taxonID	type	description	language	source
79268507FFCCFFA7CBE5D6E3FEA7FE0D.taxon	diagnosis	Diagnosis (Figs. 1 – 13). Body 13.5 – 21.0 mm long, dark-brown to black, ovate, low convex, head unarmed, male pronotum with weak node on anterior edge. Maxilla and galea reduced to small conical pieces, unarmed, terminated by pencils of sensory setae; mentum flat, ligula with apex rounded or truncate, palps globular and short; mandibles obscured by clypeus, external margin rounded. Antennae with 9 or 10 antennomeres, club with 3 lamellae, shorter than length of shaft. Clypeus subrectangular; clypeofrontal suture carinate and transverse to posteriorly arcuate. Pronotal anterior margin with weak node in male, absent to slight swelling in female, in front of a small flat area, lateral margins unguttered. Elytra with lateral margin unguttered, epipleural plane in anterior half vertical. Protarsomere 5 longer than protarsomere 1; proclaws of males asymmetrical; metafemur of male slender, of female broad with convex anterior margin; metatibia of female stout with apex dilated. Parameres with internal margin untoothed, external margin arcuate, glabrous.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCCFFA7CBE5D6E3FEA7FE0D.taxon	discussion	Remarks. In his catalogue, Krajcik (2005) listed Enracius in the tribe Oryctini; it should be in the Pentodontini.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCFFFA1CBE5D1DBFCE0FE0D.taxon	materials_examined	Type series. Holotype male (by original designation): “ Western Australia, 40 km W of Caiguna 125 ° 05´E, 32 ° 20´S [32.33 ° S, 125.08 ° E], 30. IX. 1978 ”; in “ coll. S. Gaudaire-Thore ” [label data taken from Dechambre 1999]. Paratype (Figs. 1 – 4). 1 ♂, “ 40 km W of Caiguna, W. A. 125 ° 05´E, 32 ° 20´S, 30 Sep. 1978 M. S. & B. J. Moulds ” (white label) | “ new genus, P. B. Carne det. 1983 ” (white label) | “ Enracius n. gen. ♂ longipes n. sp. mihi PARATYPE R. - P. Dechambre det. 1999 ” (white label, PARATYPE written in red) | mouthparts glued to a piece of card; in FD (images examined). Other material examined. AUSTRALIA, WESTERN AUSTRALIA: 1 ♂, same data as paratype (ANIC); 1 ♀, “ 12 km e [ast] of Eucla West. Aust. [31.64 ° S, 129.00 ° E Western Australia / South Australia border] 18. Feb. 2011 | Enracius longipes | PMH Coll # Dyn 0422 ” (PMH); 2 ♀, “ Possibly ♀ of Orthocavonus occidentalis P. B. Carne det. 1981 | W. A. M Reg. No 78 / 68 or 78 / 69 | outside N 59 or Horseshoe Cave N. E. of Madura [31.68 ° S, 127.43 ° E] W. Australia 18 – 28. iii. 1970 M. Archer et al. ” (WAM); 1 ♀, “ 10 N Rawlinna [31.00 ° S, 125.33 ° E] WA Jan 1969 J. Bywater ” (ANIC).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCFFFA1CBE5D1DBFCE0FE0D.taxon	description	Description of female (Figs. 5 – 7). Body 13.7 – 17.5 mm long, pronotum 6.1 – 7.2 mm wide at widest point, elytra 7.8 – 9.6 mm wide (n = 3); dorsal surface shiny, head, pronotum, scutellum and pygidium brown, ventral surfaces and legs brown with black suffusions, elytra dark brown to black with suture brown, clypeus with pair of black paramedian maculae. Maxillary palpomeres 1 – 3 subequal in length, palpomere 4 longest and subequal to palpomeres 2 and 3 combined, tear-drop shaped and 2.5 times longer than greatest width and bearing shallow, oval sensorium on basal half; mandibles with lateral margin bearing stout setae; mentum narrow, bearing long setae over entire surface. Antenna with 9 antennomeres, club with antennomeres 8 and 9 1.5 times width of antennomere 7 and longer than antennomeres 2 – 6 combined, scape and pedicel bearing long setae, inner margin of club setose. Clypeus transverse almost trapezoidal with anterior angles rounded, anterior margin emarginate, anterior face truncate, vertical and glabrous, surface rugose, setose; clypeofrontal ridge linear, slightly raised. Frons rugose; ocular canthi angulate to clypeus, dorsal surface rugulose, anterior margin bearing stout setae. Pronotum widest basally, subparallel on basal half, angulate medially, linear and tapering to acute anterolateral angles, broadly recurved linear, transverse anterior margin, anterior margin membranous, lateral and anterior ridges complete, basal ridge obsolete across medial two-thirds, disc evenly convex, bearing fine punctures, becoming rugulose along lateral and anterior margins, anterior margin weakly raised medially, preceding small flat area and narrow impunctate midline. Scutellum sparsely micropunctate on disc. Elytral striae weakly impressed, linear-punctate, intervals abundantly micropunctate to rugulose, laterally and apical declivity coarsely rugulose, suture punctate-striate, epipleura nonemarginate, continuous and bearing lateral setae to apical calli; pygidium almost flat, glabrous, disc punctate, becoming denser and becoming finely rugulose at basolateral angles, apical ridge setose. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum finely rugulose, bearing dense, long, light-brown setae. Abdominal sternites at midlength bearing a row of sparse setae across entire width, suture between sternites 6 and 7 wide and yellow, sternite 7 apically bearing a row of long setae and sparsely on disc. Protibia tridentate, medial and basal denticles closest; meso- and metatibia bicarinate (basal carina ill-defined in smaller specimens), apical cilia long, sharp, widely set; metafemur broad with anterior margin broadly arcuate; metatibia with concave dorsal margin apically and post-medial carina, inner surface broad and flat, metatibial inner spur 1.5 times length of outer spur, inner parallel and curved, outer spur narrow and straight, apices rounded; metatarsomeres combined 1.5 times length of metatibia. Male (Figs. 1 – 4). Description by Dechambre (1999). Males differ from females by the following characters: bicolorous brown and black (male dark brown); protarsal claws simple (male internal claw larger and thicker than external claw); anterior margin of pronotum non-tuberculate (male bears a feeble tubercle); metatarsomeres 1.5 times length of tibia (male metatarsomeres 2.0 times length of tibia).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCFFFA1CBE5D1DBFCE0FE0D.taxon	discussion	Remarks. Before his death in 1989, Carne passed some dynastines that he thought were undescribed to Dechambre (Fabien Dupuis, personal communication). Dechambre (1999) later recognised Carne in naming the new genus as Enracius. Both male specimens that make up the type series of Enracius longipes were listed by Dechambre as housed in the “ coll [ection] S. Gaudaire-Thore ” (Dechambre 1999), which was at Étigny (Yonne), France (Dechambre 2003 a, 2003 b) and had specimens named from it by Endrődi (1980) (including Aspidolea gaudairethorei Endrődi, 1980). The holotype at least should have been returned to ANIC but it is not there (https: // www. csiro. au / en / Research / Collections / ANIC / Collection-Resources / ANIC-primary _ types, accessed 19 August 2021). A third specimen collected at the same time was retained in ANIC and is, presumably, the one illustrated by Weir et al. (2019; Plate 54 I); it was loaned to and imaged by PMH. The ‘ Gaudaire-Thore collection’ was apparently a euphemism for part of Dechambre’s collection (Fabien Dupuis, Olivier Montreuil and Antoine Mantilleri, personal communications) at his family home at Étigny. After Dechambre’s death, most of his type specimens were transferred to the MNHN, as was his wish, but the Enracius longipes holotype was not part of that donation. What was left of the collection was either destroyed by dermestids or sold, and its current location (s) is unknown. However, the paratype was gifted to and is now housed in FD (Fabien Dupuis, personal communication). Given that we can unambiguously associate the name with a known form, there is no need for a designation of a neotype (International Commission on Zoological Nomenclature 1999, Article 75.3). The image of the holotype in Dechambre (1999, 2005) is obviously of a different specimen than the paratype or the male in ANIC (different arrangement of legs).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCFFFA1CBE5D1DBFCE0FE0D.taxon	distribution	Distribution and habitat. Known only from the southeastern area of Western Australia (Fig. 14) (Köppen-Geiger climate of BSk, cold semi-arid; Beck et al. 2018). The type series would have been captured at light (M. S. Moulds. personal communication), and the locality is open grassland with groups of 1 – 6 small Eucalyptus (Myrtaceae) and some sparse Acacia (Fabaceae) over limestone.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC8FFA2CBE5D0B3FBAFFEBD.taxon	materials_examined	Type series. Holotype female (Figs. 8 – 13). AUSTRALIA, WESTERN AUSTRALIA: “ Fitzgerald Riv. Res. [Fitzgerald River National Park, 34 ° S, 119.7 ° E] W. Aust. 18. xii. 1970 K. T. Richards | 119 | Agriculture (Dept) Western Australia 37851 | Dynastine genus? P. B. Carne det. 1980 ”; left antennomeres 2 – 10 on card; in WADA. Paratype. AUSTRALIA, WESTERN AUSTRALIA: 1 ♀: “ 25 miles [40 km] sth Mt. Ragged [Cape Arid National Park, 33.8 ° S, 123.5 ° E] W. Aust. 10. xii. 1974 K. T. Richards | 165 | Agriculture (Dept) Western Australia 37850 | Dynastine genus? P. B. Carne det. 1980 ”; in WADA.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC8FFA2CBE5D0B3FBAFFEBD.taxon	description	Description of holotype female (Figs. 8 – 12). Protarsomeres 2 – 5, left mesotarsomeres 3 – 5, right mesotarsomeres 4 – 5, right metatarsomeres 4 – 5 missing. Body 21.2 mm long, pronotum 9.0 mm wide at widest point, elytra 11.3 mm wide; form convex depressed, robust; dorsal surface matt, head, pronotum, elytra and pygidium black; scutellum, elytral suture, sternum, abdominal ventrites and legs dark brown to black. Maxillary palp with palpomeres 1 and 3 subequal in length, palpomere 2 longer but shorter than palpomere 4, palpomere 4 about 3 times longer than greatest width and bearing small, elongate, shallow sensorium on basal half; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, glabrous medially. Antenna with 10 antennomeres, club subequal in length to antennomeres 2 – 7 combined, scape bearing long setae, antennomeres 2 – 7 glabrous. Clypeus transverse, almost trapezoidal, anterior angles rounded, anteriorly emarginate, truncate face vertical and glabrous, surface rugulose; clypeofrontal ridge posteriorly arcuate, weakly raised. Frons rugulose; ocular canthi sub-continuous with line of clypeus, dorsal surface rugulose, anterior margin bearing row of stout setae. Pronotum widest basally, subparallel on basal half, angulate medially, linear and tapering to acute anterior angles, recurved to linear transverse anterior margin, anterior margin membranous, basal, lateral and anterior ridges complete; disc evenly convex bearing sparse micropunctures becoming larger and denser adjacent to margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals almost impunctate, apical declivity rugulose, suture punctate-striate, epipleura nonemarginate, continuous to suture, lateral setae restricted to base adjacent to mesepimeron. Pygidium moderately convex, glabrous, disc sparsely micropunctate, denser at lateral margins, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, light-brown setae. Abdominal sternites with a line of sparse setae laterally adjacent to posterior margins, suture between sternites 6 and 7 wide and glossy yellow, sternite 7 bearing narrow sulcus laterally, apical margin bearing row of long setae. Protibia tridentate, denticles equidistant; meso- and metatibia unicarinate, apical cilia short, blunt and widely set; metafemur stout, anterior margin very convex; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both broad, straight and widest pre-apically and apex at oblique angle; metatarsomeres combined longer than metatibia. Male. Unknown. Variation. Paratype length 19.1 mm, pronotal width 8.4 mm, elytral width 10.3 mm; elytra dark brown, pronotum laterally suffused brown; left meso- and metatibia present and longer than tibiae.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC8FFA2CBE5D0B3FBAFFEBD.taxon	diagnosis	Differential diagnosis. Females of Enracius richardsi can be easily differentiated from females of Enracius longipes by the following characters (E. longipes characters in brackets): body> 19 mm long (13.5 – 17.5 mm); clypeofrontal ridge posteriorly arcuate (linear); ocular canthi subcontinuous with line of clypeus (angular to clypeus); maxillary palpomeres 1 – 3 of unequal length (subequal); mentum broader; pronotum micropunctate and evenly arcuate (bearing fine punctures becoming rugulose along lateral and anterior margins, anterior margin weakly raised medially, preceding small flat area and narrow impunctate midline); elytra stria deep (weakly impressed), intervals almost impunctate (abundantly micropunctate to rugulose), epipleural lateral setae basal (lateral setae to apical calli); pygidium moderately convex (almost flat); abdominal ventrite 7 bearing pre-apical sulcus (absent); meso-and metatibial cilia short and blunt (apical cilia long, sharp); apical spurs short and stout (metatibial inner spur 1.5 times length of outer spur).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC8FFA2CBE5D0B3FBAFFEBD.taxon	etymology	Etymology. Named for WADA entomologist Kevin T. Richards (1932 – 2009) who completed a Master’s degree on potential pests in the Ord Irrigation Scheme in the Kimberley Region of Western Australia. Later in Perth, he was curator of the WADA Entomology collection of which he made many contributions. He collected both specimens of this new species.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC8FFA2CBE5D0B3FBAFFEBD.taxon	distribution	Distribution and habitat. Known only from the southern coast of Western Australia (Fig. 14) (Köppen-Geiger climate of Csb, warm-summer Mediterranean). Both specimens have incomplete tarsomeres on most legs, and the holotype was covered in a dense layer of fine undetermined matter and moth scales along margins. The collection method is not noted on their labels, although the moth scales indicate a light trap.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFCAFFADCBE5D6A9FD6BFEC5.taxon	diagnosis	Diagnosis (Figs. 15 – 49). Body 11.5 – 20.8 mm long, pronotum and elytra reddish-brown to black, legs yellowishreddish brown (almost white in live specimens); form ovate, low convex, head with recurved frontoclypeal suture often terminating in a tubercle, pronotum unarmed. Maxilla and galea reduced to small conical pieces, unarmed, terminated by pencils of sensory setae; mentum flat, ligula apex rounded, palps slender, elongate and conspicuous; mandibles obscured by clypeus, external margin rounded. Antennae with 10 antennomeres, club with 3 lamellae and shorter than length of shaft. Clypeus subrectangular; clypeofrontal suture posteriorly angulate. Pronotum with anterior margin unarmed, surface evenly convex, lateral margins guttered. Elytra with lateral margins guttered, epipleural plane in anterior half horizontal. Males with all tarsi elongate, claws enlarged, robust and slightly curved; protarsomere 5 longer than protarsomere 1; proclaws of male symmetrical; metafemur of male and female long, slender, protruding beyond elytra; metatibia of female long and slender. Parameres with internal margins untoothed, external margins preapically angulate, apices setose.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D0B3FA01FB74.taxon	materials_examined	Type series. Lectotype male of A. anthracina (designated by Allsopp 2000: 3). “ Moreton Bay, Queensland ” [about 27.5 ° S, 153.3 ° E]; in NHML (examined). Lectotype male of A. flavipes (designated by Allsopp 2000: 3). “ Rockhampton, Queensland ” [23.37 ° S, 150.53 ° E]; in NHML (examined). Holotype male of A. geotrupina (by monotypy). “ Queensland ”; in NHML (examined). Material additional to Allsopp (2000) examined. AUSTRALIA: QUEENSLAND: 1 ♂, 2 ♀, “ Barakula S. F. Chinchilla Qld. Aust. 29. Dec. 2006 B. Howton | PMH Coll # Dyn 0426, 0427, 0429 ” (PMH); 1 ♂, “ Barakula S. F. Chinchilla Qld. Aust. 30. Dec. 2006 B. Howton | PMH Coll # Dyn 0428 ” (PMH); 2 ♂, 1 ♀, “ Barakula S. F. Qld. Aust. 3. Dec- 19. Dec. 2012 C. H. Moeseneder | F. I. T. | PMH Coll # Dyn 2066, 2067 or 2018 ” (PMH); 2 ♀, “ Dunmore S. F. Chinchilla Q. Aust. 29. Dec. 2006 B. Howton | PMH Coll # Dyn 0423 or 0425 ” (PMH); 1 ♀, “ W. Mt. Garnet N. Qld. Aust. 20. Feb. 2010 J. Hasenpusch | PMH Coll # Dyn 1609 ” (PMH); 1 ♀, “ 70 km N. Mackay Midge Pt. Q 27 / 12 / 1990 M. (H). Groth | PMH Coll # Dyn 0424 ” (PMH); 2 ♂, “ Yarraman Qld. Aust. 19. Jan. 2017 R. Zeitek | PMH Coll # Dyn 1973 or 1974 ” (PMH).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D0B3FA01FB74.taxon	diagnosis	Diagnosis (Figs. 15 – 22, 41, 44 – 45). Description by Allsopp (2000). Body 11.3 – 17.0 mm long; dorsal surface shiny; distance between outer edges of canthi 1.8 – 1.9 times interocular distance; ocular canthus with lateral margin weakly arcuate then posteriorly angulate; pygidium with a few long setae at sides on anterior third; abdominal sternite 6 with posterior margin normal; metacoxal posterior margin broadly arcuate from midlength through posterolateral angle; metafemoral posterior margin weakly concave; Parameres parallel to truncate apices, apices curved slightly outwards, lateral tooth angulate and prominent	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D0B3FA01FB74.taxon	distribution	Distribution (Fig. 51). Additional material all came from within the known range of A. anthracina from eastern Queensland, through eastern New South Wales into inland Victoria. We also re-examined a male and a female in MV labelled “ Nat. Mus. Victoria W. Australia | French coll. 6.1.08 ” and, like Allsopp (2000), consider them to be A. anthracina and not the species described below. The first label is typeset and was presumably added by MV after it acquired the material sometime before 1932 (Musgrave 1932).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D53FFBEAF975.taxon	materials_examined	Type series. Holotype male (by original designation). “ Mt Spurgeon, Queensland ” [16.45 ° S, 145.18 ° E], registration number QMT 62780; in QM (examined).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D53FFBEAF975.taxon	diagnosis	Diagnosis (Figs. 23 – 27, 43, 48 – 49). Description by Allsopp (2000). Body 19.2 – 20.8 mm long; dorsal surface sericeous; distance between outer edges of canthi 1.6 times interocular distance; ocular canthus weakly arcuate then posteriorly angulate; pygidium with apical ridge setose; abdominal sternite 6 with posterior margin normal; metacoxa posterior margin sublinear then arcuate through posterolateral angle; metafemur with posterior margin sublinear; parameres tapering to acute apices, apices curved slightly outwards, with lateral tooth angulate and prominent.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFAECBE5D53FFBEAF975.taxon	distribution	Distribution (Fig. 51). Known only from high-elevation wet sclerophyll forest along the western edges of rainforest plateaus in the Wet Tropics area of northeastern Queensland (Allsopp 2000).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFB4CBE5D703FEF0FC99.taxon	materials_examined	Type series. Holotype male. AUSTRALIA, WESTERN AUSTRALIA: “ West Angelas at 23.06 ° S, 118.46 ° E, 29 June 1984, R. P. McMillan | in litter | Anomalomorpha ” | aedeagus on card; in WAM. Allotype. AUSTRALIA, WESTERN AUSTRALIA: 1 ♀: same data as holotype; in WAM.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFB4CBE5D703FEF0FC99.taxon	description	Description of holotype male (Figs. 28 – 34, 42, 46 – 47). Left mesotarsomeres 4 – 5, right mesotarsomeres 5, left metatarsomere 5, right metatarsomeres 4 – 5, left and right maxillary palpomere 4 missing, only antennal scape and pedicel present. Body 16.6 mm long, pronotum 8.4 mm wide at widest point, elytra 10.5 mm wide; form convex depressed; dorsal surface matt, head, pronotum, elytra and pygidium black; scutellum, elytral suture, margin, disc, sternum, abdominal ventrites and legs dark brown to black. Maxillary palp with palpomeres 1 and 3 subequal in length; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, sparsely setose medially. Antenna with scape bearing long setae, pedicel with sparse short setae. Clypeus transverse, almost trapezoidal, distance between outer edges of canthi 2.0 times interocular distance, anterior angles rounded, anteriorly emarginate, truncate face vertical with shallow punctures, glabrous, dorsal surface rugulose; clypeofrontal ridge posteriorly angulate, weakly raised leading to median node. Frons rugulose; ocular canthi angulate to line of clypeus then continuously arcuate, broad, dorsal surface rugulose, anterior margin bearing row of long stout setae. Pronotum widest post midlength, divergent from base and arcuate to acute anterior angles, recurved to linear transverse anterior margin, anterior margin membranous; lateral and anterior ridges complete, obsolete across base; disc evenly convex, bearing sparse micropunctures becoming larger and denser adjacent to lateral margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals impunctate, lateral and apical declivity rugulose, suture punctate-striate, epipleura non-emarginate, continuous to pygidium, lateral setae sparse and ending adjacent to pygidium. Pygidium moderately convex, disc sparsely micropunctate, denser at lateral margins where bearing sparse, long setae, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, lightbrown setae. Abdominal sternites with a line of continuous setae across midlength, sternite 7 apical margin bearing row of sparse, long setae. Protibia tridentate, denticles equidistant; meso- and metatibia bicarinate, apical cilia long, sharp, with interspersed shorter cilia; metafemur broad, anterior margin convex, posterior margin linear; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both narrow, straight and tapering to acute apex; metatarsomeres 1 – 3 combined about 0.8 times length of metatibia. Metacoxal posterior margin angulate at midlength, then linear to narrow arcuate posterolateral angle. Genitalia (in posterior view) with parameres laterally along basal half shallowly concave, midlength with low, rounded, obtusely angled process, apices tapering to acute apex. Description of allotype female (Figs. 35 – 40). Left protarsomeres 2 – 5, right protarsomeres 3 – 5, left mesotarsomeres 3 – 5, right meso-leg, metatarsomeres 3 – 5, right labial and maxillary palps missing, only antennal scape and pedicel present. Body 17.4 mm long, pronotum 8.1 mm wide at widest point, elytra 10.1 mm wide; form convex depressed; dorsal surface matt, head, pronotum, elytra and scutellum black; elytral suture, margins, sternum, abdominal ventrites, pygidium and legs brown to dark brown. Maxillary palpomeres 1 and 3 subequal in length, palpomere 2 longer but shorter than palpomere 4, palpomere 4 about 3 times longer than greatest width and bearing trace of narrow, elongate sensorium on basal two-thirds; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, sparsely setose medially (most abraded). Antennal scape bearing long setae, pedicel with sparse, short setae. Clypeus transverse, almost trapezoidal, distance between outer edges of canthi 1.9 times interocular distance, anterior angles rounded, anteriorly emarginate, truncate face vertical with shallow punctures, glabrous, dorsal surface rugulose; clypeofrontal ridge posteriorly angulate, weakly raised leading to medial node. Frons rugulose; ocular canthi angulate to line of clypeus then continuously arcuate, broad, dorsal surface rugulose, anterior margin bearing row of long, stout setae (most abraded). Pronotum widest post midlength, divergent from base and evenly arcuate to acute anterior angles (almost right-angle), recurved to linear transverse anterior margin, anterior margin membranous laterally; lateral and anterior ridges complete, obsolete across base; disc evenly convex, bearing sparse micropunctures becoming larger and denser adjacent to lateral margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals impunctate, lateral and apical declivity rugulose, suture punctate-striate, epipleura non-emarginate, continuous to pygidium, lateral setae sparse, ending adjacent to pygidium. Pygidium moderately flat, disc sparsely micropunctate, denser at lateral margins where bearing sparse, long setae, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, light-brown setae. Abdominal sternites with a line of continuous setae adjacent to posterior margin, sternite 6 posteriorly with membranous, transverse groove, sternite 7 apical margin bearing row of sparse, long setae. Protibia tridentate, denticles non-equidistant with basal denticle closer to medial than medial is to apical; meso- and metatibia bicarinate, apical cilia long, sharp with interspersed shorter cilia; metafemur broad, anterior margin convex, posterior margin linear; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both moderately broad, widest preapically, inner spur straight, outer spur slightly curved; metatarsomeres 1 – 2 combined about 0.5 times length metatibia. Metacoxal posterior margin sublinear to narrow arcuate posterolateral angle. Sexual differential diagnosis. Female differs from male by the following combination of characters: protibial denticles non-equidistant, metatibial spurs broader, metacoxa with posterior margin sublinear; abdominal sternites with setae along posterior margin, pygidium quite flat.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFB4CBE5D703FEF0FC99.taxon	diagnosis	Differential diagnosis of males (female A. monteithi unknown). Given the paucity and condition of specimens, we did not attempt cleaning. Anomalomorpha pilbara can be distinguished from its congeners by the following combination of characters: • Ocular canthus with lateral margin continuously arcuate (weakly arcuate then posteriorly angulate in A. anthracina and A. monteithi); • Distance between outer edges of canthi 1.9 times interocular distance (1.8 – 1.9 times in A. anthracina and 1.6 - 1.65 times in A. monteithi); • Pygidium with apical ridge glabrous (setose in A. anthracina and A. monteithi); • Metacoxa with posterior margin angulate at midlength, then narrowly arcuate at posterolateral angle (broadly arcuate from midlength through posterolateral angle in A. anthracina; sublinear then arcuate through posterolateral angle in A. monteithi); • Metafemur with posterior margin linear (weakly concave in A. anthracina, sublinear in A. monteithi); • Parameres with lateral tooth obtusely rounded (angulate and prominent in A. anthracina and A. monteithi); • Parameres (posterior view) tapering to acute apices that are curved inwards (parallel to truncate apices in A. anthracina; tapering to acute apices in A. monteithi; apices of both curved slightly outwards).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFB4CBE5D703FEF0FC99.taxon	etymology	Etymology. Named for the area of the type locality that is noted for its red earth, as attached to the holotype; a noun in apposition. The Pilbara Aboriginal Language Centre Wangka Maya says that the name for the Pilbara region derives from the word bilybara, meaning “ dry ” in the Nyamal and Banyjima languages (Anonymous 2021).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFC6FFB4CBE5D703FEF0FC99.taxon	distribution	Distribution (Fig. 51). The type locality is in an area classified as Köppen-Geiger climate BSh (hot semi-arid) on the edge of a larger area of BWh (hot desert). It has been modified by mining, but the surrounding area (Fig. 50) is red, rocky ranges with multiple gorges and creek lines and covered with Eucalyptus / Corymbia (Myrtaceae) with an understorey predominately of Acacia over spinifex (Triodia; Poaceae) and buffel grass (Cenchrus ciliaris Linnaeus; Poaceae).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFDEFFB1CBE5D0B3FC12FEE1.taxon	type_taxon	Type species. Anomalomorpha giveni Carne, 1957, here designated.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFDEFFB1CBE5D0B3FC12FEE1.taxon	description	Description (Figs. 52 – 63). Body 12 – 14 mm long, reddish brown, ovate, low convex, head and pronotum unarmed. Maxilla and galea reduced to small conical pieces, unarmed, terminated by pencils of sensory setae; mentum flat, apex of ligula truncate, palps globular and short; mandibles obscured by clypeus, external margin rounded. Antennae with 9 antennomeres, club with 3 lamellae and shorter than length of shaft. Clypeus rounded, glabrous; clypeofrontal suture transverse. Pronotum with anterior edge entirely narrowly membranous, surface evenly convex and micropunctate, lateral margins unguttered. Elytra with lateral margin unguttered, striae vaguely impressed, epipleural plane in anterior half vertical. Female with penultimate abdominal ventrite with row of setae removed from posterior edge. Male tarsi elongate; protarsomere 5 longer than tarsomere 1; proclaws of males symmetrical; metafemur of males and females long, slender, protruding beyond elytra; metatibia of female stout with apex dilated. Parameres with internal margin untoothed, external margin arcuate, glabrous.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFDEFFB1CBE5D0B3FC12FEE1.taxon	etymology	Etymology. The name is a based on a reversal of Roger-Paul Dechambre’s family name reflecting his use of Enracius based on a reversal of Philip Carne’s name. It is masculine.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFD9FFB0CBE5D1B6FEC9FDB9.taxon	materials_examined	Type series. Holotype male (by original designation) (Figs. 52 – 57). “ 2 m. [miles] W. Sale Vic. [Victoria] [38.12 ° S, 147.07 ° E] Sept. 1949. P. B. Carne | Holotype ♂ Anomalomorpha giveni mihi P. B. Carne det. 1952. | 486 | 21032 | Type [red-ringed circular label] | HOLOTYPE ” [red label] | tarsus glued on card; in ANIC (examined). Allotype female (Figs. 58 – 63). “ 2 m. [miles] W. Sale Vic. [Victoria] [38.12 ° S, 147.07 ° E] Sept. 1949. P. B. Carne | Holotype ♂ Anomalomorpha giveni mihi P. B. Carne det. 1954 ALLOTYPE ♀ | 486 | Allo-Type [red-ringed circular label] | ALLOTYPE ” [yellow label]; in ANIC (examined). Paratype male. “ Para-type [yellow-ringed circular label] | Canberra 3 - 50 J. Shepherd | Paratype ♂ Anomalomorpha giveni mihi P. B. Carne det. 1952 ”; in QDAF (examined).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFD9FFB0CBE5D1B6FEC9FDB9.taxon	diagnosis	Diagnosis. Description by Allsopp (2000).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFD9FFB0CBE5D1B6FEC9FDB9.taxon	distribution	Distribution (Fig. 51). Southern New South Wales, Australian Capital Territory, and Victoria (Carne 1957; Allsopp 2000).	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFD8FFB3CBE5D247FEE3FD9D.taxon	distribution	Distribution. The genus contains three species (Weir et al. 2019): A. coulonii (Burmeister, 1847), A. canei Carne, 1957 (Figs. 64 – 65) and A. mellori Carne, 1957, that occur from southern Queensland to southeastern South Australia and northern Tasmania (Carne 1957, Map 15), with A. coulonii having established in the South Island of New Zealand (Stufkens & Farrell 1980). Adults of all three species were adequately described and keyed by Carne (1957), and Dechambre (2005) illustrated adults of A. canei and A. coulonii. McQuillan (1985) described the larva of A. coulonii.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
79268507FFD8FFB3CBE5D247FEE3FD9D.taxon	discussion	Remarks. Carne (1957) named A. canei for “ Miss H. M. Cane (Mrs. B. Cameron) ” and so should have used an - ae ending (International Commission on Zoological Nomenclature 1999, Article 31.1.2). Even though it was improperly formed, the original spelling must be preserved in line with Article 31.1.3 as it should not be corrected under Article 32.5.	en	Hutchinson, Paul M., Allsopp, Peter G. (2021): Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa 5072 (5): 439-462, DOI: 10.11646/zootaxa.5072.5.2
