taxonID	type	description	language	source
65B3116808FB5E47818C063D4713BBAF.taxon	materials_examined	Material examined. Adult male from Argentina under label " Tucuman, 19 / 4 / 59, entre maderas ... ", deposited in the Museum national d'Histoire naturelle (MNHN), Paris, France. This individual was obviously among the material the original description was based on by E. H. Rapoport.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
65B3116808FB5E47818C063D4713BBAF.taxon	diagnosis	Diagnosis. Maxillary palp bifurcate, four prelabral chaetae. Dens with ~ 50 anterior chaetae. Mucro bidentate. Anterior side of manubrium with 3 + 3 chaetae. 33 / 22235 s and 11 / 111 ms on body. Ventral chaetae absent on Th. III.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
65B3116808FB5E47818C063D4713BBAF.taxon	description	Description. Maxillary outer lobe with four sublobal hairs, maxillary palp bifurcate. Labral formula as 4 / 5,5,4. Guard chaeta e 7 present on labium. Ant. III without bms and with five distal s (including one lateral), without additional s-chaetae. Th. I, II, and III without ventral chaetae. S-formula as 33 / 22235 (s), 11 / 111 (ms). Tibiotarsal tenent chaetae (1,1,1) clearly clavate. Tibiotarsi 1 - 2 with more than 24 chaetae. Ventral tube with 6 + 5 chaetae (in the adult male studied). Retinaculum with 4 + 4 teeth and two chaetae. Dens long and slender, with numerous crenulations, many (~ 50) anterior and 17 (in the adult male studied) posterior chaetae. Anterior side of manubrium with 3 + 3 chaetae. Mucro bidentate.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
65B3116808FB5E47818C063D4713BBAF.taxon	discussion	Discussion. Our redescription is based on one individual of E. Rapoport, and more material is needed to complete the understanding of B. biseta. The species obviously belongs to the genus Bagnallella by separation of two last abdominal segments and s-chaetotaxy of Abd. IV and V. After chaetotaxy of tibiotarsi, ventral tube, and dens, B. biseta appears to be a more polychaetotic species than its congeners. The close relation of B. biseta and B. tenella (sharing 3 + 3 manubrial chaetae) is doubtful due to difference in maxillary palp (bifurcate vs simple). The independence of B. koepckei and B. paronai from B. biseta call for further verification. Bagnallella paronai is not included in the key due to the incomplete diagnosis.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
65B3116808FB5E47818C063D4713BBAF.taxon	distribution	Distribution. Argentina and Chile (see Mari Mutt and Bellinger 1990 for details).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	description	Figures 9 - 14, 15 - 19, 20, 21 - 24	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	materials_examined	Material examined. Typical form: South Africa, Free State Province, Bankfontein Farm, 30.0567 ° S, 24.8942 ° E, 24. iv. 2019, Berlese-Tullgren: tree leaf litter, H. Badenhorst leg.; South Africa, Western Cape, Haarwegskloof, Swellendam, 34.3422 ° S, 20.3169 ° E, 18. vii. 2017, litter trap (G 18) with Pentameris eriostoma litter, O. Cowan leg., South Africa, SAF 583 (11. m CJ SWB); Prince Albert: Swartberg North: road to Swartberg Pass, 11 / 03 / 2019, meadow, moss, C. J. leg. Short-haired form: South Africa, Western Cape, Haarwegskloof, Swellendam, 34.3534 ° S, 20.3042 ° E, 18. vii. 2017, litter trap (A 4) with Medicago sativa litter, O. Cowan leg.; Cederberg Wilderness area, Litter trap CED 588; South Arica, Western Cape, Cederberg Wilderness area, Litter trap CED 394; Jonkershoek Nature Reserve, 33.9891 ° S, 18.9575 ° E, 05. ix. 2011, Litter trap (J 4, 124); Jonkershoek Nature Reserve, 33.9891 ° S, 18.9575 ° E, 30. vii. 2009, Litter trap, C. J. leg.; J 2, 32.1; Landdroskop, Jan. 2012, H. Basson leg.; Prince Albert, Swartberg North, Swartberg crest, 12 / 03 / 2019, SAF- 612, SAF- 618, meadow, litter and soil, L. D., C. J. and A. B. leg. Intermediate form: South Africa, Western Cape, Haarwegskloof, Swellendam, 34.3345 ° S, 20.3187 ° E, 18. vii. 2017, litter trap (R 24) with Dicerothamnus rhinocerotis litter, O. Cowan leg.; Prince Albert, Swartberg North, Swartberg crest, SAF- 612, 12 / 03 / 2019, meadow, litter, L. D., C. J. and A. B. leg.; Prince Albert, Swartberg North, Swartberg crest, SAF- 618, 12 / 03 / 2019, meadow, soil, L. D., C. J. and A. B. leg.; Prince Albert, Swartberg North, road to Swartberg Pass, SAF- 601, 12 / 03 / 2019, meadow, litter and soil, L. D., C. J. and A. B. leg.; Prince Albert, Swartberg North, Swartberg crest, SAF- 614, 12 / 03 / 2019, moss on rock, Berlese, L. D., C. J. and A. B. leg. " Clasping supermales ": SAF- 601, South Africa: Western Cape: Prince Albert: Swartberg North: road to Swartberg Pass, 12 / 03 / 2019, meadow, litter and soil, L. D., C. J. and A. B. leg. " Spiny supermales ": SAF- 554; South Africa: Western Cape: Kalk Bay: Echo Valley: Spes Bona forest, 01 / 03 / 2019, Afromontane forest, moss and lichen on rock, L. D. and A. B. leg.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	diagnosis	Diagnosis. 3 + 3 postlabial chaetae. Maxillary palp simple. Dens with four anterior and four posterior chaetae. Mucro tridentate, teeth arranged in a line. Anterior side of manubrium with 1 + 1 chaetae. 43 / 22235 s and 10 / 100 ms on body (Figs 9 - 11). No ventral chaetae on Th. III. Typical form of species with long macrochaetae (Fig. 10).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	description	Description. Maxillary outer lobe with four sublobal hairs, maxillary palp simple. Labral formula as 4 / 5,5,4. Labium with five usual papillae (- E) and labial formula A 1 B 4 C 0 D 4 E 6, guard chaeta e 7 absent, three proximal and four basomedian chaetae. Ventral side of head with 3 + 3 chaetae. PAO shorter than Ant. I width (0.6 - 0.8). Ant. I with 11 common chaetae, two ventral s-chaetae (s) and three bms, of which one dorsal long, chaeta-like (this ms was calculated together with common chaetae in first description, 12 at whole); Ant. II with three bms and one latero-distal s; Ant. III with one bms and with six distal s (including two lateral), without additional s-chaetae. Organite pin-like, not very small. Empodial appendage about half as long as Claw. Anterior and posterior furcal subcoxae with 9 - 14 and 7 - 8 chaetae, respectively. Male spurs on tibiotarsi 3 thin, stick-like. Th. I-III without ventral chaetae. Ratio manubrium: dens: mucro as 4.4 - 5.0: 3.3 - 3.8: 1; dens: Claw as 3.3 - 3.6 (for the typical form).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	discussion	Discussion. Bagnallella davidi is a rather peculiar species of the genus due to few chaetae on dens (vs many more chaetae both on anterior and posterior sides), tridentate mucro (shared with B. mishai only), and 3 + 3 postlabial chaetae (vs 4 + 4 or more in other species). The first description is almost complete and, therefore, we made very few additions concerning mouth parts. The species exhibits high variation in length of macrochaetae and show different modifications of males. All the forms (described below) can indicate either high plasticity of a single species or the complex of separate although closely related species, calling for further morphological, biological, ecological, and molecular investigations.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
A32B9BE7D66F5818AB4DBE40E47FEB79.taxon	distribution	Distribution. Eastern Cape, Amatola Mountains (type location) and widely in the Western Cape and Free State (our material) provinces of South Africa.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
0639924F46AD59378DDBBC24B98846E2.taxon	discussion	Discussion. The species can be attributed to Bagnallella by the two key characters of the genus. It is characterized by 33 / 22235 s, 11 / 111 ms, bifurcate maxillary palp, long polychaetotic dens (34 - 35 / 14), and bidentate mucro. The presence of seven ocelli and 2 + 2 chaetae on anterior side of manubrium are two unique characters among members of the genus.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
0639924F46AD59378DDBBC24B98846E2.taxon	distribution	Distribution. This species is currently only known from SE Brazil.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	description	Figures 3, 4	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	materials_examined	Material examined. New Zealand, South Island, Central Otago, Pisa Range and Old Man's Range, high alpine zone, different sites, 17.02.2014, M. Minor leg.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	diagnosis	Diagnosis. Maxillary palp bifurcate, two prelabral chaetae. Dens with 12 - 16 anterior chaetae. Mucro bidentate. Anterior side of manubrium with 1 + 1 chaetae. 33 / 22235 s and 10 / 100 ms on body. 2 + 2 ventral chaetae on Th. III.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	description	Description. Colour grey. Cuticle, ocelli, outer mouth parts, and antennae as in B. sedecimoculata. PAO as long as 0.8 - 0.9 Ant. I and as 1.4 - 1.5 as long as Claw III. Ventral side of head with 4 + 4 postlabial chaetae. Th. III with 2 + 2 ventral axial chaetae. Macrochaetae weakly differentiated, medial ones on Abd. V about as long as 0.4 - 0.5 of tergal midline. S-chaetae weakly differentiated. S-formula as 43 / 22235 (s), 10 / 100 (ms) (Fig. 4). S-chaetae on Abd. I-III in mid-tergal position. Tibiotarsi 1 - 2 with 21 chaetae, Tibiotarsi 3 with few additional chaetae. Tibiotarsal tenent chaetae not developed. Ventral tube with 4 + 4 laterodistal and usually with five posterior chaetae. Retinaculum with 4 + 4 teeth and two chaetae. Anterior furcal subcoxae with 13 - 15 chaetae, posterior ones with 7 - 9. Anterior side of manubrium with 1 + 1 distal chaetae. Dens with 12 - 16 anterior chaetae, posterior side of dens with crenulation and seven chaetae (Fig. 3). Mucro bidentate. Ratio of manubrium: dens: mucro = 6.0 - 6.7: 5.0 - 6.0: 1. Males present, with two thin spurs on Tibiotarsi I.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	discussion	Discussion. This species was named after its dubious position in generic system of Proisotoma / Cryptopygus (Deharveng 1981). It resembles B. sedecimoculata (see the Discussion below) and apparently belongs to the genus Bagnallella by separation of two last abdominal segments and s-chaetotaxy of Abd. IV and V. Our specimens from New Zealand match the first description.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
7F0F127FD59350F98D408961E0A269E3.taxon	distribution	Distribution. Bagnallella dubia was described from Marion Island and recorded in Macquarie Island (Greenslade and Wise 1986) and alpine sites of New Zealand (Babenko and Minor 2015). The species is possibly widely distributed in cold sites of high altitudes of the Southern Hemisphere. Its occurrence in Australia (Greenslade 2006) needs to be verified.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
BA7B4AC4AE5D54658A9A392254AB49C6.taxon	discussion	Discussion. It is an easily recognizable species by 43 / 22235 s, 11 / 111 ms, simple maxillary palp, dens (15 - 16 / 12 - 13), and tridentate mucro.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
BA7B4AC4AE5D54658A9A392254AB49C6.taxon	distribution	Distribution. One locality in southeastern Brazil.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
10A2360F8B8A57448C21FD4084B80CFE.taxon	diagnosis	Diagnosis. Maxillary palp bifurcate, four prelabral chaetae. Ant. I with many additional chaetae. Dens long and slender, with 20 - 30 anterior chaetae or more. Mucro bidentate. Anterior side of manubrium with 1 + 1 chaetae. 33 / 22235 s and 11 / 111 ms on body. Lateral s-chaetae on Abd. IV shifted to ventral side. Without ventral chaetae on Th. III.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
10A2360F8B8A57448C21FD4084B80CFE.taxon	discussion	Discussion. The full redescription is given by Fjellberg (2007).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
10A2360F8B8A57448C21FD4084B80CFE.taxon	distribution	Distribution and ecology. Europe. It prefers sandy places along the edge of water.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
FA6E65F7465E5545A23A15E1F95B8A1A.taxon	diagnosis	Diagnosis. Anurophorinae with all abdominal segments clearly separated and a Proisotoma - like furca: manubrium with few anterior chaetae (1 + 1 - 3 + 3), dens slender, crenulated, with rather numerous anterior and posterior chaetae, mucro clearly set off from dens, with two or three teeth. 7 + 7 - 8 + 8 ocelli in known species. With simple or bifurcate maxillary palp and four sublobal hairs, two or four prelabral chaetae. Macro s-chaetae 22235 on Abd. I-V. Tergal macro s-chaetae on abdomen situated in front of p-row of chaetae. B-row of chaetae on Ti. 1 - 2 complete (both B 4 and B 5 present). Ventral chaetae on Th. III present or absent. Sexual dimorphism present or absent.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
FA6E65F7465E5545A23A15E1F95B8A1A.taxon	distribution	Distribution and ecology of Bagnallella. The genus is distributed worldwide. More local species (B. davidi, B. douglasi, B. mishai, B. biseta, B. koepckei) are distributed in the Southern Hemisphere that indicates further relation to the " austral " genus Cryptopygus. Due to apparent ruderal B. tenella and pioneer B. ripicola, the genus also occurs in the Northern Hemisphere. The ecology of the former species is especially remarkable, as B. tenella is often recorded in mass abundances (Agrell 1939; Arle 1970; Neves and Mendonca 2016). We speculate that B. davidi has an unusual biology and ecology due to its morphological modifications (described below).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
FA6E65F7465E5545A23A15E1F95B8A1A.taxon	description	Descriptions and remarks to species of the genus. Below we give the diagnosis, description, remarks, and distribution, with, if possible, ecological remarks of all species of Bagnallella. Some sections may be omitted if the associated species have good morphological descriptions in the literature.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	description	Figure 2	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	materials_examined	Material examined. South Africa, Western Cape, Stellenbosch, Jonkershoek Nature Reserve, canyon to waterfall, SAF- 086, 34.005570 ° S, 18.992067 ° E, 15 / 03 / 2008, forest litter, Berlese, L. D. and A. B. leg.; Somerset, Helderberg, SAF- 107, SAF- 109, SAF- 116, 34.040883 ° S, 18.873649 ° E, alt. 600 m, 04 / 03 / 2009, native forest litter, L. D. and A. B. leg.; Cape Town, Wynberg, Table Mountain, second collapse, SAF- 141, 33.987637 ° S, 18.405750 ° N, alt. 725 m, 10 / 03 / 2009, native forest litter, L. D. and A. B. leg.; Constantia, Silvermine, in a small forest patch above Tokai, SAF- 235, 34.038273 ° S, 18.395478 ° E, alt. 390 m, 06 / 11 / 2010, dead wood, D. Porco leg.; Kalk Bay, Echo Valley, Spes Bona forest, SAF- 555, 01 / 03 / 2019, Afromontane forest, moss on rock, L. D. and A. B. leg. New Zealand. NZL- 049, Rotoiti: Lakes Rototongata and Rotoatua, 08 / 01 / 1996, primary forest, litter, L. D. and A. B. leg. Australia. Victoria, July 2010, University Ballarat, St. Helens, 37.629979 ° S, 143.890801 ° E, Eucalyptus plantation, moss, P. Greenslade. leg. Macquarie Island, Bauer Bay, 54.5549 ° S, 158.8760 ° E, April 2016, Turf sample, Berlese extraction, L. Phillips leg.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	diagnosis	Diagnosis. Maxillary palp bifurcate, two prelabral chaetae. Dens slender, with 16 - 20 anterior chaetae. Mucro bidentate. Anterior side of manubrium with 1 + 1 chaetae. 33 / 22235 s and 10 / 100 ms on body. Without ventral chaetae on Th. III.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	description	Description. Colour grey. Cuticle outwardly smooth. 8 + 8 ocelli, G and H smaller. PAO about as long as 0.8 Ant. I and as 1.1 - 1.3 Claw III. Maxillary outer lobe with four sublobal hairs and bifurcate maxillary palp. Labral formula as 2 / 554. Labium full set of guards (e 7 present), three proximal and four basomedian chaetae. Ventral side of head with 4 - 5 + 4 - 5 postlabial chaetae. 11 chaetae on Ant. I, with three basal micro s-chaetae (bms), of which one bms large, and two ventral s-chaetae (s), Ant. II with three bms and one laterodistal s, Ant. III with one bms and six distal s (including two lateral). Thorax without ventral axial chaetae. Macrochaetae weakly differentiated, medial ones on Abd. V as long as 0.4 - 0.5 of tergal midline. S-chaetae weakly differentiated. S-formula as 43 / 22235 (s), 10 / 100 (ms). S-chaetae on Abd. I-III in mid-tergal position. General pattern of chaetotaxy as in B. dubia (Fig. 4). Ti. 1 - 2 with 21 chaetae, Ti. 3 with few additional chaetae. Tibiotarsal tenent chaetae not developed. Ventral tube with 4 + 4 laterodistal and usually with six posterior chaetae (four in a transversal row). Retinaculum with 4 + 4 teeth and two chaetae. Anterior furcal subcoxae with 11 - 15, posterior ones with 7 - 9 chaetae. Anterior side of manubrium with 1 + 1 distal chaetae (Fig. 2). Dens with 17 - 19 (16 - 20) anterior chaetae, posterior side of dens with crenulation and seven chaetae (Fig. 2). Mucro bidentate. Ratio of manubrium: dens: mucro = 5.3 - 6.3: 5.9 - 6.7: 1.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	discussion	Discussion. Bagnallella sedecimoculata was described from New Zealand and was designated as type species for the genus Bagnallella (see the discussion to the genus above). The type specimen from New Zealand was studied (Potapov and Janion-Scheepers 2017), although only one generic character (separation of two abdominal segments) was proven. The redescription given above is based mostly on the South African material which looks conspecific to one individual in hand from New Zealand (L. D. and A. B. leg.). Bagnallella sedecimoculata resembles B. dubia and differs by ventral chaetae on Th. III (absent vs present) and a few more anterior chaetae on dens (16 - 20 vs 12 - 16). The latter character is not stable enough to separate the two species.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
758841A83C8C512DBB5161DBE84910CD.taxon	distribution	Distribution. Probably widely distributed. So far with scattered records in the South Hemisphere (New Zealand, Australia, South Africa, Macquarie Island).	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	description	Figures 5 - 8	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	materials_examined	Material examined. South Africa, Western Cape, Haarwegskloof, Swellendam, 34.3425 ° S, 20.3167 ° E, 18. vii. 2017, litter trap (R 17) with Dicerothamnus rhinocerotis litter, O. Cowan leg.; Haarwegskloof, Swellendam, 34.3444 ° S, 20.3225 ° E, 18. vii. 2017, litter trap (A 17) with Medicago sativa litter, O. Cowan leg.; Eastern Cape, Baviaanskloof, 33.7311 ° S, 23.9655 ° E, 24. iv. 2013, BAV _ F _ 49, A. Liu leg.; Free State Province, Bankfontein Farm, 30.0567 ° S, 24.8942 ° E, 24. iv. 2019, Berlese-Tullgren, tree leaf litter, H. Badenhorst leg. Cuba, Cienfuegos Province, 1984, J. Banasco-Almenteros leg. Indonesia, Lombok Island, vic. Toko Nusa Sari, marine beech, 8.7411 ° S, 116.0011 ° E, 4. IV. 2017, V. Makarov leg. Brazil, Espirito Santo, Domingos Martins, Arace, 4. VII. 2000, M. Culik leg.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	diagnosis	Diagnosis. Maxillary palp simple. Four prelabral chaetae. Dens slender, with ca 20 - 30 anterior chaetae. Mucro bidentate. Anterior side of manubrium with 3 + 3 chaetae. 33 / 22235 s and 11 / 111 ms on body. No ventral chaetae on Th. III.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	description	Description. Colour grey of different intensity. Cuticle outwardly smooth. 8 + 8 ocelli, G and H smaller. PAO (Fig. 6) about half as long as width of Ant. I and 0.8 - 0.9 as long as Claw 3. Maxillary outer lobe with four sublobal hairs and simple maxillary palp. Labral formula as 4 / 554. Labium without guards e 7, with three proximal and four basomedian chaetae. Ventral side of head with 4 - 5 + 4 - 5 postlabial chaetae. With several additional chaetae on Ant. I. Ant. I with three basal micro s-chaetae (bms), one dorsal large, Ant. II with three bms, Ant. III without bms. Thorax without ventral axial chaetae. Macrochaetae rather long, differentiated (11 / 3334), medial ones on Abd. V about as long as tergal midline. S-chaetae on tergites slightly shorter than ordinary chaetae. S-formula as 33 / 22235 (s), 11 / 111 (ms) (Figs 5, 8). S-chaetae subequal, on Abd. I-III in mid-tergal position. Micro s-chaetae on Abd. I-II in front of lateral s-chaetae, on Abd. III between medial and lateral s-chaetae (Fig. 8). Tibiotarsal tenent chaetae (1,1,1) often present, weakly clavate (see Discussion). Retinaculum with 4 + 4 teeth and 1 - 2 chaetae. Furca long. Anterior side of manubrium with 3 + 3 chaetae arranged in two lines (Fig. 7). Dens with more than 20 anterior chaetae, posterior side of dens with crenulation and 7 - 10 chaetae (Fig. 7). Mucro bidentate. Ratio of manubrium: dens: mucro = 7 - 8: 9: 1.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	discussion	Discussion. The species belongs to the genus Bagnallella by having two last abdominal segments separated and three and five s-chaetae on Abd. IV and V. It shares 3 + 3 anterior chaetae on manubrium with B. biseta, B. koepckei, and B. paronai from South America from which it differs by fewer chaetae on dens. Number of posterior chaetae on dens, clavate tibiotarsal chaetae, and chaetae on retinaculum vary. We treat all this variation within one species, but further study is needed. The independence of Proisotoma nigromaculosa (Hawaiian Islands) is doubtful. Although we have no material from Europe, Stach's (1947) concept of P. tenella based on the specimens from Poland is accepted by us (for details see Potapov 2001; Fjellberg 2007). Our tropical material fits Stach's (1947) descriptions.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
06F19A81B54A58ED88365E6555B4317F.taxon	distribution	Distribution and ecology. Widely distributed cosmopolitan species. Common in tropics. In higher latitudes only in protected soils.	en	Potapov, Mikhail, Deharveng, Louis, Janion-Scheepers, Charlene (2021): Taxonomy of the Proisotoma complex. VI. Rediscovery of the genus Bagnallella Salmon, 1951 and epitoky in Bagnallella davidi (Barra, 2001), comb. nov. from South Africa. ZooKeys 1072: 185-204, DOI: http://dx.doi.org/10.3897/zookeys.1072.71307, URL: http://dx.doi.org/10.3897/zookeys.1072.71307
