taxonID	type	description	language	source
3A1C87D99717FFD5FF1F294CB8CCA741.taxon	diagnosis	Diagnostic remarks. According to Likhitrakarn et al. (2013), the subfamily Alogolykinae seems best characterized by the absence of a clear-cut division of the solenophore or solenophore-like structure near / around the solenomere into a membranous lamina medialis or a similarly membranous lamina lateralis, or both. In Polydrepanini, this is sometimes coupled with a twisted, helicoid course of the seminal groove. In addition, the gonofemorite in Alogolykinae is often nearly lacking to very strongly abbreviated, while numerous species show adenostyles on male femora 1 and / or 2. The latter two traits are characteristic of some species of the subfamily Australiosomatinae Brölemann, 1916 as well, and are occasionally present also in the subfamily Paradoxosomatinae Daday, 1889, but in such cases the solenophore branch or branches remain free and never fully sheathe a primitively long, strong and rod-like solenomere.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99717FFD5FF1F294CB8CCA741.taxon	discussion	The attempt by Sankaran and Sebastian (2018) to denominate some membranous structures of the solenophore in their new species of Polydrepanum Carl, 1932 as the homologue to a lamina lateralis can hardly be accepted, as discussed below. Basically, the stronger / thicker the solenomere branch in both the order Polydesmida and the family Paradoxosomatidae, the less strong and intricate the support the solenophore renders to it tends to be. In contrast, the thinner / weaker the subflagelliform or filiform solenomere, the stronger, more intricate and solid the support of the solenophore tends to be.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99716FFD5FF1F2E27BD58A43D.taxon	discussion	Diagnostic remark. Based on Golovatch (2014, 2015), contrary to the Alogolykini which typically shows a nearly fully to very strongly reduced gonofemorite, coupled with a basally originating, long, thick, mostly rod-like solenomere, the tribe Polydrepanini seems to be best characterized by a clearly elongate, mostly slender, sometimes twisted / helicoid gonofemorite (fe) and a more or less thin, mostly flagelliform, suberect, distal to subapical solenomere (sl) often forming a conspicuous loop / curve (k) near the base (e. g., Figs 3 – 8).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99716FFD5FF1F2BECB810A117.taxon	description	Figs 1 – 5	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99716FFD5FF1F2BECB810A117.taxon	discussion	Remarks. Among the most important characters of this species are the presence of adenostyles only on male femora 1 (Figs 1 and 2), coupled with the gonofemorite (fe) being slender and long, devoid of any traces of torsion, while the solenophore (sph) is peculiar in shape: a single, very large, unequally bifid, pincer-shaped, lateral lobe (a), and one apicomesal spine (b) lying distal to the basal loop (k) of a free solenomere (sl). Base of sph on ventral side forming a protecting lobe (lo) to shield the basal loop (k) of sl. Distal half of both sph and sl suberect and directed ventrad (Figs 3 – 5). Because the gonopodal femorite in P. asperrimum is untwisted (Figs 3 – 5), unlike that of the type species P. tamilum (see below and Figs 17 – 19), contrary to Jeekel (1968), we are inclined to revive Grammorhabdus Carl, 1932, stat. revalid., to accommodate both G. asperrimus as its type species and a recently described species treated immediately below.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99716FFD5FF1F2DFBB853A20F.taxon	materials_examined	Type species: Polydrepanum tamilum Carl, 1932, by monotypy. Taxonomic composition. Before providing a refined generic diagnosis, the species composition of Polydrepanum must be clarified, as this genus is perhaps the most ambiguous taxonomically amongst the Alogolykinae. Even though Sankaran and Sebastian (2018) have recently reviewed the genus, provided its diagnosis, described a new species, and gave a key to species, their results require revision. The following nominate Polydrepanum species have previously been distinguished, five of which are named and the remaining three unnamed. All of them are presented below in alphabetic order, also being catalogued, rediagnosed, illustrated, and commented upon as regards their present status. All four true Polydrepanum species are keyed as well.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99711FFD2FF1F2A7DBD39A1AA.taxon	description	Figs 6 – 8	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99711FFD2FF1F2A7DBD39A1AA.taxon	discussion	Remarks. Among the most important characters of this species are the presence of adenostyles on both male femora 1 and 2, coupled with the gonofemorite being fairly slender and long, devoid of any traces of torsion, delimited from the acropodite by a very distinct postfemoral sulcus / cingulum (su), while the solenophore (sph) is peculiar in shape: two small, subdentiform, laterobasal lobes (a), and two similarly small, subdentiform, apicomesal lobes (b) lying distal to the basal loop (k) of a free solenomere (sl). Base of sph on ventral side forming an elongated and undulated protecting lobe (lo) to shield the basal sl loop. Distal half of both sph and sl subcircular and directed ventrad (Figs 6 – 8). It is the unusually elongated and undulated lo that Sankaran and Sebastian (2018) must have mistaken for a lamina lateralis. However, that homologization is hardly correct, because the same membranous structure in the undoubtedly very similar P. asperrimum is interrupted in the subbasal and middle parts, being largely traced only subapically (Figs 3 – 5). Because the gonopodal femorite in P. fissum is untwisted (Figs 6 – 8), unlike that of the type species P. tamilum (see below and Figs 17 – 19), but very much like in Grammorhabdus asperrimus, contrary to Sankaran and Sebastian (2018), we are inclined to assign P. fissum to Grammorhabdus Carl, 1932, comb. nov.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99710FFD3FF1F2A05BD2DA1A5.taxon	description	Figs 9 – 12	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99710FFD3FF1F2A05BD2DA1A5.taxon	discussion	Remarks. Among the most important characters of this species are the presence of dense metatergal setation, and of adenostyles only on male femora 2, coupled with the gonofemorite being quite slender and long, but with clear evidence of torsion, while the solenophore is also peculiar in shape and rather similar to that of P. asperrimum: a large, rounded, mesal lobe (lo), but only a single apicolateral spine (a) distal to the basal loop of a free solenomere (sl). Base of lo on mesal side forming a protecting shield for the basal sl loop. Distal half of solenophore clearly curved and directed laterad (Figs 9 – 12). Despite some critical comments following the original description of P. horridum (Bano, 1997, 1998; Bano & Murthy, 1997; Nguyen & Sierwald, 2013; Golovatch & Wesener, 2016) that suggested the erection of a new genus to accommodate this species alone, we still feel that the distinctions are not strong enough to do so, because both P. horridum and P. tamilum (cf. Figs 9 – 12 and 17 – 19) share such decisive characters as a twisted gonofemorite and the basal loop of a flagelliform solenomere (sl) protected by a large lobe (lo) of a distally strongly curved solenophore (sph). In other words, we consider P. horridum as being congeneric with P. tamilum.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99713FFDEFF1F2A08BBA2A6A6.taxon	description	Figs 13 – 15	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99713FFDEFF1F2A08BBA2A6A6.taxon	discussion	Remarks. According to Jeekel (1968, 1980 a), this species merits a genus of its own. Indeed, in addition to both male femora 1 and 2 showing adenostyles (Figs 13 and 14), the gonopodal femorite (fe) is strongly reduced in size and shows clear evidence of torsion, while the acropodite is tripartite, being divided basally into a dorsal spine (a), a coiled, ribbon-shaped, middle process (r), and the most voluminous ventral piece supporting most of the solenophore (sph), including a midway mesal spine (s) and a subapical mesal spine (b) near an unequally bifid and acuminate tip. The terminal third of sph is strongly unciform, membranous, gradually attenuating distad and directed ventromesad, with a free solenomere (sl) being flagelliform and almost as long as terminal sph third (Fig. 15). Sankaran and Sebastian (2018) transferred P. implicatum to Telodrepanum, but this was an obvious mistake for the reasons presented below. On balance, in full agreement with nearly all authors following the original description (Carl 1941), the above distinctions of P. implicatum from the true, typical Polydrepanum (see above and below) and Polydrepanini generally are sufficiently strong to warrant the erection of a new genus of Alogolykini, Carlogonopus gen. nov. (gender: masculine), to accommodate the type species alone, C. implicatus (Carl, 1941), comb, nov. We dedicate the new genus to Johann Carl (1877 – 1944), the late prominent Swiss specialist in diplopod systematics (Hollier et al. 2020) who described that species. This new genus is primarily distinguished from all contribal genera by a very short, but still quite discernible gonofemorite (fe), coupled with a flagelliform solenomere (sl) originating distally and sheathed by a prominent and complex solenophore (sph) (Fig. 15). In this respect, the position of Carlogonopus gen. nov. seems to be somewhat intermediate between the tribes Polydrepanini and Alogolykini to bridge both. However, we are inclined to treat the new genus within Alogolykini, because the gonofemorite in C. implicatus is about as short (= strongly reduced) as in Yuennanina, perhaps its closest relative, whereas subflagelliform solenomeres, albeit originating parabasally and obviously stronger, are also observed in Touranella species (Attems 1936, 1937). Bhakat (2021) has very recently described a new paradoxosomatid genus and species from Birbhum district, West Bengal state, eastern India: Manikidesmus suriensis Bhakat, 2021. The author assigned that millipede to the tribe Polydrepanini, but unfortunately the description is poor and the images are small and unclear. Because the gonofemorite appears to be considerably shortened, while the acropodite is complex and said to be bipartite, it seems better either to place it closer to Carlogonopus gen. nov. within the tribe Alogolykini or regard it provisionally as a disjunct Delarthrum. To exacerbate the situation, Manikidesmus suriensis as a name is not available yet for zoological nomenclature, having been proposed in a preprint journal issued only online, while taxonomically the name is a “ nomen nudum ” not registered in ZooBank (like the journal itself).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971DFFDEFF1F2E03BBADA58E.taxon	description	Figs 16 – 20	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971DFFDEFF1F2E03BBADA58E.taxon	discussion	Remarks. Being the type species of Polydrepanum, it shows only male femora 2 equipped with adenostyles (Fig. 16), while the gonopod demonstrates a twisted femorite and a complex solenophore (sph), the latter being divided into a small, subquadrate, lateral protecting lobe (lo) with two adjacent, curved, distofemoral processes (a and b), both latter forming a nearly complete dorsal ring and lying distal to the basal loop (k) of a free and subacuminate solenomere (sl). Base of lo on ventral side forming a protecting shield for the basal sl loop k. Distal half of solenophore strongly curved and directed dorsad (Figs 17 – 20). Because P. tamilum shows a clearly twisted gonofemorite (Figs 17 – 20), vs. an untwisted one in P. asperrimum (Figs 3 – 5) or P. fissum (Figs 6 – 8), only two latter species can be considered congeneric and belonging to Grammorhabdus. In contrast, only four species, including both P. tamilum and P. horridum, appear to represent true named Polydrepanum, as further discussed below.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971CFFDFFF1F2F97BC60A427.taxon	description	Figs 21, 22	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971CFFDFFF1F2F97BC60A427.taxon	discussion	Remarks. An unnamed species from an unspecified locality in Karnataka state, India, the left gonopod of which has been presented in a crude sketch by Bano and Murthy (1997), reproduced here in Figs 21 and 22. This species, like both following ones, is anyway to be neglected, being unnamed and based on material of unknown provenance and whereabouts, shows a clearly torsate gonofemorite, a basal loop of the solenomere, and a distinctly coiled and slender distal half of the solenophore. This condition resembles that observed in both P. tamilum and P. horridum (cf. Figs 9 – 12, 17 – 20 and 21, 22) and thus may prove to be their potential congener (see also below).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971CFFDCFF1F2807BBFDA739.taxon	description	Figs 23, 24	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971CFFDCFF1F2807BBFDA739.taxon	discussion	Remarks. An unnamed species from an unspecified locality in Karnataka state, India, the right gonopod of which has been presented in a crude sketch by Bano and Murthy (1997), reproduced here in Figs 23 and 24. Because this species, likewise to be neglected, being unnamed and based on material of unknown provenance and whereabouts, seems to show an untwisted gonofemorite, a basal loop of the solenomere, and a distinctly coiled and slender distal half of the solenophore, it does resemble Grammorhabdus asperrimus or G. fissus (cf. Figs 3 – 5, 6 – 8 and 23, 23) and thus may prove to be their potential congener.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971FFFDDFF1F2860BB26A7F1.taxon	description	Figs 25, 26	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971FFFDDFF1F2860BB26A7F1.taxon	discussion	Remarks. An unnamed species from an unspecified locality in Karnataka state, India, the right gonopod of which has been presented in a crude sketch by Bano and Murthy (1997), reproduced here in Figs 25 and 26. Because this species, albeit to be neglected in also being unnamed and based on material of unknown provenance and whereabouts, seems to show an untwisted gonofemorite, no basal loop of the solenomere, and a tripartite and complex solenophore, the distoventral half of which is distinctly enlarged, curved and acuminate, but the ventrobasal part bearing a process, it cannot be a Polydrepanum (cf. Figs 17 – 20 and 25, 26). Instead, this species so readily resembles the new Delarthrum to be described below that both these species are clearly congeneric, yet representing a genus different from Polydrepanum.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971EFFD8FF4F2A8CBCF9A45A.taxon	description	Gonofemorite elongate, slender and clearly twisted; solenophore mostly with a lateral lobe (lo) whose base on ventral side sometimes forming a protecting shield (lo) for the basal loop / curve of a long subflagelliform solenomere (sl). Distal half of solenophore elongate and unciform (Figs 9 – 12, 17 – 20).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971EFFD8FF4F2A8CBCF9A45A.taxon	synonymic_list	Polydrepanum tamilum Carl, 1932 (the type species), Polydrepanum horridum Golovatch, 1984, Polydrepanum granuliferum (Attems, 1936), comb. nov., Polydrepanum lamprum (Chamberlin, 1920), comb. nov.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971EFFD8FF4F2A8CBCF9A45A.taxon	discussion	Gyrodrepanum Carl, 1932, with the single species, G. lamprum (Chamberlin, 1920) (= Orthomorpha lampra Chamberlin, 1920, = Orthomorpha (Gyrodrepanum) bimontana Carl, 1932), described from Fiji and southern India (Carl 1932; Jeekel 1972; Bano & Murthy 1997), seems to represent yet one more true Polydrepanum, because it likewise shows an elongate, slender and clearly twisted gonofemorite, three distofemoral processes (a, b and c), a curved solenophore (sph) and, above all, a flagelliform free solenomere (sl) with a distinct basal loop and a protecting lobe (lo), however small, to shield the loop on the lateral side (Figs 46 – 50). Hindomorpha Golovatch, 1984, erected for Sundanina granulifera Attems, 1936 (= H. granulifera (Attems, 1936 )) as the type species (Golovatch 1984), is actually another Polydrepanum, because it also shows an elongate, slender and clearly twisted gonofemorite, two distofemoral processes (a and b), a curved solenophore (sph) and, above all, a flagelliform free solenomere (sl) with a distinct basal loop and a protecting lobe (lo), however inconspicuous, to shield the loop on the lateral side (Figs 44, 45). In other words, both Hindomorpha Golovatch, 1984 and Gyrodrepanum Carl, 1932 are new junior subjective synonyms of Polydrepanum Carl, 1932, both syn. nov., and both P. granuliferum (Attems, 1936) and P. lamprum (Chamberlin, 1920) are new combinations, comb. nov. ex Hindomorpha and Gyrodrepanum, respectively.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971BFFC5FF1F2D10BBFFA0E4.taxon	description	Gonofemorite elongate, slender and untwisted, its apical part typically obliquely truncate ventrad or dorsad; solenophore usually complex, protecting a long, flagelliform to somewhat ribbon-shaped solenomere (sl) devoid of a basal loop / curve (Golovatch 2014 b) (Figs 25 – 44).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9971BFFC5FF1F2D10BBFFA0E4.taxon	discussion	Species included: Delarthrum obscurum Attems, 1936, the type species of Delarthrum (Figs 27 – 30), and numerous others, including the type species of the following generic categories previously synonymized under Delarthrum (see above): Armolites spiniger (Attems, 1936), the type species of Armolites (Figs 31 – 33), Martensosoma unicolor (Attems, 1936), the type species of Martensosoma (Figs 34 – 36) (NB: although formally synonymized with Delarthrum (Golovatch 1992), Martensosoma spp. still retain two gonopodal characters that make them distinct: the femorite is devoid of any evident distal outgrowth, and the solenomere is rather ribbon-shaped than filiform; thus, Martensosoma seems best to be regarded as a defined Himalayan species-group of Delarthrum), Nepalomorpha kuznetsovi Golovatch, 1994, the type species of Nepalomorpha (Figs 37 – 39), Orophosoma hingstoni (Carl, 1935), the type species of Orophosoma (Figs 40, 41), and Parorthomorpha longiseta Golovatch, 1994, the type species of Parorthomorpha (Figs 42 – 44). All species of Delarthrum have been listed or described in Golovatch (2014 b, 2015, 2016 b), while not only Martensosoma, but also the other ex-generic categories above, currently all junior synonyms, can be considered as representing species groups of Delarthrum. Considering the removal of two Dasypharkis (see below) and the addition of a new Delarthrum from Kerala, the genus Delarthrum presently counts 55 species, the taxonomically most important of which (the type species of generic categories, presently the species groups) are being briefly reassessed and illustrated above.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99706FFC5FF1F2840BB51A1E9.taxon	discussion	Revived herewith from the synonymy with Delarthrum for both Dasypharkis rugulosa (Carl, 1932), the type species of Dasypharkis (Figs 45, 46), and D. pumila (Attems, 1944), the two from southern India and showing an elongate, slender, erect and clearly twisted gonofemorite and a fairly simple solenophore (sph) protected by its two lobes either side a flagelliform and upright solenomere (sl), the latter devoid of a basal loop / curve (Jeekel 1980 b).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC2FF1F2A9FBAEFA14D.taxon	discussion	Gonofemorite slender, long and untwisted, solenophore (sph) complex, with a large lateral lobe (lo) and two separate apicomesal spines or teeth (a and b) lying distal to basal loop (k) of a free solenomere (sl). Base of lo on ventral side forming a protecting shield for basal loop k of sl. Distal half of both sph and sl suberect and directed ventrad (Figs 3 – 8).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC2FF1F2A9FBAEFA14D.taxon	synonymic_list	Grammorhabdus asperrimus Carl, 1932 (the type species), Grammorhabdus fissus (Sankaran & Sebastian, 2018),	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC2FF1F2A9FBAEFA14D.taxon	discussion	both from southern India (Carl 1932; Sankaran & Sebastian, 2018).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC3FF1F2877BA08A15E.taxon	description	Gonofemorite slender, long, suberect and untwisted; solenophore (sph) large and cupshaped, expanded distad, with a distomesal protecting lobe (lo) to shield the basal loop (k) of a free flagelliform solenomere (sl) (Figs 55 – 57).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC3FF1F2877BA08A15E.taxon	synonymic_list	Pocockina pilifera (Pocock, 1895) (= Orthomorpha pilifera Pocock, 1895) (Figs 54 – 57), Pocockina jaegeri Golovatch, 2016 b, Pocockina schawalleri Golovatch, 2016 a,	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99701FFC3FF1F2877BA08A15E.taxon	discussion	all three from Myanmar (Jeekel 1965; Golovatch 2016 a, 2016 b).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99700FFC3FF1F2863BBE7A1A4.taxon	discussion	Gonofemorite slender, long, suberect and untwisted, with two distomesal processes (a and b), of which b is unusually large and shield-like; solenophore (sph) large and strongly coiled, with a basomesal protecting lobe (lo) to shield the basal loop of a free flagelliform solenomere (sl) (Figs 60 – 64).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99700FFC3FF1F2863BBE7A1A4.taxon	synonymic_list	Species included: T. badaga Carl, 1932, the type species (Carl 1932), as well as one more, yet undescribed species, both from southern India (Golovatch 1984).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99702FFC1FF1F2943BDAEA1EC.taxon	discussion	Gonofemorite slender, long, suberect and untwisted, with several (1 – 4) distomesal processes / spines; solenophore (sph) unusually small and strongly twisted, devoid of a basomesal protecting lobe to shield an indistinct basal loop / curve (k) of a free flagelliform solenomere (sl) (Figs 67, 68).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99702FFC1FF1F2943BDAEA1EC.taxon	synonymic_list	Xiphidiogonus spinipleurus Carl, 1932, the type species, Xiphidiogonu dravidus Carl, 1932, Xiphidiogonus hendersoni Carl, 1932,	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D99702FFC1FF1F2943BDAEA1EC.taxon	discussion	all from southern India (Carl 1932).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9970DFFCCFF1F2A8CBDD8A0CE.taxon	description	Figs 69 – 87.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9970DFFCCFF1F2A8CBDD8A0CE.taxon	materials_examined	Material examined. Male holotype (CATE- 61604 A), 2 male (CATE- 61604 B and CATE- 61604 C) and 2 female paratypes (CATE- 61604 D and CATE- 61604 E), India, Kerala state, Thrissur district, 10 ° 21 ’ 19 ’’ N, 76 ° 12 ’ 48 ’’ E, 29 m a. s. l., in June 2021 and 1 male (CATE- 61604 J) and 2 female paratypes (CATE- 61604 K and CATE- 61604 L all in 5302 B – CATE- 5302 I) from the sacred groves of Valliyur kavu, Manathavady, Wayanad district, 11 ° 48 ’ 08 ’’ N, 76 ° 01 ’ 55 ’’ E, 716 m a. s. l., M. D. Aswathy leg. Name. To emphasize the anomalous absence of adenostyles from both male legs 1 and 2.	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9970DFFCCFF1F2A8CBDD8A0CE.taxon	diagnosis	Diagnosis. Differs from congeners by the absence of adenostyles from both male legs 1 and 2, coupled with the sexually dimorphic colour pattern, the transverse tergal sulci starting with the collum, and the peculiar gonopodal conformation, primarily the presence of a small, thumb-shaped, distoventral projection (a) basal to the division of the postfemoral part into (1) the main, dorsal piece of the solenophore (sph), which is long, obliquely truncate dorsad at the base, bifid in the distal third and slightly curved mesad, and (2) a dorsal process that is split immediately at the base into a longer, stalked, leaf-shaped outgrowth (b) with peculiar, ragged, saw-like distal margins, and a much shorter, simple, dorsobasal peg (z) (Figs 84 – 87).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
3A1C87D9970DFFCCFF1F2A8CBDD8A0CE.taxon	description	Description. Length of holotype, 17.9 mm, of paratypes, 17.0 – 17.6 mm (male) or 14.0 – 14.8 mm (female). Width of midbody rings, 2.6 mm (holotype), 1.9 – 2.1 mm (male paratypes) or 2.0 – 2.3 mm (female paratypes). Coloration sex-dimorphic: in vivo, dorsum dark brown to black, venter chestnut brown (male) (Figs 69, 70) to yellowish white (female) (Fig. 71); paraterga mostly contrasting pale. Legs light brown (male) or whitish (female). Coloration after one month of preservation in 96 % ethanol somewhat darkened, but pattern evident (Figs 69 – 76). Body with 20 rings in both sexes. In width, collum <head <ring 2 <3 <4 <5 <6 <7 <8 <9 = 10, body gradually tapering thereafter (Figs 72, 73). Vertex glossy and bare; stipes, cardo and clypeus moderately setose, epicranial sulcus distinct and ending before the level of antennal sockets. Clypeal region with ca 5 + 5 setiferous pits. Antennae moderately long, only slightly clavate, extending back to ring 4 when stretched caudolaterally (Figs 72, 74). Relative antennomere lengths: 1 <6 <4 <2 <5 <3. Antennomere 6 broadest apically, post-antennal groove shallow, diameter of antennal socket and isthmus between sockets subequal, 0.24 and 0.23 mm, respectively (Fig. 77). Antennomeres 5 – 7 each with a distinct dorso-apical group of basiconic sensilla (both male and female), these being especially conspicuous on antennomere 5 (Figs 72, 74). Paraterga mostly well developed, keel-shaped, set at about upper third of body height (Figs 69 – 76). Collum ovoid, paraterga small, subtriangular and broadly rounded. Anterior margin of collum with a single transverse row of fragile short setae (Fig. 72). Anterior and posterior margins of following rings rather straight and parallel. Anterior margin with a transverse row of 2 + 2 short setae similar to those on collum. Paraterga 2 as usual, set clearly lower than 3 rd, squarish in shape, anterior corner / lobe drawn anteriad. Following paraterga 3 – 17 regularly rounded anterolaterally and increasingly drawn caudolaterad into acute triangles, but still lying within rear tergal margins (Figs 72 – 76), vs. small, spiniform and slightly drawn past rear tergal margin on rings 18 and 19 (Figs 73, 76). Lateral calluses of paraterga thin, only slightly thicker on pore-bearing rings than on poreless ones (Figs 69, 70), smooth, delimited by distinct sulci both dorsally and, to a lesser degree, ventrally; usually with a few short setae retained at lateral margin. Tegument generally smooth and shining, prozonae very delicately shagreened, metazonae mostly striate / striolate at bases of paraterga and near transverse sulci, surface being slightly undulate (Figs 72, 73). Tergal setae mostly short and inconspicuous, present both on paraterga and in anterior half of metaterga. Two deep transverse sulci present on collum (Fig. 72), followed by one prominent sulcus on metazonae 2 – 18, smooth at bottom and reaching the base of paraterga (Figs 72, 73). A faint axial line traceable on most metazonae until 18 th (Figs 72, 73). Strictures between pro- and metazonae narrow, nearly smooth, at most faintly striolate (Figs 72 – 76). Pore formula normal (5, 7, 9, 10, 12, 13, 15 – 19), ozopores small, circular, opening at ca 1 / 3 off caudolateral corner of pore-bearing calluses (Figs 74 – 76). Pleurosternal carinae decreasing in size from 2 to 4. Epiproct conical and flattened dorsoventrally, extending well beyond anal valves; tip truncate, subapical lateral incisions evident; hypoproct subtriangular, with 1 + 1 setae on minute knobs (Fig. 79). Legs moderately long and slender, clearly longer and thickened in male compared to female (Figs 70, 71), rather densely setose ventrally and with short claws; male legs with tarsal brushes absent only from last two leg-pairs (Figs 74 – 76, 78, 83). Prefemora and coxae each bearing a moderately long and stiff distoventral seta. Podomere length ratios: coxa <prefemur <tibia = postfemur <tarsus <femur (Fig. 83). Neither male leg-pair 1 nor 2 with any adenostyles, only coxae 2 as usual, with gonopores on small knobs (Figs 81, 82). Sternal lobe between male coxae 4 small, conical and densely setose. Gonopodal aperture transverse and suboval in shape, about 2 / 3 as wide as prozonite 7, its rim being not elevated, but thickened laterally (Fig. 78). Gonopods high, complex, in situ held parallel to each other (Figs 78, 80). Coxite darkened, about as long as femorite, subcylindrical, setose distoventrally (Figs 84 – 87); cannula a short curved tube, as usual. Prefemoral part short, globose, densely setose as usual, with a particularly long seta distoventrally. Femorite (fe) relatively stout and short, somewhat enlarged distad, subclavate, almost as long as acropodite (= postfemoral part), subcylindrical, without evidence of torsion; a small, thumb-shaped, distoventral projection (a) basal to acropodite bifurcation into a longer, larger, more slender, subacuminate, ventral, solenophore branch (sph), rather slightly curved dorsad and bearing both a strong spine (k) in distal third and a free solenomere (sl), and a shorter, more elaborate, dorsal process divided immediately at base into a longer, stalked, leaf-shaped outgrowth (b) with peculiar, ragged, saw-like distal margins, and a much shorter, simple, dorsobasal peg (z). Seminal groove first running all along mesal side of femorite, then turning slowly towards bifurcation point before moving onto a free, flagelliform sl tightly attached to mesal side of sph, with only a short tip of sl remaining exposed over sph. Division of fe and acropodite through a transverse sulcus, or of sph into laminae, wanting (Figs 84 – 87).	en	Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha, Sudhikumar, Ambalaparambil Vasu (2021): Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae). Zootaxa 5068 (4): 485-516, DOI: https://doi.org/10.11646/zootaxa.5068.4.2
