taxonID	type	description	language	source
0627421AE7064948FF50200E6104FAE1.taxon	type_taxon	Type species: Jethsura ferruginea Cameron. Monotypic and original designation.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7064948FF50200E6104FAE1.taxon	diagnosis	Generic diagnosis. Jethsura is distinguished by the combination of the following characters: 1) mandible slightly narrowed to moderately wide and nearly unidentate, with the ventral tooth greatly reduced and represented only by a small notch; 2) dorsal face of propodeum usually with carinae obsolete (moderately to weakly-developed in J. pyriformis); 3) gastrocoelus and thyridium of T 2 varying from subobsolete to moderately well-developed but always narrower than interthyridial interval; 4) males without tyloids; and most notably, 5) female with T 6 anteriorly constricted. The latter character is unique among phaeogenine genera, and thus makes the females instantly recognizable. Males of J. pyriformis superficially resemble a few species of Stenodontus, particularly several undescribed species in Mexico and the southwestern U. S. A. Jethsura pyriformis males can be readily distinguished from these species by the broader, bidentate mandible as opposed to the narrower, sickle-shaped, unidentate mandible in Stenodontus.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	description	Figs 1 – 2, 12 urn: lsid: zoobank. org: act: 66 A 98690 - 15 A 1 - 4345 - 8144 - 4 FC 0 E 8 E 24747	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	diagnosis	Diagnosis. Jethsura euthenia is similar to J. gonawindua, both being overall black and with the mesosoma not laterally flattened. The black mandible with a dark reddish apex is sufficient to distinguish J. euthenia from the latter species which has a brown to light brown mandible with a dark brown apex. Additionally the head morphology differs significantly between the two species. In anterior view, the head in J. euthenia is slightly longer and tapers ventrally and the clypeus and supraclypeal area are slightly convex in profile. In males, however, the differences in head morphology are less pronounced.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	description	Description. Female (Fig. 1 A – H). Body length: 6.9 – 8.1 mm; fore wing length: 4.4 – 4.9 mm. Color. Primarily black with few yellow and yellowish-white markings and partially brownish legs. Head black, except for mandibular brown with black base and dark reddish-brown apex, paraocular area with narrow light yellowish mark medially dorsal to antennal base and at dorsolateral corner (25 % of specimens with venterolateral corner brownish). Antenna black, except for pedicel apically brownish. Mesosoma black, except for dorsal margin of pronotum and tegula yellowish-white. Fore leg: coxa varying from entirely black to black and brownish anteriorly; trochanter black, except for apex brownish laterally; trochantellus brown; femur anteriorly brownish-yellow, except for basal 0.3 dark brown, posteriorly dark brown, except for apical 0.2; tibia brownish-yellow ventrally, darker dorsally; tarsus reddish-brown except tarsomere 5 dark brown to dark reddish-brown. Mid leg: coxa varying from entirely black to black and brownish anteriorly; trochanter black, except for apex anteriorly brownish; trochantellus brown; femur dark brown, except for anteriorly with apical 0.2 brownish-yellow, posteriorly varying from entirely dark brown to apical 0.1 brownish-yellow; tibia brownish-yellow, dark dorsally; tarsus reddish-brown except tarsomere 5 dark brown to dark reddish-brown. Hind leg: coxa black; trochanter black; trochantellus dark brown; femur dark brown to black except for apical 0.1 brownish; tibia brownish-yellow becoming brown dorsally; tarsomeres 1 – 4 reddish-brown with brownish-yellow bases; tarsomere 5 dark brown. MS 1 black. T 2 - 7 black with lateral and posterior margins light brownish. Wing: membrane clear; basal 0.1 of wing with veins white, remaining sections brown. Head. Slightly tapering ventrally. Mandible moderately wide (MI: ~ 3.2 ×). Clypeus and supraclypeal area slightly convex in lateral view. Clypeus smooth and moderately to coarsely punctate with punctures separated by 0.5 – 1.8 × their diameter. Supraclypeal area smooth and densely punctate with punctures varying from subadjacent to separated by 0.3 × their diameter, 50 % of specimens with dorsal punctures confluent forming rugulae submedially. Gena smooth and finely punctate with punctures separated by 1.5 – 2.5 × their diameter. Supra-antennal area smooth and finely to moderately punctate with punctures subadjacent to separated by 0.3 × their diameter, 20 % of specimens with some punctures confluent and forming transverse rugulae. Vertex smooth and finely, sparsely punctate with punctures separated by 1.0 – 3.0 × their diameter, denser medially. Hypostomal and occipital carinae meeting before mandible. Antenna with 24 flagellomeres. Mesosoma. Of normal width, not laterally flattened. Pronotum varying from smooth and finely punctate dorsally to primarily rugulose and smoother anterodorsally. Propleuron smooth with fine, dense, shallow punctation. Mesonotum smooth with anterior 0.1 very finely, densely punctate, remainder moderately punctate with punctation varying considerably in size and density. Scutellum smooth and sparsely punctate with punctures separated by 1.0 – 2.0 × their diameter. Mesopleuron smooth and varying from densely, coarsely punctate with punctures subadjacent to densely punctate-rugulose. Speculum smooth and finely to moderately punctate. Ventral division of metapleuron smooth and sparsely to densely punctate. Propodeum: of normal width, length and width nearly equal; evenly rounded mediolaterally; dorsal face smooth and varying from densely, coarsely punctate to punctate-rugulose laterally becoming smoother medially; posterior face varying from rugulose to punctate-rugulose; carinae obsolete, except for pleural carina, posterior transverse carina, and median longitudinal carina posterior to posterior transverse carina subobsolete. Legs of normal length. Hind femur normal, not anteroposteriorly flattened. Metasoma. MS 1 smooth, postpetiole varying from moderately to densely, coarsely punctate, medially nearly impunctate. T 2 smooth and with moderately dense punctation varying from fine to coarse. Gastrocoelus well-developed. Thyridium well-developed; narrow, 0.7 – 0.8 × as wide as interthyridial width; distant from T 2 anterior margin by 0.2 × tergite length. T 3 – 7 smooth and varying from moderately to finely punctate with punctation becoming sparser and finer posteriorly. Male (Fig. 2 A – C). Body length: 6.9 – 8.1 mm; fore wing length: 4.4 – 4.9 mm. As in female except for: postpetiole of MS 1 moderately to densely punctate, 10 % of specimens with some punctures confluent becoming irregular longitudinal rugulae; T 2 – 7 densely punctate with punctures separated by 0.3 – 0.5 their diameter and becoming sparser posteriorly. Antenna with 26 flagellomeres.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	materials_examined	Material examined. Holotype: Colombia • 1 ♀; Boyacá, SFF Iguaque, Cabaña Mamarramos; 2855 m; 5 ° 25 ’ N, 73 ° 27 ’ W; 23. v. – 08. vi. 2000; P. Reina; P. Reina; M. 149; IAVH. Paratypes: Colombia • 36 ♂♂, 3 ♀♀; same collection data as holotype; 19. iv. – 0.6. v. 2000; M. 55; IAVH • 10 ♂♂, 4 ♀♀; same collection data as preceding; EMUSENT 00004089, EMUSENT 00004021, EMUSENT 00003574, EMUSENT 00004386, EMUSENT 00004082, EMUSENT 00004382, EMUSENT 00003759, EMUSENT 00003999, EMUSENT 00004201, EMUSENT 00003513, EMUSENT 00003545, EMUSENT 00004267, EMUSENT 00003884, EMUSENT 00003966 • 38 ♂♂, 5 ♀♀; same collection data as holotype; 06 – 23. v. 2000; M. 74; IAVH • 3 ♂♂, 3 ♀♀; same collection data as preceding; EMUSENT 00003632, EMUSENT 00003932, EMUSENT 00003674, EMUSENT 00003725, EMUSENT 00004360, EMUSENT 00003944 • 19 ♂♂, 5 ♀♀; same collection data as holotype; 23. v. – 08. vi. 2000; M. 149; IAVH • 6 ♂♂, 2 ♀♀; same collection data as preceding; EMUSENT 00003788, EMUSENT 00004334, EMUSENT 00004206, EMUSENT 00004419, EMUSENT 00004385, EMUSENT 00003572, EMUSENT 00004261, EMUSENT 00004477 • 1 ♂; same collection data as holotype; SFF Iguaque, La Planada; 2850 m; 23. v. – 08. vi. 2000; M. 152; IAVH • 19 ♂♂, 6 ♀♀; same collection data as holotype; 08 – 25. vi. 2000; M. 184; IAVH • 1 ♀; same collection data as preceding; EMUSENT 00003663 • 7 ♂♂; same collection data as holotype; 13 - 30. vii. 2000; M. 380; IAVH • 10 ♂♂; same collection data as holotype; 30. vi-. 17. vii. 2000; M. 390; IAVH • 12 ♂♂, 7 ♀♀; same collection data as holotype; 01 – 23. ix. 2000; M. 614; IAVH • 1 ♂♂, 1 ♀; same collection data as preceding; EMUSENT 00003460, EMUSENT 00003733 • 31 ♂♂, 8 ♀♀; same collection data as holotype; 23. ix. – 11. x. 2000; M. 752; IAVH • 1 ♀; same collection data as preceding; EMUSENT 00004447 • 3 ♂♂; same collection data as holotype; 04 – 21. xii. 2002; M. 1080, IAVH • 6 ♂♂, 2 ♀♀; Boyacá, SFF Iguaque; 5 ° 25 ’ 12 ’’ N, 73 ° 27 ’ 24 ’’ W; 2855 m; 16. iii. – 01. iv. 2000; P. Reina; IAVH • 21 ♂♂, 9 ♀♀; Boyacá, SFF Iguaque, Cabaña Mamorramos; 5 ° 25 ’ 12 ’’ N, 73 ° 27 ’ 24 ’’ W; 2855 m; 17. viii. – 15. ix. 2000; P. Reina; IAVH • 12 ♂♂, 2 ♀♀; same collection data as preceding; 25. vi – 13. vii. 2000; IAVH • 5 ♂♂, 5 ♀♀; same collection data as preceding; 28. ii. – 16. iii. 2000; M. 4; IAVH • 2 ♂♂, 3 ♀♀; same collection data as preceding; 01 – 18. iv. 2000; M. 24; IAVH.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	distribution	Distribution. Known only from type locality (Fig. 12).	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	etymology	Etymology. Noun in apposition from Greek meaning abundance due to the large number (relatively) of specimens collected.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE705494CFF5023E6621CFE37.taxon	discussion	Comments. At the type locality, J. euthenia can be locally abundant as one Malaise trap yielded 46 males and 9 females within a three-week period.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	description	Figs 3 – 4, 12 urn: lsid: zoobank. org: act: 0 AB 93066 - 4 A 28 - 45 B 8 - A 593 - E 272 FE 2 A 3168	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	diagnosis	Diagnosis. Similar to J. euthenia, although differentiated from the latter based on the primarily brown to light brown mandible, flatter clypeus and supraclypeal area in lateral view, and subquadrate head.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	description	Description. Female (Fig. 3 A – H). Body length: 6.9 mm; fore wing length: 4.0 mm. Color. Primarily black with a few yellow and yellowish-white markings and partially brownish legs. Head black, except for mandibular brown to light brown with black base and dark brown apex, paraocular area with narrow light yellowish mark medially dorsal to antennal base and at dorsolateral corner (25 % of specimens with venterolateral corner brownish). Antenna black, except for pedicel apically brownish. Mesosoma black, except dorsal margin of pronotum and tegula yellowish-white. Fore leg: coxa varying from entirely dark brown to black, 50 % of specimens with brownish mark anterolaterally; trochanter dark brown to black with brownish apex; trochantellus brown; femur anteriorly brownish-yellow, except for basal 0.3 dark brown, posteriorly dark brown, except for apical 0.2 brownish-yellow; tibia brownish-yellow ventrally becoming brownish dorsally; tarsus dark reddish-brown, except for tarsomere 5 dark brown. Mid leg: coxa varying from black to dark brown; trochanter dark brown to black with brownish apex; trochantellus brown; femur dark brown except for anteriorly with apical 0.2 brownish-yellow, posteriorly varying from entirely dark brown to apical 0.1 brownish-yellow; tibia brownish-yellow ventrally becoming brownish dorsally; tarsus dark reddish-brown, except for tarsomere 5 dark brown. Hind leg: coxa and trochanter black; trochanter black; trochantellus brown; femur dark brown to black except for apical 0.1 brownish; tibia brownish-yellow ventrally; tibia brown to reddish-brown; tarsus varying from reddish-brown to dark reddish-brown, 50 % of specimens with tarsomeres 1 – 3 paler basally. MS 1 black. T 2 – 7 black with lateral and posterior margins brownish, 50 % of specimens with anterior margins of T 4 – 6 with brownish semi-circular marks. Wing: membrane clear; basal 0.1 of wing with veins white, remaining sections brown. Head. Subquadrate. Mandible slightly narrowed (MI: ~ 3.5 ×). Clypeus and supraclypeal area nearly flat in lateral view. Clypeus smooth and coarsely, sparsely punctate with punctures denser along margins. Supraclypeal area smooth and moderately punctate with 0.2 – 0.8 × their diameter. Gena smooth and finely punctate with punctures separated by 1.0 – 3.0 × their diameter, denser posteriorly. Supra-antennal are smooth and moderately punctate with punctures separated by 0.5 – 1.5 × their diameter. Vertex smooth and finely, sparsely punctate with punctures separated by 1.0 – 3.0 × their diameter, denser medially. Hypostomal and occipital carinae meeting before mandible. Antenna with 27 flagellomeres. Mesosoma. Of normal width, not laterally flattened. Dorsal 0.6 of pronotum smooth and finely, sparely punctate with punctures denser at margins, anterior 0.4 rugulose. Propleuron smooth and finely, sparsely punctate with punctures separated by 2.0 – 4.0 × their diameter. Mesonotum smooth, very finely, sparsely punctate anteromedially, remainder moderately punctate with punctures separated by 0.2 – 1.0 × their diameter, sparser laterally. Scutellum smooth and moderately punctate with punctures separated by 1.0 – 1.5 × their diameter. Mesopleuron smooth, dorsal 0.3 moderately punctate with anterodorsal corner rugulose, ventral 0.7 with punctation coarser and denser with some punctures forming rugulae. Speculum smooth and finely punctate. Ventral division of metapleuron smooth and coarsely, densely punctate with punctures sparser anteriorly and posteriorly at margin denser with some punctures confluent forming rugulae. Propodeum: of normal width, nearly equal in length and width; evenly rounded mediolaterally; dorsal face smooth and densely coarsely punctate laterally with punctures adjacent to subadjacent becoming sparser medially, posterior face rugulose; carinae obsolete except for pleural carina, posterior transverse carina, and median longitudinal carina posterior to posterior transverse carina subobsolete. Legs of normal length. Hind femur normal, not anteroposteriorly flattened. Metasoma. MS 1 smooth and postpetiole moderately punctate with punctate denser anteriorly and laterally and sparser medially and posteriorly. T 2 smooth and varying from finely to moderately punctate becoming sparser and finer near posterior margin. Gastrocoelus well-developed. Thyridium well-developed; narrow, 0.6 × as wide as interthyridial width; distant from T 2 anterior margin by 0.2 × tergite length. T 3 – 7 smooth and varying from moderately to finely punctate with punctation becoming sparser and finer posteriorly. Male (Fig. 4 A – C). Body length: 6.6 – 7.0 mm; fore wing length: 4.3 – 4.4 mm. As in female except for: metasoma more densely punctate with punctures separated by 0.5 – 1.0 × their diameter. Antenna with 26 – 27 flagellomeres.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	materials_examined	Material examined. Holotype: Colombia • 1 ♀; Magdalena, PNN Santa Marta, El Ramo; 2500 m; 10 ° 48; N, 73 ° 39 ’ W; 29. i. – 15. ii. 2001; J. Cantillo; M. 1283; IAVH. Paratypes: Colombia • 1 ♀; same collection data as holotype; IAVH • 1 ♂, 2 ♀♀; same collection data as holotype; 30. iii. – 05. v. 2001; 2200 m; M. 1749; IAVH • 2 ♀♀; same collection data as preceding; EMUSENT 00004004, EMUSENT 00004180 • 1 ♀; Magdalena, PNN Santa Marta, El Ramo; 2400 m; 10 ° 48 ’ N, 73 ° 39 ’ W; 25. v. – 09. vi. 2000; I. Uribe; IAVH • 1 ♂; Magdalena, PNN Santa Marta, El Ramo, La Estacion; 2200 m; 10 ° 46 ’ N, 73 ° 39 ’ W; 31. vii – 15. viii. 2000; J. Cantillo; M. 603; IVAH • 1 ♂; same collection data as preceding; 15 – 31. viii. 2000; M. 606; IAVH • 1 ♂; same collection data as preceding; 15 – 31. viii. 2000; M. 606; EMUSENT 00004559 • 2 ♂♂; same collection data as preceding; 31. viii. – 15. ix. 2000; M. 633; IAVH • 3 ♂♂; same collection data as preceding; EMUSENT 00005148, EMUSENT 00004832, EMUSENT 00004769. Non-type material: 1 ♂; Boyaca, SFF Iguaque, Lagunillas; 2855 m; 5 ° 25 ’ N, 73 ° 27 ’ W; 28. iii. - 16. iv. 2001; P. Reina; M. 1515; IAVH.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	distribution	Distribution. All but one specimen was collected at the type locality in the Sierra Nevada de Santa Marta National Natural Park (PNN Santa Marta) (Fig. 12). The only specimen not from this locality was apparently collected in the Iguaque Flora and Fauna Sanctuary (SFF Iguaque) which was extensively sampled by Malaise trapping for several years. Due to the geographic isolation of the Sierra Nevada de Santa Marta mountain range and the high levels of endemicity in the native fauna (Todd & Carriker 1922; Hernández-Camacho et al. 1992), the record from SFF Iguaque is suspect and has been excluded from the distribution map. It may be the result of mislabeling during mounting or mishandling the Malaise trap samples. Alternatively, the specimen may have indeed been collected in SFF Iguaque which further sampling of the region could corroborate.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE7024941FF5022AB64B0FEDB.taxon	etymology	Etymology. The species name refers to the type locality and honors the indigenous Kogi community that live within the Sierra Nevada de Santa Marta National Natural Park. Specifically Gonawindua is the place name used for Pico Cristóbal Colón and a general term for the surrounding region by the Kogi people.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	description	Figs 5 – 6, 11	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	diagnosis	Diagnosis. Among Jethsura species, J. pyriformis is immediately recognizable by the overall brownish-red color, whereas at least the head and mesosoma are black in other species. Jethsura pyriformis shares a striking superficial resemblance to an undescribed Stenodontus species that it co-occurs with in southeastern Arizona. Jethsura pyriformis can be differentiated from this undescribed Stenodontus by the anterior constriction of T 6 and the shorter first flagellomere in females, and the bidentate mandible with a highly reduced ventral tooth in both sexes. In Stenodontus, the mandible is always unidentate without a trace of a ventral tooth.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	description	Description. Female (Fig. 5 A – H). Body length: 8.0 – 9.9 mm; fore wing length: 4.4 – 5.3 mm. Color. Overall brownish-red with highly variable yellowish marks on mesosoma and black along mesosomal sclerite margins. Head brownish-red, except for dark red mandibular apex, medial paraocular area varying from yellowish to brownish-red and supra-antennal area infrequently (5 % of specimens) dark brown to black medially. Antenna: scape and pedicel brownish-red; F 1 / 2 usually anteriorly brownish-red and brown posteriorly, infrequently entirely black; F 2 / 3 – F 6 / 8 black; F 7 / 8 – 12 / 13 yellow-white to yellow; F 13 / 14 – 28 / 29 black. Mesosoma brownish-red, except the following areas yellowish to varying degrees: anterior margin of pronotum with yellowish color stronger anteriorly, posterodorsal corner of pronotum, subalar ridge, and scutellum; approximately 10 % of specimens with these areas entirely brownish-red. Legs brownish-red, except for approximately 20 % of specimens with fore and middle coxae yellowish anteriorly and tarsomere 5 of fore, middle and hind legs dark brown. Metasoma brownishred. Wing: membrane clear; basal 0.1 – 0.2 of wing with veins white, remaining sections brown. Head. Head subquadrate. Mandible moderately wide (MI: ~ 3.0 ×). Clypeus smooth and coarsely punctate with punctures separated by 0.5 – 1.0 × their diameter. Supraclypeal area densely, coarsely punctate with punctures separated by 0.2 – 0.5 × their diameter laterally, sparser and finer medially. Gena smooth and finely punctate with punctures separated by 1.5 – 3.0 × their diameter, denser posteriorly. Supra-antennal area smooth and densely, finely punctate with punctures separated by 0.2 – 0.8 × their diameter. Vertex smooth and finely densely punctate medially becoming sparse laterally. Hypostomal and occipital carinae meeting at or slightly before mandibular base. Antenna with 28 – 29 flagellomeres. Mesosoma. Of normal width, not laterally flattened. Dorsal 0.5 of pronotum smooth and densely punctate at margins, ventral 0.5 rugulose-punctate. Propleuron smooth and finely, densely punctate with punctures subadjacent. Mesonotum smooth, anterior 0.3 finely, densely punctate becoming denser and indistinct anteriorly, remainder densely punctate with punctures separated by 0.2 – 1.0 × their diameter. Scutellum smooth and moderately punctate with punctures separated by 0.8 – 1.5 × their diameter. Mesopleuron densely punctate with punctures adjacent to subadjacent, some confluent forming transverse rugulae, sparser on speculum. Ventral division of metapleuron densely punctate with punctures adjacent to subadjacent. Propodeum: of normal width, slightly wider than long; weakly rounded mediolaterally; dorsal face densely punctate to partially rugulose-punctate laterally becoming smooth or coriaceous anteromedially; posterior face densely rugulose-punctate; pleural carina well-developed, remainder obsolete to subobsolete except for median longitudinal carina well-developed posteriorly and posterior transverse carina well-developed laterally. Legs of normal length. Hind femur normal, not anteroposteriorly flattened. Metasoma. MS 1 smooth and varying from sparsely punctate to moderately punctate, usually with postpetiole sparser. T 2 smooth and densely punctate with punctures separated by 0.5 – 1.0 × their diameter, posterior margin impunctate. T 3 – 7 densely punctate, except for impunctate posterior margins and becoming smoother at posterior tergites. Gastrocoelus varying from shallow to well-developed. Thyridium subobsolete; narrow, 0.4 – 0.5 × as wide as interthyridial width; distant from T 2 anterior margin by 0.2 – 0.3 × tergite length. Male (Fig. 6 A – C). Body length: 7.3 – 9.6 mm; fore wing length: 4.6 – 5.9 mm. As in female, except for: head yellow with supra-antennal area black medially, gena varying from yellow with posterior margin reddish-brown to nearly entirely reddish-brown, and occiput black; yellowish-white markings on mesosoma much more extensive with the entirety of ventral and dorsal margins of pronotum, subalar ridge, and tegula yellowish-white; propleuron and anterior 0.2 of mesopleuron usually yellowish-white and propodeum usually with posterior submedial yellowish-white marks; black areas around mesosomal sutures more extensive and pronotum medially black; head wider than in female; clypeus narrower than female.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	materials_examined	Material examined. Non-type material: See supplementary file.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	distribution	Distribution. Widely dispersed in the Nearctic region from Newfoundland, Canada to Saltillo, Mexico and from the eastern seaboard to central Colorado and southeastern Arizona (Fig. 11). The presence of J. pyriformis in the Palearctic should be considered with caution. Two specimens identified by Erich Diller were reported from Hirvensalmi and Tuusula, Finland by Ranin (1982). Only the former specimen could be found at the Finnish Museum of Natural History (J. Paukkunen pers. comm.) and its identity as J. pyriformis was confirmed via examination of the specimen images. Given the occurrence of J. pyriformis as far north as Newfoundland, Canada, it is plausible that it may occur in Finland, although the lack of further specimens (E. Diller pers. comm.) collected in recent decades may indicate that the Finnish specimens were part of an adventive population that has since been extirpated.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70F4943FF5022C761EBF846.taxon	discussion	Comments. Jethsura pyriformis exhibits considerable chromatic variation in both males and females but particularly in the former. In males, the black areas along the mesosomal sutures and the yellowish-white markings are especially variable. The extent of these black and yellowish-white markings is negatively correlated. For instance, one exceptional male has a predominantly black mesosoma with the typical yellowish-white markings replaced by brownish-red markings. More frequently, males are overall lighter in color with yellowish-white markings more extensive and occasionally with yellowish-white marks appearing in irregular places like the third lateral area of the propodeum. Despite the range of J. pyriformis spanning over 25 degrees of latitude, no geographic signal in either color or size was observed.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	description	Figs 7 – 8, 11 urn: lsid: zoobank. org: act: 82464 EA 0 - 0266 - 49 CA- 89 F 5 - 07 BB 00 DC 0182	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	diagnosis	Diagnosis. Easily differentiated from other known Jethsura species by the black head and mesosoma and brownish-red metasoma.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	description	Description. Female (Fig. 7 A – H). Body length: 8.0 – 8.1 mm; fore wing length: 4.8 – 4.9 mm. Color. Black head and mesosoma with limited yellowish-white to yellowish marks and brownish-red metasoma. Head black, except for mandible dark brownish-red with darker base and paraocular area yellowish-white medially from antenatal base to median ocellus and posterodorsal corner, in one specimen the left side with paraocular area venteromedially ventral to antennal base yellowish and venterolateral corner with faint impression of yellowish mark. Antenna black. Mesosoma black, except for the following areas yellowish-white: medial 0.3 of ventral margin of pronotum, posterior 0.8 of dorsal margin of pronotum, tegula, and small mark on subalar ridge. Legs: coxae, trochanters, and trochantelli dark brown to black; fore femur anteriorly with basal 0.3 dark brown and apical 0.7 brownish-yellow, posteriorly dark brown; mid and hind femur dark brown except, for brown apex; fore and mid tibiae anteriorly brownish-yellow, posteriorly yellowish-brown to brown; hind tibia reddish-brown at base becoming dark brown apically; tarsi with tarsomeres 1 – 4 reddish-brown and tarsomere 5 dark reddish-brown, tarsomeres 1 – 4 of hind tarsus lighter basally. Metasoma reddish-brown, except for holotype with large mottled brownish mark on T 2. Wing: membrane clear; basal 0.1 – 0.3 of wing with veins white, remaining sections brown. Head. Slightly tapering ventrally. Mandible moderately wide (MI: ~ 3.1 ×). Clypeus smooth and coarsely punctate with punctures separated by 0.5 – 1.5 × their diameter. Supraclypeal area densely punctate with punctures adjacent forming transverse rugulae dorsally, punctures becoming coarser and sparser laterally. Gena smooth and finely punctate with punctures separated by 2.0 – 4.0 × their diameter, denser posteriorly. Supra-antennal are smooth and moderately punctate with punctures separated by 0.5 – 1.5 × their diameter. Vertex smooth and finely punctate with punctures separated by 1.0 – 3.0 × their diameter, denser and coarser medially. Hypostomal and occipital carinae meeting immediately before mandibular base. Antenna with 26 flagellomeres. Mesosoma. Of normal width, not laterally flattened. Pronotum smooth and moderately punctate with punctures sparser anterodorsally, coarser ventrally. Propleuron smooth and finely, densely punctate with punctures separated by 0.5 – 1.0 × their diameter. Mesonotum smooth and sparsely punctate with punctures separated by 1.0 – 3.0 × their diameter, anterior 0.1 with punctation denser. Scutellum smooth and sparely punctate with punctures separated by 2.0 – 4.0 × their diameter. Mesopleuron smooth and densely punctate, dorsal 0.3 with punctures separated by 0.5 – 1.0 × their diameter. Speculum finely and sparely punctate, ventral 0.6 denser with punctures adjacent to subadjacent and some confluent forming longitudinal rugulae. Ventral division of metapleuron smooth coarsely, densely punctate with punctures subadjacent, posteriorly some punctures confluent and forming rugulae. Propodeum: of normal width, nearly equal in length and width; dorsal face moderately convex, surface sculpture smooth and coarsely, densely punctate laterally with punctures adjacent and some confluent forming rugulae, especially posteriorly, punctation sparser medially; posterior face punctate-rugose; carinae obsolete, except for pleural carina posterior transverse carina well-developed, median longitudinal carina posterior to posterior transverse carina subobsolete. Legs of normal length. Hind femur normal, not anteroposteriorly flattened. Metasoma. MS 1 smooth, postpetiole moderately, coarsely punctate laterally and impunctate to sparsely punctate medially. T 2 smooth and finely to moderately punctate with punctures separated by 1.0 – 4.0 × their diameter, sparser posteriorly. Gastrocoelus shallow, subobsolete. Thyridium essentially obsolete, only indicated by faint impression; narrow, 0.2 – 0.3 × as wide as interthyridial width; distant from T 2 anterior margin by 0.2 × tergite length. T 3 smooth and finely, sparsely punctate. T 4 – 7 smooth and nearly impunctate except for very fine, shallow, sparse punctures. Male (Fig. 8 A – C). Body length: 8.0 mm; fore wing length: 5.5 mm. As in female, except for: yellowish-white or yellowish areas on head and mesosoma of female white in male; white marks on paraocular areas posteroventrally and posterodorsally; MS 1 smooth and finely, densely punctate with punctation on postpetiole sparser medially; T 2 – 7 finely, densely punctate with punctures separated by 0.5 – 1.0 × their diameter. Antenna with 29 flagellomeres.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	materials_examined	Material examined. Holotype: Mexico • 1 ♀; Durango, 10 mi W El Salto; 9000 ft.; 12. iv. 1964; W. R. M. Mason; CNCI. Paratypes: Mexico • 1 ♀; same collection data as holotype; 23. vi. 1964 • 1 ♂; same collection data as holotype; 07. vi. 1965.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	distribution	Distribution. Known only from type locality (Fig. 11).	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70C4945FF5023E66118FF47.taxon	etymology	Etymology. Formed by rubri- (Latin) meaning red and cauda (Latin) meaning tail and refers to the distinctive red metasoma.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	description	Fig. 9, 12 urn: lsid: zoobank. org: act: 8007935 D-B 445 - 499 E- 93 D 2 - 888 B 5 EAE 9 AF 5	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	diagnosis	Diagnosis. Jethsura serpentina and a putative undescribed species from Valle de Frailejón in Chingaza National Natural Park (Jethsura sp. X) are both characterized by an elongated habitus, broad mandible, and laterally compressed mesosoma. As such, they are easily differentiated from other Jethsura species. Jethsura serpentina is distinguished from Jethsura sp. X by the entirely smooth and coarsely punctate mesopleuron. A few punctures are nearly confluent and form very weak rugulae but the mesopleuron is never longitudinally rugulose as in Jethsura sp. X.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	description	Description. Female (Fig. 9 A – H). Body length: 8.9 – 9.4 mm; fore wing length: 5.1 – 5.4 mm. Color. Primarily black with few yellow and yellowish-white markings and partially brownish legs. Head black, except for mandibular apex dark brownish-red, light yellowish along either entire length of medial paraocular area (25 % of specimens) or interrupted ventral to antenatal base (50 % of specimens) or absent ventral to antennal base (25 % of specimens), and light yellowish ovoid to subtriangular marks at dorsolateral and venterolateral corners of paraocular area. Antenna: scape and pedicel black; flagellum black to dark brown basally, apical ¾ reddish-brown ventrally. Mesosoma black, except for the dorsal margin of pronotum, tegula, and subalar prominence yellowishwhite, 25 % of specimens with anterolateral yellowish-white marks on scutellum. Legs: coxae and trochanters black, 75 % of specimens with fore and mid coxae brownish anterolaterally; femora dark reddish-brown with bases and apices light brown and apical 0.3 – 0.4 of fore and mid femora yellowish-brown anteriorly; tibiae brownish-red to reddish-brown with fore and mid tibiae lighter anteriorly; fore and mid tarsi brownish-red to reddish-brown; hind tarsomeres 1 – 4 reddish-brown to brownish-red and tarsomere 5 dark reddish-brown with bases yellowish-white becoming reduced toward apical tarsomeres. MS 1 black. T 2 – varying from primarily black with minor, mottled light reddish-brown marks laterally to narrowly black dorsally and laterally light reddish-brown with mottled black marks. Wing: membrane clear; basal 0.2 of wing with veins white, remaining sections brown. Head. Distinctly longer than wide, not tapering ventrally. Mandible broad (MI: ~ 2.1 ×). Clypeus and supraclypeal area slightly convex in lateral view. Clypeus smooth and coarsely, sparsely punctate with punctures separated by 1.0 – 4.0 × their diameter. Supraclypeal area smooth and coarsely punctate with 0.5 – 2.0 × their diameter. Gena smooth and finely, sparsely punctate with punctures separated by 3.0 – 6.0 × their diameter. Supra-antennal are smooth and finely punctate with punctures separated by 0.5 – 1.0 × their diameter. Vertex smooth and finely, sparsely punctate with punctures separated by 3.0 – 5.0 × their diameter. Antenna with 23 – 24 flagellomeres. Mesosoma. Narrow, elongated and laterally flattened. Dorsal 0.6 of pronotum smooth and finely, sparely punctate with punctures denser at margins, anterior 0.4 rugulose. Propleuron smooth and finely, sparsely punctate with punctures separated by 3.0 – 6.0 × their diameter. Mesonotum smooth, very finely, sparsely punctate anteromedially, punctation denser and coarser laterally and especially dorsomedially. Scutellum smooth and sparsely punctate with punctures separated by 2.0 – 5.0 × their diameter. Mesopleuron smooth and punctation varying from sparse and fine anterodorsally becoming coarse and dense ventrally and posteroventrally to relatively coarse throughout with some punctures forming weak rugulae and denser ventrally and posteroventrally. Speculum smooth with at most a few scattered punctures. Ventral division of metapleuron smooth and coarsely, sparsely punctate anterodorsally becoming densely punctate posteroventrally. Propodeum: narrow and elongate, 1.4 × as long as wide; dorsal face strongly rounded transversely, surface sculpture smooth dorsally becoming coarsely punctate laterally and rugulose-punctate posterolaterally; posterior face densely rugulose-punctate; carinae obsolete. Legs short. Hind femur anteroposteriorly flattened. Metasoma. MS 1 anteriorly smooth and impunctate, postpetiole smooth and with a few scattered punctures medially becoming sparsely to moderately punctate laterally. T 2 smooth and finely to moderately punctate with punctures separated by 0.5 – 2.0 × their diameter becoming sparser laterally and posteriorly. T 3 + smooth and densely, finely punctate becoming obsolete posteriorly. Gastrocoelus shallow to moderately deep. Thyridium narrow, 0.5 × as wide as interthyridial width; distant from T 2 anterior margin by 2.0 – 2.5 × thyridial length. Male. Unknown.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	materials_examined	Material examined. Holotype: Colombia • 1 ♀; Boyaca, SFF Iguaque, Lagunillas; 3380 m; 5 ° 25 ’ N, 73 ° 27 ’ W; 09 – 24. ii. 2001; P. Reina; M. 1272; IAVH. Paratypes: Colombia • 1 ♀; same collection data as holotype • 1 ♀; same collection data as holotype; EMUSENT 000003704 • 1 ♀; same collection data as holotype; 09 – 28. iii. 2001; M. 1511.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	distribution	Distribution. Known only from type locality (Fig. 12).	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	etymology	Etymology. The species name is from the Latin adjective “ sepentinus ” meaning serpent-like and refers to the somewhat snake-like habitus.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
0627421AE70A4946FF5027E06195FAB3.taxon	discussion	Comments. The male of J. serpentina is unknown but could presumably be easily associated with the female at the type locality since J. serpentina and the putative undescribed species, Jethsura sp. X, are allopatric. Also, presumably the male of J. serpentina would be considerably elongated as in the female which would readily distinguish it from J. euthenia males.	en	Claridge, Brandon (2021): Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini). Zootaxa 5071 (2): 223-241, DOI: https://doi.org/10.11646/zootaxa.5071.2.3
