identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
11718260FF92060AFDA0F84AFC5AC49F.text	11718260FF92060AFDA0F84AFC5AC49F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megastilicus Casey 1889	<div><p>Genus Megastilicus Casey, 1889</p> <p>Figs 1–2</p> <p>Megastilicus Casey, 1889: 183 (original description).</p> <p>Megastilicus – Casey 1905: 227 (description and key). — Wickham 1900: 3 (note); 1911: 15 (checklist). — Leng &amp; Wickham 1920: 24 (catalogue). — Holmquist 1928a: 84 (note); 1928b: 335 (note). — Park 1929: 199 (record/note); 1935: 217 (biology). — Blackwelder 1939: 100, 119 (key and list). — Herman 1970: 3 (note). — Moore &amp; Legner 1979: 107–108 (description and drawing). — Newton et al. 2020: 327, 387 (key and note). — Bousquet et al. 2013: DwC-A database (checklist). — Parker 2016: 78 (note).</p> <p>Type species</p> <p>Megastilicus formicarius Casey, 1889.</p> <p>Diagnosis</p> <p>Body robust, antennae short and stout. Body covered with short, black, stout setae. Dorsal surface of head, pronotum and elytra granulate. Head orbicular in shape.</p> <p>Redescription</p> <p>BODY. Medium size beetle; body robust, somewhat flattened. Colour of specimens from pale brown/sandy to brown/reddish brown. Head, pronotum and elytra covered with short, black setae. Dorsal surface of body densely granulato-reticulate, abdomen smoother, and not granulate.</p> <p>HEAD. Orbicular, with rounded hind angles; widest in middle, tapering towards antennal insertions anteriorly and towards neck posteriorly; disc of head without punctation, with leather-like integument. Head wider and longer than pronotum. Antennae short, reaching only to anterior part of pronotum, compact, mostly transverse, with well-developed stems in between, pubescent. Labrum large, almost completely covering mandibles from above, notched, broadly emarginate in middle at apex, with two distinct lobes, each bearing long and pale macroseta and comb of setae near emargination; emargination bidentate; weakly sclerotised, connected with head by membrane. Mandibulae without prostheca, with two teeth on right and one on left mandibula. Maxillary palpomere 3 about twice as long as palpomere 2; palpomere 4 elongate and thin, around ⅓ of palpomere 3 length and ⅓ of palpomere 3 width. Labial palpomere 1 shorter and thinner than palpomere 2; palpomere 2 slightly elongate, expanded toward apex, twice as long as palpomere 1; palpomere 3 thin and long, longer than half of palpomere 2 and ¼ of palpomere 2 width. Mentum transverse. Submentum with pair of setae on sides. Ligula bilobed. Gular sutures fused. Eyes small, from ⅕ to ⅙ of head length. Temples long, more than half of head length. Shallow longitudinal depression on each side of head ventrally. Neck distinct and narrow, about ⅕ of head width.</p> <p>THORAX. Pronotum longer than wide, anterior angles obtuse, widest in middle. Superior marginal line deflexed, becoming indistinct close to apical portion of prosternum, but reaching prosternum. Basisternum without macrosetae or microsculpture, but surface wrinkled. Furcasternum triangular, acute. Pronotal disc dull, with leather-like integument, without punctures. Elytra longer and wider than pronotum, without punctures, without epipleural ridge; humeral angles indistinct, rounded; surface dull with leather-like integument, with unorganised rows of small, black macrosetae. Anterior margin of scutellum rounded. Posterior (?) scutellar ridge present. Hind wings fully developed.</p> <p>LEGS. Long and slender, uniformly covered with glossy black macrosetae. All tarsi 5-segmented. Tarsomeres 1–4 of protarsus not inflated. Protibia with three fully developed protibial combs. One pair of empodial setae on each tarsus, equal to or slightly shorter than claws (never shorter than half of claw length). Meso and metatarsomere 1 longer than 2. Metatarsi almost as long as metatibiae, with tarsomeres 1–4 decreasing gradually in length, metarsomere 1 shorter than 2 and 3 combined, slightly longer than tarsomere 5.</p> <p>ABDOMEN. Sternite III with medial longitudinal keel. Abdomen widest at tergite V. Abdominal segments III to VII with pair of paratergites on each side; setae along length of abdomen. Tergite/sternite IX or X with dense setation. [Similar structure for female.]</p> <p>AEDEAGUS. With parameres reduced and fused to median lobe; ventral process slightly longer than uneverted internal sac; in parameral view with apex of ventral process acute, sides sinuate; dorsal plate rather large and relatively weakly sclerotised (Figs 1D–E, 2D–E).</p></div> 	http://treatment.plazi.org/id/11718260FF92060AFDA0F84AFC5AC49F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Żyła, Dagmara;Koszela, Katarzyna	Żyła, Dagmara, Koszela, Katarzyna (2021): Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species. European Journal of Taxonomy 778: 138-147, DOI: https://doi.org/10.5852/ejt.2021.778.1575
11718260FF90060DFDCEFDD2FE44C61D.text	11718260FF90060DFDCEFDD2FE44C61D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megastilicus formicarius Casey 1889	<div><p>Megastilicus formicarius Casey, 1889</p> <p>Fig. 1</p> <p>Type material</p> <p>Lectotype (here designated) USA • ♂; “Drac. Mass. / Type USNM 38356 / Megastilus formicarius Csy ”; NMNH.</p> <p>Paralectotypes USA • 1 spec.; “Drac. Mass. / Casey bequest 1925 / formicarius-5 PARATYPE USNM 38356 ”; NMNH • 1 spec.; “Casey bequest 1925 / formicarius-2 PARATYPE USNM 38356 ”; NMNH • 1 spec. with ant; “Casey bequest 1925 / formicarius-3 PARATYPE USNM 38356 ”; NMNH • 1 ♂; “Mass / Casey bequest 1925 / formicarius-6 PARATYPE USNM 38356 ”; NMNH.</p> <p>Additional studied material</p> <p>USA • 1 spec.; “Pen Mar Pa”; NMNH • 1 spec.; “ Harrisonville Mo. / Aug 2015 ”; NMNH • 1 spec.; “ Cumberland Co. / Maine 1916”; NMNH • 1 spec.; “ Newark NJ”; NMNH • 1 spec.; “ Linell? Coll Chittenden ”; NMNH • 1 ♂; “ CypMills Tex / Coll Chittenden ”; NMNH • 1 spec.; “ Sherborn, Massachusetts / IV-5 ’-25 / C.A. Frost /John George Gehring collection”; MCZ • 1 spec. [with Formica exsectoides]; “Wickham collection 1933 / Iowa City / Wickham IV-24-98 ”; MCZ • 2 specs; “ Natick / V-12-12 / Mass”; MCZ • 9 specs; “ Drac. Mass / 4/4/-86 ”; MCZ • 1 spec. [with Formica exsectoides]; “ Newfoundl’d, N.J. / X.17.1913 / C.A. Frost / C.A. Frost Collection 1962”; MCZ • 1 spec.; “ Chocorua N.H. / 5-2-06 / F.C. Bowditch Collection ”; MCZ • 1 spec.; “ Palos Park / Ill. / 4.14.34 / c. Seevers”; AMNH • 1 spec.; “ Natik Mass. Frost. 12.5 / Massachusetts / N.am / Mus 8 crm”; AMNH • 1 spec.; “ Crawfordsville Ind. 4.XII.50 ”; AMNH.</p> <p>Diagnosis</p> <p>Head longer than wide.Antennomere 4 elongate. Teeth on labrum blunt. Eyes smaller than in M. iowaensis sp. nov., ca ⅙ of head length. Median emargination of posterior margin of sternum VIII deeper and broader than in M. iowaensis sp. nov.</p> <p>Redescription</p> <p>BODY. Colour of specimens from pale brown/sandy to brown (Fig. 1A–B). Head and tip of abdomen darker. Body length from tip of head to tip of abdomen 5.5–6.0.</p> <p>HEAD. Width 1.0–1.1, length 1.0–1.5. Antennomere 1 elongate, widest at tip; antennomeres 2 and 3 elongate; antennomere 4 weakly elongate; antennomeres 5 to 10 gradually becoming strongly transverse; antennomere 11 elongate, 1.5 × as long as antennomere 10. Antennomeres 1 and 2 sparsely pubescent; antennomeres 3 and 4 more densely pubescent, 5 to 11 densely covered with tomentose pubescence. Maxillary palpomere 2 short and slender, with only few setae, maxillary palpomere 3 slightly expanded towards apex with denser setation, maxillary palpomere 4 glossy. Eyes about ⅙ of head length, shifted anteriorly, ¼ to ⅕ of temple length.</p> <p>THORAX. Pronotum width 0.9–1.0, length 1.0–1.1. Elytra width 1.4–1.5, length 1.3–1.4.</p> <p>LEGS. Two long setae on outer margin of metacoxa. Seta closer to abdomen longer and thicker than seta more distant from abdomen.</p> <p>ABDOMEN. Sternum VII with apical margin straight. Tergum VIII with rounded posterior margin. Male with sternum VIII with moderately deep and broad median emargination of posterior margin; depth of emargination of ca ⅕ of sternum length; sides of emargination rounded (Fig. 1C); surface right anterior to emargination without setae, while margins with dense setation.</p> <p>AEDEAGUS. With ventral process in parameral view narrower in middle; lateral plates of internal sac fused to sides of median lobe and indistinctive (Fig. 1D–E).</p> <p>Occurrence</p> <p>The species has previously been recorded from the Canadian provinces of Manitoba, Newfoundland and Labrador, and Ontario and the following US states: Illinois, Indiana, Iowa, Massachusetts, New Jersey and Ohio (A. Newton, unpublished database). New state records include: Missouri, New Hampshire, Pennsylvania and Texas.</p></div> 	http://treatment.plazi.org/id/11718260FF90060DFDCEFDD2FE44C61D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Żyła, Dagmara;Koszela, Katarzyna	Żyła, Dagmara, Koszela, Katarzyna (2021): Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species. European Journal of Taxonomy 778: 138-147, DOI: https://doi.org/10.5852/ejt.2021.778.1575
11718260FF96060EFDA7FEEBFC31C43C.text	11718260FF96060EFDA7FEEBFC31C43C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megastilicus iowaensis Żyła & Koszela 2021	<div><p>Megastilicus iowaensis sp. nov.</p> <p>urn:lsid:zoobank.org:act: 9603D57D-FAA2-4DAD-B68F-C91647D98CD0</p> <p>Fig. 2</p> <p>Diagnosis</p> <p>Head as long as wide.Antennomere 4 more transverse compared to elongated in Megastilicus formicarius. Teeth on labrum pointed. Eyes larger than in M. formicarius, ca ⅕ of head length. Median emargination of posterior margin of sternum VIII shallower and narrower than in M. formicarius.</p> <p>Etymology</p> <p>The species name ʻ iowaensis ʼ is derived from the name of the US state Iowa where this species has been collected.</p> <p>Type material</p> <p>Holotype USA • ♂ with ant [identified as Formica subsericea]; “ Iowa City Wickham IV 10? / WICKHAM Collection 1933”; NMNH.</p> <p>Paratypes USA • 1 ♀; “ Iowa City Iowa IV 10 Wickham / WICKHAM Collection 1933”; NMNH • 1 ♀ with ant; “ Iowa City Wickham IV-13 -?8 / WICKHAM Collection 1933”; NMNH • 1 ♀ with ant; “ Iowa City Wickham IV-29-68? / Megastilicus formicarius Csy. / Host ant: Formica subsericea / 9650 / Percy Gardner Bolster Coll’n / M. formicarius Casey 9650 / MCZ ENT 00744565 ”; MCZ.</p> <p>Description</p> <p>BODY. Colour of specimens pale brown/sandy (Fig. 2A–B). Head darker. Body length from tip of head to tip of abdomen 6.0.</p> <p>HEAD. Width 1.0, length 1.1–1.2. Antennomere 1 elongate, widest at tip; antennomeres 2 and 3 elongate; antennomeres 4 to 10 gradually becoming strongly transverse; antennomere 11 elongate, 1.5 × as long as antennomere 10. Antennomeres 1 and 2 sparsely pubescent, antennomeres 3 and 4 more densely pubescent, antennomeres 5 to 11 densely covered with tomentose pubescence. Emargination on anterior margin of labrum shallow. Maxillary palpomere 2 slender and short, with only few setae, maxillary palpomere 3 slightly expanded towards apex with denser setation, maxillary palpomere 4 glossy. Eyes about ⅕ of head length, shifted anteriorly, ⅓ of temple length.</p> <p>THORAX. Pronotum width 0.9, length 1.0–1.1. Elytra width 1.4–1.5, length 1.3–1.4.</p> <p>LEGS. Two long setae on outer margin of metacoxa. Seta closer to abdomen shorter and thinner than seta more distant from abdomen.</p> <p>ABDOMEN. Tergum VIII with slightly pointed posterior margin. Male with sternum VIII with shallow and broad median emargination of posterior margin; depth of emargination of ca ⅐ of sternum length; sides of emargination rounded (Fig. 2C); surface anterior to emargination without setae, while margins with dense setation.</p> <p>AEDEAGUS. With ventral process in parameral view widened in middle; lateral plates of internal sac welldeveloped and attached to sides of median lobe (Fig. 2D–E).</p> <p>Occurrence</p> <p>The new species has so far only been recorded from Iowa State.</p></div> 	http://treatment.plazi.org/id/11718260FF96060EFDA7FEEBFC31C43C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Żyła, Dagmara;Koszela, Katarzyna	Żyła, Dagmara, Koszela, Katarzyna (2021): Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species. European Journal of Taxonomy 778: 138-147, DOI: https://doi.org/10.5852/ejt.2021.778.1575
11718260FF94060EFF44FE99FAEDC7AC.text	11718260FF94060EFF44FE99FAEDC7AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megastilicus Casey 1889	<div><p>Key to the species of Megastilicus Casey, 1889</p> <p>1. Head longer than wide; antennomere 4 elongate; teeth on labrum blunt; eyes about ⅙ of head length, ¼ to ⅕ of temple length; median emargination of posterior margin of sternum VIII of ca ⅕ of sternum length, ventral process of aedeagus narrower in middle (parameral view); lateral plates of internal sac indistinctive..................................................................................... M. formicarius Casey, 1889</p> <p>– Head as long as wide; antennomere 4 transverse; teeth on labrum pointed; eyes about ⅕ of head length, ⅓ of temple length; median emargination of posterior margin of sternum VIII of ca ⅐ of sternum length; ventral process of aedeagus widened in middle (parameral view); lateral plates of internal sac distinctive.......................................................................................................... M. iowaensis sp. nov.</p></div> 	http://treatment.plazi.org/id/11718260FF94060EFF44FE99FAEDC7AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Żyła, Dagmara;Koszela, Katarzyna	Żyła, Dagmara, Koszela, Katarzyna (2021): Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species. European Journal of Taxonomy 778: 138-147, DOI: https://doi.org/10.5852/ejt.2021.778.1575
