taxonID	type	description	language	source
03AA87D3FFF48927FF21D9ECFC0CFDDE.taxon	type_taxon	Type-species. Cormocephalus rubriceps Newport, 1843 (by subsequent designation of Attems 1930: 61).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48927FF21D9ECFC0CFDDE.taxon	distribution	Range. All tropical and subtropical regions with temperate climate.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Figs 2 – 31	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Locus typicus: Saint Vincent Island, Lesser Antilles.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	materials_examined	Studied material. Holotype of C. guildingii (No MYRI- 016 - 01 in OMNH), Figs 2 – 5 (digital photos examined by the first author; also figs 28 – 31 in Schleyko 2018). Holotype of C. impressus (= C. guildingii) (No 453 in GMNH), Figs 6, 7 (digital photos examined by the first author). Mexico (Figs 13, 14, 16 – 21), South of Jalisco State, Chamela Biological Research Station of UNAM, tropical deciduous forest at coastal plain, inside the epiphytic bromeliad genus Tillandsia L.: 1 ad. (spm 1, No EBCh-CHI- 0001 in EBCh), 1 ad (spm 2, No EBCh-CHI- 0002 in EBCh), 19 º 20 ’ - 19 º 34 ’ N, 104 º 58 ’ - 105 º 04 ’ W, 26. II. 1989, col. E. Ramírez; 1 ad. (spm 3, No EBCh-CHI- 0003 in EBCh), 19 º 29 ’ 59.29 ” N, 105 º 02 ’ 36.86 ” W, 15. IX. 2010, col. F. Cupul; 1 ad. (spm 4, No EBCh-CHI- 0004 in EBCh), 19 º 29 ’ 56.30 ” N, 105 º 02 ’ 31.32 ” W, 15. IX. 2012, col. F. Cupul. Jamaica (Figs 15, 22 – 27): 1 ad. (ca. 37 mm), Rc 7687, JBS 1, St. Catherine Parish, Caymanas area, hills north of Caymanas Estate, N slope, 4.7 road km from A 1 at Ferry Town, 18 ° 02.63 ’ N, 076 ° 53.86 ’ W, alt. 40 – 60 m, 24. V. 1999, col. I. V. Muratov & G. Rosenberg; 1 sad. (ca. 27 mm), No Rc 7690, JBS 159, Clarendon Parish, Portland Ridge, Trail 27, primary forest, limestone, red soil, flat, 17 ° 44.41 ’ N, 077 ° 08.69 ’ W, alt. 80 m, 12. X. 1999, col. I. V. Muratov & G. Rosenberg; 1 ad. (ca. 31 mm), Rc 7688, JBS 161, Clarendon Parish, Portland Ridge, by gun club gate, secondary forest and costal scrub, limestone, orange soil, 0 – 20 ° N slope, 17 ° 45.33 ’ N, 077 ° 10.28 ’ W, alt. 5 – 20 m, 12. X. 1999, col. I. V. Muratov & G. Rosenberg; 1 sad. (ca. 30 mm), Rc 7691, JBS 54, St. Thomas Paris, Pera, forested hillside near sugar cane plantation, 17 ° 52.92 ’ N, 076 ° 17.56 ’ W, alt. 0 m, 2. VI. 1999, col. I. V. Muratov & G. Rosenberg; 2 ad. (ca. 32 mm), Rc 7689, JBS 85, St. Andrew Parish, Lucky Valley, around walls of abandoned estate, 2.0 road km N on E side of Mammee River, secondary forest, 17 ° 58.57 ’ N, 076 ° 40.64 ’ W, alt. 360 m, 25. IX. 1999, col. I. V. Muratov & G. Rosenberg.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Hispaniola (= Haiti) Island (Figs 28 – 31), Dominican Republic: 1 sad. (ca. 25 mm) Rc 7074, St. Cristobal [San Cristóbal Province], “ La Cueva ” Colonia, 97 H 6, alt. 550 m, 03.1997, col. I. V. Muratov & G. Robinson.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	materials_examined	Other personally examined material of C. guildingii. 1 ad. (CIRAD), Lesser Antilles, Martinique Island, Morne Aca, Le Marin, 22.11.2017, lat. 14.4614, long. - 60.9002, alt. 213 m, col. Mathieu Coulis (Figs 8, 9, digital photos examined by the first author; also figs 1 – 3 on page 19 in Iorio & Coulis 2019) 1 (s) ad (ATESB), Lesser Antilles, Guadeloupe island group, La Désirade Island about 8 km off the eastern end of Guadeloupe Island, сol. Karl Questel (Figs 10 – 12, digital photos examined by the first author). Type series of C. bonaerius Attems, 1928 (= C. guildingii synonymy by Schileyko 2018): 1 ad. lectotype (designated by Schileyko & Stagl 2004) + 1 sad. paralectotype (No 919 in NHMW), 10 sad. paralectotypes (No 921 in NHMW), Lesser Antilles, Bonnaire Island near Curaçao, leg. & don. A. Gabriel (all specimens personally examined by the first author, see pp. 106 – 109 and figs 22 – 25 in Schileyko & Stagl 2004).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	diagnosis	Composite diagnosis of C. guildingii. Maximal body length 46.5 mm, color of live specimens from yellow to dark brownish (head and both body ends visibly darker than middle portion of body and legs) with some purplish reflections on tergites (Figs 8, 9, fig. 1 at page 19 in Iorio & Coulis 2019); antennae pale blue (Fig. 9). Antennae of 17 (rarely 15 – 19) articles, 5 – 7 basal ones practically glabrous. Cephalic plate (Figs 10, 13, 22, fig. 22 in Schileyko & Stagl 2004) with shortened anteriorly (as long as 1 / 2 – 2 / 3 of cephalic plate) paramedian sutures, their posterior ends crossed by transverse suture; basal plates well-developed. Forcipular coxosternite (Figs 7, 11, 14, 23, 24, 29) with two complete (or slightly shortened posteriorly) longitudinal sutures which much converging anteriorly, meeting each other in a short semilunar suture plus complete (rarely somewhat shortened) transverse suture approximately at the level of condyles. Forcipular tooth-plate typically with 4 (Figs 4, 7, 11, 14, 23) (occasionally / abnormally with 3, Fig. 15) teeth, lateral one clearly isolated; their basal sutures form very obtuse angle or practically straight line. Tergite 1 with complete paramedian sutures (Fig. 13), tergite 21 in most cases with complete median suture (Figs 12, 21 and fig. 22 in Schileyko & Stagl 2004); number of laterally marginated posterior tergites (Fig. 21) varies from 4 to 10, but often only tergite 21 has this margination complete and definite. Sternites 2 – 20 with complete paramedian sutures (Fig. 16), presternites as paired triangles in sternites 1 – 20 (Figs 6, 15, 16, 23). Coxopleuron without welldeveloped process, its rounded median corner with 0 – 2 small apical spines (Figs 17, 18, 19, 26); coxal pore-field oval, slightly longer than sternite 21. Ultimate legs “ truly pincer-shaped ” (sensu Schileyko, Vahtera & Edgecombe 2020; Figs 5, 8, 9, 17, 21, 27, 31; fig. 25 in Schileyko & Stagl 2004); prefemur (Figs 12, 17, 20, 21, 25, 26, 27, 31) with 0 – 4 spines ventrally and ventro-laterally, 0 – 5 ones medially and ventro-medially plus 1 – 3 dorso-medially (of these, 1 or 2 at the position of corner spine, Figs 5, 12, 21, 25, 27, 30). Prefemur ventro-medially with spineless area bordered by very low U-shaped ridge (Figs 20, 26); prefemur, femur and tibia apically with dorso-medial longitudinal depression or sulcus (Figs 5, 21, 25); ventral surface of (at least) tarsus 1 swollen distally (Figs 5, 17, 27, fig. 25 in Schileyko & Stagl 2004); pretarsus enlarged, approximately twice as long as tarsus 2.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Composite description of Mexican specimens (Figs 13, 14, 16 – 21). Length of body 35 – 46.5 mm. Color in ethanol: the whole body and legs light yellow, tergites 10 – 19 may be indistinctly marginated by accumulations of the small granules of pale grey pigment. Antennae composed of 15 – 17 cylindrical articles, reaching the middle of tergite 3 when reflexed. 6 or 7 basal articles practically glabrous with scattered long setae and the subsequent articles densely covered by short setae. Cephalic plate: oval, convex anteriorly and relatively narrow (considerably narrower than tergite 1, Fig. 13); its posterior margin with rounded corners, covered by tergite 1. Anteriorly incomplete paramedian sutures visibly diverge forwards, nearly as long as 2 / 3 of the cephalic plate (or slightly shorter); their posterior ends are crossed by a transverse suture forming well-developed basal plates (Fig. 13). Maxillae 2 with a well-developed dorsal brush. Pretarsus approximately as long as 1 / 4 length of article 3 of telopodite with a claw-shaped tip; accessory spines absent. Dorsal spur of telopodite article 2 not visible. Forcipular coxosternite (Fig. 14) with two complete longitudinal sutures, which much converging anteriorly and meet each other in a short semilunar suture, the latter encircling a minor coxosternal median diastema. Longitudinal sutures are crossed by a complete and branching transverse suture (Fig. 14); chitin-lines absent. Tooth-plates slightly higher than wide (or, less often, as long as wide), definitely narrowing anteriorly. Tooth-plate with 4 teeth (Fig. 14), the lateral tooth is the shortest one and is clearly isolated; two medial teeth are the longest ones and have a common base, being fused to a varying degree. A single minute seta in a small rounded depression directly under tooth margin. The basal sutures of the tooth-plates form a very obtuse angle (Fig. 14) or a practically straight line. Process of the trochanteroprefemur with apical and two medial tubercles, considerably longer than the tooth-plate (Fig. 14); this process is with a basal suture and a characteristic additional transverse suture just below the former. Tarsungulum of normal length, its interior surface with two sharp longitudinal ridges. Tergites: tergite 1 with complete (rarely shortened anteriorly) paramedian sutures (Fig. 13) which are practically parallel or slightly converging anteriorly. Tergites 2 – 20 with complete paramedian sutures, other tergal sutures absent. Tergites 11 – 20 with nearly complete (= slightly shortened posteriorly) definite lateral margination (Fig. 21), only tergite 21 is definitely and completely marginated. Tergite 21 (Fig. 21) with complete median suture, considerably (1.5 times) wider than long and slightly broadened towards the posterior margin; the latter convex apically. Sternites 2 – 20 with complete paramedian sutures. Sternite 21 (Figs 17 – 19) with a poorly-developed longitudinal median depression in its anterior half, trapeziform (somewhat longer than wide (spm 1, 2) or vice versa (spm 3, 4) and distinctly narrowed towards the slightly concave (spm 1, 2) or practically straight (spm 3, 4) posterior margin. Presternites (Figs 14, 16) are well-developed as paired triangles in sternites 1 – 20. Spiracles small, the first pair triangular and the others praсtically round. The spiracle atrium is tightly closed by the two (upper and lower) cuticular folds, so the inner portion of the atrium is not visible; the third (median) fold is very strongly reduced. Legs: pretarsus of legs 1 – 20 with two well-developed accessory spines. Ultimate LBS: coxopleuron visibly longer than sternite 21, with a rounded median corner at the position of the coxopleural process (Figs 17 – 19); coxopleural surface without setae. Coxal pores numerous, coxal pore-field oval, slightly longer than sternite 21; it does not reach closely to the lateral margin of the coxa and considerably less so to the caudal one. 0 – 2 small apical spines positioned at the rounded median corner of the coxopleuron and a small lateral spine (Fig. 18) may be present at its posterior margin. Ultimate legs (Figs 17, 21) “ truly pincer-shaped ” (sensu Schileyko, Vahtera & Edgecombe 2020), 6.9 – 8.3 mm long, much shortened and broadened (with prefemur ratio of length: width 1 – 1.4). Prefemur (Fig. 20) with 0 – 4 / 5 ventral (including 0 or 1 ventro-lateral) spines, 0 – 5 medial + ventro-medial ones and 1 – 3 dorso-medial (including 1 or 2 ones at the position of the corner spine). Prefemur ventro-medially with a spineless area bordered by a low U-shaped ridge. Prefemur (Fig. 21), femur and tibia distinctly flattened dorsally, ventral surface of tibia and tarsus 1 somewhat swollen distally. Distal end of prefemur, femur and tibia with a well-developed dorso-medial longitudinal depression which is approximately as long as 1 / 4 of length of the corresponding article (Fig. 21). Pretarsus enlarged, twice as long as tarsus 2 but somewhat shorter than the tarsal articles taken together; ventral surface of pretarsus forms a sharp ridge, accessory spines absent. Variability. Right antenna of spm 1 has 13 articles (Fig. 13) most probably having been regenerated. The specimens studied demonstrate considerable variation in spination of the coxopleuron. Spm 1 (46 mm long) lacks any coxopleural spines (Fig. 19), in spm 2 (46.5 mm long) the right coxopleuron is also spineless whereas the left one has 1 apical spine only (Fig. 17), in spm 3 (35 mm long) each coxopleuron is with 2 apical plus 1 lateral spine (Fig. 18), in spm 4 (40 mm long) the right coxopleuron is with 2 apical plus 1 lateral spine and the left one with 1 apical plus 1 lateral one. The only other difference between these specimens is the number of spines of the ultimate prefemur: spm 1 has 2 distal dorso-medial plus 1 ventral spine, spm 2 has left leg with 2 distal dorso-medial and right leg with 1 distal dorso-medial plus 1 medial spine (Fig. 17), spm 3 has left leg with 2 distal dorso-medial plus 7 ventral spines and right leg with 1 distal dorso-medial plus 5 ventral spines (Fig. 20). Spm 4 has both these legs with 2 distal dorso-medial plus 5 ventral spines.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	discussion	Remarks on Mexican specimens. Studied specimens are typical representatives of this species but show considerable intraspecific variability of the spine numbers on the coxopleuron and ultimate prefemur. Thus the taxonomic weight of both these characters should be decreased in the species of the guildingii - subgroup. Composite description of Jamaican specimens (Rc 7687 – 7691, Figs 15, 22 – 27)	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Length of body up to 37 mm. Color in ethanol: the whole body and legs light yellow. The whole body (head, forcipular segment, all tergites and sternites) is covered by numerous very short setae. Antennae composed of 16 or 17 cylindrical articles, practically reaching the posterior margin of tergite 2 when reflexed. 5 or 6 basal articles practically glabrous with scattered long setae and the subsequent articles densely covered by short setae. Cephalic plate (Fig. 22): oval and relatively narrow (considerably narrower than tergite 1), its posterior margin with rounded corners, covered by tergite 1. Paramedian sutures visibly diverge forwards, approximately as long as 1 / 2 of cephalic plate (or slightly longer), their posterior ends are crossed by a transverse suture forming welldeveloped basal plates (Fig. 22). Maxillae 2 with well-developed dorsal brush. Pretarsus approximately as long as 1 / 4 length of article 3 of telopodite (Fig. 24) with claw-shaped tip; of two accessory spines the dorsal one is more slim and much adpressed to the pretarsus being very poorly recognizable. Telopodite article 2 with a clearly visible dorso-apical spur. Forcipular coxosternite (Figs 15, 23, 24) with two shortened posteriorly (sometimes slightly exceeding the mid-point of the coxosternite) longitudinal sutures, which much converging anteriorly and meet each other in a semilunar suture, the latter encircling a minor coxosternal median diastema. These longitudinal sutures are crossed by a practically complete, branching transverse suture; chitin-lines absent. Tooth-plates approximately as long as wide, definitely narrowing anteriorly. Tooth-plate (Fig. 23) with 4 teeth (in sad. Rc 7690 occasionally / abnormally 3 teeth, Fig. 15), of these the lateral one is clearly isolated and visibly shorter than the medial teeth, the latter have a common base being fused to a variable degree (to practically fully fused in Rc 7687, Fig. 24). A single clearly visible seta occurs in a rounded depression directly under the tooth margin. The basal sutures of the tooth-plates form a practically straight line. Process of trochanteroprefemur (Figs 15, 23, 24) with one apical and one medial tubercle, considerably longer than the tooth-plate. This process with a basal suture and a characteristic additional transverse suture just below the former. Tarsungulum of normal length, its interior surface with two sharp longitudinal ridges Tergites: tergite 1 with complete paramedian sutures which slightly converging anteriorly (Fig. 22). Tergites 2 – 20 with complete paramedian sutures, other tergal sutures not visible. Tergites 10 / 13 – 20 with incomplete (somewhat shortened posteriorly) and more or less definite lateral margination, only tergite 21 definitely and completely marginated. Tergite 21 (Fig. 25) without a median suture, an incomplete shallow median longitudinal sulcus (not suture) may be visible in the middle of this tergite (Rc 7687, 7688); tergite 21 considerably (approximately 1.5 – 2 times) wider than long and slightly broadened towards the posterior margin, the latter convex apically. Sternites 2 – 20 with complete paramedian sutures. Sternite 21 (Fig. 26) with a very poorly-developed longitudinal median depression in the anterior half, trapeziform (approximately as long as wide) and distinctly narrowed towards the practically straight posterior margin. Presternites well-developed as paired triangles (Figs 15, 23) in sternites 1 – 20. Spiracles small, the first pair somewhat elongated and the others praсtically round. The spiracle atrium is tightly closed by the two (upper and lower) cuticular folds, the third (median) fold is very much reduced being practically invisible. Legs: pretarsus of legs 1 – 20 with two well-developed accessory spines. Ultimate LBS: coxopleuron (Fig. 26) visibly longer than sternite 21, with a rounded median corner at the position of the coxopleural process; coxopleural surface without setae. Coxal pores numerous, of various sizes; coxal pore-field oval, slightly longer than sternite 21, it does not reach slightly to the lateral margin of the coxa and is considerably distant from the caudal one. Two small apical spines are positioned at the rounded median corner of the coxopleuron (Fig. 26) plus a small spine at its posterior margin. Ultimate legs (Figs 25, 26, 27) “ truly pincer-shaped ”, ca 8 mm long (when body ca 32 mm), much shortened and broadened (with prefemur ratio of length: width 1 – 1.4). Prefemur with 5 longitudinal rows of very small spines: dorso-medial one of 3 spines, medial one of (1) 2, ventro-medial of 2, ventral of 1 – 3 and ventro-lateral row of 2 spines. Two most apical dorso-medial spines (they have common base) form a kind of corner spine (Fig. 25). Prefemur ventro-medially with a spineless area bordered by very low U-shaped ridge (Fig. 26). Prefemur, femur and tibia definitely flattened dorsally, ventral surface of tarsus 1 and tarsus 2 somewhat swollen distally. Distal end of prefemur, femur and tibia with a shallow dorso-medial longitudinal depression which is as long as 1 / 3 – 1 / 4 of length of the corresponding article (Fig. 25). Pretarsus enlarged and much elongated (Fig. 27), on average nearly twice as long as the tarsal articles taken together; ventral surface of pretarsus forms a well-developed, very sharp ridge, accessory spines absent.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	discussion	Remarks on Jamaican specimens. Schileyko (2018: 72) considered C. bonaerius Attems, 1928 to be a junior synonym of C. guildingii basing, in particular, on re-investigation of Jamaican material; for data on the type series of C. bonaerius see Schileyko & Stagl (2004: 106).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	description	Description of specimen from Hispaniola (1 sad. Rc 7074, Figs 28 – 31) Length ca. 25 mm (Fig. 28). The whole body with well-developed small spines. 17 antennal articles, of these 6 or 7 basal ones with some long setae. Head with incomplete (somewhat shortened anteriorly) paramedian sutures. Forcipular coxosternite (Fig. 29) with two complete typical and much converging anteriorly longitudinal sutures plus an incomplete transverse suture; right tooth plate broken, left one with 3 teeth (but the most medial one is apparently a result of regeneration of two original teeth). Tergite 1 with incomplete (somewhat shortened anteriorly) paramedian sutures, about 10 posterior tergites marginated laterally, tergite 21 without a median suture (Fig. 30). Sternites 2 – 20 with complete paramedian sutures, sternite 21 trapeziform, nearly as long as wide, with a welldeveloped longitudinal median sulcus / depression in the posterior half, its posterior margin practically straight. Presternites are well-developed as paired triangles in sternites 1 – 20 (the largest ones in sternite 1), the presternites are better developed on midbody segments. All legs with two well-developed pretarsal accessory spines. Ultimate LBS: coxopleuron (Fig. 31) with a very short and apically rounded process which has two apical spines, a single spine at the posterior coxopleural margin, the pore field slightly longer than sternite 21. Ultimate prefemur (Fig. 31) with 2 ventro-lateral spines, 2 ventral, 2 ventro-medial, 2 medial and 3 dorso-medial ones (of these 2 at the position of the prefemoral corner spine); prefemur and femur apically with a short characteristic dorso-medial longitudinal depression, pretarsus much longer than tarsus 2 and slightly shorter than tarsal articles taken together.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF48928FF21DAFBFB7DFCDF.taxon	discussion	Remarks on specimen from Hispaniola. Schileyko (2018: 76) mentioned this specimen as “ Cormocephalus sp. ” noting that it “ … is very similar to C. guildingii but has tergite 21 without median suture ”; apart from this difference this subadult fits well in the expanded and improved new concept of C. guildingii. Range of C. guildingii (see also Chagas et al. 2014: 139) N America: W Mexico; Antilles: from Cuba (Kraepelin 1904: 251), Cayman Islands and Jamaica to Bonnaire Island near Curaçao; Range of former C. impressus: Northern and North-Western regions of South America: Northern Venezuela (Caracas and Puerto Cabello), Ecuador, Western Colombia (Quindío Department and Eastern Cordillera), Northern, Central and Southern Peru (Fig. 1). Remarks on Range of C. guildingii Until recent years C. guildingii was known as an endemic to the Antilles (Schileyko et al. 2018: 561, Iorio & Coulis 2019: 18); this study confirms its occurrence in the Dominican Republic (Rc 7074 from San Cristóbal Province). However, since the synonymy of C. impressus under C. guildingii is confirmed below, we have transferred all the records of that former species to C. guildingii. Also, the studied material from Western Mexico allowed to expand the distribution area of C. guildingii (Fig. 1) to the most southern part of North America (Cupul-Magaña 2009: 90). Kraepelin (1903: 181) mentioned a questionable specimen of Cupipes (= Cormocephalus) impressus from Paraguay (without locality) and in 1904 recorded this species (p. 251) from they same country based on specimen (s) kept in MNHN. However, Paraguay lies outside the proven boundaries of the distribution of Cormocephalus guildingii which seems to be replaced in eastern and southern parts of Brazil by C. andinus. Bücherl (1939: 249) gave a very short description of C. bonaerius (= C. guildingii, see Schileyko 2018) mentioning it from “ Guyanas ” (Guyana, Surinam and French Guiana) without any localities or specimen data, and in the faunistic paper of 1941 he mentioned (p. 299) C. (C.) impressus var. neglectus (= C. impressus) from the river Madeira in the Brazilian State Mato Grosso. However, at present there is no confirmed information (i. e. data on specific specimens + localities) on the occurrence of C. guildingii in these regions. Schileyko et al. (2018: 561) erroneously stated the type locality of C. guildingii as “ Hispaniola Island, Greater Antilles ”. Remarks on C. guildingii Until now this species had not been described in detail (and not illustrated adequately), except for the original description of Otostigma cormocephalinum Pocock, 1888, Kraepelin’s (1903: 181) description of Cormocephalus impressus (both are synonyms of C. guildingii) and the data of Schileyko (2018: 74 – 75, figs 28 – 31).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFFA892BFF21DD61FB2AFC03.taxon	description	Figs 33 – 38	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFFA892BFF21DD61FB2AFC03.taxon	materials_examined	Studied material. Panama, IBISCA project, Colón Province, San Lorenzo Forest, tree canopies, 1 sad. (Rc 7155), 2004, col. I. Tuf. Brazil, Amazonas, Reserva Florestal A. Ducke, near [about 25 km N of] Manaus, 59 59 ’ W, 2 55 ’ S, primary tropical rainforest forest on Terra firme, 1 ad. (Rс 6483), 06 - 011. III. 1998, col. S. I. Golovatsch.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFFA892BFF21DD61FB2AFC03.taxon	description	Description of ad. Rc 6483 [data on sad. Rc 7155 in square brackets where it differs]. Length ca. 30 [15] mm. Coloration in alcohol uniformly yellow [greyish] with head visibly darker [head, body (including sternites) and ultimate legs with small aggregations of dark pigment, Figs 33, 34]; numerous small setae at all body surface. Head (Fig. 35) with much shortened anteriorly (as long as ½ [1 / 3] of head), but well-developed [indefinite] paramedian sutures; very short antennae (left 15, right 17 articles [both 17]) hardly extending to the posterior margin of tergite 1 [2] when reflexed. Forcipular coxosternite (Figs 36, 33) with two much shortened posteriorly (as long as 1 / 2 [1 / 3] of coxosternite) and converging anteriorly typical longitudinal sutures; transverse suture absent. Tooth-plate with 4 teeth, of these the most lateral and the most median ones are approximately of the same length, being much shorter than 2 remaining teeth; long and pointed trochanteroprefemoral process with one lateral tubercle. Tergites 1 – 20 with complete paramedian sutures; tergite 21 (Fig. 38), with complete median suture [plus a median depression in the posterior half], only this tergite is marginated laterally. Sternites 2 – 20 with complete paramedian sutures disposed in paramedian sulci; sternite 21 (Fig. 38) trapeziform, slightly [noticeably] longer than wide, its posterior margin rounded [straight]; sternite 21 with a short, shallow median sulcus / depression in the posterior third. Legs with 2 short [minute] but clearly recognizable [hardly recognizable] pretarsal accessory spines. Ultimate LBS (Fig. 37): coxal pore field as long as [very slightly longer than] sternite 21, coxopleuron slightly longer than sternite 21, with rounded posterior margin, lacking either any process or spines [the only minute spine at the very inner corner of the coxopleuron]. Ultimate legs (Figs 37, 38): prefemur, femur and tibia much flattened dorsally. These articles are each with characteristic dorso-medial longitudinal depression apically; tibia and tarsus 1 somewhat swollen / bulbous ventrally. Prefemur with the only spine at the position of the corner spine (Fig. 38) [prefemur (Fig. 34) with 3 unusually large dorso-medial spines — 2 of them disposed at a remarkably enlarged corner spine — plus 1 ventro-medial and 1 medial one, both of the latter are disposed at the distal margin of the prefemur, forming a transverse row together with the corner spine; no any ventral spines].	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFFA892BFF21DD61FB2AFC03.taxon	discussion	Remarks. Adult spm Rc 6483 has been mentioned by Schileyko (2002) as C. brasiliensis Humbert & Saussure, 1870 but the present re-investigation shows that it is C. ungulatus, because it shares with the latter such important peculiarities of the forcipular coxosternite as very short paramedian sutures plus a total absence of a transverse one. This specimen also has no paired dorso-distal “ bifid spines ” on the ultimate femur (!), which should be diagnostic for C. brasiliensis (see Chamberlin 1914: 183) and being very unusual (if not unique) within Cormocephalus; Lewis (1989: 1006) wrote that C. brasiliensis “ was described on the basis of a defective specimen ”.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	description	According to the faunistic data of Kraepelin (1904: 251), Brölemann (1909: 9), Bücherl (1939: 251, 1941: 300, 1974: 103) and Chagas et al. (2014: 139) this species has been recorded from French Guiana (Ouanary at Oyapock river), the Brazilian states Pernambuco (Recife) and Amazonas (Manaus), from a few places in Colombia and (with no definite localities) from “ Antilles ”, Haiti, Venezuela, Ecuador, Peru, Bolivia, Paraguay and Argentina. We add to this list Panama, confirm occurrence of this species in the Brazilian State Amazonas but do not confirm it for the Brazilian State Rondonia (because spm Rc 7239 recorded by Schileyko (2002) as C. ungulatus has now been reidentified as C. andinus). Such a disjunct distribution of C. ungulatus seems to be quite unnatural, possibly being a result of insufficient data. We suppose that this species has a transbrazilian range, being distributed sympatrically with C. andinus in the western section of Amazonia. According to the literature data the distribution areas of C. ungulatus and C. guildingii seem to overlap in Antilles, Venezuela, Ecuador and Peru.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	discussion	Discussion on C. ungulatus and its relationship to C. guildingii Kohlrausch (1878: 23) described the genus Cupipes, which contained 6 species (of them 3 from the Neotropics, including the new species С. microstoma, synonymised to C. impressus in 1903 by Kraepelin). Cupipes ungulatus was described by Meinert (1886: 187) from the very distant parts of the Neotropics — a couple of localities in Hispaniola Island (syntypes CHIL- 1169, CHIL- 1170 in MCZ) and Pernambuco (syntype CHIL- 1168 in MCZ), one of the most eastern Brazilian States. The short original description is not detailed and contained mainly the general peculiarities of the former genus Cupipes but no real diagnostic characters at the species level. Hence, it does not allow us to distinguish Cormocephalus ungulatus from C. guildingii. In 1893 Pocock gave a short note (with no morphological details) on Cupipes ungulatus judging that it is “ … closely related to C. Guildingii. But it undoubtedly differs in having all the tergites except the last immarginate ”. The first adequately detailed description of C. ungulatus is that of Kraepelin (1903: 177) who described a few / some (“ … die oben beschriebenen von Panama ”) adult (“ Lange 40 mm ”) specimens which should differ somewhat (“ passt nicht vollig ”) from “ Die Exemplare Meinert’s vom Haiti and Pernambuco ”. Kraepelin (1903) was the first to mention the presence and configuration of the forcipular coxosternite’s sutures as a diagnostic character of Cormocephalus species. This author wrote (p. 177) that in C. ungulatus the corresponding longitudinal sutures are developed “ only in the anterior third ” but said nothing about the corresponding transverse suture (so it would be logical to suggest that the latter was absent in his material). All Kraepelin’s specimens had each forcipular tooth plate with typical « 1 + 3 » teeth; he also noted (p. 178) “ the pale ” juvenile specimen from Colombia which should fit well to the description of above-mentioned specimens from Panama. Brölemann (1898: 318) mentioned a single immature (21 mm long) specimen of Cupipes sp. from “ Bas Sarare (Venezuela) ” giving a drawing (corresponding fig. 3) in lateral view of its posterior end of the body — the “ possible new species ” should be characterized by a total absence of spines on both the ultimate prefemur and the coxopleuron. In 1905 he described (p. 65) this specimen as Cupipes ungulatus var. Venezueliana (using on the same page the trinominal “ C. ungulatus venezuelianus ”, see also Schileyko 2014: 186). A description of this doubtful form (Schileyko and Stagl (2004) noted, that its taxonomical position is “ too unclear to put in any group ”) is not detailed enough and lacks data on sutures of the forcipular coxosternite, being, perhaps, based on the abnormal specimen of Cormocephalus ungulatus. However, as an absolutely spineless ultimate prefemur is not characteristic for Neotropical representatives of this genus (except for the doubtful Cupipes lineatus biminensis Chamberlin, 1952 and Cormocephalus C. impulsus Lewis, 1989) we prefer to keep C. venezuelianus as an independent species until its type-series is re-examined.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	materials_examined	Data on C. ungulatus given by Chamberlin (1914, 1918, 1921, 1922, 1925, 1944) are just faunistic records, which contain neither any morphological data, nor drawings although this author “ worked with material deposited at the MCZ ” (Martínez-Muñoz & Perez-Gelabert 2018: 76). The only worthwhile data on C. ungulatus presented by Chamberlin were two short notes in his papers of 1914 and 1957, where he mentioned that the number of spines on the ultimate prefemur varies in this species considerably and (1914: 184) “ it may prove not possible to segregate the forms [of this species] on the basis of this character ”.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	discussion	In 1955 Chamberlin described Cormocephalus mundus from two adult (55 – 60 mm) plus one “ immature ” syntypes from Southern Peru (Abancay and Ayacucho). According to the original description (which lacks drawings), “ C. mundus ” differs from C. ungulatus only by body size, so we feel confident to synonymise this very doubtful form (see also Kraus 1957: 383). Thus Cormocephalus mundus Chamberlin, 1955 is a junior synonym of C. ungulatus (Meinert, 1886). Later (1957: 31) Chamberlin described “ Cormocephalus (Cupipes) tingonus ” from a single adult (“ Length, 40 mm ”) specimen from Central Peru. According to the original description (which lacks drawings but is adequately detailed) the new form differs from C. ungulatus by having somewhat longer typical paired longitudinal sutures of the forcipular coxosternite plus a slightly differing number and disposition of the spines on the ultimate prefemur (a character mentioned by Chamberlin himself as very variable, see above). Taking into consideration both these minor differences and the geographical aspect we consider Cormocephalus (Cupipes) tingonus Chamberlin, 1957 to be a junior synonym of C. ungulatus (Meinert, 1886). It is interesting, that Chamberlin (1957: 31) wrote about his C. tingonus: ” … two [(!)] last dorsal plates [= tergites] with a median longitudinal sulcus [= suture?] ”; the presence of a single (!) median suture on any tergites except for the first and the ultimate is not known at present for any scolopendromorph.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	diagnosis	Attems (1930: 62) reasonably synonymised Cupipes to Cormocephalus but just copied (p. 101) Kraepelin’s description of C. ungulatus (including the data on sutures of the forcipular coxosternite). Bücherl (1939: 251) provided a kind of diagnosis for Cormocephalus ungulatus mentioning it from “ Pernambuco, Amazonas, Venezuela ” but gave no localities or data on studied specimens, so (possibly), he had no actual material at his disposal. In 1942 Bücherl described in detail (p. 123) “ Cormocephalus (C.) impressus unimarginatus n. subsp. ” from an unknown number of specimens (“ 28 – 29 mm long, No. 141 in Museu Nacional do Rio de Janeiro ”) from South-Eastern Brazil (Parque Nacional da Chapada dos Veadeiros in Goiás State), which is quite close to Pernambuco — one of the localities mentioned in the original description of C. ungulatus. Bücherl’s C. impressus unimarginatus fits well to our composite diagnosis of C. guildingii but definitely differs by having 3 + 3 teeth on the forcipular tooth plates (fig. 5 in Bücherl 1942) and the total absence of “ un sulco horizontal no coxosternum forcipular ” — the diagnostic character of C. ungulatus — so we presume that the mentioned above material from Goiás represents the latter species.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	discussion	In their faunistic account Martínez-Muñoz and Perez-Gelabert (2018: 76) wrote that Chamberlin (1918) “ reexamined ” Meinert’s Haitian syntypes of C. ungulatus as C. guildingii “ leaving the syntype from Pernambuco as the only name bearer for C. ungulatus ” (p. 83). In fact, Chamberlin (1918: 156) simply noted these syntypes as “ Cupipes guildingi [sic!] (Newport) ” but gave no reasons for this. Also there is no published evidence (morphological data, drawings etc.) of that “ reexamination ”; however, as the formal act of synonymisation took place, we re-validate C. ungulatus basing on the set of diagnostic characters (see below).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	diagnosis	Summing up all these, short and much scattered, morphological descriptions we can state that, at present, C. ungulatus is the closest relative of C. guildingii differing from the latter by the total absence of a forcipular transverse suture (this reliable character seems to be the main diagnostic one) and much shortened (about 1 / 3 of coxosternite length) corresponding longitudinal sutures, which are complete (or nearly so) in C. guildingii. Two other characters are: the number of laterally marginated tergites — examined specimens of C. ungulatus actually show only the ultimate tergite marginated (compare to composite diagnosis of C. guildingii above, Fig. 21) — and the absence of ventral spines on the ultimate prefemur. However, both of these characters are not stable in Cormocephalus and should not be used as diagnostic on their own.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFF88935FF21DBF4FD6AFE77.taxon	discussion	It should be noted also, that only a few specimens of C. ungulatus are known from the literature, of them only Kraepelin’s (1903) specimen from Panama and our Rc 6483 and Rc 7155 (which seems to be immature due to its small length and, possibly, newly molted) have been described in enough detail to distinguish this species from C. guildingii. Thus, we prefer to keep C. ungulatus (Meinert, 1886) as an independent species until re-examination of the type series (which is not available at the present).	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFE68936FF21DD4CFB46FE2F.taxon	discussion	The genus Cormocephalus s. str. occurs in the New World in the Neotropical region only (see Appendix 1) being represented there solely by the main part of the gervaisianus species group (i. e. by the majority of species of the former genus Cupipes Kohlrausch, 1878 in the sense of Kraepelin 1903). We consider these ten (or approximately so; see below) species to form the general Neotropical clade, which is well circumscribed morphologically. The present study allows to correct and much extend the group of four closely related Antillean species mentioned for the first time by Schileyko (2018: 72 – 75). The species investigated personally (including the recently studied and re-examined material) — C. guildingii, C. ungulatus, C. andinus, C. lineatus (Figs 43 – 47) and Cormocephalus sp. (Rc 7468, see above) — definitely form a well-defined subclade within the general clade of Neotropical Cormocephalus. We name this new subclade as the “ guildingii - subgroup ” after its oldest member; it is unequivocally united by nine diagnostic synapomorphies (see paragraph “ Diagnosis of the guildingii - subgroup ” below). The guildingii - subgroup may also include two questionable New World members of the gervaisianus - group, namely C. brasiliensis (see above) and C. amazonae (Chamberlin, 1914), the latter is known from a single specimen (most likely of C. andinus) with abnormally developed forcipular tooth-plates. Also under question remain five poorly described Venezuelan species of González-Sponga (2000) (C. glabratus, C. edithae, C. abundantis, C. facilis and C. maritime; see also Schileyko 2014: 188). According to the accompanying drawings, two of these forms are not members of the guildingii- subgroup: C. glabratus has no sutures on the forcipular coxosternite while C. abundantis has an unusually short ultimate pretarsus; C. edithae may be a junior synonym of C. ungulatus as it has a forcipular coxosternite with the typical paired longitudinal sutures only. However we are not able to analyze these questionable forms in more detail here as the corresponding types are not currently available.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFE68936FF21DD4CFB46FE2F.taxon	diagnosis	At present at least three members of the general Neotropical clade can not be included in the new subclade / subgroup because of some disparity in the diagnostic peculiarities. They are: the questionable C. venezuelianus (see above), the well-described C. impulsus (US Virgin Islands, Lesser Antilles) and C. lineatus biminensis, poorly described from three small (20 – 24 mm long) immature (?) specimens (South Bimini, Bahama Islands). These three species distinctly do not fit to the diagnosis of the new subgroup, in particular by the total (!) absence of spines on the ultimate prefemur, which condition, however, may be a subject of rare individual variation or an abnormality. Hence the type series should be re-examined to draw any conclusions about the relations of these forms with the guildingii – subgroup and about the validity of both C. venezuelianus and C. lineatus biminensis.	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
03AA87D3FFE0893CFF21DFD6FCC8FE05.taxon	synonymic_list	Cormocephalus guildingii Newport, 1845 (W Mexico, Antilles from Cuba Cayman Islands and Jamaica to Bonnaire Island near Curaçao, Northern Venezuela, Ecuador, Western Colombia, Northern, Central and Southern Peru) Cormocephalus andinus Kraepelin, 1903 (South-Eastern Mexico, Bolivia, Central and Southern Peru, Ecuador (including Galapagos Islands), Brazil: Amazonas and Pará) Cormocephalus ungulatus Meinert, 1886 (Haiti, Panama, French Guiana, Venezuela, Brazil: Pernambuco and Amazonas (possibly transbrazilian distribution), Colombia, Ecuador, Peru, Bolivia, Paraguay, Argentina) Cormocephalus lineatus Newport, 1845 (Lesser Antilles)? Cormocephalus brasiliensis Humbert & Saussure, 1870 (Brazil: Amazonas, Venezuela?)? Cormocephalus amazonae Chamberlin, 1914 (Brazil: Amazonas)	en	Schileyko, Arkady A., Cupul-Magaña, Fabio G. (2021): Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus). Zootaxa 5071 (3): 301-325, DOI: https://doi.org/10.11646/zootaxa.5071.3.1
