taxonID	type	description	language	source
5038879DCE2A550590A12A4CFDDAFCFE.taxon	description	Figs 1 – 8	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE2A550590A12A4CFDDAFCFE.taxon	discussion	Remark. As noted above, the original description of this species was rather superficial and incomplete, especially as regards the crucial gonopodal structure (Pocock 1892). Brief redescription. Body with 20 rings (19 + T), length ca 10.5 mm. Coloration mostly dark grey-brown with contrasting pallid legs, tegument heavily coated with an earth crust (Figs 4 – 6). Head fully covered from above by a flabellate anterior brim of a dome-shaped collum, the latter with 5 + 5 equal, distinct, rounded lobulations at anterior margin. Antennae C-shaped, short and clavate, antennomere 5 being the largest and subequal in length to 2 nd. Paramedian tubercles / crests (PM) especially strongly developed, high, each tightly fused medially almost all along and only apically slightly divided, inclined anteriad on collum and rings 2 – 4, subvertical until ring 16, thereafter declined increasingly caudad (Figs 3 & 4). PM mostly thickened and vaguely bilobate apically, only last few PM trilobate (Fig. 4). Paraterga very low, oblique and strongly declined, vaguely bi- or trilobate at lateral margin (Figs 3 – 6). Pore formula normal (5, 7, 9, 10, 12, 13, 15 – 19), ozopores borne on distinct porosteles on rings 5, 7, 9, 10, 12, 13 and 15, but opening flush on dorsal surface at base of caudal lobulation on rings 16 – 19 (Figs 3 – 6). Surface below PM microgranulate, with neither DL nor i, nor Am, nor Cm discernible beneath an earth crust. PM 19 subhorizontal, fused medially and forming a deep caudal notch, tip of epiproct being slightly visible from above. Legs robust, densely setose, each prefemur with a particularly long distoventral seta; claw small and simple (Figs 5 & 6). Gonopods (Figs 7 & 8) relatively simple, in situ held almost parallel to each other, with only tips of telopodites (= solenomeres), both slightly curved ventrad and poorly bifid, crossing medially; each gonopod consisting of a large, globose, densely microsetose and micropapillate coxite and a unipartite, rather long, ribbon-shaped, distally attenuating telopodite / solenomere curved caudad, strongly sunken inside a deep gonocoel and terminating the seminal groove on top.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE2A550590A12A4CFDDAFCFE.taxon	distribution	Distribution. Pundaluoya, Sri Lanka.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE2A550390A12981FC67FE1E.taxon	type_taxon	Type-species: Pyrgodesmus obscurus Pocock 1892, by monotypy.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE2A550390A12981FC67FE1E.taxon	synonymic_list	Other species included: Pyrgodesmus permutatus (Attems, 1936), comb. nov.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE2A550390A12981FC67FE1E.taxon	diagnosis	Pyrgodesmus seems to be particularly similar to Klimakodesmus in most somatic characters, as correctly noted by Carl (1932) and recently reinstated by Aswathy et al. (2021). Yet both genera are distinct primarily in gonopodal structure: hypertrophied coxites, a deep gonocoel and ribbon-shaped, unipartite, distally attenuating, but not acuminate telopodites (Figs 5 – 8, 29 & 30), vs. moderately incrassate, apparently smaller, subspherical or squarish coxites, each of which supports a strong and unipartite telopodite represented solely by a slender, strongly exposed, increasingly attenuating and acuminate solenomere. In addition, PM crests, including the one on the penultimate ring that overhangs and conceals the epiproct from above, are much higher and stronger, whereas DL reduced, in Pyrgodesmus compared to Klimakodesmus species (Aswathy et al. 2021).	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE28550290A12B40FDD2FCE6.taxon	description	Figs 9 – 30	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE28550290A12B40FDD2FCE6.taxon	discussion	Remark. As noted above, the original description and illustrations of this species were most detailed (Silvestri 1920), albeit misidentified as belonging to Pyrgodesmus obscurus. Silvestri’s iconography is reproduced in full (Figs 9 – 30). Brief redescription. Body length ca 11.5 mm. Coloration mostly black-brown, regardless of tegument being heavily coated with an earth crust (Figs 9 – 12) or clean (Figs 13 – 18). All main characters as in P. obscurus, except as follows. Paramedian tubercles / crests (PM) slightly less strongly developed, high, largely fully separated and not fused medially even at base (except for PM 19), inclined anteriad on collum and on rings 2 and 3 (Figs 9 – 18). Surface below PM regularly microgranulate, with neither DL nor i, nor Am, nor Cm discernible beneath an earth crust (Figs 9 – 12), but readily visible in a clean animal (Figs 13 – 18). Tip of epiproct invisible from above between basally fused PM 19 (Fig. 26). Gonopods (Figs 29 & 30) much as in P. obscurus, but gonocoel slightly less deep, coxites apparently neither conspicuously microsetose nor micropapillate, telopodites / solenomeres more clearly exposed, but stout, their tips being somewhat curved dorsad.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
5038879DCE28550290A12B40FDD2FCE6.taxon	distribution	Distribution. Paradenyia, Sri Lanka.	en	Golovatch, Sergei I. (2021): Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida). Zootaxa 5068 (4): 572-578, DOI: https://doi.org/10.11646/zootaxa.5068.4.6
