taxonID	type	description	language	source
03FB8799FF8AD5780580FF6E0C11F82C.taxon	etymology	Etymology: — clavulus (Latin) refers to basidiomata of this species look like a tack nailed to the ground. Holotype: — CHINA, Yunnan Province, Kunming city, Yeya Lake, 28 July 2014, broadleaf forests, dominated by Quercus, 25 ° 07 ′ 34 ″ N; 102 ° 51 ′ 42 ″ E, 2126 m asl., leg. LHJ 14072805 (RITF 3334). Diagnosis: — Basidiomata medium-sized to large; pale to pale yellow pileus centre with small brown spots, margin with tuberculate striation up to the 1 / 3 of the radius; lamellae with some small pale yellow spots; white to yellowish spore print; subglobose to broadly ellipsoid spores with short or long ridges; hymenial cystidia on lamellae sides that are mainly subclavate or fusiform and apically often obtuse or acute; pileocystidia near the pileus margin always long and one-celled, turning light purple in SV. Description: — Basidiomata medium-sized to large, 70 – 92 mm in diam., broadly convex to applanate with a slightly depressed centre; margin incurved, rarely cracked, tuberculate striation up to the 1 / 3 of the radius; cuticle smooth, glabrous, viscous when humid, not peeling readily; pale (2 A 2) to pale yellow (4 A 3) with small brown spots in the cap centre; margin white (1 A 1), buttercup yellow (4 A 5) in some part. Lamellae 2 – 4 mm deep, relatively dense, adnate to slightly free, white (1 A 1) to pale (2 A 2), with some small pale yellow (4 A 3) spots; lamellulae rarely or absent; occasionally forked and slightly interveined near the stipe. Stipe 80 – 95 × 10 – 25 mm, central, subcylindrical to cylindrical, slightly expanded towards the base, surface dry, rugulose longitudinally, white to whitish with or brownish spot, pithy and hollow. Context 2 – 3 mm thick at pileus centre, white to cream, unchanging when bruised; taste mild to slightly acrid; odour indistinct. Spore print white to yellowish. Spores (5.6 –) 7.0 – 7.8 – 8.6 (– 9.0) × (5.4 –) 6.5 – 7.2 – 8.0 (– 8.3) μm, Q = (1.0 –) 1.03 – 1.09 – 1.14 (– 1.24), subglobose to broadly ellipsoid; ornamentation of large, distant to moderately distant (3 – 6 in a 3 μm diam. circle) amyloid warts, up to 1.6 μm high, isolated or occasionally fused (0 – 2 fusions in the circle), often forming short or long ridges [1 – 3 fusions in the circle], line connections absent; suprahilar spot indistinct. Basidia (30.6 –) 43.6 – 48.8 – 54 (– 57.7) × (9.6 –) 11.5 – 12.9 – 14.2 (– 15.5) μm, mostly 4 - spored, some 2 - and 3 - spored, subclavate to clavate, some with granulose contents, rarely containing oil droplets; basidiola clavate, ca. 10 – 18 μm wide. Hymenial cystidia on lamellae sides moderately numerous, ca. 1000 / mm 2, (45.2 –) 48 – 53.4 – 58.8 (– 65) × (5.8 –) 7.2 – 10 – 13 (– 16) μm, subclavate or fusiform, apically often obtuse or acute, sometimes moniliform, occasionally with 2 – 8 μm long appendage, thin-walled; contents abundant granulose or flocculent, turning to blackish-grey in SV. Hymenial cystidia on lamellae edges often longer and narrower, (47 –) 52.5 – 58.8 – 65.2 (– 75) × (5.4 –) 6 – 7.8 – 9.6 (– 13.7) μm, narrowly clavate to subcylindrical, apically obtuse or mucronate, sometimes with 2 – 5 μm long appendage, contents granulose, turning to blackish-grey in SV. Marginal cells (9.3 –) 11.8 – 15 – 18.3 (– 21.5) × (4 –) 4.8 – 5.6 – 6.5 (– 7.3) μm, clavate or subcylindrical, sometimes flexuous. Pileipellis orthochromatic in Cresyl blue, not sharply delimited from the underlying context, 185 – 255 μm deep, two – layered; suprapellis 65 – 75 μm deep, composed of ascending or erect hyphae terminations; subpellis 120 – 180 μm deep, composed of strongly gelatinised horizontally narrow hyphae. Hyphal terminations near the pileus margin rarely branched, sometimes flexuous, thin – walled; terminal cells (12 –) 13.7 – 24.7 – 35.7 (– 60.7) × (2.5 –) 3.2 – 4.3 – 5.3 (– 7) μm, mainly clavate, occasionally subcylindrical, apically obtuse; subterminal cells often narrower, ca. 2 – 5 μm wide, usually unbranched. Hyphal terminations near the pileus centre less flexuous; shorter terminal cells (8.9 –) 13.4 – 20.6 – 27.8 (– 37.6) × (2.7 –) 3.6 – 4.6 – 5.6 (– 9.3) μm, lageniform or subcylindrical, apically obtuse or occasionally attenuated; subterminal cells often shorter, usually unbranched, ca. 2.5 – 6 μm wide. Pileocystidia near the pileus margin always long one-celled, (38.7 –) 76 – 129 – 182 (– 200) × (3.9 –) 4.4 – 5.8 – 7 (– 9) μm, cylindrical to clavate, apically usually obtuse or occasionally mucronate, contents dense granulose, turning to brownish – black in SV. Pileocystidia near the pileus centre often shorter, similar in shape and contents, (38.2 –) 59 – 74 – 89 (– 100) × (5 –) 5.5 – 6.4 – 7.3 (– 9) μm, contents turning to brownish – black in SV. Cystidioid hyphae in subpellis and context, contents granulose. Oleiferous hyphae in context, with oil contents. Additional specimens examined: — CHINA, Guizhou Province, Zunyi city, Wuchuan County, Fenshui Town, Jinhua Village, 22 September 2014, 28 ° 53 ′ 29 ″ N, 107 ° 59 ′ 7 ″ E, 1072 m asl., leg. LHJ 14092226 (RITF 3300); ibid., Southwest Guizhou Buyi and Miao Autonomous Prefecture, Wangmo County, Wangmo Nature Reserve, 23 August 2013, 25 ° 12 ′ 34 ″ N; 106 ° 15 ′ 48 ″ E, 1272 m asl., leg. Wu 481 (RITF 3216); Yunnan Province, Kunming city, Qiongzhu Temple, 28 July 2012, 25 ° 03 ′ 55 ″ N; 102 ° 37 ′ 28 ″ E, 2153 m asl., leg. R 5 (RITF 3204).	en	Chen, Bin, Song, Jie, Zhang, Jin-Hua, Liang, Jun-Feng (2021): Morphology and molecular phylogeny reveal two new species in Russula sect. Ingratae from China. Phytotaxa 525 (2): 109-123, DOI: 10.11646/phytotaxa.525.2.2
03FB8799FF87D5750580FF040C95F81F.taxon	etymology	Etymology: — multilamellula (Latin), referring to the species has many lamellulae. Holotype: — CHINA. Guizhou Province, Zunyi city, Daozhen County, Yuxi Town, 24 September 2014, on the ground in Pinus forest, 28 ° 53 ′ 26.22 ″ N, 107 ° 36 ′ 31.05 ″ E, 819 m asl., leg. LHJ 14092423 (RITF 3346). Diagnosis: — Basidiomata medium – sized; brownish orange to hazel centre, margin satin white to yellowish-white with brownish tinge; lamellulae often present, usually irregular in length and often anastomosing with lamellae; lily white stipe with brownish spot at the base, slightly curved towards centre; subglobose to broadly ellipsoid spores with short or long ridges; hymenial cystidia apically mainly obtuse, contents granulose or crystalline, turning to yellowishbrown in SV; marginal cells sometimes flexuous; Hyphal terminations near the pileus margin sometimes flexuous, terminal cells apically usually obtuse, occasionally attenuated; pileocystidia always one-celled, apically often obtuse, contents turning light purple in SV. Description: — Basidiomata medium-sized, 50 – 75 mm in diam., hemispheric to planoconvex with a slightly depressed centre above the stipe; margin incurved, tuberculate striation up to the 1 / 3 of the radius; cuticle smooth, glabrous, slightly viscid when wet, peeling readily; brownish orange (6 C 8) to hazel (6 E 7) at the centre, margin satin white (2 A 1) to yellowish-white (3 A 2) with brownish tinge. Lamellae adnate to adnexed, relatively dense, up to 6 mm deep, lily white (1 A 1) to satin white (2 A 1); lamellulae often present, (0) 1 – 3 between each pair of the lamellae, usually irregular in length and often anastomosing with lamellae; furcations absent or rare; edge entire and concolour. Stipe 50 – 60 × 12 – 16 mm, central to subcentral, subcylindrical to cylindrical, slightly curved towards centre, slightly inflate near the base, rugulose longitudinally, surface dry, lily white (1 A 1) with brownish spot at the base, pithy and hollow. Context 2 – 3 mm thick at pileus centre, white to cream, without colour changing when bruised; tastes lightly bitter; odour indistinct. Spore print white to cream. Spores (5.6 –) 6.3 – 6.9 – 7.4 (– 8) × (5 –) 5.8 – 6.4 – 7 (– 7.3) μm, Q = (1.0 –) 1.01 – 1.07 – 1.13 (– 1.25), subglobose to broadly ellipsoid; ornamentation of large, moderately distant to dense (4 – 7 in a 3 μm diam. circle) amyloid warts, up to 1.5 μm high, occasionally fused in long chains (0 – 2 fusions in the circle), often forming short or long ridges (2 – 3 fusions in the circle), line connections absent; suprahilar spot large, inamyloid. Basidia (38.8 –) 45.3 – 52.9 – 60.4 (– 85) × (5.6 –) 10.8 – 13.7 – 16.6 (– 18.2) μm, mostly 4 - spored, some 2 - and 3 - spored, subclavate or clavate; basidiola subcylindrical or clavate, ca. 7 – 15 μm wide. Hymenial cystidia on lamellae sides moderately numerous, ca. 1000 / mm 2, (61.6 –) 73.5 – 84.5 – 95.5 (– 115) × (6.9 –) 10.2 – 12.6 – 15 (– 18.8) μm, clavate or subcylindrical, apically obtuse and mucronate, often with 2 – 6 μm long appendage, thin-walled, contens granulose or crystalline, turning to yellowish-brown in SV. Hymenial cystidia on lamellae edges (43 –) 54.7 – 67 – 79.3 (– 88.5) × (5 –) 8.6 – 11.3 – 14 (– 20) μm, often smaller, clavate or fusiform, apically mainly obtuse, occasionally mucronate, sometimes with 2 – 5 μm long appendage, contents granulose or crystalline, thin-walled, turning to yellowish-brown in SV. Marginal cells (10 –) 12.3 – 16.7 – 21 (– 28) × (3 –) 4.4 – 5.4 – 6.5 (– 8) μm, narrowly clavate or fusiform, often flexuous. Pileipellis orthochromatic in Cresyl blue, not sharply delimited from the underlying context, 180 – 260 μm deep, two-layered, weakly gelatinized; suprapellis 120 – 160 μm deep, consisting of erect or repent hyphal terminations; subpellis 80 – 120 μm deep, consisting of horizontally orientated, relatively dense, intricate, 2 - 6 μm wide hyphae. Hyphal terminations near the pileus margin occasionally branched, sometimes flexuous, thin-walled; terminal cells (12.9 –) 17.3 – 24.6 – 32 (– 40.3) × (3 –) 3.5 – 4 – 4.7 (– 5.6) μm, mainly cylindrical, rarely lageniform or fusiform, apically usually obtuse but occasionally attenuated; subterminal cells often shorter and wider, ca. 3 – 7 μm wide, unbranched or occasionally branched. Hyphal terminations near the pileus centre less flexuous; shorter terminal cells (8.4 –) 9.5 – 13.6 – 17.6 (– 23.6) × (2.4 –) 3.4 – 4.3 – 5.3 (– 7.4) μm, mainly cylindrical and clavate, apically usually obtuse and rarely attenuated; subterminal cells usually wider, ca. 3 – 6 μm wide, unbranched or rarely branched. Pileocystidia near the pileus margin always one – celled, (40.7 –) 45.3 – 59.3 – 73.3 (– 90) × (3.6 –) 4.2 – 5 – 6 (– 7) μm, narrowly clavate to subcylindrical, often flexuous, apically usually obtuse, with granulose contents, turning light purple in SV. Pileocystidia near the pileus centre often longer and wider, (53.5 –) 61.5 – 77.4 – 93.4 (– 120) × (4.8 –) 5.6 – 9.2 – 12.9 (– 18.8) μm, usually clavate or subcylindrical, often flexuous, apically obtuse and with similar contents, turning light purple in SV. Cystidioid hyphae in subpellis and context, with granulose contents. Oleiferous hyphae abundant in subpellis, with oil contents. Additional specimens examined: — CHINA, Guizhou Province, Tongren city, Mount Fanjing National Nature Reserve, 11 July 2012, 27 ° 56 ′ 36 ″ N; 108 ° 36 ′ 47 ″ E, 810 m, asl., leg. Wuxingliang 03 (RITF 3260).	en	Chen, Bin, Song, Jie, Zhang, Jin-Hua, Liang, Jun-Feng (2021): Morphology and molecular phylogeny reveal two new species in Russula sect. Ingratae from China. Phytotaxa 525 (2): 109-123, DOI: 10.11646/phytotaxa.525.2.2
