identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
04D6F12E943D5710A5862DFCA1508456.text	04D6F12E943D5710A5862DFCA1508456.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zospeum simplex Inäbnit, Jochum & Neubert, sp. nov.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Zospeum simplex 
Inaebnit
, Jochum &amp; Neubert, sp. nov.
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            <p>Figures 1, 3</p>
            <p>Type specimens.</p>
            <p> Holotype: NMBE 568060, Jama Dobravljevac, 25.08.2019, leg. R. Slapnik &amp; J.  Valentinčič ; Paratypes: NMBE 568061-568063; SMF 349425, 4 shells; RSC 3760, 6 shells; Jama Dobravljevac, 25.08.2019, leg. R. Slapnik &amp; J.  Valentinčič . </p>
            <p>Specimens examined.</p>
            <p> NMBE 568054,  Špilja Dahna, 03.09.2009, leg. A. Schoenhoffer; NMBE 568055-568058, Jama u kamenolomu, 24.08.2019, leg. R. Slapnik &amp; J.  Valentinčič ; NMBE 568059,  Vranjača , 24.08.2019, leg. R. Slapnik &amp; J.  Valentinčič . </p>
            <p>Diagnosis.</p>
            <p>Shell usually ca. 1.3 mm in height, transparent, conical, peristome thickened, roundish, with a differentiated parietal shield, lamellae not present.</p>
            <p> Measurements (n = 9): Shell height: 1.26-1.42 mm (mean: 1.378  ± 0.047 mm); shell width: 0.93-1.04 mm (mean: 0.976  ± 0.035 mm); aperture height: 0.54-0.67 mm (mean: 0.6  ± 0.037); aperture width: 0.54-0.65 mm (mean: 0.601  ± 0.033 mm); number of whorls: 5-5.5. </p>
            <p>Description.</p>
            <p>Shell conical, translucent when fresh; suture deep; aperture somewhat roundish to reniform; parietal shield clearly differentiated from the rest of the lip, straight and thin; no lamellae present.</p>
            <p> Differing from  Z. pretneri and  Z. tholussum by its broader shell and the differentiated parietal shield; differs from  Z. manitaense by the absence of a visible parietalis in the aperture; barely differs from  Z. aff. troglobalcanicum morphologically, on average with reduced shell broadness and a slightly deeper suture (see Remarks). </p>
            <p>Distribution.</p>
            <p> Known from four caves (Jama Dobravljevac,  Špilja Dahna, Jama u kamenolomu,  Vranjača ) in the municipality of Tomislavgrad in Bosnia-Herzegovina. </p>
            <p>Etymology.</p>
            <p> Named  Zospeum simplex (= simple, unsophisticated) due to the lack of any form of shell sculpture or lamellae. </p>
            <p>Remarks.</p>
            <p> Difficult to separate from  Z. troglobalcanicum without genetic data (which is not uncommon in  Zospeum ; see  Inäbnit et al. 2019). Both species have a nondescript shell without prominent shell sculpture or lamellae within the aperture.  Absolon’s (1916) description of  Z. troglobalcanicum consisted out of a photograph depicting multiple specimens haphazardly clustered together in various positions and a legend that established the name and type locality. The lack of a written characterisation of the species in the original description and the fact that the specimens in the photograph  weren’t depicted in any standardised position makes a characterisation of the species fairly challenging (putative syntype specimen, collected by K. Absolon from the type locality, was only located very recently by AJ in Vienna (NHMW Mol.Coll.Edlauer 32.749) and  couldn’t be studied yet). From the photograph in Absolon (1916), the species can be characterised as similar to  Z. manitaense in shell shape, without any visible lamella in the aperture and with a comparatively large parietal shield. The larger parietal shield might serve as a distinguishing character between  Z. simplex and  Z. troglobalcanicum , though the illustration of a topotypic specimen in Bole (1974; fig. 3h) might indicate that this character is variable within the population. The two specimens we preliminarily assigned to  Z. troglobalcanicum (Fig. 3, NMBE 568052;  Inäbnit et al. 2019: fig. 7u) only have a small parietal shield. As of now, the shell height:shell width ratio seems to be the most effective way of separating the two specimens from  Z. simplex (  Z. simplex : generally higher than 1.3 (one exception);  Z. aff. troglobalcanicum : below 1.3), but that might just be due to the low sample sizes. Investigation of the inner aspects of the shells will be presented in a later work. </p>
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	https://treatment.plazi.org/id/04D6F12E943D5710A5862DFCA1508456	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Inaebnit, Thomas;Jochum, Adrienne;Slapnik, Raijko;Neubert, Eike	Inaebnit, Thomas, Jochum, Adrienne, Slapnik, Raijko, Neubert, Eike (2021): New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175-193, DOI: http://dx.doi.org/10.3897/zookeys.1071.66417, URL: http://dx.doi.org/10.3897/zookeys.1071.66417
