taxonID	type	description	language	source
9D67398B495E5D1D8ACF25BCE423826F.taxon	description	Figs 1, 2	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
9D67398B495E5D1D8ACF25BCE423826F.taxon	diagnosis	Diagnosis. Differs from all other congeners by the presence of elongate scandent shoots that exceed four meters in length, coriaceous leaves that are velutinous on lower surface, and a corolla tube that is broadly ampliate on the dorsal surface.	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
9D67398B495E5D1D8ACF25BCE423826F.taxon	description	Description. Scandent subshrub with elongate horizontal shoots, sparingly branched, to 4 m long, to 1.5 cm in diameter, subwoody; internodes 3.8 - 8 cm long, subquandrangular, brown velutinous. Leaves opposite, isophyllous, coriaceous; blade symmetric, ovate to broadly ovate, 9.3 - 12.5 x 5 - 6.7 cm, base truncate to slightly cordate, apex acute, margin serrulate, adaxially light green, densely pilose, abaxially uniformly dark red to green with red venation, densely villous, lateral veins 6 - 9, primary and secondary veins occasionally red. Inflorescence reduced to a single axillary flower (rarely 2 - 3); peduncles absent or highly reduced (<2 mm); bracts absent or caducous, 2 x 3 mm. Flowers resupinate, subtended by elongate pedicels, 1.2 - 4.5 cm long, densely pilose, oriented horizontal relative to shoot, becoming more pendent during anthesis; calyx lobes 5, nearly free, mostly equal in size and shape, dorsal lobe slightly smaller, lobes appressed to flower when immature and spreading during anthesis, mostly green with red margins, 1.6 - 3.0 x 1.0 - 1.3 cm, broadly ovate, apex acute to acuminate, margin with 5 - 10 deep serrations (ca. 4 mm long teeth), densely pilose on both surfaces; corolla tubular, broadly ampliate on dorsal surface (not ampliate on ventral surface), posture horizontal relative to calyx, corolla tube 2.4 - 2.6 cm long, outer surface densely pilose, uniformly bright yellow on the inside and white suffused with yellow on the outside, lobes 3 - 5 x 4 - 6 mm. Androecium of 4 stamens, filaments connate at the base and forming a filament curtain for 3 - 4 mm, free portion of filaments 2 - 2.5 cm long, glabrous; anthers longer than broad, ca. 2 x 1.5 mm, dehiscing by longitudinal slits; staminode lanceolate 3 - 5 x 1 - 2 mm; nectary a bilobed dorsal gland, sometimes appearing truncate, glabrous; ovary superior, densely pilose, 2 - 4 x 2 mm, style ca. 2.0 cm long, glabrous, stigma included and shallowly bifid. Immature fruit cone-shaped, densely pilose, 1.4 x 0.9 cm. Mature fruit not observed.	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
9D67398B495E5D1D8ACF25BCE423826F.taxon	etymology	Etymology. The specific epithet is in reference to Hans Wiehler (1930 - 2003). Wiehler was a practicing Mennonite from East Prussia (now Poland) and immigrated to the USA in the 1950 s. He attained a Bachelor's degree from the Eastern Mennonite College in 1954 and a Bachelor of Divinity degree in 1956 from Goshen College in Goshen, Indiana (Clark 2003). He eventually left the Mennonite church and pursued his passion for botany. Wiehler earned a Master's degree from Cornell and obtained his Ph. D. in Botany from the University of Miami. Wiehler's advanced degrees focused on the taxonomy and classification of Gesneriaceae. Wiehler was one of the first botanists hired by the Marie Selby Botanical Gardens where he served as the associate editor and business manager of the garden's journal, Selbyana (1975 - 1981). He left Selby in 1982 and established the Gesneriad Research Foundation (GRF) in Sarasota, Florida where he directed annual seminars that were attended by horticulturists, taxonomists, students and plant enthusiasts. Wiehler also organized and directed 14 study trips to South and Central America, including the 1993 expedition that resulted in the discovery of Glossoloma wiehlerii. The first author met Hans Wiehler in 1994 and corresponded with him regularly until he died in 2003. Wiehler's passion for Gesneriaceae was contagious.	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
9D67398B495E5D1D8ACF25BCE423826F.taxon	materials_examined	Comments. Most Glossoloma are terrestrial woody subshrubs with an unbranched primary stem. An epiphytic habit is unusual in Glossoloma, especially with elongate or scandent stems. Glossoloma chrysanthum, G. penduliflorum, G. scanden, and G. wiehlerii are the only known species of Glossoloma with an epiphytic habit and elongate scandent stems. Some species, such as G. altescandens (Mansf.) J. L. Clark or G. herthae (Mansf.) J. L. Clark are facultative epiphytes, but their stems are erect and non-scandent. The population of Glossoloma wiehlerii from the type locality was observed to have stems that exceed four meters in length. The longest recorded stem in the genus is G. chrysanthum that exceeded five meters in length (Fig. 3 E). Glossoloma chrysanthum is endemic to Venezuela and is differentiated from G. wiehlerii by a corolla tube that is apically constricted (Fig. 3 A) vs. apically ampliated (Fig. 1 A). In addition, G. wiehlerii differs by the presence of a velutinous indument on the lower leaf surface (vs. hirsute to pilose in G. chrysanthum) and coriaceous leaves (vs. papyraceous in G. chrysanthum). The mature resupinate flowers of G. wiehlerii are inflated on the upper surface (i. e. ampliate or gibbous) and straight on the lower surface (i. e. not ampliate or gibbous). Glossoloma penduliflorum is readily differentiated from all other members of the genus by the presence of solitary axillary flowers with elongate slender pedicels that are 10 - 15 cm long, the longest known pedicels for any member of Glossoloma. Glossoloma scandens differs from G. penduliflorum by the presence of three flowers per axil and relatively short pedicels (<1 cm long).	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
9D67398B495E5D1D8ACF25BCE423826F.taxon	materials_examined	Specimens examined. ECUADOR. Pichincha: canton Quito, distrito Metropolitano de Quito, Chillogallo, road San Juan-Chiriboga, near San Juan, 0.416333 ° N, 78.6580 ° W, 3004 m alt., 20 Feb 2020, Tobar & Gavilanes 3521 (QCNE); distrito Metropolitano de Quito, Nono, comunidad de Alaspungo, Bosque Protector Pacaya, 0.002320 ° N, 78.631260 ° W, 3000 m alt., 15 Jul 2019, Tobar, Marcayata & Gavilanes 3399 (QCNE, US).	en	Clark, John L., Tobar, Francisco (2021): Glossoloma wiehlerii (Gesneriaceae), a new species from the northwestern Andes of Ecuador. PhytoKeys 186: 1-9, DOI: http://dx.doi.org/10.3897/phytokeys.186.73554, URL: http://dx.doi.org/10.3897/phytokeys.186.73554
