taxonID	type	description	language	source
76711CAE00E15BA89DA4CBA67C3FF66F.taxon	description	Figures 1, 2, 3, 4, 5, 6, 7	en	Fuentes-Reines, Juan Manuel, Suarez-Morales, Eduardo, Silva-Briano, Marcelo (2021): A new species of Esola Edwards, 1891 (Crustacea, Copepoda, Harpacticoida, Laophontidae) from the Caribbean coast of Colombia. ZooKeys 1074: 1-15, DOI: http://dx.doi.org/10.3897/zookeys.1074.73030, URL: http://dx.doi.org/10.3897/zookeys.1074.73030
76711CAE00E15BA89DA4CBA67C3FF66F.taxon	etymology	Etymology. The species is named after Dr John B. Wells as a tribute for his longstanding, solid contributions to the taxonomic knowledge of harpacticoid copepods (Grieve et al. 2019; Huys 2021). It is a noun in apposition, gender masculine.	en	Fuentes-Reines, Juan Manuel, Suarez-Morales, Eduardo, Silva-Briano, Marcelo (2021): A new species of Esola Edwards, 1891 (Crustacea, Copepoda, Harpacticoida, Laophontidae) from the Caribbean coast of Colombia. ZooKeys 1074: 1-15, DOI: http://dx.doi.org/10.3897/zookeys.1074.73030, URL: http://dx.doi.org/10.3897/zookeys.1074.73030
76711CAE00E15BA89DA4CBA67C3FF66F.taxon	description	Description of female. Habitus (Figs 1 A, 5 B). Body roughly cylindrical in dorsal view, pro-some gradually tapering anteriorly. Total body length measured from anterior margin of rostrum to posterior margin of caudal rami ranging from 616 to 658 μm (average length = 640 μm, n = 8; holotype length = 658 μm). Rostrum triangular in lateral view, rounded in dorsal view (Figs 5 A, 6 C). Genital double-somite with dorsal suture, somite completely fused ventrally (Fig. 1 B). Cephalothorax slightly wider than free prosomites, with anterodorsal row of spinules. Cephalosome with dense pattern of small spinules (Figs 5 A, 6 C). Ventral surface of urosome smooth except for penultimate urosomite furnished with row of spinules along posterior margin. Urosomites (Figs 6 B, D, 7 A) with dense pattern of dorsal and lateral small spinules. Anal somite short, with rounded anal operculum furnished with 12 - 16 large spinules (Figs 5 C, 6 A). Caudal rami (Figs 1 D, 5 C, 6 A, B) subrectangular, length / width ratio = 2.6. Proximal half distinctly expanded, distal half narrow, concave. Outer margin furnished with spinules. Rami with 7 setal elements. Seta I small, spiniform. Seta II and III smooth and closely set, almost equally long, seta V about twice as long as seta IV, both lightly pinnate and with spinules at insertion point. Seta VI about 1 / 2 length of seta VII. Antennule (Figs 2 A, 5 D, E) 6 - segmented. First segment with rows of small spinules along outer and distal margins, second segment longest, cylindrical, about 3.3 times as long as wide; third segment slender, about 3.4 times as long as wide, armed with aesthetasc fused with 1 smooth seta). Acrothek set on apical pedestal. Segmental antennule armature as follows, s = setae, ae = aesthetascs: 1 (1 s) - 2 (8 s) - 3 (6 s) - 4 (2 + ae) - 5 (1 s) - 6 (9 + acrothek). Antenna (Figs 2 B, 5 F, G) 3 - segmented, comprising coxa, allobasis, free 1 - segmented ENP and 1 - segmented EXP. Coxa small, with row of spinules. Allobasis elongate, with pinnate abexopodal seta inserted near distal margin. EXP slender, about 5 times longer than wide, with 4 well-developed pinnate setae (2 lateral, 2 apical). ENP slightly longer than allobasis, subquadrate, 1.1 times as long as wide, with row of spinules on inner and outer margins; lateral armature arising from distal half, consisting of small, slender seta flanked by 2 pinnate spines. Subdistal part with thick spine; apical armature consisting of 4 setae. Mandible (Fig. 2 C) with gnathal blade narrow, bearing several multicuspidate teeth plus short pinnate seta. Palp slender 2 - segmented, first segment (basis) cylindrical, armed with single seta; second segment (endopod) subquadrate, with 3 subequally long apical setae. Maxillule (Fig. 2 D). Precoxal arthrite with 8 distal spines / setae, inner margin with single pinnate seta. Coxa with cylindrical endite carrying slender seta. Basal endite cylindrical, unfused, armed distally with 2 setae and 1 spine. Endopod incorporated in basis, forming small peduncle armed with single seta. Exopod 1 - segmented, with plumose apical seta. Maxilla (Fig. 2 E). Syncoxa with row of long slender spinules on outer margin plus rounded smooth protuberance. Three armed syncoxal endites (1 - 3 in Fig. 2 F) including small first endite, with single seta, second and third endites each armed with 2 stout elements. Allobasis transformed into strong, slightly curved, distally pinnate claw with single proximal slender seta. Endopod represented by 2 setae. Maxilliped (Figs 2 F, 5 H) slender, represented by cylindrical syncoxa with proximal patch of small spinules and short setules along outer margin, segment armed with 2 equally long distal plumose setae. Basis with rows of small, strong spinules along inner margin plus small spinule patches on medial and subdistal position of outer margin, basis about 1.4 times as long as syncoxa. ENP short, forming long, curved claw. Slender seta inserted at base. P 1 (Fig. 3 A) with dense cuticular ornamentation on precoxa, coxa, and basis. Basis with short pinnate seta on anterior surface; with outer basipodal seta and distal seta, surface furnished with spinules. EXP short, 2 - segmented, reaching proximal 1 / 4 of ENP 1 length. EXP 1 with 1 outer spine, EXP 2 with 2 outer spines and 1 outer geniculate seta, and 2 apical geniculate setae. ENP 1 about 8 times as long as wide, ENP 2 short, 1.6 times as long as wide, with long distal claw and slender accessory seta. Intercoxal sclerite narrow, smooth. P 2 (Fig. 3 B) Coxa furnished with long spinules along outer margin. Basis with short bipinnate outer spine. ENP 2 - segmented, shorter than EXP, almost reaching insertion of EXP 2 inner seta; ENP 1 with 1 inner seta, ENP 2 with 2 inner, 2 apical, and 1 outer seta. EXP 3 - segmented, segments with outer spinules; EXP 1 smooth, EXP 2 with inner seta, EXP 3 with 7 elements (3 outer spines, 2 apical setae, 2 inner elements). Intercoxal sclerite narrow, smooth. P 3 (Fig. 3 C) Coxa as in P 2. Basis with lightly plumose outer seta. ENP 2 - segmented, shorter than EXP, not reaching insertion of inner seta of P 3 EXP 2; ENP 1 with single inner seta, ENP 2 with 6 elements, 2 inner, 2 apical, and 2 outer setae. EXP 3 - segmented, outer margin of segments furnished with spinules; EXP 1 armed with outer spine, inner margin smooth; EXP 2 with inner seta; EXP 3 armed with 7 elements (3 outer spines, 2 apical setae, 2 inner setae). Intercoxal sclerite narrow, smooth. P 4 (Fig. 3 D) Coxa and basis as in P 3. ENP 2 - segmented, shorter than EXP, reaching tip of EXP 2; ENP 1 with short, slender inner seta arrowed in Fig. 3 D), ENP 2 with 5 elements (2 inner, 2 apical, 1 outer). EXP 3 - segmented, segments furnished with small spinules on outer margins (Fig. 6 F). EXP 1 lacking inner seta, EXP 2 with inner seta; EXP 3 with 7 elements (3 outer spines, 2 apical setae, 2 inner elements). Intercoxal sclerite narrow, smooth. Armature formula P 2 - P 4 P 5 (Figs 1 E, F, 7 A-D) slender, with EXP and baseoendopod covered with small spinules. Setophore armed with long seta. Endopodal lobe short, reaching insertion of outer seta of EXP, with 1 short and 1 long pinnate setae apically, plus 2 long inner setae. EXP elongate, about 6 times as long as wide, armed with 6 elements (each arrowed in Fig. 1 E, F).	en	Fuentes-Reines, Juan Manuel, Suarez-Morales, Eduardo, Silva-Briano, Marcelo (2021): A new species of Esola Edwards, 1891 (Crustacea, Copepoda, Harpacticoida, Laophontidae) from the Caribbean coast of Colombia. ZooKeys 1074: 1-15, DOI: http://dx.doi.org/10.3897/zookeys.1074.73030, URL: http://dx.doi.org/10.3897/zookeys.1074.73030
76711CAE00E15BA89DA4CBA67C3FF66F.taxon	description	Description of male. Habitus (Fig. 4 A) as in female except for urosome segmentation. Total body length ranging from 476 to 518 μm (average = 492 μm; n = 10; allotype = 476 μm). Antennule (Fig. 3 E) 7 - segmented, subchirocer. Geniculation between segments 5 and 6. Segment 1 with row of spinules / setules on anterior and distal margins, with 2 spinous processes on inner margin (arrows in Fig. 3 E). Segment 2 longest; segment 4 shortest; segment 5 bearing strong spiniform process proximally (arrowed in Fig. 3 E) and very long distal aesthetasc. Segment 7 with 6 apical setae plus acrothek. Armature formula as follows: 1 (1 s) - 2 (9 s) - 3 (4 s) - 4 (2 s) - 5 (7 s, 1 pinnate + 1 spine + 2 s + ae) - 6 (1 s + 2 process) - 7 (6 s + acrothek). Mouthparts, P 1, P 2 and P 4 as in female. P 3 (Figs 2 G, 4 C, D) with 3 - segmented EXP and ENP. EXP as in female. ENP 1 and ENP 2 with inner seta, ENP 2 bearing distinctively dentate apophysis (arrowed in Figs 2 G, 4 D), ENP 3 with 2 lateral and 2 apical setae. P 5 (Fig. 4 E) with basoendopod lacking endopodal lobe. EXP relatively smaller than in female, about 3.6 times as long as wide, armed with 5 setae (2 outer pinnate, 1 plumose apical, 1 pinnate inner distal, and 1 smooth inner proximal). P 6 (Fig. 4 F) represented by small subrectangular segment armed with 1 small lateral and subequally long apical seta. Anal operculum (Fig. 4 B) stronger than in female, with 8 - 10 large spinules and small spinules on surface. Caudal ramus subrectangular, lacking bulbiform expansions as those observed in the female. Distal part about 1.5 times as long as wide.	en	Fuentes-Reines, Juan Manuel, Suarez-Morales, Eduardo, Silva-Briano, Marcelo (2021): A new species of Esola Edwards, 1891 (Crustacea, Copepoda, Harpacticoida, Laophontidae) from the Caribbean coast of Colombia. ZooKeys 1074: 1-15, DOI: http://dx.doi.org/10.3897/zookeys.1074.73030, URL: http://dx.doi.org/10.3897/zookeys.1074.73030
76711CAE00E15BA89DA4CBA67C3FF66F.taxon	distribution	Distribution and ecology. The new species is currently known only from the type locality, Rodadero Bay, Caribbean coast of Colombia. It was found in a mangrove system at a depth of 0.70 m, where the water temperature varies seasonally between 30 and 32 ° C, salinity is 36.1 psu, and pH is 8.3. It is likely that it has a wider distributional range in similar habitats of the western Caribbean region.	en	Fuentes-Reines, Juan Manuel, Suarez-Morales, Eduardo, Silva-Briano, Marcelo (2021): A new species of Esola Edwards, 1891 (Crustacea, Copepoda, Harpacticoida, Laophontidae) from the Caribbean coast of Colombia. ZooKeys 1074: 1-15, DOI: http://dx.doi.org/10.3897/zookeys.1074.73030, URL: http://dx.doi.org/10.3897/zookeys.1074.73030
