taxonID	type	description	language	source
024FDF7CFFAAE41CFF7B55B1FE1BFF24.taxon	materials_examined	Type species: Kribiodosis fasciata Kieffer, 1921 [= Kribiodosis clavigera Kieffer, 1921] Other included species: Kribiodosis cantonensis Tang sp. n. These two species conformed largely to the generic diagnosis of Freeman (1958), with the following emendations.	en	Han, Wu, Liu, Jie, Luo, Yifan, Tang, Hongqu (2021): No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses. Zootaxa 5072 (6): 560-574, DOI: 10.11646/zootaxa.5072.6.4
024FDF7CFFAAE41CFF7B55B1FE1BFF24.taxon	description	Male. Coloration. Thorax generally brown in color, with variable extent and depth of pigment, scutal vittae dark or dark brown. Wing slightly brown, membrane weakly punctate. Leg brown or dark brown, with pale bands at the apices of all femorae and tibiae. Abdomen generally yellow, with banded intersegmental conjunctives on TI-VII. Segment VIII and IX usually wholly dark brown, gonocoxite and gonostylus pale yellow. Head. Antenna with 13 segments (if counting as 12 - segmented, then the boundary of last segmentation is indistinct). Frontal tubercle absent. Palp segment normal, the terminal segment usually no longer than the preterminal segment. Thorax. With scutal tubercle (K. cantonensis sp. n.) or without (K. clavigera) a tubercle. Acrostichals absent, dorsocentrals and scutellars few (usually no more than five). Prealars and supraalars both absent. Humeral pits present. Legs. Fore tibia with slender round scale bearing thin and pointed spur. Tibial combs of mid and hind tibiae separate (K. clavigera) or fused (K. cantonensis sp. n.); if separated, with only one hooked long spur in the smaller comb. Middle section of fourth tarsus of male fore leg with three dark brown pseudospurs, longer than the diameter of fourth tarsus, projecting inwardly. Pulvilli distinct. Wing. Membrane without setae or microtrichia. Anal lobe absent. Costa ending abruptly at apex of R 4 + 5, proximal to wing apex; R 2 + 3 ending near proximal 1 / 4 – 1 / 5 between R 1 and R 4 + 5. FCu clearly distal to RM. R bare, R 1 and R 4 + 5 with a few setae. Squama bare. Abdomen. Segment VIII clearly tapered anteriorly. Hypopygium. Anal tergite bands strong, meeting in median area. T IX with 1 – 2 median anal setae and 4 – 8 marginal setae on each side. Anal point short, parallel-sided, distally rounded. Superior volsella thin triangular, with narrow apex, inner margin with 2 – 3 basal setae. Inferior volsella setose, apex with 10 – 12 strong setae, extending beyond apex of anal point. Gonostylus relatively short, with a slightly triangular apex, inner margin with 6 – 8 nearly equally spaced long setae. Female. Coloration. As male except wing somewhat smoky, and abdominal segments IV – VIII nearly wholly dark brown. Head. Antenna with 4 or 5 segments. Frontal tubercle absent. Thorax and wing as male. Legs without pseudospurs on the fore tarsomere 4 on fore leg. Color as male. Genitalia. S VIII with 14 – 16 setae arising from the large pale pits on each side. Gonocoxapodeme straight, fused medially. Notum distinctly longer than the rami. Dorsomesal lobe prominent, expanded posterior medially; ventrolateral lobe absent or reduced into a narrow strip. Apodeme lobe lamellar, with slightly expanded apex. Labium apparently bare. Postgenital plate triangular. T IX with setae biserial, the upper row with 2 setae, and the lower with 6 – 8 setae.	en	Han, Wu, Liu, Jie, Luo, Yifan, Tang, Hongqu (2021): No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses. Zootaxa 5072 (6): 560-574, DOI: 10.11646/zootaxa.5072.6.4
024FDF7CFFADE411FF7B5625FE08FE78.taxon	materials_examined	Material examined. Holotype, male. CHINA: Guangdong Province, Guangzhou City, Conghua District, Lyutian Town, Dongkeng Reservoir, 23 ° 53.287 ′ N, 113 ° 55.069 ′ E, alt., 388 m, 23. iii. 2015 (emerged 01. iv. 2015), H. Q. Tang; paratype, 2 males, 22. iv. 2020; 1 female, 27. vi. 2020, as previous, light trap, H. Q. Tang.	en	Han, Wu, Liu, Jie, Luo, Yifan, Tang, Hongqu (2021): No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses. Zootaxa 5072 (6): 560-574, DOI: 10.11646/zootaxa.5072.6.4
024FDF7CFFADE411FF7B5625FE08FE78.taxon	etymology	Etymology. From the type locality, Canton, now Guangzhou. Male (n = 2 – 3) (Fig. 2 – 4) Total length 3.14 – 3.63 mm. Wing length 1.55 – 1.63 mm. Coloration (Figs. 2 – 3). Thorax generally brown, with light brown humeral area, scutellum, pleurae and preepisternum. Scutal vittae dark brown. Leg generally brown except pale connection areas in femur-tibia knees and tibia-first tarsus joints in all legs (Fig. 3 A, C – D). Tarsomere 4 in fore legs with three dark pseudospurs (Figs. 3 B, 4 B). Abdomen tergite predominately yellow, with brown conjunctive from T I – VII. T VIII and T IX nearly completely brown. T III with a pair of brown spots subapically (Fig. 2 D). Gonocoxite and gonostylus yellow (Figs. 2 E – F). Head. Antennal flagellomere 1 – 12, 555 μm; flagellomere 13, 600 μm, apex with two long setae (Fig. 2 A), AR 1.08. Palpomere lengths (in μm): 30 – 35; 35; 120 – 125; 105 – 120; 200. Temporals 8 – 12. Clypeus with 9 – 10 setae. Thorax (Figs. 2 C, 4 C). Antepronotals and acrostichals absent, scutum with a prominent scutal tubercle, 15 – 20 μm high; dorsocentrals 3 – 4, usually 1 above the humeral area, 1 below the scutal tubercle, and 2 posterior, 2 – 3 tiny pale pits usually occurred in the middle section. Prealars and supraalars both absent, scutellum with 2 setae. Tiny trans-oval remnants of humeral pits are present. Wing (Fig. 2 B). VR 1.29 – 1.31. Surface without macrochaetae. R without seta; R 1 with 4 – 5 setae. R 2 + 3 almost fused with R 1 in the end; R 4 + 5 with 2 – 3 setae distally. Squama bare. Legs (Fig. 3). Total length of fore leg 5.10 mm, about 6.5 times as long as the thorax height. Fore tibia with a slender, offset spur, 15 – 20 μm long (Fig. 4 A). Mid and hind tibia with one fused comb, bearing a distal hooked spur, 70 – 80 μm long. LR 1 2.48; LR 2 0.91; LR 3 1.21. BV 1 2.15; BV 2 2.85; BV 3 2.48. SV 1 1.24; SV 2 2.59; SV 3 1.85. BR 1 3.0; BR 2 2.4; BR 3 2.8 (n = 1). Hypopygium. (Figs. 2 E – F; 4 D – E). Anal tergite bands well-developed, meeting in the middle, with 1 – 2 anal median setae. Distal margin with 6 – 8 setae at each side. Gonocoxite 90 – 105 μm long, with a longitudinal row of five long setae in the medio-ventral section. Gonostylus 88 – 98 μm long, inner margin with 6 – 8 evenly distributed long setae. Superior volsella (Fig. 4 F) base without microtrichia, bearing 2 – 3 inner setae from the tubercle base, distal digitus long and slender, distal gradually attenuated. Inferior volsella slightly bullous distally, with 12 – 14 setae. HR 1.03 – 1.07; HV 3.59 – 3.92. Female (n = 1) (Fig. 5) Total length 2.33 mm. Wing length 1.85 mm. Coloration (Fig. 5 A). Similar to male except the abdomen with brown patch, more extensive in anterior 1 / 5 – 1 / 3 of each segment. Tarsomere 4 of fore legs without dark brown pseudospurs. Head. Eyes bare, with clear dorsal extension, separated by 1.3 times their terminal width. Antenna with five flagellomeres (Fig. 5 B), each length (in μm): 150, 105, 110, 65, 130; AR 0.31. Palpomere lengths (in μm): 40; 30; 120; 100; N / A (absent). Temporals 6. Clypeus with 16 setae. Thorax. Antepronotals and acrostichals both absent, scutal tubercle present; dorsocentrals 4, usually 2 above the humeral area, 1 below the scutal tubercle, and 1 posterior, 2 – 3 tiny pale pits usually in the middle and posterior section. Prealars and supraalars both absent, scutellum 2. Larger trans-oval remnants of humeral pits present, 30 μm long and 15 μm wide. Wing. VR 1.30. Surface without macrochaetae. R without seta, R 1 with 9 setae; R 2 + 3 indistinct, almost fused with R 1 all through the length; R 4 + 5 with 5 – 6 setae distally. Squama bare. Legs. Total length of fore leg 6.03 mm, about 8.0 times as long as the thorax height. Fore tibia with a slender spur, 50 μm long. Mid and hind tibia with two fused combs, with a distal – hooked spur, 70 – 75 μm long. LR 1 3.08; LR 2 1.0; LR 3 no data. BV 1 2.05; BV 2 3.0; BV 3 no data. SV 1 1.03; SV 2 2.42; SV 3 no data. Bristles of legs unmeasurable. Genitalia (Fig. 5 C). S VIII without distinct oval depression, with 16 – 18 setae. Coxosternapodeme V-shaped, fused in the middle. Notum 150 μm long, about three times as long as the rami. Lateral sternite IX with three setae. Dorsomesal lobe prominent, expanded post-medially; ventrolateral lobe reduced into a small strip, hidden below the well-developed dorsomesal lobe. Apodeme lobe without microtrichia. Two spermatheca oval, 70 μm in the long axis. Postgenital plate triangular shaped. Cercus ship-shaped, 80 μm long. Pupa. Possibly under the description of Zavreliella shidai Cao & Tang, 2017 in error (see discussion). Larva. Unknown.	en	Han, Wu, Liu, Jie, Luo, Yifan, Tang, Hongqu (2021): No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses. Zootaxa 5072 (6): 560-574, DOI: 10.11646/zootaxa.5072.6.4
