taxonID	type	description	language	source
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	description	Figs 1 A – C, 3, 6 A	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	diagnosis	Diagnosis This species can be diagnosed by the following characters combined: 1) labrum less than ⅓ as long as wide, with anterior margin straight; 2) pronotal setose region anteriorly expanded inward onto pronotal disc to middle of dorsal eye; 3) punctation absent on head and pronotum; 4) punctures on surface of elytral glabrous region elongate transverse or oblique, several narrow longitudinal stripes shown on elytral glabrous region, micro-reticulation on elytral glabrous region weak and subtle, very elongate (Fig. 3 A, G); 5) in lateral view elytral pubescent area in basal ½ distinctly narrower than on pronotum, expanded to similar width apically (Fig. 1 C); 6) elytral apex rounded, epipleural angle acute; 7) female elytra similar to those of male; 8) male protarsus as in Fig. 3 B – C, adhesive pallet with few adhesive setae and large suction discs, ⅔ length of protibia; 9) aedeagus as in Fig. 3 D, median lobe shorter than parameres, extending to ¾ of their length, strongly laterally expanded in apical ⅓; 10) female reproductive tract as in Fig. 6 A, spermatheca elongate, reniform; fertilization duct broad and flattened, forming several compact hairpin hoops. This species is a member of Group 2 in Vazirani’s (1984) key (Group V in the key of Ochs 1928). The few adhesive setae and large suction discs on the male protarsus and characteristic elytral surface sculpture can easily distinguish this species from other known Chinese species of this group. This species is similar to P. coomani (Peschet, 1925), but can still be easily seperated by its narrow setose region at the elytral base and by the median lobe strongly laterally expanded in apical ⅓.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	etymology	Etymology This species is named after the type locality ʻHainan Provinceʼ.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	materials_examined	Material examined Holotype CHINA – Hainan • ♂; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 华立 中采 [Hua Lizhong leg.]; SYSU. Paratypes CHINA – Hainan • 5 ♂♂, 2 ♀♀; same collection data as for holotype; SYSU • 1 ♂, 2 ♀♀; 海南尖峰 岭五分区 [Hainan, Jianfengling Mt., fifth division,]; 5 Jul. 1981; 张新府采 [Zhang Xinfu leg.]; SYSU • 1 ♂; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 7 Jul. 1981; 黄云采 [Huang Yun leg.]; SYSU • 1 ♂, 3 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 7 Jul. 1981; 陈宁美采 [Chen Ningmei leg.]; SYSU • 1 ♂, 1 ♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 7 Jul. 1981; 刘元福采 [Liu Yuanfu leg.]; SYSU • 9 ♂♂, 8 ♀♀; 海南尖峰岭三分区 [Hainan, Jianfengling Mt., third division]; 7 Jul. 1981; 李宗冉采 [Li Zongran leg.]; SYSU • 10 ♂♂, 6 ♀♀; 海 南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 7 Jul. 1981; 梁少营采 [Liang Shaoying leg.]; SYSU • 1 ♂, 5 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 何国锋采 [He Guofeng leg.]; SYSU • 4 ♂♂, 6 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 雷群邦采 [Lei Qunbang leg.]; SYSU • 1 ♂; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 李逸昭采 [Li Yizhao leg.]; SYSU • 1 ♀, 海南尖峰岭 五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 陈润吉采 [Chen Runji leg.]; SYSU • 2 ♂♂, 2 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 29 Jul. 1983; 陈焕强 采 [Chen Huanqiang leg.]; SYSU • 1 ♂, 3 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 29 Jul. 1983; 江世贵采 [Jiang Shigui leg,]; SYSU • 1 ♂, 3 ♀♀; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Sep. 1983; 保护站采 [Reserve station leg.]; SYSU • 1 ♀; 海南尖峰 岭天池 [Hainan, Jianfengling Mt., Tianchi]; 8 Sep. 1983; 保护站采 [Reserve station leg]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	description	Description MEASUREMENTS. Holotype: 7.0 mm in length, 3.0 mm in width; male: 6.9 – 9.0 mm in length, 3.0 – 3.8 mm in width; female: 7.0 – 8.5 mm in length, 3.2 – 3.4 mm in width. HABITUS AND COLOUR (Fig. 1 A – C). Medium-sized species, oblong-oval in shape, widest just before middle, moderately convex in lateral view. Dorsal side black with strong bronze sheen, stronger on elytron than on head and pronotum. Pronotum and elytra with yellow lateral borders. Ventral side mostly black, with legs and rear of abdomen ferruginous, epipleuron yellow. HEAD. Labrum wide and short, ⅓ as long as wide, with anterior margin straight; anterior surface smooth; posterior surface with strong punctate-tomentose. Micro-reticulation on surface of clypeus distinct with polygonal meshes. Micro-reticulation on frons and vertex rather distinct, with meshes transverse or oblique (Fig. 3 E), gradually obliterated backwards. Punctation absent on head. THORAX. Pronotum regularly attenuated from base to apex, with margins weakly convex. Surface of pronotal glabrous region with weak and subtle micro-reticulation formed by elongate transverse or oblique meshes (Fig. 3 F); punctation absent on pronotum. Lateral pubescence anteriorly expanded inward onto pronotal disc to middle of dorsal eye, posteriorly narrower and parallel to lateral borders. Scutellum wide and short, ½ as long as wide, with surface smooth. Lateral pubescence on elytra anteriorly strongly reduced, distinctly narrower than on basal pronotum until middle of elytra, then abruptly expanded and reaching suture just before truncature (Fig. 3 A). Pronoto-elytral glabrous region irregular oval, terminating posteriorly with obtuse and wide angle. Surface of elytral glabrous region covered by distinct and rather dense punctation formed by elongate transverse or oblique strioles and with several narrow longitudinal stripes, micro-reticulation invisible under optical microscope, but visible under SEM, similar to that on pronotum (Fig. 3 G). Strioles at outer side of elytra much longer than that at inner side. Truncature of elytra straight; epipleural angle almost 90 °, acute; sutural angle almost 90 °, narrowly rounded. MALE GENITALIA (Fig. 3 D). Median lobe shorter than parameres, extending ¾ of their length, basal half parallel, strongly laterally expanded in apical ⅓, apical part regularly narrowed toward apex, apex point shortly narrowed. FEMALE REPRODUCTIVE TRACT (Fig. 6 A). Gonocoxosternite elongate, apically narrowly rounded, anterior process elongate; laterotergite elongate, slightly curved apically; vagina elongate; spermatheca elongate, reniform; fertilization duct broad and flattened, forming several compact hairpin hoops; accessory gland short and slender, inserting dorsally on bursa. SEXUAL DIMORPHISM. This species exhibits very little sexual dimorphism. Protibia of both sexes rather short and wide, basal third regularly expanded from base to apex, apical two-thirds subparallel, anteroexternal angle rather obliterated. Protarsus in male expanded, sub-oval, slightly more than ½ as long as protibia, slightly wider (Fig. 3 B), inner margin straight, external margin convex, ventral side with large sucking disc on top of adhesive setae (Fig. 3 C), while not expanded and without adhesive setae in female.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEAFFFE557ACE1D3347FE99.taxon	distribution	Distribution Only known from type locality Jianfengling Mt, Hainan Province, China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	description	Figs 1 D – F, 4, 6 B	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	diagnosis	Diagnosis This species can be diagnosed by the following characters combined: 1) labrum less than ⅓ as long as wide, with anterior margin straight; 2) pronotal setose region anteriorly expanded inward onto pronotal disc to external third of dorsal eye; 3) dorsal surface with strong and dense micro-punctation, much stronger on elytron (Fig. 4 E – G); 4) pronoto-elytral glabrous region regular oval, terminating posteriorly with an obtuse but not very wide angle, slightly wider in female (Fig. 4 A); 5) elytral apex straight, epipleural angle obtuse; 6) male protarsus as in Fig. 4 B – C, adhesive pallet with dense adhesive setae and small suction discs, distinctly longer than protibia; 7) aedeagus as in Fig. 4 D, median lobe broad, shorter than parameres, extending 4 / 5 of their length; 10) female reproductive tract as in Fig. 6 B, spermatheca elongate, C-shaped; fertilization duct slender, forming several loose hairpin hoops. This species is also a member of Group 2 in Vazirani’s (1984) key. It can be further assigned to the wui - subgroup proposed by Mazzoldi (1998) by the very large protarsus in the male. This species closely resembles Patrus wangi (Mazzoldi, 1998) described from Zhejiang and Anhui. These two species both have a well-developed protarsus which is distinctly longer than the protibia. However, it can be easily distinguished from the latter by the shape of the protarsus in the male (Fig. 4 B) and of the elytral glabrous region in the female. Besides, it is larger in size, the truncature of the protibia is more oblique and the median lobe of the aedeagus lacks the hook-like process (Fig. 4 D). The new species is also similar to P. wui (Ochs, 1932). Compared to P. wui, this species is larger in size, the protarsus is also larger, longer than the protibia, the glabrous region is less broad posteriorly, and the median lobe of the aedeagus is broader.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	etymology	Etymology This species is named after the type locality ʻJiangxi Provinceʼ.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	materials_examined	Material examined Holotype CHINA – Jiangxi • ♂; Jiangxi Prov. Pingxiang City, Luxi County, Wugongshan; 27 ° 47 ′ 53 ″ N, 114 ° 14 ′ 83 ″ E; alt. 520 m; 22 – 24 Sep. 2016; Ruijuan Zhang and Yudan Tang leg.; SYSU. Paratypes CHINA – Jiangxi • 30 ex.; same collection data as for holotype; SYSU • 2 ♀♀; Jiangxi Prov. Shangyou County, Guanggushan (光姑山) Nature Reserve; 25 ° 54 ′ 55 ″ N, 114 ° 03 ′ 09 ″ E; alt. 183 m; 19 – 21 Sep. 2016; Ruijuan Zhang and Yudan Tang leg.; SYSU • 14 ex.; 江西靖安县观音岩 [Guanyinyan, Jiangxi]; 29.04 N, 115.14 E; alt. 690 m; 20 Jul. 2014; 林仁超采 [Lin Renchao leg.]; SYSU • 1 ♀; 江西靖安县 三爪仑乡白水洞景区 [Jiangxi, Jing’an, Sanzhualun, Baishuidong Scenic Area]; 29.04 N, 115.11 E; alt. 660 m; 22 Jul. 2014; 林仁超采 [Lin Renchao leg.]; SYSU • 1 ♂; 江西井冈山白银湖 [Jiangxi, Jinggangshan, Baiyinhu]; alt. 880 m; 23 Apr. 2011; 采集人: 赵爽 [Zhao Shuang leg.]; SYSU • 1 ♂, 3 ♀♀; 江西井冈山湖羊塔 [Jiangxi, Jinggangshan, Huyangta]; 28 Apr. 2011; 采集人: 贾凤龙 [Jia Fenglong leg.]; SYSU • 1 ♂; 江西井冈山 [Jiangxi, Jinggang Shan]; 1 Apr. 2011; 李韵采 [Li Yun leg.]; SYSU • 1 ♂; W. Jiangxi, Ji’an City, Jinggang Shan, Jingzhu Shan; 26 ° 29 ′ 45 ″ N, 114 ° 04 ′ 45 ″ E; alt. 1130 m; 2 Aug. 2014; Chen, Hu, Lv and Yu leg.; in the stream; SHNU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	description	Description MEASUREMENTS. Holotype: 8.5 mm in length, 3.9 mm in width; male: 8.4 – 9.8 mm in length, 3.9 – 4.0 mm in width; female: 7.0 – 8.6 mm in length, 3.1 – 3.9 mm in width. HABITUS AND COLOUR (Fig. 1 D – F). Medium-sized species, oblong-oval in shape, widest just before middle, moderately convex in lateral view. Dorsal side black with strong bronze sheen. Pronotum and elytra with yellow lateral borders. Ventral side mostly reddish brown to dark brown, with legs and epipleura yellow to reddish yellow, last abdomen ferruginous. HEAD. Labrum wide and short, ⅓ as long as wide, with anterior margin straight; anterior surface smooth; posterior surface with strong punctate-tomentose. Clypeus slightly shorter than labrum, with anterior margin straight. Fronto-clypeal suture distinct. Surface of clypeus with weak micro-reticulation anteriorly, while distinct posteriorly, among meshes of micro-reticulation with dense irregular punctation. Surface on frons and vertex with strong micro-reticulation formed by isodiametric polygonal meshes, gradually obliterated backwards. Amongst meshes of the micro-reticulation strong irregular-shaped punctation (Fig. 4 E). THORAX. Pronotum regularly attenuated from base to apex, with margins weakly convex. Surface of pronotal glabrous region with distinct micro-reticulation formed by isodiametric polygonal meshes and strong irregular punctation (Fig. 4 F), much of which merges in rows to form distinct wrinkles. Lateral pubescence anteriorly expanded inward onto pronotal disc to external third of eye, posteriorly weakly reduced backwards in concave line. Scutellum wide and short, less than ½ as long as wide, with surface smooth. Pronoto-elytral glabrous region regular oval, terminating posteriorly with obtuse but not very wide angle (Fig. 4 A). Lateral pubescent band in male anteriorly as wide as pronotal one, subparallel to elytral margin backwards until middle of elytron, then regularly expanded, reaching suture at about onetenth of distance from truncature; in female anteriorly as wide as pronotal one, parallel to elytral margin backwards until ⅔ of elytron, then regularly expanded, reaching suture at about 1 / 12 of distance from truncature, so that elytral glabrous region in female slightly wider. Surface of elytral glabrous region covered by weak micro-reticulation formed by transverse or oblique and slightly elongate polygonal meshes and by strong punctation formed by short transverse or oblique strioles (Fig. 4 G). Truncature of elytra straight; epipleural angle obtuse, narrowly rounded, sutural angle almost 90 °, narrowly rounded. MALE GENITALIA (Fig. 4 D). Median lobe shorter than parameres extending 4 / 5 of their length, basal third parallel, then slightly narrowed until apical third, sub-apex parallel, apical part regularly narrowed, apex point shortly narrowed. FEMALE REPRODUCTIVE TRACT (Fig. 6 B). Gonocoxosternite elongate, apically narrowly protruding, anterior process short; laterotergite elongate, slightly narrowed towards apex; vagina elongate; spermatheca elongate, C-shaped, strongly curved; fertilization duct slender, forming several loose hairpin hoops; accessory gland short and slender, inserting dorsally on bursa. SEXUAL DIMORPHISM. Male slightly larger in size than female; protibia in male short and wide, short triangular, strongly expanded exteriorly towards apex, apex of protibia obliquely truncate, anteroexternal angle obtuse, rounded (Fig. 4 B), while shorter and less expanded, and apex straight-truncate in female. Protarsus of male large, strongly expanded, distinctly longer than protibia, slightly narrowed towards apex, ventral side with dense adhesive setae and small suction discs (Fig. 4 C); elytral glabrous region slightly broader posteriorly in female.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFEFFFF0557ACCC73391F9C1.taxon	distribution	Distribution Known from Pingxiang, Shangyou, Jingxi and Jinggangshan in Jiangxi Province, China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE1FFF5555BCB0F33C0F8AC.taxon	description	Figs 2, 5, 6 C	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE1FFF5555BCB0F33C0F8AC.taxon	diagnosis	Diagnosis This species can be diagnosed by the following characters combined: 1) labrum less than ⅓ as long as wide, with anterior margin straight; 2) pronotal setose region anteriorly expanded inward onto pronotal disc to external third of dorsal eye; 3) dorsal surface with strong and dense micro-punctation, much stronger on elytron (Fig. 5 F – H); 4) pronoto-elytral glabrous region regularly oval in male, terminating posteriorly with obtuse but not very wide angle, with two postero-lateral expansions in females (Fig. 5 A – B); 5) elytral apex straight, epipleural angle obtuse; 6) male protarsus as in Fig. 5 C – D, adhesive pallet with dense adhesive setae and small suction discs, almost as long as protibia; 7) aedeagus as in Fig. 5 E, median lobe shorter than parameres, extending 4 / 5 length, apical part notably narrowed, sub-apically slightly constricted; 10) female reproductive tract as in Fig. 6 C, spermatheca elongate, C-shaped; fertilization duct rather slender and flattened, forming several loose hairpin hoops. This species is also a member of the wui - subgroup. It closely resembles Patrus wangi (Mazzoldi, 1998) and P. wui (Ochs, 1932). All these three species show a distinct sexual dimorphism in the shape of the elytral glabrous region. Two postero-lateral expansions are shown in female of both, the new species and P. wangi but much weaker in this species than in P. wangi so that the lateral pubescence on the elytra is visible from above throughout (Figs 2 D, 5 B). Besides, the protarsus in the male is less developed, at most as long as protibia in this species (Fig. 5 C – D). The elytral glabrous region in the female is much broader posteriorly than in male P. wui, whereas postero-lateral expansions are not present in this species. The male genitalia of these three species are also distinctly different from each other.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE1FFF5555BCB0F33C0F8AC.taxon	etymology	Etymology This species is named after the type locality ʻShangchuan Islandʼ. Material examined Holotype CHINA – Guangdong • ♂; 广东省台山市上川岛猕猴省级自然保护区 [Guangdong, Taishan City, Shangchuan Island Macaque Provincial Nature Reserve]; 21 ° 44 ′ 51 ″ N, 119 ° 49 ′ 36 ″ E; alt. 50 m; 17 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU. Paratypes CHINA – Guangdong • 13 ♂♂, 12 ♀♀; same collection data as for holotype; SYSU • 5 ♂♂, 3 ♀♀; 广东省台山市上川岛猕猴省级自然保护区 [Guangdong, Taishan City, Shangchuan Island Macaque Provincial Nature Reserve]; 21 ° 44 ′ 36 ″ N, 112 ° 50 ′ 33 ″ E; alt. 10 m; 18 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU • 1 ♂; 广东省台山市上川岛猕猴省级自然保护区石 背潭 [Guangdong, Taishan City, Shangchuan Island Macaque Provincial Nature Reserve, Shibeitang]; 21 ° 46 ′ 1 ″ N, 112 ° 49 ′ 2 ″ E; alt. 20 m; 19 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE1FFF5555BCB0F33C0F8AC.taxon	description	Description MEASUREMENTS. Holotype: 7.7 mm in length, 3.6 mm in width; male: 7.1 – 8.6 mm in length, 3.2 – 3.6 mm in width; female: 6.9 – 7.6 mm in length, 2.9 – 3.4 mm in width. HABITUS AND COLOUR (Fig. 2 A – E). Medium-sized species, oval in shape, widest just before middle, moderately convex in lateral view. Dorsal side black with strong bronze sheen. Pronotum and elytra with yellow lateral borders. Ventral side mostly black, with legs and epipleura yellow to reddish yellow. HEAD. Labrum wide and short, slightly less than ⅓ as long as wide, with anterior margin straight; anterior surface smooth; posterior surface strongly punctate-tomentose. Clypeus slightly shorter than labrum, with anterior margin slightly concave. Fronto-clypeal suture distinct. Surface of clypeus with strong micro-reticulation formed by isodiametric polygonal meshes, among meshes with strong and dense punctation. Surface of frons and vertex with strong micro-reticulation formed by isodiametric polygonal meshes, gradually obliterated backwards. Amongst meshes of micro-reticulation strong irregular-shaped punctation (Fig. 5 F). THORAX. Pronotum regularly attenuated from base to apex, with margins weakly convex. Surface of pronotal glabrous region with strong irregular-shaped punctation, some of which merges in rows to form distinct wrinkles (Fig. 5 G), micro-reticulation on median disc strong, formed by isodiametric polygonal meshes, obliterated towards margins. Lateral pubescence anteriorly expanded inward onto pronotal disc to external third of dorsal eye, posteriorly weakly reduced backwards in concave line. Scutellum wide and short, less than ½ as long as wide, with surface smooth. Pronoto-elytral glabrous region in male regularly oval. Lateral pubescent area in male anteriorly as wide as on pronotum, parallel to elytral margin backwards until the middle of elytron, then regularly expanded, reaching suture at about ⅛ to truncature (Fig. 5 A). Pronoto-elytral glabrous region in female in characteristic shape, with weak postero-lateral expansion at each side (Fig. 5 B). Lateral pubescence in female anteriorly two-fifths slightly narrowed backwards, then regularly narrowed in concave line, apical fifth suddenly expanded, reaching suture at same position as in male. Surface of elytral glabrous region covered by distinct micro-reticulation formed by transverse or oblique and slightly elongate polygonal meshes and by strong punctation formed by short transverse or oblique strioles (Fig. 5 H). Truncature of elytra straight; epipleural angle obtuse, narrowly rounded, sutural angle almost right angle, narrowly rounded. FRONT LEGS. Protibia in male short and wide, short triangular, strongly expanded exteriorly towards apex, apex of protibia obliquely truncate, antero-external angle obtuse, narrowly rounded (Fig. 5 C). Protibia in female slenderer than in male. Protarsus of male large, strongly expanded, almost as long as protibia, basal four-fifths subparallel, apical fifth regularly narrowed (Fig. 5 D). MALE GENITALIA (Fig. 5 E). Median lobe notably shorter than parameres, basal third parallel, then slightly narrowed backwards until apical third, slightly expanded subapically, apical part regularly narrowed, apex form obtuse angle. FEMALE REPRODUCTIVE TRACT (Fig. 6 C). Gonocoxosternite elongate, apically narrowed, anterior process short and broad; laterotergite elongate, slightly narrowed towards apex; vagina elongate; bursa rather broad; spermatheca elongate, C-shaped, strongly curved; fertilization duct rather slender and flattened, forming several loose hairpin hoops; accessory gland short and slender, inserting dorsally on bursa. SEXUAL DIMORPHISM. Protibia in male short and wide, short triangular, strongly expanded exteriorly towards apex, apex of protibia obliquely truncate, antero-external angle obtuse, narrowly rounded (Fig. 5 C), while slender and less expanded, and apex straight-truncate in female. Protarsus of male large, strongly expanded, almost as long as protibia, basal four-fifths subparallel, apical fifth regularly narrowed, ventral side with dense adhesive setae and small suction disks (Fig. 5 D); elytral glabrous region regularly oval in male and with two weak postero-lateral expansion in female.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE1FFF5555BCB0F33C0F8AC.taxon	distribution	Distribution Only known from the type locality Shangchuan Island, Guangdong Province, China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF45549CC933472FB75.taxon	materials_examined	Material examined CHINA – Hainan • 2 ♀♀; 广东海南 [Guangdong Hainan]; 5 Jun. 1957; 赵养昌采 [Zhao Yangchang leg.]; SYSU • 2 ♀♀; 海南岛尖峰岭 [Hainan Is., Jianfengling Mt.]; 19 Dec. 1963; 彭统序采 [Peng Tongxu leg.]; SYSU • 2 ex.; 广东海南三亚 [Guangdong, Hainan, Sanya]; alt. 10 m; 5 Apr. 1960; 采集 者: 李学忠 [Li Xuezhong leg.]; IZCAS. – Jiangxi • 1 ♂; 江西南昌 [Jiangxi, Nanchang]; Jul. 1963; 马 恩沛采 [Ma Enpei leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF45549CC933472FB75.taxon	distribution	Distribution Distributed in the Oriental and Australian Regions, known from Fujian, Guangdong, Guangxi, Hong Kong, Taiwan in China. New for Hainan and Jiangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF45540C9AC325AF9AF.taxon	materials_examined	Material examined CHINA – Henan • 1 ♀; 河南信阳鸡公山 [Henan, Xinyang, Jigongshan Mt.]; Aug. 1936; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF45540C9AC325AF9AF.taxon	distribution	Distribution Known from China (Fujian, Guangdong, Guangxi, Guizhou, Hong Kong, Hubei, Hunan, Jiangxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang), Japan and Vietnam. New for Henan.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF7554DCBC53407FDE8.taxon	materials_examined	Material examined CHINA – Beijing • 1 ex.; 北京白石桥 [Beijing, Baishiqiao]; 5 Apr. 1955; 张毅然采 [Zhang Yiran leg.]; IZCAS. – Hainan • 11 ex.; Hainan Is. S China, Ka-chek, K’ieng-tong District; 4 May 1932; F. K. To leg.; SYSU • 1 ♂; 海南 [Hainan]; 16 Jul. 1957; 利翠英采 [Li Cuiying leg.]; SYSU • 1 ex.; Hainan, Fanyang; 15 Apr. 1936; G. Ros leg.; IZCAS. – Shaanxi • 4 ex.; 陕西西安浐灞 [Shaanxi, Xi’an, Chanba]; 21 May 2010; 许升金采 [Xu Shengjin leg.]; SYSU. – Tianjin • 2 ex.; 天津八里台 [Tianjin, Balitai]; 1 May 1955; 刘英采 [Liu Ying leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE5FFF7554DCBC53407FDE8.taxon	distribution	Distribution Distributed in China (Fujian, Guangdong, Guangxi, Guizhou, Hebei, Hong Kong, Hubei, Jiangsu, Liaoning, Shandong, Shanghai, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang), Japan, Korea, Vietnam and Laos. New for Beijing, Hainan, Shaanxi and Tianjin.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF75524C9F236DBF8D8.taxon	materials_examined	Material examined CHINA – Yunnan • 5 ex. 云南西双版纳勐阿 [Yunnan, Xishuangbanna, Meng’e]; 1050 – 1080 m; 1958. VIII. 22; 采集人: 王书永 [leg. Wang Shuyong]; IZCAS • 5 ex.; 云南西双版纳勐遮 [Yunnan, Xishuangbanna, Mengzhe]; alt. 1200 m; 8 Apr. 1958; 采集人: 王书永 [Wang Shuyong leg.]; IZCAS • 1 ♀; 云南屏边 [Yunnan, Pingbian]; alt 1300 m; 19 – 26 Jun. 1956; 黄克仁采 [Huang Keren leg.]; IZCAS • 8 ex.; 云南思茅附近 [Yunnan, near Simao]; 26 Mar. 1957; A. 孟恰茨基采 [Менчацкий leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF75524C9F236DBF8D8.taxon	distribution	Distribution Widely distributed in the Oriental and Australian Regions, known from Jiangxi, Taiwan and Xizang. New for Yunnan.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF7552BCFC13671FBE6.taxon	materials_examined	Material examined CHINA – Guangxi • 5 ♂♂, 3 ♀♀; 广西百寿 [Guangxi, Baishou]; 27 Jul. 1952; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF7552BCFC13671FBE6.taxon	distribution	Distribution Known from southern China (Guizhou, Hainan, Yunnan and Xizang) and northern Vietnam. New for Guangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF65542CA7F339BFE2C.taxon	materials_examined	Material examined CHINA – Heilongjiang • 1 ♂; 黑龙江密山 [Heilongjiang, Mishan], 1 Oct. 1951; SYSU • 2 ♂♂, 2 ♀♀; Heilongjiang Prov., Yichun, Wuying, Wuying National Forest Park; 11 Jul. 2004; Li-Zhen Li leg; SHNU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE6FFF65542CA7F339BFE2C.taxon	distribution	Distribution Narrowly distributed in China (Beijing and Liaoning) and Japan. New for Heilongjiang.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE7FFF6553CCF433390FB02.taxon	materials_examined	Material examined. CHINA – Guangxi • 11 ex.; 广西临桂黄沙 [Guangxi, Lingui, Huangsha]; 13 Aug. 2006; 黄国华采 [Huang Guohua leg.]; SYSU • 1 ♂; 广西容县 [Guangxi, Rongxian]; 10 Oct. 1962; 古德祥采 [Gu Dexiang leg.]; SYSU • 6 ex.; Guangxi Prov., Jiuwangdashan, Yangmeiao; 25 ° 11 ′ 42 ″ N, 108 ° 38 ′ 51 ″ E; alt. 1183 m; 20 Jul. 2015; Rencao Lin and Yudan Tang leg.; SYSU. – Hubei • 1 ♂; 湖北英山吴家山 [Hubei, Yingshan County, Wujiashan Mout.]; alt. 800 – 1000 m; 12 Jul. 2006; 采集人: 石磊 [Shi Lei leg.]; SYSU. – Hunan • 1 ♀; 湖南古丈县高望界国家自然保护区 [Hunan, Guzhang County, Gaowangjie National Reserve]; 28 ° 39.898 ′ N, 110 ° 3.575 ′ E; 1053 m; 21 Jun. 2017; F. - L. Jia leg.; SYSU. – Zhejiang • 3 ♀♀, 2 ♂♂; Zhejiang prov., West Tianmushan Mout.; 22 Aug. 2001; L-Z Li leg.; SYSU • 13 ♀♀, 6 ♂♂; Zhejiang Prov., Lin’an City, West Tianmushan MT.; alt. 300 – 400 m; 6 Jun. 2007; Tang Liang leg.; SHNU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE7FFF6553CCF433390FB02.taxon	distribution	Distribution Known from China (Fujian, Guangdong, Guizhou, Hong Kong, Jiangsu, Jiangxi, Shanghai, Sichuan, Yunnan), Russia (Far East) and Vietnam. New for Guangxi, Hubei, Hunan and Zhejiang.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE7FFE9555AC9BA33C5FE73.taxon	materials_examined	Material examined CHINA – Guangdong • 1 ♂, 3 ♀♀; 广东珠海横琴岛 [Guangdong, Zhuhai, Hengqin Island]; 12 Jul. 2005; 贾粤采 [Jia Yue leg.]; SYSU. – Inner Mongolia • 4 ♀♀; China, Nei Mongol, Hulunbeier, Huihe; 25 Jul. 2013; Shi Li, Jin Chunyun and De chaoqun leg.; SYSU • 2 ♂♂, 5 ♀♀; 中国内蒙古呼伦贝尔 中蒙边境国门外 [China, Inner Mongolia, Hulunbeier, near the border of China and Mongolia]; 29 Aug. 2014; 贾凤龙采 [Fenglong Jia leg.]; SYSU • 1 ♀; Inner Mongolia, Hanma Management Centre, a stream by monitoring station; 30 Jul. 2014; Shi Li, Zhu Lixuan, Gao Xuefeng and Chen chao leg.; SYSU • 1 ♀; 内蒙古海拉尔市 [Inner Mongolia, Hailaer City]; 23 – 26 Jul. 2003; 采集人: 贾凤龙 [Jia Fenglong leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE7FFE9555AC9BA33C5FE73.taxon	distribution	Distribution Widely distributed in the Palaearctic Region. Known from Heilongjiang, Jilin, Liaoning, Xinjiang in China. New for Guangdong and Inner Mongolia.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFE7FFE9555AC9BA33C5FE73.taxon	discussion	Remarks As it is a typical northern species, the occurrence of this species from Guangdong, such a southern province is quite questionable. However, the material we examined did not differ from material from the northern provinces of China. Although it is possible that the four specimens from Guangdong were wrongly labelled, we have treated the material from Guangdong as G. minutus here.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE95564CCAA360AFC39.taxon	materials_examined	Material examined CHINA – Xinjiang • 2 ♂♂; 新疆博湖 [Xinjiang, Bohu]; 7 Aug. 2006; 赵玲采 [Zhao Ling leg.]; SYSU • 1 ♂; 新疆塔域哈尔墩河坝 [Xinjiang, Tacheng, Haerdun Heba]; 4 Aug. 2006; 赵玲采 [Zhao Ling leg.]; SYSU • 25 ♂♂, 7 ♀♀; 新疆额敏河 [Xinjiang, Eminhe County]; 25 Jul. 1955; 马世骏, 夏凯龄, 陈永林采 [Ma shijun, Xia kailin, Chen Yonglin leg.]; IZCAS • 1 ♀; 新疆富蕴苏普特 [Xinjiang, Fuyun County, Supute]; 47.07 ° N, 89.46 ° E; alt. 1065 m; 10 Jul. 2009; 采集人: 王志良 [Wang zhiliang leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE95564CCAA360AFC39.taxon	distribution	Distribution Widely distributed in the Palaearctic Region, known from Inner Mongolia and Xizang in China. New for Xinjiang.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE95569C95035F7FA28.taxon	materials_examined	Material examined CHINA – Beijing • 2 ex.; 北京钓鱼台 [Beijing, Diaoyutai]; 8 May 1956; 张毅然采 [Zhang Yiran leg.]; IZCAS • 5 ex.; 北京碧云寺 [Beijing, Biyun Temple]; 24 Aug. 1956; 张毅然采 [Zhang Yiran leg.]; IZCAS • 1 ♂; 北京动物园 [Beijing Zoo]; 10 May 1955; 张毅然采 [Zhang Yiran leg.]; IZCAS • 1 ex.; 北京圆明园 [Beijing, Yuanmingyuan]; 17 May 1955; 张毅然采 [Zhang, Yiran leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE95569C95035F7FA28.taxon	distribution	Distribution Known from China (Jilin, Liaoning), Belarus (Ryndevich et al. 2014), Finland, Korea, Russia and Sweden (Holmen 1987). New for Beijing.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE9554ECB4035BFF860.taxon	materials_examined	Material examined CHINA – Yunnan • 1 ♂; 云南维西塔城吧珠村 [Yunnan, Weixi, Tacheng, Baxi Village]; 19 Aug. 2013; 陈红星采 [Chen Hongxing leg.].	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF8FFE9554ECB4035BFF860.taxon	distribution	Distribution Endemic to China (Sichuan). New for Yunnan.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85541CC02334DFCC6.taxon	materials_examined	Material examined CHINA – Guangxi • 2 ♀♀; 广西兴安县猫儿山 [Guangxi, Xing’an County, Maoershan Mout.]; 29 May 2010; 赵爽采 [Zhao Shuang leg.]; SYSU. – Guizhou • 1 ♂; 贵州, 江口梵净山 [Guizhou, Jiangkou County, Fanjingshan Mout.]; alt. 530 m; 15 Jul. 1988; 王书永采 [Wang Shuyong leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85541CC02334DFCC6.taxon	distribution	Distribution Endemic to China (Fujian, Guangdong). New for Guangxi and Guizhou.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85548CE043504FB05.taxon	description	Fig. 7	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85548CE043504FB05.taxon	materials_examined	Material examined CHINA – Yunnan • 1 ♀; 云南景东 [Yunnan, Jingdong]; alt. 1200 m; A. 孟恰茨基采 [A. Менчацкий leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85548CE043504FB05.taxon	distribution	Distribution Known from Laos. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85501C83835DEF86A.taxon	distribution	Fig. 8	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85501C83835DEF86A.taxon	materials_examined	Material examined CHINA – Yunnan • 2 ♂♂; Yunnan Prov., Baoshan City, Baihuangling, Jiujiezi; 25 ° 18 ′ N, 98 ° 47 ′ E; alt. 2016 m; 19 Jun. 2015; Renchao Lin and Yudan Tang leg.; SYSU • 1 ♀; 云南楚雄西面山上 [Yunnan, Chuxiong, Ximianshan Mt.]; alt. 2500 m; 19 May 1956; 克雷让诺夫斯基采 [Klejanovsky leg.]; IZCAS • 1 ♀; 云南楚雄- 弥渡 [Yunnan, Chuxiong-Midu]; 19 May 1956; B. 波波夫采 [В. Попов leg.]; IZCAS • 18 ex.; 云南西双版纳勐宋 [Yunnan, Xishuangbanna, Mengsong]; alt. 1600 m; 26 Apr. 1958; 王书 永采 [Wang Shuyong leg.]; IZCAS • 42 ex.; 云南西双版纳勐混 [Yunnan, Xishuangbanna, Menghun]; alt. 1200 – 1400 m; 24 May 1958; 洪淳培采 [Hong Chunpei leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF9FFE85501C83835DEF86A.taxon	distribution	Distribution Known from Vietnam. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFBFFEA552ACC403305FD8C.taxon	materials_examined	Material examined CHINA – Jiangsu • 3 ♂♂, 5 ♀♀; 江苏南京 [Jiangsu, Nanjing]; Jul. 2009; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFBFFEA552ACC403305FD8C.taxon	distribution	Distribution Endemic to China (Fujian, Guangdong, Shanghai, Zhejiang). New for Jiangsu.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFBFFED5524CFCA3433FD4D.taxon	description	(Figs 9 A – 10)	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFBFFED5524CFCA3433FD4D.taxon	materials_examined	Material examined Type material INDIA – Sikkim • 1 ♂, 1 ♀; mounted together. “ Sikkim Kunleong, 1904 – 215 ” [register says: ‘ received in exchange from Régimbart’], “ Orectochilus murinus Rég. ♂ ♀ ”, “ CO-TYPE ”, “ NHMUK 014433292 ”; BMNH. Additional material CHINA – Yunnan • 2 ♀♀ 云南省宁蒗彝族自治县高峰村 [Yunnan Prov., Ninglang Yi-Nationality Autonomous County, Gaofeng Village]; alt. 2249 m; 27 ° 13 ′ 32 ″ N, 100 ° 52 ′ 13 ″ E; 24 May 2019; 梁祖 龙, 杨圳铭采 [Liang Zulong and Yang Zhenming leg.]; SYSU • 6 ♂♂, 9 ♀♀; Yunnan Prov., Nabanhe N. R. Nabancun; alt. 720 m; 6 May 2009; Jia-Yao Hu and Zi-wei Yin leg.; SHNU. SIKKIM • 1 ♀; “ Environs de Kurseong, R. P. Bretaudeau, Fry Coll, 1905 – 100 70770 ”, “ Orectochilus murinus Rég ”, “ NHMUK 014433293 ”; BMNH; • 1 ex.; “ Environs de Kurseong, R. P. Bretaudeau, 99.105 ”, “ O. murinus R named by Regimbart ”, “ Received with this name from Heyne C. O. W. [C. O. Waterhouse], “ NHMUK 014433294 ”; BMNH • 1 ex.; “ India Or., Sikkim, Fry Coll. 1905.100 ”, “ Orectochilus murinus Rég. Sikkim nec. Africa ” [from Sikkim not Africa], “ NHMUK 014433295 ”; BMNH.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFBFFED5524CFCA3433FD4D.taxon	distribution	Distribution Known from Indochina, Sikkim and India. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFDFFEC551FCC403253F94F.taxon	materials_examined	Material examined CHINA – Hainan • 9 ex.; 海南尖峰岭三分区 [Hainan, Jianfengling Mt., third division]; 7 Jul. 1981; 何国锋采 [He Guofeng leg.]; SYSU • 1 ♂; 海南尖峰岭 [Hainan, Jianfengling Mt.]; 9 Jul. 1981; 饶漱 静采 [Rao Shujing leg.]; SYSU • 5 ex.; 海南尖峰岭三分区 [Hainan, Jianfengling Mt., third division]; 7 Jul. 1981; 李宗冉采 [Li Zongran leg.]; SYSU • 9 ex.; 海南尖峰岭三分区 [Hainan, Jianfengling Mt., third division]; 31 Jul. 1983; 陈焕强采 [Chen Huanqiang leg.]; SYSU • 7 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 雷群邦采 [Lei Qunbang leg.]; SYSU • 3 ex.; 海 南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jun. 1981; 雷群邦采 [Lei Qunbang leg.]; SYSU • 1 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 22 Feb. 1982; 黄治河采 [Huang Zhihe leg.]; SYSU • 5 ex.; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 21 Feb. 1982; 黄治河采 [Huang Zhihe leg.]; SYSU • 1 ex.; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 8 Nov. 1983; 黄治河采 [Huang Zhihe leg.]; SYSU • 4 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 张军雄采 [Zhang Junxiong leg.]; SYSU • 1 ex.; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 7 Jul. 1981; 李胜文采 [Li Shengwen leg.]; SYSU • 3 ex.; 海南尖峰岭五分 区 [Hainan, Jianfengling Mt., fifth division]; 22 Feb 1982; 龙建国采 [Ling Jianguo leg.]; SYSU • 2 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 10 Feb. 1984; 陆高平采 [Lu Gaoping leg.]; SYSU • 3 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 26 Feb. 1982; 张昆 智采 [Zhang Kunzhi leg.]; SYSU • 1 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; Aug. 1981; 华立中采 [Hua Lizhong leg.]; SYSU • 1 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 22 Feb. 1982; 龙永成采 [Long Yongcheng leg.]; SYSU • 1 ex.; 海南尖峰岭五分 区 [Hainan, Jianfengling Mt., fifth division]; 8 Aug. 1981; 邝永泉采 [Kuang Yongquan leg.]; SYSU • 1 ex.; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Aug. 1981; SYSU • 1 ex.; 海 南尖峰岭三分区 [Hainan, Jianfengling Mt., third division]; 13 Oct. 1983; 保护站采 [Reserve station leg.]; SYSU • 1 ex.; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 21 Feb. 1982; 陈振耀采 [Chen Zhenyao leg.]; SYSU • 1 ex.; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 24 Feb. 1982; 陈振耀采 [Chen Zhenyao leg.]; SYSU • 22 ex.; 海南尖峰岭, 第三, 五分区 [Hainan, Jianfengling Mt., third and fifth division]; 9 May 2011; 赵爽采 [Zhao Shuang leg.]; SYSU. – Henan • 1 ♀; 河南信阳 鸡公山 [Henan, Xinyang City, Jigongshan Mt.]; Aug. 1936; SYSU. – Hubei • 3 ♂♂, 1 ♀; 湖北英山吴 家山 [Hubei, Yingshan County, Wujiashan Mt.]; 12 Jul. 2006; 石磊采 [Shi Lei leg.]; SYSU. – Jiangxi • 26 ex.; 江西龙南九连山 [Jiangxi, Longnan County, Jiulianshan Mout.]; 12 – 13 Aug. 2008; 贾凤龙 采 [Jia Fenglong leg.]; SYSU • 1 ♂, 1 ♀; Jiangxi Prov. Longnan County, Jiulianshan Nature Reserve; 24 ° 34 ′ 54 ″ N, 114 ° 26 ′ 21 ″ E; alt. 425 m; 23 – 24 Apr. 2017; Wang Shishuang leg.; SYSU. – Shaanxi • 1 ♂; 陕西西安楼观台 [Shaanxi, Xi’an City, Louguantai]; 24 Jul. 2001; 赵玲采 [Zhao Lin leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFDFFEC551FCC403253F94F.taxon	distribution	Distribution Endemic to China (Fujian, Guangdong). New for Hainan, Henan, Hubei, Jiangxi and Shaanxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFDFFEF5513CBE63340FE99.taxon	materials_examined	Material examined CHINA – Guangdong • 21 ex.; 广东韶关南岭国家自然保护区 ， 溪流 [Guangdong, Shaoguan, Nanling National Natural Reserve, in the stream]; 24 ° 56 ′ 18 ″ N, 113 ° 0 ′ 13 ″ E; alt. 1210 m; 21 Jul. 2020; leg. Liang Zulong, Yang Zhenming; SYSU. – Guangxi • 5 ♂♂, 9 ♀♀; Guangxi Prov., Jiuwandashan, Yangmeiao; 25 ° 11 ′ 42 ″ N, 108 ° 38 ′ 51 ″ E; 20 Jul. 2015; Renchao Lin and Yudan Tang leg.; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFDFFEF5513CBE63340FE99.taxon	distribution	Distribution Endemic to China (Shaanxi, Sichuan). New for Guangdong and Guangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFEFFEF551DCC303528FB69.taxon	materials_examined	Material examined CHINA – Chongqing • 1 ♂; 重庆, 巫山县 [Chongqing, Wushan County]; alt. 200 – 1800 m; 16 Aug. 2006; 张小蓉采 [Zhang Xiaorong leg.]; SYSU. – Guangdong • 1 ♂; 广东连县大东山 [Guangdong, Lianxian County, Dadongshan Mout.]; 13 Jul. 1998; 蔡于琛采 [Cai Yuchen leg.]; SYSU • 1 ♂; 广 东连县大东山 [Guangdong, Lianxian County, Dadongshan Mout.]; 13 Jul. 1998; 陈翠萍采 [Chen Cuiping leg.]; SYSU. – Henan • 2 ♀♀; 河南信阳鸡公山 [Henan, Xinyang City, Jigongshan Mount.]; Aug. 1926; SYSU. – Hunan • 1 ♀; Hunan Province, Yanling Co., Nanfengmian, mixed forest, shrub, flower sifted and beating; 26 ° 16 ′ 32 ″ N, 113 ° 59 ′ 34 ″ E; alt. ca 1380 m; 27 May 2014; Peng, Shen, Yu and Yan leg.; SYSU • 1 ♀; Hunan Province, Guidong Co., Bamian Shan; 26 ° 17 ′ 59 ″ N, 113 ° 59 ′ 22 ″ E; alt. 1030 m; 31 May 2014; Peng, Shen, Yu and Yan leg.; light trap; SYSU. – Jiangxi • 1 ♀; W Jiangxi, Ji’an City, Jinggang Shan, Jingzhu Shan; 26 ° 29 ′ 45 ″ N, 114 ° 04 ′ 45 ″ E; alt. 1130 m; 2 Aug. 2014; Chen, Hu, Lv and Yu leg.; in the stream; SHNU. – Jiangsu • 1 ♀; 江苏南京 [Jiangsu, Nanjing]; Jul. 2009; SYSU. – Sichuan • 1 ♂; 四川省青川县青溪镇河口里村 [Sichuan Prov., Qingchuan County, Qingxi Town, Hekouli]; alt. 1127 m; 32 ° 31 ′ 31 ″ N, 104 ° 50 ′ 22 ″ E; 11 May 2019; 梁祖龙, 杨圳铭采 [Liang Zulong and Yang Zhenming leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFEFFEF551DCC303528FB69.taxon	distribution	Distribution Endemic to China. Known from Shanghai and Zhejiang. New for Chongqing, Guangdong, Henan, Hunan, Jiangxi, Jiangsu and Sichuan.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFEFFEF551DC84235A5F89C.taxon	description	Fig. 11	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFEFFEF551DC84235A5F89C.taxon	materials_examined	Material examined CHINA – Hainan • 1 ♂; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; 21 Mar. 1983; 梁少营 采 [Liang Shaoying leg.]; SYSU • 7 ex.; 海南岛尖峰岭 [Hainan, Jianfengling]; 18 Aug. 1963; SYSU • 3 ex.; 海南尖峰岭 [Hainan, Jianfengling]; 4 Feb. 1981; 陈振耀 [Chen Zhenyao leg.]; SYSU • 2 ♂♂; 海南陵水吊罗山 [Hainan, Lingshui, Diaoluoshan Mt.]; 26 Dec. 1983; 李济才采 [Li Jicai leg.]; SYSU. VIETNAM • 2 ♀♀; “ Tonkin: Hoabinh. A. de Cooman. B. M. 1940 – 13 ”; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFEFFEF551DC84235A5F89C.taxon	distribution	Distribution Known from India. New for China and Vietnam.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFFFFE15561CC4134CBFD7F.taxon	materials_examined	Material examined CHINA – Hainan • 1 ♂; 海南尖峰岭五分区 [Hainan, Jianfengling Mt., fifth division]; 8 Jul. 1981; 饶漱靖采 [Rao shujing leg.]; SYSU • 1 ♀; 海南尖峰岭天池 [Hainan, Jianfengling Mt., Tianchi]; Jul. 1981; SYSU. – Jiangxi • 1 ♂, 1 ♀; 江西井冈山弯坑 [Jiangxi, Jinggangshan Mt.]; 29 Apr. 2011; 采 集人: 赵爽 [Zhao Shuang leg.]; SYSU • 4 ♂♂, 1 ♀; 江西井冈山白银湖 [Jiangxi, Jinggangshan Mt.]; alt. 880 m; 23 Apr. 2011; 采集人: 赵爽 [Zhao Shuang leg.]; SYSU • 1 ♂; 江西井冈山白银湖 [Jiangxi, Jinggangshan Mt., Baiyinhu Lake]; alt. 800 m; 27 Apr. 2011; 采集人: 贾凤龙 [Jia Fenglong leg.]; SYSU • 2 ♀♀; 江西井冈山大井- 龙潭风景区 [Jiangxi, Jinggangshan Mt., Dajing-Longtang Scenic Area]; 3 Oct. 2010; 赵爽采 [Zhao Shuang leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFFFFFE15561CC4134CBFD7F.taxon	distribution	Distribution Endemic to China (Guangdong). New for Hainan and Jiangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF0FFE1551FCFBE35DEF892.taxon	description	Fig. 12	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF0FFE1551FCFBE35DEF892.taxon	materials_examined	Material examined CHINA – Guangdong • 1 ♂, 2 ♀♀; 深圳市大鹏半岛坪头岭 [Shenzhen, Dapeng Peninsula, Pingtouling Mt.]; 22 ° 39 ′ 48 ″ N, 114 ° 25 ′ 11 ″ E; alt. 259 m; 25 Jul. 2019; 梁祖龙, 杨圳铭, 姜卓寅, 郭光宇, 纪新元采 [Liang Zulong, Yang Zhenming, Jiang Zuoyin, Guo Guangyu and Ji Xinyuan leg.]; SYSU • 1 ♀; 广东省 深圳市大鹏半岛田头山 [Shenzhen, Dapeng Peninsula, Tiantoushan Mt.]; 22 ° 40 ′ 12 ″ N, 114 ° 24 ′ 44 ″ E; alt. 208.4 m; 30 Jul. 2019; 杨圳铭, 姜卓寅, 郭光宇, 纪新元采 [Yang Zhenming, Jiang Zuoyin, Guo Guangyu, Ji Xinyuan leg.]; SYSU • 7 ♂♂, 5 ♀♀; 广东省台山市上川岛猕猴省级保护区 [Guangdong Prov., Taishan City, Shangchuan Island Macaque Provincial Nature Reserve]; 21 ° 44 ′ 36 ″ N, 112 ° 50 ′ 33 ″ E; alt. 10 m; 18 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU • 2 ♂♂, 1 ♀; 广东省台山市上川岛猕猴省级保护区 [Guangdong Prov., Taishan City, Shangchuan Island Macaque Provincial Nature Reserve]; 21 ° 44 ′ 51 ″ N, 112 ° 49 ′ 36 ″ E; alt. 50 m; 17 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU • 1 ♀; 广东省台山市上川岛猕猴省级保护区石背潭 [Guangdong Prov., Taishan City, Shangchuan Island Macaque Provincial Nature Reserve, Shibeitang]; 21 ° 46 ′ 1 ″ N, 112 ° 49 ′ 2 ″ E; alt. 20 m; 19 Jan. 2019; 贾凤龙, 梁祖龙采 [Jia Fenglong and Liang Zulong leg.]; SYSU • 1 ♂; 广东肇庆鼎湖山 [Guangdong, Zhaoqing, Dinghushan Mt.]; 19 Jun. 1958; 利翠英 采 [Li Cuiying leg.]; SYSU • 1 ♂, 4 ♀♀; 广东肇庆鼎湖山 [Guangdong, Zhaoqing, Dinghushan Mt.]; 13 Oct. 1958; SYSU • 1 ♂, 3 ♀♀; 广东鼎湖山 [Guangdong, Dinghushan Mt.]; 8 Jul. 1964; 吴思添采 [Wu Sitian leg.]; SYSU • 1 ♀; 肇庆鼎湖山 [Zhaoqing, Dinghushan Mt.]; 3 Nov. 1974; 春强采 [Chun Qiang leg.]; SYSU • 1 ♀; 广东高要县鼎湖山 [Guangdong, Gaoyao County, Dinghushan Mt.]; 15 Jul. 1964; 何目秀采 [He Muxiu leg.]; SYSU • 1 ♀; 和平县九连山 [Heping County, Jiulianshan Mt.]; 26 Aug. 1986; 陈振耀采 [Chen Zhenyao leg.]; SYSU • 1 ♀; 广东连县大东山 [Guangdong, Lianxian County, Dadongshan Mt.]; 4 Sep. 1994; 彭红采 [Peng Hong leg.]; SYSU • 1 ♂, 2 ♀♀; 广东高明鹿田 [Guangdong Gaoming, Lutian]; 14 May 2004; 贾凤龙, 方小端采 [Jia Fenglong and Fang Xiaoduan leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF0FFE1551FCFBE35DEF892.taxon	distribution	Distribution Known from Vietnam. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF1FFE35522CC4135A4FDA4.taxon	description	Figs 13 – 14	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF1FFE35522CC4135A4FDA4.taxon	materials_examined	Material examined Syntype INDIA – Tamil Nadu • 1 ♂, 3 ♀♀; Madras; 1908; Maurice Régimbart leg.; MNHN EC 11801 to EC 11804. CHINA – Yunnan • 1 ♀; Yunnan Prov., Xishuangbanna, Botanical garden; 5 – 6 Jul. 2003; Hu and Tang leg; SHNU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF1FFE35522CC4135A4FDA4.taxon	distribution	Distribution Known from India and Pakistan. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF2FFE25547CFDC34C7FDCF.taxon	materials_examined	Material examined CHINA – Guangdong • 1 ♂; 广东肇庆鼎湖山 [Guangdong, Zhaoqing, Dinghushan Mt.]; 19 Aug. 1958; 利翠英采 [leg. Li Cuiying.]; SYSU • 1 ♀; 广东连县大东山 [Guangdong, Lianxian County, Dadongshan Mt.]; 13 Aug. 1998; 谭乐采 [Tan Le leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF2FFE25547CFDC34C7FDCF.taxon	distribution	Distribution Endemic to China. Known from Hainan. New for Guangdong.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF3FFE55544CF673312FDCF.taxon	materials_examined	Material examined CHINA – Guangxi • 1 ♂, 4 ♀♀; 广西隆林 [Guangxi, Longlin]; 26 May 1977; 黄治河采Huang Zhihe [leg.]; SYSU • 1 ♂, 1 ♀; 广西靖西底定 [Guangxi, Jingxi City, Diding Village]; 8 Aug. 2010; 黄建华 采 [Huang Jianhua leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF3FFE55544CF673312FDCF.taxon	distribution	Distribution Known from China (Guizhou, Yunnan), Vietnam and Laos. New for Guangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE556E1CF673501FC3A.taxon	description	Fig. 15	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE556E1CF673501FC3A.taxon	materials_examined	Material examined CHINA – Yunnan • 1 ♀; 云南小勐养 [Yunnan, Xiaomengyang]; alt. 810 m; 25 Mar. 1957; 蒲富基采 [Pu Fuji leg.]; IZCAS.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE556E1CF673501FC3A.taxon	distribution	Distribution Known from India. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE556F1C9523338F9EB.taxon	materials_examined	Material examined CHINA – Fujian • 1 ♂; 福建, 建阳黄坑桂林 [Fujian, Jianyang, Huangkeng, Guilin]; alt. 270 – 390 m; 5 Apr. 1960; 马成林采 [Ma Chenlin leg.]; IZCAS. – Hainan • 1 ex.; 海南尖峰岭 [Hainan, Jianfengling Mt.]; 13 Oct. 1983; 利翠英采 [Li Cuiying leg.]; SYSU • 1 ex.; 海南尖峰岭 [Hainan, Jianfengling Mt.]; 19 Dec. 1983; 利翠英采 [Li Cuiying leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE556F1C9523338F9EB.taxon	distribution	Distribution Endemic to south China. Known from Guangdong. New for Fujian and Hainan.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE45576CB0335C5FC97.taxon	materials_examined	Material examined CHINA – Guangxi • 1 ♂, 3 ♀♀; Guangxi Prov., Jinxiu County; 24 Jul. 2011; alt. 850 – 950 m; PENG Zhong leg.; SHNU. – Jiangxi • 1 ♂, 1 ♀; 江西井冈山香洲 [Jiangxi, Jinggangshan Mt., Xiangzhou]; 13 Sep. 2011; 采集人: 杨立军 、 李锦伟 、 成牧春 [Yang Lijun, Li Jinwei and Cheng Muchun leg.]; SYSU. – Zhejiang • 1 ♂, 1 ♀; Zhejiang Prov., Longwang Shan; 4 Oct. 2003; Li-Zhen Li leg; SHNU • 1 ♀; Zhejing Prov., Mt. Longwang; alt. 300 – 500 m; 24 Apr. 2004; Huang and Chi leg.; SHNU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE45576CB0335C5FC97.taxon	distribution	Distribution Endemic to China (Fujian, Guangdong, Hong Kong). New for Guangxi, Jiangxi and Zhejiang. Patrus procerus (Régimbart, 1884) comb. nov.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE45576CB0335C5FC97.taxon	distribution	Fig. 16	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE45576CB0335C5FC97.taxon	materials_examined	Material examined CHINA – Guangxi • 1 ♂, 2 ♀♀; 广西防城板八乡 [Guangxi, Fangcheng, Banbaxiang]; 4 Jun. 2000; 李 文柱采 [Li Wenzhu leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF4FFE45576CB0335C5FC97.taxon	distribution	Distribution Known from Cambodia. New for China.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF5FFE75548CECF362AFD1E.taxon	materials_examined	Material examined CHINA – Guangdong • 10 ex. 广东珠海横琴岛 [Guangdong, Zhuhai, Hengqin Island]; 12 Jul. 2005; 采集人: 贾粤 [Jia Yue leg.]; SYSU • 2 ♂♂, 1 ♀; 广东省肇庆市丹霞山 [Guangdong, Zhaoqing, Dinghushan Mt.]; 11 Jun. 2018; 梁祖龙采 [Liang Zulong leg.]; SYSU • 24 ex.; 深圳坪山区马峦山 [Shenzhen, Pingshan District, Maluanshan Mt.]; 22 ° 38 ′ 42 ″ N, 114 ° 19 ′ 38 ″ E; alt. 311 m; 27 Jul. 2019; 杨 圳铭, 姜卓寅, 郭光宇, 纪新元采 [Yang Zhenming, Jiang Zuoyin, Guo Guangyu and Ji Xinyuan leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF5FFE75548CECF362AFD1E.taxon	distribution	Distribution Widespread in the Oriental Region, known from Hainan, Hong Kong, Jiangsu, Macao in China. New for Guangdong.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF6FFE75547CFB633B5FB56.taxon	materials_examined	Material examined CHINA – Jiangsu • 1 ♀, 1 ♂; 江苏南京 [Jiangsu, Nanjing]; Jul. 2009; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF6FFE75547CFB633B5FB56.taxon	distribution	Distribution Endemic to China. Known from? Guangdong, Guizhou, and Hong Kong. New for Jiangsu.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF7FFE65557CC41335CFD04.taxon	materials_examined	Material examined CHINA – Guangdong • 1 ♀; 广东肇庆鼎湖山 [Guangdong, Zhaoqing, Dinghushan Mt.]; Jul. 1973; SYSU. – Jiangxi • 2 ♂♂, 2 ♀♀; 江西三清山 [Jiangxi, Sanqingshan Mt.]; 15 Aug. 2006; 贾凤龙采 [Jia Fenglong leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF7FFE65557CC41335CFD04.taxon	distribution	Distribution Endemic to China (Anhui, Zhejiang). New for Guangdong and Jiangxi.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF7FFE6558ECFBC348AF90F.taxon	materials_examined	Material examined CHINA – Guangdong • 15 ex.; 广东肇庆黑石顶 [Guangdong, Zhaoqing, Heishiding]; 25 Jul. 2018; 贾凤龙采 [Jia Fenglong leg.]; SYSU • 4 ex.; 广东封开县黑石顶 [Guangdong, Fengkai, Heishiding]; 23 ° 31 ′ N, 111 ° 52 ′ E; 20 – 22 Sep. 2014; 贾凤龙, 唐余丹采 [Jia Fenglong, Tang Yudan leg.]; SYSU • 7 ex.; 广东省封开县黑石顶 [Guangdong, Fengkai, Heishiding]; 23 ° 31 ′ N, 111 ° 52 ′ E; 20 – 22 Sep. 2014; 贾 凤龙, 林仁超, 唐余丹采 [Jia Fenglong, Lin Renchao, Tang Yudan leg.]; SYSU • 2 ex.; 广东封开黑石 顶 [Guangdong, Fengkai, Heishiding]; 11 Jul. 2000; 罗菁采 [Luo Jing leg.]; SYSU • 2 ♂♂; 广东封 开黑石顶 [Guangdong, Fengkai, Heishiding]; 5 Jul. 1987; 钟声雄采 [Zhong Shengxiong leg.]; SYSU • 1 ♂; 广东封开黑石顶 [Guangdong, Fengkai, Heishiding]; 3 Jul. 1987; 陈邵峰采 [Chen Shaofeng leg.]; SYSU • 1 ♂, 2 ♀♀; 广东鼎湖山 [Guangdong, Dinghushan Mt.]; 22 Jul. 1964; 吴思添采 [Wu Sitian leg.]; SYSU • 1 ♂; 广州河南 [Guangzhou, Henan]; 15 Oct. 1992; 邝骉采 [Kuang Biao leg.]; SYSU • 1 ♀; 广东乳源南岭半山腰保护站 [Guangdong, Ruyuan, Nanling Hillside Reserve Station]; alt. 1000 m; 23 Aug. 2010; 赵爽采 [Zhao Shuang leg.]; SYSU • 1 ♀; 广东高明杨梅镇 [Guangdong, Gaoming, Yangmei Town]; May 2004; 采集人: 贾凤龙 [Jia Fenglong leg.]; SYSU • 2 ♂♂, 2 ♀♀; 广东 高明杨梅镇 [Guangdong, Gaoming, Yangmei Town]; 23 – 26 Apr. 2006; 采集人: 贾凤龙 [Jia Fenglong leg.]; SYSU • 1 ♂, 1 ♀; 广东高明杨梅镇 [Guangdong, Gaoming, Yangmei Town]; 20 – 27 Apr. 2006; 袁采亚采 [Yuan Caiya leg.]; SYSU • 1 ♂; 广东高明杨梅镇 [Guangdong, Gaoming, Yangmei Town]; 20 – 27 Apr. 2006; 胡剑采 [Hu Jian leg.]; SYSU.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
56557D6EFFF7FFE6558ECFBC348AF90F.taxon	distribution	Distribution Endemic to south China. Known from Fujian. New for Guangdong.	en	Liang, Zulong, Angus, Robert B., Jia, Fenglong (2021): Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae). European Journal of Taxonomy 767: 1-39, DOI: https://doi.org/10.5852/ejt.2021.767.1481
