mm in Pholeuodromus), body pubescence (short, recumbent vs long, erect in Blattochaeta), shape of antennae (elongate, thin, including distal antennomeres vs short, distal antennomeres wide, barely longer than wide in Proleonhardella), length of antennae (exceeding the middle of the body vs reaching, but not exceeding the middle of the body in Augustia and not reaching the middle of the body in Pholeuodromus), presence/absence of mesosternal carina (present vs absent in Augustia), shape of mesosternal carina (with no concavity on its anterior border, not atrophied vs with a concavity on its anterior border in Blattochaeta and atrophied posteriorly in Pholeuodromus), shape of lateral pronotal margins (arcuate vs weakly convex in Augustia), position of maximum pronotal length (sub-basally vs at base in Pholeuodromus), ratio of pronotum width to elytral width (pronotum slightly narrower than elytra vs pronotum as wide as elytra in Blattochaeta and Pholeuodromus and pronotum clearly narrower than elytra in Augustia), shape of elytra (more rounded, gradually narrowed distally vs less rounded, more pronouncedly attenuated distally in Augustia), shape of aedeagus (long, elongate vs short, wide in Blattochaeta and Augustia and mostly wide in Proleonhardella), shape of basal bulb (elongate, narrow vs short, rounded in Proleonhardella, Blattochaeta and Augustia) and its basal projection (long vs short in Proleonhardella and Augustia), and shape of parameral apex (narrow vs widened in Blattochaeta) (Jeannel 1910, 1924, 1930, 1931, 1934; Reitter 1910; Breit 1913; Zariquiey 1927; Knirsch 1928; Guéorguiev 1976).</p> <p>Etymology</p> <p>This genus is named after the late Academician Božidar Ćurčić, a well-known Serbian biospeleologist and zoologist.</p> <p>Description</p> <p>HABITUS. A small-sized elliptical leptodirine with short and wide head, transverse pronotum and elongate obovoid elytra. Blind, reddish-brown, body shiny, densely pubescent, dorsoventrally convex, finely punctate. Pubescence composed of short yellow hairs, on pronotum and elytra recumbent, while on head erect. Legs and antennae long and slender, densely pubescent. Microsculpture composed of isodiametric meshes.</p> <p>HEAD. Anophthalmous, of almost equal length and width. Antennae inserted medially on head, elongate, thin, apically widened and flattened, exceeding middle of body, reaching basal third of elytral length. Antennomere I shorter than antennomere II. Antennomere III shorter than antennomere II and longer than antennomere IV. Antennomeres IV–VI of similar length. Antennomere VII elongate, apically widened. Antennomere VIII short, elongate, oval. Ultimate antennomere slender, ovoid, about as long as antennomeres IX and X combined. Occipital carina present.</p> <p>THORAX. Pronotum almost twice as wide as long, with arcuate and well-rounded lateral margins, slightly narrower than elytra, widest slightly prior to pronotal base. Mesosternal carina well-developed, with no furrow, high, obtuse-angled, with an apical tooth.</p> <p>ELYTRA. Elongate, much longer than pronotum, rounded medially, regularly arcuate distally, not attenuated. Scutellar striae absent. A part of pygidium not covered by elytra.</p> <p>LEGS. Extended and slender. Fore tarsi tetramerous. Male protarsi dilated. Tibiae with spines on external edges. No comb on external edges or apical parts of protibiae. Meso- and metatibiae with no apical baskets.</p> <p>ABDOMEN. Median lobe of aedeagus slender, rounded sub-terminally, with an elongate triangular apex. Basal bulb elongate, narrow, with a long sub-triangular basal projection. Each paramere longer than median lobe, thin, sub-terminally widened, with three apical setae.</p> <p>GONOSTYLI. Elongate, slender, almost straight.</p> <p>Distribution</p> <p>The new genus is currently known to inhabit deep soil on Mt Bobija and the Simina Jama Pit on Mt Povlen in the surroundings of the town of Ljubovija, western Serbia. It is probable that it might be present both in the soil and caves of the adjacent areas.</p></div> 	http://treatment.plazi.org/id/F2440A25FF8A4B3FD939BF5DFC86FA35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF8B4B32DB82B8E5FAB2FC81.text	F2440A25FF8B4B32DB82B8E5FAB2FC81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leonhardella Reitter 1903	<div><p>Key to the leptodirine leiodid genera of the phyletic series of “ Leonhardella ”</p> <p>(modified after Guéorguiev 1976) (Fig. 1)</p> <p>1. Body of bathyscioid form, oval or ovoid, wide and short................................................................ 2</p> <p>– Body of elliptical or pholeuonoid form............................................................................................ 6</p> <p>2. Mesosternal carina absent (Fig. 1A). Cavernicolous, Mt Čvrsnica, southwestern Bosnia and Herzegovina.............................................................................................. Augustia Zariquiey, 1927</p> <p>– Mesosternal carina present (Fig. 1B)................................................................................................ 3</p> <p>3. Antennae short, not reaching middle of body................................................................................... 4</p> <p>– Antennae much longer, reaching middle of body............................................................................. 5</p> <p>4. Pubescence fine and recumbent, except in both lateral exterior border of elytra and apical elytral half, where long hairs occur. Antennae very short, barely exceeding pronotal base. Longer species (TL 2.5 mm). Mesosternal carina rounded. Basal lamina of tegmen of aedeagus without tooth. Cavernicolous, region of Kuči, vicinity of the city of Podgorica, eastern Montenegro..................................................................................................................................... Weiratheria Zariquiey, 1927</p> <p>– Pubescence entirely normal and recumbent. Antennae somewhat longer, but not reaching middle of body. Shorter species (TL 1.85 mm). Mesosternal carina triangular. Basal lamina of tegmen of aedeagus with a pronounced tooth basally. Cavernicolous, vicinity of the town of Karystos, island of Euboea, southern Greece.......................................................................... Henrotiella Perreau, 1999</p> <p>5. Pubescence short and recumbent. Anterior border of mesosternal carina without concavity (Fig. 1C). Protarsi weakly dilated in males. Shorter species (TL 1.3–3.5 mm). Cavernicolous and endogean, central, eastern and southern Bosnia and Herzegovina, southwestern and western Serbia and eastern Montenegro...................................................... Proleonhardella Jeannel, 1910</p> <p>– Pubescence long and erect. Anterior border of mesosternal carina with a deep concavity (Fig. 1D). Protarsi not dilated in males. Longer species (TL 4.0– 5.5 mm). Cavernicolous, western, southwestern and eastern Montenegro, southern Bosnia and Herzegovina and southern Croatia............................................................................................................................................. Blattochaeta Reitter, 1910</p> <p>6. Body of elliptical form (Fig. 1E). Pronotum regularly or almost regularly arcuate, slightly narrower than elytra or as wide as elytra......................................................................................................... 7</p> <p>– Body of pholeuonoid form (Fig. 1F). Pronotum campanuliform or strongly sinuated backwards, clearly narrower than elytra.............................................................................................................11</p> <p>7. Body shorter (TL 1.8–2.0 mm). Antennae almost reaching middle of body. Elytra very attenuated apically, with sparse pubescence. Ventral border of mesosternal carina triangular and deeply grooved. Endogean and cavernicolous, southern Croatia and western Bosnia and Herzegovina....................................................................................................................................... Anisoscapha Müller, 1917</p> <p>– Body longer (TL 2.5–5.2 mm). Antennae reaching middle of body. Elytra regularly arcuate apically, with dense pubescence. Ventral border of mesosternal carina not grooved..................................... 8</p> <p>8. Body shorter (TL 2.5–3.0 mm). Mesosternal carina not atrophied. Protarsi dilated in males (Fig. 1G)........................................................................................................................................... 9</p> <p>– Body longer (TL 3.8–5.2 mm). Mesosternal carina atrophied posteriorly. Protarsi not dilated in males (Fig. 1H)............................................................................................................................... 10</p> <p>9. Body very elongate and narrower. Pubescence long and erect. Median lobe more elongate, thin. Basal bulb small, with a short rounded basal projection (Fig. 1I). Paramerae distally widened. Cavernicolous, southern and southeastern Bosnia and Herzegovina and western Montenegro........................................................................................................................... Anillocharis Reitter, 1903</p> <p>– Body less elongate and wider. Pubescence short and recumbent. Median lobe less elongate, wide. Basal bulb elongate, with a long sub-triangular basal projection (Fig. 1J). Paramerae distally narrow. Endogean and cavernicolous, Mts Bobija and Povlen, western Serbia........................................................................................................................................... Bozidaria Ćurčić &amp; Pavićević gen. nov.</p> <p>10. Body shorter (TL 3.8–4.6 mm). Pronotum as wide as elytra. Lateral pronotal margins regularly arcuate. Anterior border of mesosternal carina with no concavity. Paramerae with three setae. Endogean, central and southern Bosnia and Herzegovina.................... Pholeuodromus Breit, 1913</p> <p>– Body longer (TL 5.0– 5.2 mm). Pronotum slightly narrower than elytra. Lateral pronotal margins weakly sinuate in basal third. Anterior border of mesosternal carina with a deep concavity. Paramerae with four setae. Cavernicolous, southeastern Bosnia and Herzegovina and northern Montenegro................................................................................................................... Blattodromus Reitter, 1904</p> <p>11. Antennomere VIII subglobular (Fig. 1K). Elytral punctuation strong and deep. Pubescence long, usually double, with bristle-like setae. Cavernicolous and endogean, western Serbia, southern, southeastern, central, western and eastern Bosnia and Herzegovina and northwestern Montenegro......................................................................................................... Pholeuonopsis Apfelbeck, 1901</p> <p>– Antennomere VIII elongate (Fig. 1L). Elytral punctuation fine and shallow. Pubescence short, with no bristle-like setae......................................................................................................................... 12</p> <p>12. Body shorter (TL 2.0 mm). Pronotum elongate, very narrow basally. Mesosternal carina low. Paramerae with two setae (Fig. 1M). Cavernicolous, vicinity of the town of Ključ, western Bosnia and Herzegovina................................................................................. Deelemaniella Perreau, 2002</p> <p>– Body longer (TL 3.2–4.5 mm). Pronotum as long as wide or transverse, wider basally. Mesosternal carina high or atrophied. Paramerae with three setae (Fig. 1N)..................................................... 13</p> <p>13. Mesosternal carina high. Protarsi somewhat dilated in males. Tibiae with no external spur. Cavernicolous, northwestern, western and northern Montenegro and southeastern and southern Bosnia and Herzegovina......................................................................... Leonhardella Reitter, 1903</p> <p>– Mesosternal carina atrophied. Protarsi not dilated in males. Tibiae with external spur. Cavernicolous, Mt Durmitor, northern Montenegro................................. Tartariella Nonveiller &amp; Pavićević, 1999</p></div> 	http://treatment.plazi.org/id/F2440A25FF8B4B32DB82B8E5FAB2FC81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF864B36DA9EBE98FAD5FDE8.text	F2440A25FF864B36DA9EBE98FAD5FDE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bozidaria serbooccidentalis Curcic & Pavicevic 2021	<div><p>Bozidaria serbooccidentalis Ćurčić &amp; Pavićević gen. et sp. nov.</p> <p>urn:lsid:zoobank.org:act: A2B51421-4CD9-4696-BF46-7EACBC395C4E</p> <p>Figs 2–3</p> <p>Diagnosis</p> <p>The genus is currently monotypic and therefore a differential diagnosis for Bozidaria serbooccidentalis gen. et sp. nov. cannot be provided.</p> <p>Etymology</p> <p>The species is named after western Serbia, where its type locality and known localities are situated.</p> <p>Type material</p> <p>Holotype SERBIA • ♂; western Serbia, town of Ljubovija, Mt Bobija; alt. 1000 m; 19 Apr. 1980; Guido Nonveiller leg.; traps for endogean fauna baited with rotten meat; IZFB-21/1.</p> <p>Paratypes SERBIA • 1 ♂, 2 ♀♀; same collection data as for holotype; SBS- 21/1 to 21/3 • 1 ♀; same collection data as for holotype; IZFB-21/2 • 2 ♂♂, 2 ♀♀; same collection data as for holotype; CDP- 21/1 to 21/4 • 10 ♂♂, 14 ♀♀; western Serbia, town of Ljubovija, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.62789&amp;materialsCitation.latitude=44.14228" title="Search Plazi for locations around (long 19.62789/lat 44.14228)">Mt Povlen</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.62789&amp;materialsCitation.latitude=44.14228" title="Search Plazi for locations around (long 19.62789/lat 44.14228)">Debelo Brdo</a> saddle, village of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.62789&amp;materialsCitation.latitude=44.14228" title="Search Plazi for locations around (long 19.62789/lat 44.14228)">Gornje Košlje</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.62789&amp;materialsCitation.latitude=44.14228" title="Search Plazi for locations around (long 19.62789/lat 44.14228)">Simina Jama Pit</a>; 44°08′32.2″ N, 19°37′40.4″ E; 20 May–5 Nov. 2017; Miloš Kuraica leg.; pitfall trapping; IZFB-21/3 to 21/26 • 2 ♂♂, 1 ♀; same locality as for preceding; 31 Dec. 2010; Iva Njunjić leg.; pitfall trapping; SBS- 21/4 to 21/6 • 18 ♂♂, 39 ♀♀; same collection data as for preceding; CDP-21/5 to 21/61 (Fig. 2).</p> <p>Description</p> <p>HABITUS. Body elliptical, TL R 2.51–2.80 mm (R 2.51–2.73 mm in males, R 2.67–2.80 mm in females), reddish-brown in colour, shiny, pubescent, with a fine punctuation (Fig. 2A).</p> <p>HEAD. Short, wide, slightly wider than long (HL/HW M 0.97), without eyes (Fig. 2A). Antennae long and slender, apically gradually widened and flattened, ending slightly after basal third of elytra in males or slightly prior to basal third of elytra in females. Antennomere II longer than antennomere I (A1/A2 M 0.795). A3/A2 M 0.67. A3/A5 M 1.24. Antennomeres IV–VI of similar length, of which IV narrowest and VI widest. Antennomere VII apically widened. A7/A6 M 1.56. Antennomere VIII slightly longer than half of antennomere VII (A7/A8 M 1.53), oval, somewhat elongate (A8LW M 1.52). Antennomere IX somewhat elongate, gradually widened distally (A9LW M 1.47). A9/A8 M 1.52 in males, M 1.42 in females. Antennomere X slightly longer than wide (A10LW M 1.24), more widened apically. Antennomere XI slender, ovoid, apically pointed, more elongate in males (A11LW M 2.45) than in females (A11LW M 1.97), as long as preceding two antennomeres combined or slightly shorter than the latter. Occipital carina present. Hairs yellow, erect. Microsculpture composed of small isodiametric meshes.</p> <p>THORAX. Pronotum transverse, widest sub-basally, almost twice as wide as long (PL/PW M 0.59) (Fig.2A). Lateral pronotal margins arcuate, rounded medially, sub-parallel prior to hind pronotal angles. Pronotal base more than twice as long as anterior pronotal margin (PB/AM M 2.13). Both anterior pronotal margin and pronotal base convex medially, the latter less pronouncedly. PL+EL/AL M 1.63 in males, M 1.89 in females. Fore angles prominent, obtuse, rounded, hind angles sharp, rounded, prominent, directed backwards. Microsculpture of pronotum composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 2B). Pronotal disc weakly convex. Mesosternal carina high, obtuse-angled, anterior margin strongly convex, posterior margin barely convex, almost straight, with hairs and teeth (Fig. 2C). Mesosternal carina with an apical tooth. Ventral border of mesosternal carina not grooved.</p> <p>ELYTRA. Elongate (EL/EW M 1.38 in males, M 1.40 in females), more than 2.5 times as long as pronotum (EL/PL M 2.65), obovoid, sub-parallel below humeral angles, weakly narrowed basally, rounded medially, narrowed apically (Fig. 2A). Apex rounded. Sutural striae absent. Scutellum small, triangular. Elytra widest between basal third and mid-length. Microsculpture composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 2D). Elytral disc convex. Pygidium not completely covered by elytra.</p> <p>LEGS. Elongate and thin, with hairs (Fig. 2A). Tibiae with a few spines laterally.Anterior tarsi tetramerous in both genders, somewhat dilated in males (P1LW M 1.825 in males, M 1.88 in females).</p> <p>ABDOMEN. Median lobe of aedeagus elongate, thin, sub-parallel, sub-apically rounded (Figs 2E, 3A). Apex elongate, triangular. Basal bulb relatively narrow, elongate, with a sub-triangular basal projection. Copulatory piece weakly chitinised, consisting of a basal phanera, median paired stripes and apical paired sclerotizations. Median lobe proximally straight, distally relatively curved, gradually narrowed distally in lateral view (Fig. 3B). Basal bulb narrow in lateral view. Parameres slender, thin, longer than median lobe, sub-terminally widened, terminally narrowed, apex slightly dilated, proximally arcuate and distally straight in lateral view, with three setae: one apical terminal, one apical inner and one sub-apical inner (Fig. 2F). Two apical parameral setae close-set. Parameral apices directed inwards. Parameres basally slightly curved, distally relatively straight in lateral view, sub-terminally widened, terminally narrowed in lateral view (Fig. 3B).</p> <p>GONOSTYLI. Almost straight, elongate, thin, with one apical seta, three inner setae and one outer seta (Fig. 2G).</p> <p>SPERMATHECA. Small, curved, widest in proximal third, apically sub-spherical (Fig. 2H).</p> <p>FEMALE ABDOMINAL STERNITE VIII. Large, transverse, setose both medially and distally, with a narrow, pointed anterior process. Microsculpture consisting of transverse polygonal meshes (Fig. 2I).</p> <p>Bionomy, distribution and type locality</p> <p>The type specimens were collected in traps for endogean fauna (cans) baited with rotten meat placed in the deep soil on Mt Bobija, near the town of Ljubovija, western Serbia, as well as by pitfall trapping with rotten meat as bait in the deep, totally dark parts of the Simina Jama Pit, village of Gornje Košlje, Debelo Brdo saddle, Mt Povlen, near the town of Ljubovija, western Serbia (Fig. 9). The type locality on Mt Bobija is located on its northern slope, at an altitude of 1000 m a.s.l., in a beech forest, close to several streams. The entrance of the Simina Jama Pit is situated at 920 m a.s.l., the total length of its investigated channels is 270 m, while its depth is 56 m. It starts with a 31-m long vertical passage, which splits into two horizontal channels – left and right (Anđelić et al. 2011). Beetle specimens were found at the end and in the middle of the left horizontal channel with a clay muddy substrate and rocks, on the vertical limestone walls and floor with a high level of humidity (presence of trickling water). The places where the specimens were found in the pit are shown in Fig. 4. It is assumed that the species is actually endogean, as is the case with some other leiodid taxa (e.g., Magdelainella spp.), which inhabit the soil beneath deeply sunken rocks and forest detritus, but can also be found in caves and pits (Pavićević et al. 2012).</p> </div>	http://treatment.plazi.org/id/F2440A25FF864B36DA9EBE98FAD5FDE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF834B2EDA52BC44FE20FE41.text	F2440A25FF834B2EDA52BC44FE20FE41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proleonhardella (Proleonhardella) tarensis Curcic & Pavicevic 2021	<div><p>Proleonhardella (Proleonhardella) tarensis Ćurčić &amp; Pavićević sp. nov.</p> <p>urn:lsid:zoobank.org:act: E1B53D50-0C09-4E5C-AFB1-AC9F6EBBD426</p> <p>Figs 5–6</p> <p>Diagnosis</p> <p>Proleonhardella (Proleonhardella) tarensis sp. nov. is most closely related to P. (P.) hirtella (from several caves and pits near the towns of Prijepolje (southwestern Serbia) and Pljevlja (northern Montenegro)), P. (P.) weiratheri (Reitter, 1913) (from the Vrteljka Cave, village of Đipi, Mt Sjemeć, near the town of Višegrad, eastern Bosnia and Herzegovina) and P. (P.) neumanni (Apfelbeck, 1901) (from a small unnamed cave, village of Podromanija, near the town of Sokolac, eastern Bosnia and Herzegovina) (Fig. 9) (Perreau 2000; Pavićević et al. 2012). Another congener from Serbia, P. (P.) remyi (from caves and pits in the area of Kamena Gora and near the towns of Prijepolje, Priboj (southwestern Serbia) and Pljevlja (northern Montenegro)), is of bathyscioid shape (P. (P.) tarensis sp. nov. is more elongate and of oval shape), it is significantly longer than the new species (TL R 3.0– 3.5 mm vs 2.185 –2.435 mm in P. (P.) tarensis sp. nov.) and has a quite different shape of aedeagus (stout, with a rounded apex, longer than parameres vs elongate, with a pointed apex, shorter than parameres in P. (P.) tarensis sp. nov.), indicating that these two species are not closely related (Jeannel 1934; Ćurčić et al. 2008a).</p> <p>The new species differs from its closest congeners in the TL R (2.185 –2.435 mm vs 1.6–1.8 mm in P. (P.) hirtella and 1.6–2.0 mm in P. (P.) weiratheri), antennal length (ending prior to basal third of elytra vs reaching only basal quarter of elytra in P. (P.) hirtella and exceeding middle of body in P. (P.) weiratheri), A1/A2 M (0.76 vs 0.80 in P. (P.) neumanni), A7/A6 M (1.66 vs 1.50 in P. (P.) hirtella), A7/A8 R (1.75–2.00 vs 3.00 in P. (P.) hirtella), A11/A9+A10 M (0.90 vs 1.00 in P. (P.) weiratheri), shape of antennomere VIII (oval and slightly longer than wide in males and spherical in females vs as long as wide in males and almost transverse in females in P. (P.) weiratheri), shape of antennomeres IX and X (slightly longer than wide vs as long as wide in P. (P.) hirtella), position of maximum width of pronotum (sub-basally vs at base in P. (P.) hirtella and P. (P.) weiratheri), shape of mesosternal carina (obtuse-angled vs almost right-angled in P. (P.) neumanni), shape of elytra (narrowed basally vs parallel basally in P. (P.) hirtella and P. (P.) weiratheri and clearly sinuate basally in P. (P.) neumanni), EL/EW (R 1.31–1.39 vs M 1.75 in P. (P.) weiratheri), position of maximum width of elytra (slightly after basal third vs prior to middle in P. (P.) neumanni), and shape of aedeagus (less elongate, with wider apex and larger basal bulb vs more elongate, with narrower apex and smaller basal bulb in P. (P.) hirtella) (Apfelbeck 1901; Reitter 1913; Jeannel 1924, 1934).</p> <p>Etymology</p> <p>The species is named after Mt Tara in western Serbia, where its type locality is situated.</p> <p>Type material</p> <p>Holotype SERBIA • ♂; western Serbia, town of Bajina Bašta, Mt Tara, village of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.553217&amp;materialsCitation.latitude=43.90853" title="Search Plazi for locations around (long 19.553217/lat 43.90853)">Kaluđerske Bare</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.553217&amp;materialsCitation.latitude=43.90853" title="Search Plazi for locations around (long 19.553217/lat 43.90853)">Pit 4-1-3-27</a>; 43°54′30.712″ N, 19°33′11.585″ E; 5 Jul. 2014; Fabrizio Bosco leg.; pitfall trapping; IZFB-21/27.</p> <p>Paratypes SERBIA • 3 ♂♂, 3 ♀♀; same collection data as for holotype; IZFB-21/28 to 21/33 • 1 ♀; same collection data as for holotype; SBS-21/7 • 1 ♂, 2 ♀♀; western Serbia, town of Bajina Bašta, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.515638&amp;materialsCitation.latitude=43.877693" title="Search Plazi for locations around (long 19.515638/lat 43.877693)">Mt Tara</a>, village of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.515638&amp;materialsCitation.latitude=43.877693" title="Search Plazi for locations around (long 19.515638/lat 43.877693)">Šljivovica</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.515638&amp;materialsCitation.latitude=43.877693" title="Search Plazi for locations around (long 19.515638/lat 43.877693)">Sovljačka Pećina Cave</a>; 43°52′39.7″ N, 19°30′56.3″ E; 7 May 2003; Dragan Pavićević leg.; pitfall trapping; SBS- 21/8 to 21/10 • 2 ♂♂, 3 ♀♀; same collection data as for preceding; CDP-21/62 to 21/66 (Fig. 5).</p> <p>Description</p> <p>HABITUS. Body oval, relatively elongate, TL R 2.185 –2.435 mm (R 2.185–2.32 mm in males, 2.435 mm in females), colour brownish-red (one teneral female specimen yellowish), shiny, pubescent and with a fine punctuation (Fig. 5A).</p> <p>HEAD. Short, wide, slightly longer than wide (HL/HW R 1.00–1.06), anophthalmous (Fig. 5A). Antennae long and narrow, ending prior to basal third of elytra, apically widened and flattened. Antennomere II longer than antennomere I (A1/A2 M 0.76). Antennomeres III–VI small, narrow, of similar shape and length. A3/A2 M 0.58. A3/A5 M 1.19. Antennomere VII apically widened, obovoid. A7/A6 M 1.66. Antennomere VIII half as long as antennomere VII, oval and slightly longer than wide in males (A8LW M 1.31), while somewhat shorter (A7/A8 M 1.75), nearly as long as wide (A8LW M 1.04) and spherical in females. Antennomeres IX and X slightly longer than wide (A9LW M 1.28 and A10LW M 1.21, respectively), apically widened. A9/A8 M 1.60 in males, 2.00 in females. Antennomere XI ovoid, twice as long as wide in males, somewhat shorter in females (A11LW M 1.92), slightly shorter than two preceding antennomeres combined (A11/A9+A10 M 0.90). Occipital carina present. Hairs yellow, erect. Microsculpture composed of small isodiametric meshes.</p> <p>THORAX. Pronotum transverse, almost twice as wide as long (PL/PW M 0.59), widest sub-basally (Fig. 5A). Lateral pronotal margins arcuate, most rounded medially, almost sub-parallel prior to hind pronotal angles. Anterior pronotal margin somewhat convex medially, base almost straight, around twice as long as anterior pronotal margin (PB/AM M 1.985). PL+EL/AL M 1.685 in males, 1.94 in females. Fore angles prominent, obtuse, rounded, hind angles sharp, rounded, prominent, directed backwards. Microsculpture of pronotum composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 5B). Pronotal disc weakly convex. Mesosternal carina high, obtuse-angled, anterior margin convex, posterior margin straight, setose and with unpronounced teeth (Fig. 5C). Mesosternal carina with an apical tooth.</p> <p>ELYTRA. Elongate (EL/EW M 1.39 in males, 1.31 in females), more than twice as long as pronotum (EL/ PL M 2.46), obovoid, sub-parallel below humeral angles, conspicuously narrowed basally, rounded medially, attenuated apically (Fig. 5A). Apex rounded. Sutural striae absent. Scutellum small, triangular. Elytra widest slightly after basal third. Microsculpture composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 5D). Elytral disc gently convex apically, more steeply distally. Pygidium completely covered by elytra.</p> <p>LEGS. Moderately elongate and thin, with hairs (Fig. 5A). Tibiae with a few spines. Anterior tarsi tetramerous in both genders, dilated in males (P1LW M 1.50 in males, 2.00 in females).</p> <p>ABDOMEN. Median lobe of aedeagus elongate, thin, sub-parallel, sub-terminally somewhat widened, then narrowed apically (Figs 5E, 6A). Apex triangular. Basal bulb large, elongate, with a sub-triangular basal projection. Copulatory piece weakly chitinised, consisting of a basal phanera, median paired stripes and distal paired sclerotizations. Median lobe proximally weakly curved, distally more curved, gradually narrowed distally in lateral view (Fig. 6B). Basal bulb relatively narrow in lateral view. Parameres slender, thin, longer than median lobe, sub-terminally widened, terminally narrowed, basally arcuate and distally straight in lateral view, with three setae: one apical terminal, one apical inner and one sub-apical inner (Fig. 5F). Two apical parameral setae close-set. Parameral apices directed inwards. Parameres basally slightly curved, distally relatively straight, sub-terminally widened, terminally narrowed, apically directed downwards in lateral view (Fig. 6B).</p> <p>GONOSTYLI. Straight, elongate, thin, with one apical seta, three inner setae and one outer seta (Fig. 5G).</p> <p>SPERMATHECA. Small, hook-like, widest in middle, apically sub-spherical (Fig. 5H).</p> <p>FEMALE ABDOMINAL STERNITE VIII. Large, transverse, setose in distal half, with a small, narrow anterior process. Microsculpture consisting of transverse polygonal meshes (Fig. 5I).</p> <p>Comparisons</p> <p>Proleonhardella (P.) hirtella, P. (P.) weiratheri and P. (P.) tarensis sp. nov. are somewhat elongate and their aedeagus is narrower than in the remaining congeners, suggesting their specific position within the genus. Based on these features, they are similar to Bozidaria gen. nov., but are much shorter (TL R 1.6– 1.8 mm in P. (P.) hirtella, 1.6–2.0 mm in P. (P.) weiratheri and 2.185 –2.435 mm in P. (P.) tarensis sp. nov. vs 2.51–2.80 mm in B. serbooccidentalis gen. et sp. nov.), share other characteristics of Proleonhardella and additionally differ from the new genus in the shape of the antennae, the median lobe of the aedeagus, the basal bulb and its basal projection.</p> <p>Proleonhardella (P.) tarensis sp. nov. and its closest relatives (P. (P.) hirtella, P. (P.) weiratheri and P. (P.) neumanni) share the presence of elongate, somewhat convex elytra, which are more than twice as long as the pronotum. Furthermore, the new species, P. (P.) hirtella and P. (P.) weiratheri have a somewhat elongate body shape, while the body shape in the remaining Proleonhardella taxa is more or less bathyscioid. These three species have an elongate aedeagus, contrary to other known congeners, in which the aedeagus is more or less short (Jeannel 1924, 1934). The shape of the aedeagus of P. (P.) neumanni wasn’t mentioned in the description of the species or elsewhere (Apfelbeck 1901; Jeannel 1924).</p> <p>Bionomy, distribution and type locality</p> <p>The type specimens were gathered using pitfall traps with rotten meat as bait in Pit 4-1-3- 27 in the village of Kaluđerske Bare, as well as in the Sovljačka Pećina Cave in the village of Šljivovica (Fig. 9). Both localities are situated on Mt Tara, near the town of Bajina Bašta, western Serbia. Beetles were found in the inner (from the middle to the innermost point), totally dark parts of the cave sites. The entrance of Pit 4-1-3-27 is situated at 868 m a.s.l., the total length of its investigated channels is 28 m, while its depth is 22 m. After a short vertical passage, the pit opens into a large chamber which contains big rocks at its lowest part. At this point another vertical passage starts, at the end of which is situated a small, moist semicircular chamber with a clay substrate and rocks (Bosco 2016). Beetle specimens were found in the inner part of the larger chamber, among rocks, and in the smaller chamber with a clay substrate and rocks, on the floor and vertical limestone walls with a high level of humidity (presence of trickling water). The entrance of the Sovljačka Pećina Cave is situated at 1080 m a.s.l. and its total length is 43 m (Bosco 2016). The cave is located in a coniferous forest in a valley where the Sovljak stream runs. It is entirely horizontal and consists of a single channel which is oriented to the left. Its height is slightly decreasing towards the end. Beetle individuals were found in the inner part of the cave, on the floor among rocks, both on limestone and clay substrate, where a high level of humidity (presence of trickling water) was evident. Images of the cave localities and the places where the specimens were found in the caves are shown in Figs 7–8. The new species is most probably endogean and is likely to be found outside caves as well – in the deep soil strata and other speleological sites in the surroundings.</p></div> 	http://treatment.plazi.org/id/F2440A25FF834B2EDA52BC44FE20FE41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
F2440A25FF9A4B2FDB82BCD9FAB2FCFA.text	F2440A25FF9A4B2FDB82BCD9FAB2FCFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proleonhardella Jeannel 1910	<div><p>Key to the taxa of the genus Proleonhardella Jeannel, 1910</p> <p>(modified after Jeannel 1924) (Figs 9–10)</p> <p>1. Body more elongate, elliptical. Pronotum as wide as elytra, well constricted basally. Pronotal lateral margins well-rounded backwards. Mesosternal carina very low. Elytral punctuation rough and deep (subgenus Pholeuonillus Breit, 1913). Endogean, Mt Treskavica, southern Bosnia and Herzegovina....................................................................... P. (Pholeuonillus) adolfi (Reitter, 1911)</p> <p>– Body less elongate, oval. Pronotum narrower than elytra, weakly constricted basally. Pronotal lateral margins weakly rounded backwards. Elytral punctuation fine. Mesosternal carina elevated, angled (subgenus Proleonhardella Jeannel, 1910)....................................................................................... 2</p> <p>2. Elytra shorter, more convex, less than twice as long as pronotum (Fig. 10A)................................. 3</p> <p>– Elytra longer, less convex, more than twice as long as pronotum (Fig. 10B).................................. 6</p> <p>3. Body of oval shape, longer (TL 1.8–2.2 mm). Antennae short, not reaching middle of body. Antennomere III not longer than antennomere V. Antennomere VIII globular in males (Fig. 10C). Cavernicolous, Mts Bjelašnica and Igman, central Bosnia and Herzegovina [P. (Proleonhardella) matzenaueri (Apfelbeck, 1907)]....................................................................................................... 4</p> <p>– Body of subglobular shape, shorter (TL less than 1.8 mm). Antennae long, reaching middle of body. Antennomere III longer than antennomere V. Antennomere VIII slightly elongate in males (Fig. 10D)......................................................................................................................................... 5</p> <p>4. Body of almost regular oval shape, longer (TL 2.0– 2.2 mm). Elytra not widened towards middle. Cavernicolous, Mt Bjelašnica, central Bosnia and Herzegovina......................................................................................................... P. (Proleonhardella) matzenaueri matzenaueri (Apfelbeck, 1907)</p> <p>– Body narrower anteriorly, shorter (TL 1.8–2.0 mm). Elytra clearly widened medially. Cavernicolous, Mt Igman, central Bosnia and Herzegovina.......................................................................................................................................................... P. (Proleonhardella) matzenaueri ottonis Müller, 1917</p> <p>5. Body more elongate and convex, longer (TL 1.5 mm). Elytral punctuation finer and denser.Antennae shorter, with apical antennomeres thicker. Antennomere VIII barely longer than wide, antennomere IX as long as wide and antennomere X transverse in females. Cavernicolous, Mt Treskavica, southern Bosnia and Herzegovina............................................ P. (Proleonhardella) leonhardi (Breit, 1913)</p> <p>– Body less elongate and convex, shorter (TL 1.3 mm). Elytral punctuation stronger and less dense. Antennae longer, with apical antennomeres less thick. Antennomeres VIII, IX and X longer than wide in females. Cavernicolous, village of Trnovo, vicinity of the city of Sarajevo, central Bosnia and Herzegovina...................................................... P. (Proleonhardella) apfelbecki Jeannel, 1924</p> <p>6. Body longer (TL 3.0– 3.5 mm). Cavernicolous, Kamena Gora and vicinity of the towns of Prijepolje, Priboj and Pljevlja, southwestern Serbia and northern Montenegro............................................................................................................................................ P. (Proleonhardella) remyi Jeannel, 1934</p> <p>– Body shorter (TL less than 2.435 mm)............................................................................................. 7</p> <p>7. Elytra parallel in basal half (Fig. 10E).............................................................................................. 8</p> <p>– Elytra narrowed in basal half (Fig. 10F)........................................................................................... 9</p> <p>8. Pubescence long. Punctuation less fine. Apical antennomeres more widened. Pronotum less rounded. Apex of median lobe of aedeagus less bent ventrally. Elytral apex wide, obtuse and oblique. Cavernicolous, vicinity of the towns of Prijepolje and Priboj (southwestern Serbia) and Pljevlja (northern Montenegro).................................................. P. (Proleonhardella) hirtella Jeannel, 1934</p> <p>– Pubescence short. Punctuation finer. Apical antennomeres less widened. Pronotum more rounded. Apex of median lobe of aedeagus more bent ventrally. Elytral apex attenuated. Cavernicolous, Mt Sjemeć, eastern Bosnia and Herzegovina......... P. (Proleonhardella) weiratheri (Reitter, 1913)</p> <p>9. A1/A2 M 0.80. Mesosternal carina almost right-angled. Elytral lateral margins below humeral angles clearly sinuate. Maximum width of elytra prior to middle. Cavernicolous, village of Podromanija, near the town of Sokolac, eastern Bosnia and Herzegovina......................................................................................................................................... P. (Proleonhardella) neumanni (Apfelbeck, 1901)</p> <p>– A1/A2 M 0.76. Mesosternal carina obtuse-angled. Elytral lateral margins below humeral angles sub-parallel. Maximum width of elytra slightly after basal third. Cavernicolous, Mt Tara, western Serbia.................................................... P. (Proleonhardella) tarensis Ćurčić &amp; Pavićević sp. nov.</p></div> 	http://treatment.plazi.org/id/F2440A25FF9A4B2FDB82BCD9FAB2FCFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ćurčić, Srećko;Pavićević, Dragan;Vesović, Nikola;Vrbica, Maja;Kuraica, Miloš;Marković, Đorđe;Petković, Matija;Lazović, Vladimir;Pantelić, Dejan;Bosco, Fabrizio	Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan, Bosco, Fabrizio (2021): On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia. European Journal of Taxonomy 782 (1): 55-81, DOI: https://doi.org/10.5852/ejt.2021.782.1589, URL: http://dx.doi.org/10.5852/ejt.2021.782.1589
