taxonID	type	description	language	source
7D6487FFFFF4FFE92FD134A0C4004510.taxon	materials_examined	Type species Zercon obtusus Koch, 1839: 13, sensu Karg, 1960: 440.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF4FFE42FD13795C4F8458B.taxon	description	(Figs 1 – 5; 58 – 62) Following the classification of Chant & McMurtry (2004), the new species Amblyseius azaliae sp. nov. belongs to the obtusus species group with the ventrianal shield pentagonal, seta z 4 short and J 2 present; and sub-group pamperisi having spermatheca with calyx swollen basally, then narrowing and finally flaring distally. Female (n = 3) Diagnosis. The species is distinguished by dorsal shield smooth with seven pairs of pores, female idiosomal setal pattern: 10 A: 9 B / JV – 3: ZV, peritreme extends beyond seta j 1, sternal shield with three pairs of setae, posterior margin of sternal shield concave, fixed digit of chelicera with 11 teeth and movable digit with three, spermatheca tubular with constriction at the distal part of calyx and then flared, all macrosetae on leg IV are pointed with longest one on genu of leg IV. Dorsum (Fig. 1). Dorsal shield 400 (396 – 401) long and 275 (273 – 276) wide, smooth, with sigillum mostly on prodorsal area, dorsum with seven pairs of pores (gd 1, gd 2, gd 4, gd 5, gd 6, gd 8 and gd 9); 17 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j 1 38 (36 – 39), j 3 52 (51 – 53), j 4 7 (6 – 8), j 5 5 (4 – 6), j 6 6 (6 – 8), J 2 5 (5 – 6), J 5 5 (5 – 6), z 2 8 (8 – 10), z 4 8 (7 – 9), z 5 5 (4 – 6), Z 1 8 (8 – 9), Z 4 113 (111 – 113), Z 5 238 (237 – 239), s 4 75 (74 – 76), S 2 11 (10 – 12), S 4 10 (9 – 11), S 5 10 (9 – 11), r 3 13 (12 – 14), R 1 14 (13 – 15). All setae smooth, setae j 1, j 3, s 4, and Z 4 long, seta Z 5 longest, all other setae medium to minute. Peritreme (Fig. 1). Extending beyond the bases of j 1. Venter (Figs 2; 59). Sternal shield square shaped, posterior margin of sternal shield concave. Sternal shield 73 (72 – 74) long and 86 (84 – 87) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on distinct metasternal plate 27 (26 – 28) long with conspicuous poroids. Distances between st 2 - st 2 83 (82 – 85), st 5 - st 5 72 (71 – 74). Genital shield smooth, broad at the base and narrower at genital opening with anterior irregular circular flap. Ventrianal shield smooth, pentagonal 123 (122 – 125) long, 83 (82 – 84) wide at level of ZV 2 and 64 (62 – 65) wide at anus level with three pairs of pre-anal setae, JV 1 15 (14 – 16), JV 2 18 (17 – 19), ZV 2 16 (15 – 18) and one pair of crescentic pores gv 3 17 (16 – 19) apart at the level JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 15 (14 – 16), ZV 3 14 (12 – 15), JV 4 12 (11 – 15) and JV 5 88 (86 – 89) and smooth. Two pairs of metapodal shields present. Primary inguinal sigilla 26 (25 – 27) long and 6 (5 – 7) wide, secondary ones 12 (10 – 12) long. Chelicera (Figs 3; 58). Fixed digit 36 (35 – 38) long with 11 teeth and a distinct pillus dentilis; movable digit 40 (39 – 41) long with three teeth. Spermatheca (Figs 4; 61 – 62). Calyx tubular 19 (18 – 20) long, arms of the calyx flared distally at vesicle, with narrow constriction. Atrium covers almost width of calyx or incorporated into the basis of calyx; major duct clearly visible, minor duct indistinct. Legs (Figs 5; 60). The genu, tibia and tarsus of leg IV with pointed macrosetae of following lengths: genu 111 (109 – 114), tibia 75 (74 – 76) and tarsus 70 (69 – 71). Genu and tibia of leg I, II and III with macrosetae of the following length respectively, SgeI 40, StiI 30, SgeII 30, StiII 25, SgeIII 50, StiIII 30. Chaetotactic formula of genu II 1 2 / 1, 1 / 1 1 and genu III 1 1 / 1, 2 / 1 1. Length of leg I 375 (373 – 376), leg II 255 (253 – 256), leg III 278 (276 – 280) and leg IV 354 (351 – 355).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF4FFE42FD13795C4F8458B.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8371 / 2019) collected from Azalea, (Rhododendron sp.), at Upper Shillong: 25 ° 32 ' 09 " N, 91 ° 49 ' 29 " E, 1589 m AMSL, Ri-Bhoi, Meghalaya on 2 nd May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Two paratype females (Acarol. lab / BCKV / 8372 – 8373 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF4FFE42FD13795C4F8458B.taxon	etymology	Etymology. The specific name azaliae refers to the host plant azalea (Rhododendron sp.) of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF4FFE42FD13795C4F8458B.taxon	discussion	Remarks. Amblyseius azaliae is different from all species under pamperisi sub-group by having its unique type of spermatheca, distal part of of calyx is constricted followed by flared arms. However, this new species is very close to A. orientalis Ehara, 1959; A. kulini Gupta, 1978 c; A. raoiellus Denmark & Muma, 1989; A. pamperisi Papadopoulis, 1997; and A. euterpes Gondim Jr. & Moraes, 2001; by having similar type of spermatheca, leg IV with three setaceous macrosetae. However, the new species differs from these close species by the length of dorsal setae, number of teeth in fixed and movable digit of chelicera, length of leg macrosetae and shape of spermatheca. All of them have tubular spermathecae, but spermateca of the new species is very specific with a constriction in calyx and this combination of all characters makes it unique from all the other species of this group. All the differences between the new species and its close species are presented in Table 2.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF9FFE62FD137F9C3654543.taxon	description	(Figs 6 – 10; 63 – 65) Following the classification of Chant & McMurtry (2004) the new species Amblyseius meghalayensis sp. nov. belongs to the obtusus species group with the ventrianal shield pentagonal, seta z 4 shorter and J 2 present; and subgroup obtusus, having calyx of spermatheca with annulated stalk, flaring distally. Female (n = 4). Diagnosis. The new species Amblyseius meghalayensis is distinguished by having dorsum smooth with six pairs of pores, posterior margin of the sternal shield concave, fixed digit of chelicerae with eight teeth, and movable digit with five teeth, leg IV with three pointed macrosetae. Dorsum (Fig. 6). Dorsal shield 330 (328 – 332) long and 202 (200 – 203) wide, smooth, with sigillum mostly on prodorsal area, prodorsum wide, with six pairs of pores (gd 2, gd 4, gd 5, gd 6, gd 8 and gd 9); 17 pairs of setae on dorsal shield and two pair of setae on unsclerotised cuticle: j 1 29 (28 – 30), j 3 42 (41 – 44), j 4 5 (4 – 6), j 5 5 (4 – 6), j 6 5 (4 – 6), J 2 5 (4 – 6), J 5 5 (4 – 6), z 2 5 (4 – 6), z 4 5 (4 – 6), z 5 5 (4 – 6), Z 1 8 (7 – 9), Z 4 106 (105 – 108), Z 5 182 (182 – 184), s 4 84 (83 – 85), S 2 8 (8 – 10), S 4 8 (8 – 10), S 5 8 (8 – 10), r 3 14 (13 – 15), R 1 13 (12 – 14). All setae smooth, setae j 1, j 3, s 4, and Z 4 long while seta Z 5 longest, and all other setae medium to minute. Peritreme (Fig. 6). Extending beyond the bases of j 1. Venter (Figs 7; 63 – 65). Sternal shield nearly square, posterior margin of sternal shield concave, lateral margins of both sides are indented inwardly with a distinct groove at the middle of anterior margin. Sternal shield 66 (65 – 67) long, and 69 (67 – 70) wide at the level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 2 5 (25 – 27) long. Distances between st 2 - st 2 and st 5 - st 5 62 (61 – 64) and 69 (68 – 71) respectively. Genital shield smooth, broad at the base and narrow at genital opening with irregular circular flap that reaches posterior margin of sternal shield. Ventrianal shield smooth, pentagonal 110 (108 – 111) long, 85 (84 – 86) wide at level of ZV 2 and 56 (54 – 57) wide at anus level with three pairs of pre-anal setae, JV 1 15 (15 – 18), JV 2 15 (15 – 18), ZV 2 15 (15 – 18) and one pair of pre-anal pores gv 3 29 (28 – 32) apart at the level of JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 15 (15 – 18), ZV 3 10 (9 – 11), JV 4 8 (8 – 10) and JV 5 72 (71 – 74); JV 5 long and smooth. Two pairs of metapodal shields present; Primary inguinal sigilla 16 (15 – 18) long, 5 (4 – 6) wide, and secondary ones 10 (9 – 11). Chelicera (Fig. 8). Fixed digit 39 (38 – 40) long with eight teeth and movable digit 35 (33 – 37) long with five teeth. Spermatheca (Figs 9; 65). Calyx elongated, 15 (14 – 16) long; annulated distally and broad near atrium, atrium covers almost the width of calyx or incorporated in the basis of calyx, major duct flattened, clearly visible while minor indistinct. Legs (Fig. 10). The genu, tibia and tarsus of leg IV with pointed macrosetae of following lengths: genu 88 (88 – 90), tibia 68 (67 – 70) and tarsus 85 (84 – 87). Chaetotactic formula of genu II 0 2 / 2, 1 / 1 1 and genu III 0 2 / 1, 1 / 1 1. Length of leg I 395 (393 – 398), leg II 315 (314 – 319), leg III 306 (304 – 308) and leg IV 355 (352 – 356). Leg II and III with macrosetae SgeII 50 (48 – 51), SgeIII 47 (46 – 50), StiIII 43 (43 – 45) long.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF9FFE62FD137F9C3654543.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8374 / 20) collected from Cogon grass (Imperata cylindrica), at Jaintia Hills: 25 ° 20 ' 31 " N, 91 ° 53 ' 19 " E, 1496 m AMSL, West Jaintia Hills, Meghalaya on 5 th May, 2019 and deposited in the NZC, Zoological Survey of India, Kolkata. Three paratype females (Acarol. lab / BCKV / 8374 - 8375 / 2019) with same collection data as holotype and deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF9FFE62FD137F9C3654543.taxon	etymology	Etymology. The specific name meghalayensis refers to the type locality Meghalaya of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFF9FFE62FD137F9C3654543.taxon	discussion	Remarks. Amblyseius meghalayensis is distinguished from all other species of the obtusus sub-group by its unique tubular spermatheca which is annulated distally towards vesicle. However, this new species is close to A. abbasovae Wainstein & Beglyarov, 1971; A. isuki Chant & Hansell, 1971; A. guntheri McMurtry & Schicha, 1987; by having annulated calyx of spermatheca, dorsum smooth, seta Z 4 and Z 5 very long, leg IV with three setaceous macrosetae. The new species differs from these close species by the setal length, ventrianal shield, number of teeth in fixed and movable digit of chelicerae, annulation pattern of spermatheca, length of leg macrosetae etc. All the differences between the new species and its close species are presented in Table 3.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFDFFE02FD13756C39F4408.taxon	materials_examined	Type species: Seiulus finlandicus Oudemans, 1915: 183. In the classification of Chant & McMurty (2007) both the species Euseius fascae sp. nov. and E. dwakiensis sp. nov. belong to the tribe Euseiini Chant & McMurtry (2005), subtribe Euseiina Chant & McMurtry (2005) and genus Euseius, with 19 pairs of dorsal setae, and the female idiosomal setal pattern: 10 A: 9 B / JV- 3: ZV.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFDFFE32FD1366EC5F646AB.taxon	description	(Figs 11 – 15; 66 – 69) Female (n = 13). Diagnosis. The species is distinguished by having the dorsum strongly reticulated, lateral margin at waist region irregular, peritreme extends near about the bases of j 3, dorsum with five pairs of pores, all ventral shields smooth, fixed digit of chelicerae with three distinct anterior teeth and a pilus dentilis, spermatheca tubular, annulated at the distal part of calyx, leg IV with three pointed macrosetae. Dorsum (Figs 11; 66). Dorsal shield 298 (296 – 299) long and 198 (195 – 200) wide, strongly reticulated, with sigillum mostly on prodorsal area, prodorsum wide, irregular lateral margin at waist region with five pairs of pores (gd 2, gd 5, gd 6, gd 8 and gd 9); 17 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j 1 28 (28 – 30), j 3 28 (28 – 29), j 4 15 (13 – 15), j 5 15 (13 – 15), j 6 15 (13 – 15), J 2 15 (13 – 15), J 5 5 (5 – 7), z 2 22 (20 – 23), z 4 22 (20 – 23), z 5 15 (13 – 15), Z 1 15 (13 – 15), Z 4 20 (18 – 20), Z 5 58 (57 – 60), s 4 30 (28 – 30), S 2 23 (20 – 23), S 4 23 (20 – 23), S 5 30 (28 – 30), r 3 16 (16 – 18), R 1 13 (13 – 15). All setae smooth, seta Z 5 slightly serrated, setae j 1, j 3 and s 4 long, seta Z 5 longest, all other setae medium to small. Peritreme (Figs 11; 66). Extending near about the bases of j 3. Venter (Figs 12; 69). Sternal shield rectangular, posterior margin forms a distinct loop. Sternal shield ornamented with lateral stripes, 53 (53 – 55) long and 72 (70 – 74) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 18 (18 – 20) long and conspicuous poroids on distinct metasternal plate. Distances between st 2 - st 2 61 (60 – 63), st 5 - st 5 70 (70 – 73). Two pairs of metapodal shields present. Primary inguinal sigilla 18 (17 – 20) long and 3 (3 – 4) wide, secondary ones 4 (3 – 5). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap touching sternal loop anteriorly. Ventrianal shield smooth, slightly sclerotised at anterior part 88 (85 – 88) long, 46 (44 – 47) wide at level of ZV 2 and 66 (65 – 68) wide at anus level with three pairs of pre-anal setae, JV 1 17 (16 – 18), JV 2 18 (17 – 20), ZV 2 18 (18 – 20) and one pair of large elliptical pre-anal pores gv 3 26 (25 – 28) apart below JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 16 (15 – 18), ZV 3 8 (8 – 10), JV 4 10 (10 – 12) and JV 5 30 (30 – 33); seta JV 5 long and smooth. Chelicera (Fig. 13). Fixed digit 24 (23 – 25) long with three anterior teeth and distinct pilus dentilis, movable digit 20 (20 – 21) long with two teeth. Spermatheca (Figs 14; 68). Calyx tubular, elongated 16 (15 – 18) long, annulated distally, narrow at middle and gradually widening thereafter towards conspicuous atrium, minor duct and major duct visible. Leg (Figs 15; 67). The genu, tibia and tarsus of leg IV with pointed macrosetae of following lengths: genu 30 (30 – 32), tibia 30 (28 – 30) and tarsus 50 (48 – 50). Chaetotactic formula of genu II 1 2 / 1. 1 / 1 1 and genu III 1 2 / 1. 1 / 1 1. Length of leg I 310 (310 – 314), leg II 275 (273 – 276), leg III 250 (248 – 250) and leg IV 329 (325 – 330).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFDFFE32FD1366EC5F646AB.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8376 / 20) collected from needlewood tree (Schima wallichii Dc. Korth), at Barapani: 25 ° 40 ' 52 " N, 91 ° 54 ' 41 " E, 973 m, AMSL, Ri-Bhoi, Meghalaya on 3 rd May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. 12 paratype females (Acarol. lab / BCKV / 8377 – 8380 / 2019) with same collection data as holotype and deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFDFFE32FD1366EC5F646AB.taxon	etymology	Etymology. The specific name fascae refers to the unique annuli present at the basal part of calyx of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFDFFE32FD1366EC5F646AB.taxon	discussion	Remarks. The new species Euseius fascae is different from all the species in the genus Euseius by having unique type of spermatheca, calyx annulated basally and flared at atrium. It is close to E. insanus (Khan & Chaudhri, 1969); E. bandispermathecae Pramanik & Karmakar, 2016 and E. astrictus Karmakar & Bhowmik, 2018 by having same pattern of ventral plates and annulation or constriction in calyx. However, this new species differs from the close species by dorsal setae, lateral margin of waist region, position of R 1, denticles in chelicerae, macrosetae on leg IV, tip of macrosetae and shape or annulation pattern of spermatheca. All the differences between the new species and close species are presented in Table 4.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFEFFFD2FD134CDC6FC47DB.taxon	description	(Figs 16 – 20; 70 – 74) Female (n = 6). Diagnosis. The new species is characterised by dorsum strongly reticulated, seta R 1 on the dorsal shield, waist region irregular, ventrianal shield vase-shaped, fixed digit of chelicerae with five anterior teeth, with pilus dentilis, spermatheca with a cap-like structure on calyx, narrow at the base of vesicle, leg IV with three macrosetae with rounded tips. Dorsum (Figs 16; 70). Dorsal shield 328 (326 – 330) long and 230 (229 – 232) wide, strongly reticulated, with sigillum mostly on prodorsal area, prodorsum wide, irregular lateral margin at waist region with five pairs of pores (gd 2, gd 5, gd 6, gd 8 and gd 9); 18 pairs of setae on dorsal shield and one pair of setae on unsclerotised cuticle: j 1 33 (30 – 33), j 3 24 (23 – 25), j 4 10 (10 – 11), j 5 10 (10 – 12), j 6 13 (12 – 14), J 2 14 (13 – 15), J 5 4 (3 – 5), z 2 19 (18 – 20), z 4 21 (20 – 23), z 5 14 (13 – 15), Z 1 14 (13 – 15), Z 4 19 (18 – 20), Z 5 60 (60 – 63), s 4 28 (25 – 29), S 2 20 (19 – 20), S 4 23 (23 – 25), S 5 29 (28 – 31), r 3 14 (13 – 15), R 1 14 (13 – 15). All setae smooth, setae j 1, j 3 and s 4 long, seta Z 5 longest, all other setae medium to small. Peritreme (Fig. 16; 70). Extending to the level near about the base j 3. Venter (Figs 17; 71). The sternal shield rectangular, posterior margin of sternal shield with a distinct loop. Sternal shield 62 (60 – 63) long and 77 (74 – 78) wide at the level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 16 (15 – 18) long with conspicuous poroids on distinct metasternal plate. Distances between st 2 - st 2 60 (58 – 60), st 5 - st 5 71 (70 – 73). Two pairs of metapodal shields present. Primary inguinal sigilla 23 (23 – 25) long and 4 (3 – 5) wide, secondary ones 6 (5 – 8). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap touching sternal loop anteriorly. Ventrianal shield vase-shaped, smooth, 90 (89 – 92) long, 40 (40 – 43) wide at level of ZV 2 and 65 (65 – 67) wide at anus level with three pairs of pre-anal setae, JV 1 18 (17 – 20), JV 2 22 (20 – 23), ZV 2 18 (18 – 20) and one pair of large elliptical pre-anal pores gv 3 28 (27 – 30) apart below JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 18 (17 – 20), ZV 3 9 (8 – 10), JV 4 9 (8 – 10) and JV 5 31 (30 – 33); seta JV 5 long and smooth. Chelicera (Figs 18; 74). Fixed digit 25 (23 – 25) long with five anterior teeth and distinct pilus dentilis, movable digit 22 (20 – 23) long with a single tooth. Spermatheca (Figs. 19; 72). Calyx elongated 22 (19 – 23) long with cap like structure. Distal part of calyx narrow towards the vesicle with divergent arms. Atrium incorporated at the base of calyx, major and minor duct distinctly visible. Legs (Figs 20; 73). The macrosetae on genu, tibia and tarsus of leg IV with bulbous tips, of following lengths: genu 48 (46 – 49), tibia 35 (33 – 36) and tarsus 68 (65 – 69). Chaetotactic formula of genu II 1 2 / 1. 1 / 1 1 and genu III 1 1 / 1. 2 / 1 1. Length of leg I 375 (373 – 376), leg II 255 (253 – 256), leg III 278 (276 – 280) and leg IV 354 (351 – 355).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFEFFFD2FD134CDC6FC47DB.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8381 / 20) collected from Carambola, (Averrhoa carambola L.), at Dwaki: 25 ° 131 " N, 91 ° 58 ' 16 " E, 520 m AMSL, West Jaintia Hills, Meghalaya on 2 nd May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Five paratype females (Acarol. lab / BCKV / 8382 - 8383 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFEFFFD2FD134CDC6FC47DB.taxon	etymology	Etymology. The specific name dwakiensis refers to the type locality “ Dwaki ” from where this new species was collected.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFFEFFFD2FD134CDC6FC47DB.taxon	discussion	Remarks. The new species, Euseius dwakiensis is different from all the members of Euseius by having its spermatheca with cap or flap-like structure at atrium. However, the species is close to E. coccosocius Ghai & Menon, 1967; E. elinae Schicha, 1977 a; E. victoriensis Schicha, 1977; by the similar pattern of dorsum and ventral plates. The new species differs from these close species by the position of dorsal seta R 1, number of teeth in fixed and movable digit of chelicerae, leg macrosetae and spermatheca. All the differences between new species and close species are presented in Table 5.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE3FFFF2FD13239C5D1468F.taxon	materials_examined	Type species Okiseius subtropicus Ehara, 1967 a: 77.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE3FFFF2FD13239C5D1468F.taxon	materials_examined	Type species: Amblyseius (Kampimodromus) maritimus Ehara, 1967 b: 224. Following the classification of Chant & McMurty (2007) Okiseius jainticus sp. nov. and Okiseius unisetatus sp. nov. belong to tribe Kampimodromini Kolodochka, 1998, and subtribe Kampimodromina Kolodochka, 1998, without a distinct notch or incision in the lateral margin of the dorsal shield at the level of seta s 4. Chant & McMurtry (2003) proposed two species groups based on the point of insertion of seta R 1: the subtropicus species group with this seta inserted on dorsal shield with a distinct notch at the lateral margin of dorsal shield below R 1 of females and the maritimus species group with seta R 1 inserted on lateral integument of females without notch at the lateral margin. The new species O. jainticus sp. nov. is grouped under maritimus species group while O. unisetatus sp. nov. is grouped under subtropicus species group.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE2FFF92FD134E8C3AB4647.taxon	description	(Figs 21 – 25; 75 – 78) Female (n = 7). Diagnosis. The species Okiseius jainticus sp. nov. belongs to the subgroup maritimus, by having R 1 off the dorsal shield. This new species is distiguished from all the six species under this subgroup (Chant & McMurtry, 2003) by having all setae serrated except j 4, j 5, j 6, J 5, and z 5 which are smooth and all the setae arise from distinct tubercles, seta S 5 serrated, both the setae r 3 and R 1 are off the dorsal shield and each with a distinct depression at the lateral margin of dorsal shield, one pair of long narrow metapodal shields, fixed digit of chelicerae with six teeth, movable digit with three teeth, leg IV with three distinct macrosetae with bulbous tips. In addition, the genu of leg IV having all setae rod-like except macroseseta which has bulbous tip, tibia and basitarsus of leg IV having rod-like setae but ventral setae are pointed. Dorsum (Figs 21; 75). Dorsal shield 333 (332 – 335) long and 193 (192 – 195) wide, strongly reticulated, prodorsum wide, smooth lateral margin with seven pairs of pores (gd 1, gd 2, gd 4, gd 5, gd 6, gd 8 and gd 9); 15 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j 1 26 (25 – 28), j 3 38 (38 – 40), j 4 9 (8 – 10), j 5 9 (8 – 10), j 6 25 (24 – 25), J 5 5 (4 – 5), z 2 34 (33 – 35), z 4 43 (42 – 45), z 5 13 (13 – 15), Z 1 38 (38 – 41), Z 4 50 (49 – 53), Z 5 70 (69 – 72), s 4 48 (46 – 49), S 2 49 (48 – 50), S 5 15 (14 – 17), r 3 41 (39 – 42), R 1 28 (26 – 29). All setae serrated, except j 4, j 5, j 6, J 5 and z 5 which are smooth. Peritreme (Fig. 21). Extending beyond the bases of j 1. Venter (Figs 22; 78). All shields smooth. Sternal shield longer than width, posterior margin of sternal shield wavy. Sternal shield 66 (63 – 67) long and 76 (74 – 77) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 17 (15 – 18) long and conspicuous poroids on distinct metasternal plate. Distances between st 2 - st 2 59 (58 – 60), st 5 - st 5 65 (64 – 68). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap and almost straight posteriorly. Secondary ones fused anteried form a single pair 42 (40 – 44) long. Ventrianal shield smooth, pentagonal 105 (103 – 105) long, 56 (54 – 57) wide at level of ZV 2 and 60 (59 – 62) wide at anus level with three pairs of pre-anal setae, JV 1 14 (13 – 15), JV 2 14 (13 – 15), ZV 2 13 (12 – 14) and one pair of pre-anal pores gv 3 26 (24 – 27) apart at the level JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 18 19 (18 – 20), ZV 3 10 (9 – 10), JV 4 15 (13 – 15) and JV 5 49 (48 – 50); seta JV 5 long and serrated. Chelicera (Fig. 23). Fixed digit 25 (24 – 25) long with six teeth and a distinct pilus dentilis, movable digit 25 (24 – 25) long with three teeth. Spermatheca (Figs 24; 77). Calyx short 10 (7 – 11) long, arms diverging distally, atrium nodular, minor duct and major duct distinctly visible. Legs (Figs 25; 76). The genu, tibia and tarsus of leg IV with bulbous macrosetae of following lengths: genu 29 (28 – 30), tibia 24 (23 – 25) and tarsus 51 (50 – 53). Chaetotactic formula of genu II 1 2 / 1. 1 / 1 1 and genu III 1 1 / 1. 2 / 1 1. Length of leg I 282 (280 – 284), leg II 250 (246 – 250), leg III 236 (235 – 238) and leg IV 342 (341 – 345).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE2FFF92FD134E8C3AB4647.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8384 / 20) collected from Golden Himalayan raspberry (Rubus elypticus), at West Jaintia Hills, Meghalaya 25 ° 21 ' 44 " N, 91 ° 52 ' 40 " E, 1933 m AMSL on 1 st May, 2019; and deposited in the NZC, Zoological Survey of India, Kolkata. Six paratype females (Acarol. lab / BCKV / 8385 - 8387 / 2019) collected from undetermined climber at Mawkdok: 25 ° 24 ' 30 " N, 91 ° 46 ' 44 " E, 1806 m AMSL, East Khasi Hills, with same data as holotype and deposited in the Acarological Laboratory of BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE2FFF92FD134E8C3AB4647.taxon	etymology	Etymology. The specific name jainticus refers to the Jaintia Hills, the habitat of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE2FFF92FD134E8C3AB4647.taxon	discussion	Remarks. Okiseius jainticus is close to O. maritimus (Ehara, 1967 a); O. alniseius Wainstein & Beglyarov, 1972; O. juglandis Wang & Xu, 1985; O. tibetagramins Wu, 1987; O. tribulation Walter, 1999 and Okiseius wongi Denmark & Kolodochka, 1996, by the similar type of dorsal and ventral pattern, presence of depression at level r 3, R 1 and S 5. The new species differs from all these close species by the length of dorsal setae, serration of dorsal setae, number of metapodal shields, denticles on fixed and movable digit of chelicerae, number of rod-like setae on leg IV etc. All the differences between the new species and its close species are presented in Table 6.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	description	(Figs 26 – 30; 79 – 84)	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	diagnosis	Female (n = 3). Diagnosis. The new species is characterised by strongly reticulated dorsum, all setae including J 5 and S 5 thick and serrated, arising from distinct tubercles, a small depression present on lateral margin of dorsal shield at level r 3, distinct notch present at below level of R 1, posterior margin of sternal shield concave, ventrianal shield vase-shaped, only one pair of elongated, inwardly curved metapodal shields, fixed digit of chelicerae with seven teeth including pilus dentilis, leg IV with three distinct rod-like macrosetae, all the dorsal setae are rod-like while all the ventral are pointed. All these characters make the species different from all the ten species under this subgroup.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	description	Dorsum (Figs 26; 79 – 80). Dorsal shield 308 (307 – 310) long and 175 (174 – 176) wide, strongly reticulated, with five pairs of pores (gd 2, gd 5, gd 6, gd 8 and gd 9); 16 pairs of setae on dorsal shield and one pair of setae on unsclerotised cuticle: j 1 25 (24 – 26), j 3 21 (20 – 23), j 4 15 (15 – 17), j 5 15 (15 – 17), j 6 26 (25 – 27), J 5 6 (5 – 7), z 2 25 (24 – 26), z 4 25 (24 – 26), z 5 19 (18 – 20), Z 1 32 (31 – 33), Z 4 47 (46 – 49), Z 5 47 (46 – 48), s 4 28 (27 – 29), S 2 36 (35 – 38), S 5 19 (18 – 21), r 3 25 (24 – 26), R 1 26 (25 – 28). All setae serrated, only seta j 3 weakly serrated. Peritreme (Figs 26; 79 – 80). Extended beyond the bases of j 1. Venter (Figs 27; 81). All shields are smooth. Sternal shield is longer than width, posterior margin concave. Sternal shield 64 (62 – 65) long and 62 (61 – 63) wide at the level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on metasternal plate on unsclerotised membrane at the margin of genital flap 16 (15 – 18) long with a conspicucous poroid on metasternal plate. Distances between st 2 - st 2 44 (42 – 48), st 5 - st 5 57 (56 – 60). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap and almost straight posteriorly. One pair of elongated narrow inwardly curved metapodal shields present 39 (37 – 40) long. Ventrianal shield smooth, pentagonal 113 (110 – 115) long, 50 (48 – 50) wide at level of ZV 2 and 58 (56 – 59) wide at anus level with three pairs of pre-anal setae, JV 1 12 (11 – 14), JV 2 8 (7 – 10), ZV 2 9 (8 – 10) and one pair of pre-anal pores 18 (17 – 19) apart closely placed on longitudinal line below JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 12 (11 – 13), ZV 3 9 (8 – 10), JV 4 11 (10 – 12) and JV 5 32 (31 – 33); the latter one is long and serrated. Chelicera (Figs 28; 83). Fixed digit 18 (17 – 20) long with six teeth and a distinct pilus dentilis, movable digit 16 (15 – 17) long with single tooth. Spermatheca (Figs 29; 84). Calyx short 4 (3 – 5) long with conspicuous atrium, minor duct and major duct visible. Legs (Figs 30; 82). The genu, tibia and tarsus of leg IV with macrosetae of blunt tips, of following lengths: genu 11 (10 – 12), tibia 16 (15 – 17) and tarsus 19 (17 – 20). All the dorsal setae have blunt tips while the ventral and lateral have pointed tips. Chaetotactic formula of genu II 1 2 / 1. 1 / 1 1 and genu III 1 1 / 1. 2 / 1 1. Length of leg I 280 (280 – 284), leg II 250 (246 – 250), leg III 235 (235 – 238) and leg IV 342 (341 – 345).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8389 / 20) collected from Litchi, (Litchi chinensis Sonn.) at Dwaki: 25 ° 13 ' 1 " N, 91 ° 58 ' 16 " E, 520 m AMSL, West Jaintia Hills, Meghalaya on 2 May 2019; and deposited in the NZC, Zoological Survey of India, Kolkata, two paratype females (Acarol. lab / BCKV / 8389 – 8390 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	etymology	Etymology. The specific name unisetatus refers to the uniform serration pattern of dorsal setae of new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFE7FFFB2FD13581C2C940C9.taxon	discussion	Remarks. The new species is close to O. subtropicus Ehara, 1967 b, O. merenoi Schicha, 1987; O. eharai Liang & Ke, 1982; O. sikkimensis Gupta, 1986; O. yazuliensis Gupta, 1986 and O. himalayana Gupta, 1986 by having similar type of heavily sculptured dorsum, depression at the level r 3, and rod-like shape of leg macrosetae. However, the new species differs from these close species by the length of dorsal setae, serration pattern of dorsal setae, denticles of chelicerae, number of metapodal shields, number of rod-like setae in leg IV and shape of spermatheca. In O. subtropicus all dorsal setae serrated except J 5, with a distinct notch at level between R 1 and S 5; fixed digit of chelicerae with three teeth; metapodal shields fused, very long, about 130 long. The new species also differs from O. merenoi by all setae serrated except J 5 and S 5; setae j 6, S 5 and R 1 shorter than the new species; fixed digit of chelicerae with two teeth; two pairs of metapodal shield while in new species all dorsal setae are serrated, fixed digit with six teeth and with single metapodal shield. In new species j 1 longer and s 4 shorter, all dorsal setae serrated, fixed digit of chelicerae with six teeth while in O. himalayana j 1 shorter and s 4 longer, J 5 smooth, fixed digit of chelicerae with three teeth. All the differences between the new species and its close species are presented in Table 7.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF62FD13707C3934481.taxon	materials_examined	Type species Gamasus plumifer Canestrini & Fanzago, 1876: 130, designated by Vitzthum, 1941.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF62FD13707C3934481.taxon	materials_examined	Type species Phytoseius (Dubininellus) corniger Wainstein, 1959: 1361, by original designation. Chant & McMurtry (1994) recognised three species groups based on presence or absence of setae J 2 and R 1: the horridus species group, Denmark (1966) with setae J 2 and R 1 absent, the plumifer species group Chant & Yoshida- Shaul (1992) characterised with setae J 2 and R 1 present, and purseglovei species group Chant & Yoshida-Shaul (1992) with seta J 2 absent and R 1 present. Both the new species P. clavus and P. aonlae belong to horridus species group with setae J 2 and R 1 absent, ventral setal pattern JV- 3, 4: ZV (Chant & McMurtry 1994).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF12FD136FFC7A4422F.taxon	description	(Figs 31 – 37; 85 – 87) Female (n = 4). Diagnosis. The new species is characterised by dorsum strongly reticulated, all the dorsocentral setae arising from distinct pin head-shaped tubercles, ventrianal shield sole-shaped with three pairs of pre-anal setae, fixed digit of chelicerae with six teeth including pilus dentilis and movable digit with two teeth, all setae including three macrosetae of leg IV rod-like and blunt. Dorsum (Figs 31; 85 – 86). Dorsal shield 328 (326 – 329) long and 146 (145 – 147) wide, strongly reticulated, with five pairs of pores (gd 2, gd 5, gd 6, gd 8 and gd 9); 15 pairs of setae on the dorsal shield: j 1 32 (29 – 33), j 3 37 (35 – 38), j 4 8 (8 – 9), j 5 8 (8 – 9), j 6 8 (8 – 9), J 5 22 (21 – 23), z 2 21 (20 – 23), z 3 33 (32 – 33), z 4 20 (20 – 21), z 5 8 (8 – 9), Z 4 68 (67 – 70), Z 5 78 (77 – 80), s 4 72 (72 – 75), s 6 91 (89 – 92), r 3 48 (47 – 50). All setae serrated except the setae j 4, j 5, j 6, and z 5 which are smooth and pointed. Peritreme (Fig. 31; 85 – 86). Extended upto the bases of j 1. Venter (Fig. 32). All the shields are smooth. The sternal shield square-shaped, posterior margin of sternal shield indistinct. Sternal shield 60 (58 – 60) long and 77 (74 – 78) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 23 (22 – 25) long. Distances between st 2 - st 2 58 (58 – 60), st 5 - st 5 62 (60 – 65). Base of genital shield broad, posterior margin little convex with a notch at lateral margin below ST 5. Metapodal shields not visible, genital shield smooth, broad at base and narrow at genital opening with semicircular flap. Ventrianal shield smooth, sole-shaped 84 (83 – 85) long, 32 (31 – 33) wide at level of ZV 2 and 49 (48 – 50) wide at anus level with three pairs of pre-anal setae, JV 1 14 (13 – 15), JV 2 12 (11 – 13), ZV 2 10 (10 – 11) and one pair of pre-anal pores on longitudinal line below JV 2, 10 (9 – 12) apart. Membrane surrounding ventrianal shield with three pairs of setae ZV 1 16 (15 – 18), ZV 3 9 (8 – 10), and JV 5 66 (65 – 68); seta JV 5 long and serrated. Chelicera (Fig. 33). Fixed digit 28 (26 – 30) long with two teeth and distinct pilus dentilis, movable digit 28 (26 – 29) long with two teeth. Spermatheca (Fig. 34). Calyx short 8 (7 – 9) long, funnel-shaped, wide at base of vesicle, atrium indistinct, minor duct not visible while the major duct long and very distinct. Legs (Fig. 35). The genu, tibia and tarsus of leg IV with rod-like macrosetae with blunt tip of following lengths: genu 22 (21 – 24), tibia 35 (34 – 36) and tarsus 35 (34 – 36). Chaetotactic formula of genu II 1 2 / 1. 2 / 0 1 and genu III 1 2 / 1. 2 / 0 1. Length of leg I 243 (242 – 245), leg II 220 (217 – 222), leg III 236 (235 – 238) and leg IV 388 (387 – 390). Male (n = 1). Dorsum. Dorsal shield 228 long and 142 wide strongly reticulated, prodorsum wide, smooth lateral margin with 15 pairs of setae on the dorsal shield: j 1 22, j 3 28, j 4 8, j 5 7, j 6 7, J 5 14, z 2 16, z 3 29, z 4 19, z 5 7, Z 4 32, Z 5 35, s 4 50, s 6 47, r 3 36. All setae serrated, setae j 4, j 5, j 6, and z 5 are smooth. Peritreme. Extended beyond the bases of j 1. Venter (Fig. 36). Sternogenital shield smooth with five pairs of setae and three pairs of distinguishable poroids. Ventrianal shield 100 long, 111 wide at level of ZV 2, 52 wide at level of anus, with three pairs of pre-anal setae, JV 1 7, JV 2 7, ZV 2 7 and a pair of pores below JV 2, 13 apart. Unsclerotised membrane surrounding the ventrianal shield with one pair of setae JV 5 23 long and serrated. Chelicera (Fig. 37). Spermatodactyl with an elongated shaft 20 long terminating with a wide toe. Leg. Leg IV with three macrosetae of blunt tip of following lengths: genu 14, tibia 14 and tarsus 27. Chaetotactic formula of genu II 1 2 / 1. 2 / 0 1 and genu III 1 2 / 1. 2 / 01. Length of leg I 224, leg II 206, leg III 191 and leg IV 302.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF12FD136FFC7A4422F.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8391 / 2019) deposited in the NZC, Zoological Survey of India, Kolkata), collected from Rhododendron, (Rhododendron sp.), at Upper Shillong: 25 ° 32 ' 9 " N, 91 ° 49 ' 29 " E, 1589 m AMSL, Upper Shillong, Meghalaya on 2 May 2019; two paratype females (Acarol. lab / BCKV / 8391 – 8392 / 2019) and one paratype male (Acarol. lab / BCKV / 8393 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF12FD136FFC7A4422F.taxon	etymology	Etymology. The specific name clavus refers to the “ pin-shaped ” prodorsal setae of this new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFEBFFF12FD136FFC7A4422F.taxon	discussion	Remarks. The new species Phytoseius clavus is notable from all the species under horridus species group by all the dorsocentral setae arising from distinct pin head-shaped tubercles and all setae including three macrosetae of leg IV rod-like and blunt. However, this new species is close to P. corniger Wainstein, 1959; P. nipponicus Ehara, 1962; P. meyerae Gupta, 1977; P. domesticus Rather, 1985 and P. dorsospinosus Pramanik & Karmakar, 2016 by having similar pattern of dorsal shield ornamentation, dorsocentral setae smooth, sole-shaped ventrianal shield with three pairs of setae. The new species differs from these close species by the length, shape and pattern of dorsal shield setae, serration of setae z 2 and z 4, number of metapodal shield, denticles in fixed and movable digit of chelicerae, macrosetae on leg IV and shape of spermatheca. All the differences between the new and close species are given in Table 8.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFECFFCC2FD13048C44A4543.taxon	description	(Figs 38 – 42; 88 – 91) Female (n = 8). Diagnosis. The species is distinguished by having the dorsum strongly reticulated forming a pattern like leopard skin, all dorsal setae serrated except the setae j 4, j 5, j 6, J 5, z 2, and z 4, setae s 4, s 6, Z 4 and Z 5 are long, serrated with distinct longitudinal groove covering near about 1 / 3 rd part of longitudinal line of setae. Ventrianal shield sole-shaped with three pairs of pre-anal setae arranged along the anterior lateral margins of ventrianal shield. Fixed digit of chelicerae with three teeth and pilus dentilis, movable digit with single tooth, leg IV with three macrosetae with bulbous tips, a hyaline cap visible covering the tip. Dorsum (Figs 38; 88). Dorsal shield 315 315 (314 – 316) long and 148 149 (147 – 150) wide, strongly reticulated like leopard skin, with five pairs of pores (gd 4, gd 5, gd 6, gd 8 and gd 9); 15 pairs of setae on the dorsal shield: j 1 30 (28 – 31), j 3 35 (33 – 36), j 4 8 (7 – 9), j 5 8 (7 – 9), j 6 8 (7 – 9), J 5 6 (5 – 7), z 2 16 (15 – 19), z 3 35 (32 – 36), z 4 13 (12 – 15), z 5 8 (7 – 9), Z 4 84 (82 – 85), Z 5 82 (79 – 83), s 4 136 (135 – 138), s 6 70 (67 – 71), r 3 48 (45 – 49). All setae serrated, setae j 4, j 5, j 6, J 5, z 2, z 4, and z 5 smooth and having longitudinal groove all along the length of the setae s 4, s 6, Z 4, and Z 5. Peritreme (Fig. 38). Extended beyond the bases of j 1. Venter (Fig. 39). All shields are smooth. The sternal shield square-shaped, posterior margin of sternal shield nearly straight. Sternal shield 51 (48 – 52) long and 69 (66 – 71) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 16 (15 – 18) long. Distances between st 2 - st 2 57 (55 – 59), st 5 - st 5 62 (60 – 65). The base of the genital shield broad indented posterior lateral margin below st 5. One pair of elongated, narrow inwardly curved metapodal shields present 35 (32 – 36) long and 3 (2 – 4) wide. Genital shield smooth broad at the base and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, sole-shaped 103 (102 – 105) long, 35 (33 – 36) wide at level of ZV 2 and 53 (51 – 54) wide at anus level with three pairs of pre-anal setae, JV 1 12 (11 – 13), JV 2 12 (11 – 13), ZV 2 12 (11 – 13) and one pair of pre-anal pores at the level of JV 2, 9 (8 – 10) apart. Membrane surrounding ventrianal shield with three pairs of setae ZV 1 13 (11 – 14), ZV 3 8 (9 – 10), and JV 5 51 (50 – 53); the later one is long and serrated. Chelicera (Figs 40; 90). Fixed digit 19 (18 – 22) long with two teeth and a distinct pilus dentilis, movable digit 19 (18 – 22) long with single tooth. Spermatheca (Figs 41; 91). Calyx short 8 (7 – 9) long, cup-shaped, atrium nodular from where minor and major duct arises, major duct flattened and distinctly visible. Legs (Figs 42; 89). The genu, tibia and tarsus of leg IV with bulbous macrosetae of following lengths; genu 13 (12 – 15), tibia 54 (52 – 55) and tarsus 28 (25 – 29). Chaetotactic formula of genu II: 1 2 / 1, 2 / 0 1 and genu III 1 2 / 1, 2 / 0 1. Length of leg I 245 (242 – 245), leg II 220 (217 – 220), leg III 235 (235 – 238) and leg IV 388 (387 – 390).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFECFFCC2FD13048C44A4543.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8394 / 2019) collected from Aonla, (Phyllanthus emblica), at Dwaki: 25 ° 13 ’ 1 ” N, 91 ° 58 ’ 16 ” E, 520 m AMSL, Dwaki, Meghalya, on 2 May 2019; deposited in the NZC, Zoological Survey of India, Kolkata. two paratype females (Acarol. lab / BCKV / 8395 / 2019) collected from same host and location and five paratype females (Acarol. lab / BCKV / 8396 – 98 / 2019) collected from same host at Keibul Lamjao National Park: 24 ° 28 ’ 24 ” N, 93 ° 49 ’ 1 ” E, 781 m AMSL Bishnupur, Manipur, deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFECFFCC2FD13048C44A4543.taxon	etymology	Etymology. The specific name aonlae refers to the host plant from where the species was collected.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFECFFCC2FD13048C44A4543.taxon	discussion	Remarks. The new species is different from all the other species of the horridus group. Phytoseius aonlae sp. nov. is close to P. crinitus Swirski & Shechter, 1961; P. woodburyi De Leon, 1965; P. neoferox Ehara & Bhandhufalck, 1977; and P. litoralis Silva et al., 2013 by having similar dorsal setal pattern. However, the new species differs from its close species by the dorsal setal length; length and shape of macrosetae in leg IV; denticles in fixed digit of chelicerae; shape of ventrianal and genital shield; and shape of spermatheca. All the differences between the new species and its close species are described in Table 9.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFCD2FD13619C543426B.taxon	materials_examined	Type species: Anthoseius hebetis De Leon, 1959: 258.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFCD2FD13619C543426B.taxon	materials_examined	Type species: Typhlodromus fleschneri Chant, 1960: 60. Typhlodromus rhenanus group, Chant, 1959 b: 62. Type species – Seiulus rhenanus Oudemans, 1905: 78. Chant & McMurtry (1994) recognised nine species groups viz., arizonicus, transvaalensis, singularis, daresalaami, rickeri, religiosus, rhenanus, egypticus and bergi species groups under the subgenus Anthoseius based on presence or absence of setae S 4, JV 3 and JV 4, morphology of dorsal setae and their length, shape of female ventrianal shield and the point of insertion of r 3 and R 1. Among these species groups the present species Typhlodromus (Anthoseius) barapanicus sp. nov., T. (Anthoseius) campana sp. nov., and T. (Anthoseius) cherrapunjiensis sp. nov. belong to rhenanus species group with setae r 3 and R 1 inserted on the lateral integument of the adult female, with the setae on the dorsal shield in the z - Z and s - S series shorter than the distances between their bases and with the female ventrianal shield pentagonal in shape.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFC82FD1308DC3C6437B.taxon	description	(Figs 43 – 47; 92 – 95) Female (n = 6). Diagnosis. The species is notable by having dorsum strongly reticulated; all the dorsal setae serrated except j 4, j 5, j 6, J 2, J 5, and z 5; seta Z 5 serrated with bulbous tip; posterior margin of sternal shield forms an indistinct loop touching the genital cover flap, ventrianal shield pentagonal with four pairs of pre-anal setae, fixed digit of chelicerae with four teeth while movable digit with single tooth, spermatheca long tubular, basal half thick walled with thin walled cervix narrowed at the base of atrium, leg IV with one macroseta with bulbous tip. Dorsum (Figs 43; 92). Dorsal shield 335 (334 – 338) long and 175 (174 – 177) wide, strongly reticulated, with six pairs of pores (gd 1, gd 2, gd 5, gd 6, gd 8 and gd 9); 18 pairs of setae on the dorsal shield and two pairs on unsclerotised cuticle: j 1 22 (20 – 23), j 3 17 (16 – 20), j 4 10 (9 – 11), j 5 10 (9 – 11), j 6 12 (11 – 13), J 2 14 (13 – 15), J 5 9 (8 – 10), z 2 12 (11 – 13), z 3 25 (24 – 26), z 4 14 (13 – 15), z 5 11 (10 – 13), Z 4 23 (23 – 25), Z 5 39 (38 – 40), s 4 25 (23 – 26), s 6 25 (22 – 25), S 2 18 (16 – 19), S 4 20 (19 – 20), S 5 24 (23 – 25), r 3 15 (14 – 16), R 1 15 (14 – 16). All setae are serrated and with pointed tip except j 4, j 5, j 6, J 2, J 5, and z 5 which are smooth, setae Z 5 serrated with bulbous tip. Peritreme (Figs 43; 92). Extended beyond the bases of j 1. Venter (Fig. 44; 93). All the shields are smooth. The sternal shield is rectangular with posterior lateral lobe and posterior margin forms a loop touching the genital cover flap. Sternal shield 75 (74 – 76) long and 53 (52 – 54) wide at the level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 13 (13 – 15) long. Distances between st 2 - st 2 53 (52 – 54), st 5 - st 5 55 (52 – 54). The base of the genital shield broad, flat posteriorly. Two pairs of metapodal shields present. The primary inguinal sigilla 27 (25 – 28) long and 4 (3 – 5) wide, secondary ones 7 (7 – 9). Genital shield smooth, broad at the base and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, pentagonal, 121 (120 – 123) long, 81 (80 – 83) wide at level of ZV 2 and 73 (71 – 74) wide at anus level with four pairs of pre-anal setae, JV 1 8 8 (7 – 9), JV 2 8 8 (7 – 9), JV 3 8 (7 – 9), ZV 2 8 (7 – 9) and one pair of pre-anal pore gv 3 15 (14 – 16) apart just below JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 9 (8 – 10), ZV 3 8 (8 – 10), JV 4 10 (8 – 10) and JV 5 29 (28 – 30); the later one is long serrated and pointed. Chelicera (Fig. 45). Fixed digit 30 (30 – 33) long with four teeth and distinct pilus dentilis, movable digit 30 (28 – 30) long with single tooth. Spermatheca (Figs 46; 95). Calyx tubular, 35 (33 – 35) long. Distal half of calyx thick walled and proximal part thin walled with a neck connected to atrium, minor and major duct visible. Legs (Figs 47; 94). The basitarsus of leg IV with single macroseta with bulbous tip, macrosetae on genu and tibia of leg IV with pointed tip and of following lengths: genu 13 (11 – 13), tibia 15 (13 – 15) and basitarsus 28 (25 – 28). Chaetotactic formula of genu II: 0 2 / 1, 2 / 1 1 and genu III 1 2 / 1, 2 / 01. Length of leg I 305 (300 – 305), leg II 218 (213 – 219), leg III 213 (212 – 218) and leg IV 288 (283 – 288).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFC82FD1308DC3C6437B.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8399 / 2019) collected from Broom grass, (Thysanolaena latifolia Roxb.), at Barapani: 25 ° 40 ' 52 " N, 91 ° 54 ' 41 " E, 973 m AMSL, Barapani, Meghalaya on 3 rd May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Three paratype females (Acarol. lab / BCKV / 8399 – 8400 / 2019) with same collection data as holotype and two paratype females (Acarol. lab / BCKV / 8401 / 2019) collected from Loktak, 24 ° 33 ' 1.0 " N, 93 ° 47 ' 20 " E, 768 m AMSL, Manipur, from same host and deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFC82FD1308DC3C6437B.taxon	etymology	Etymology. The specific name barapanicus refers to the type locality “ Barapani ” of Meghalaya from where this new species was collected.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD0FFC82FD1308DC3C6437B.taxon	discussion	Remarks. The new species Typhlodromus (Anthoseius) barapanicus is different from all the members of rhenanus group by having unique type of spermatheca. The new species is close to T (A.) muliebris Van der Merwe, 1968; T (A.) paganus Van der Merwe, 1968; and T. (A.) carambolae Karmakar & Bhowmik, 2018 by having similar pattern of dorsum and venter. Typhlodromus (A.) barapanicus sp. nov. differs from these close species by the dorsal and ventral setal length, form of setae, posterior margin of sternal shield, shape of spermatheca and macrosetae of leg IV. All the differences between the new species and close species are given in Table 10.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD5FFCB2FD13189C5584543.taxon	description	(Figs 48 – 52; 96 – 98) Female (n = 3). Diagnosis. The species is distinguished by having dorsum strongly reticulated, all the dorsal setae short, thin, smooth and pointed except Z 5 smooth with bulbous tip; posterior margin of sternal shield slightly wavy, fixed digit of chelicerae with six teeth while movable digit with three teeth, spermatheca bell-shaped, basitarsus of leg IV with bulbous macroseta while the same on tibia and tarsus are pointed. Dorsum (Fig. 48). Dorsal shield 350 (349 – 353) long and 165 (164 – 167) wide, strongly reticulated, with six pairs of pores (gd 2, gd 4, gd 5, gd 6, gd 8 and gd 9); 18 pairs of setae on dorsal shield and two pairs on unsclerotised dorsal cuticle: j 1 15 (14 – 16), j 3 18 (17 – 19), j 4 12 (11 – 14), j 5 13 (11 – 14), j 6 16 (15 – 17), J 2 15 (14 – 16), J 5 8 (6 – 8), z 2 13 (12 – 14), z 3 16 (14 – 17), z 4 15 (14 – 16), z 5 14 (13 – 15), Z 4 25 (22 – 26), Z 5 31 (29 – 34), s 4 23 (21 – 24), s 6 16 (14 – 17), S 2 22 (20 – 24), S 4 25 (24 – 26), S 5 18 (17 – 19), r 3 20 (19 – 22), and R 1 18 (16 – 19). All setae smooth and pointed, seta Z 5 smooth and bulbous. Peritreme (Fig. 48). Extended beyond the bases of j 1. Venter (Fig. 49). All shields are smooth. The sternal shield rectangular, posterior margin of sternal shield slightly wavy. Sternal shield 69 (68 – 71) long and 64 (63 – 65) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st 4) on unsclerotised membrane 20 (19 – 22) long. Distances between st 2 - st 2 58 (56 – 59), st 5 - st 5 56 (55 – 57). The base of the genital shield broad, flat posteriorly. Two pairs of metapodal shields present. The primary inguinal sigilla 21 (20 – 22) long and 5 (4 – 6) wide, secondary ones 14 (13 – 15) long. Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, pentagonal 112 (111 – 114) long, 90 (89 – 91) wide at level of ZV 2 and 65 (64 – 68) wide at anus level with four pairs of pre-anal setae, JV 1 10 (9 – 11), JV 2 10 (9 – 11), ZV 2 12 (11 – 13), JV 3 12 (11 – 13) and one pair of pre-anal pores gv 3 22 (20 – 23) apart below JV 2. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 10 (9 – 11), ZV 3 12 (12 – 14), JV 4 14 (13 – 15) and JV 5 38 (37 – 40); JV 5 long, smooth with bulbous tip. Chelicera (Figs 50; 97). Fixed digit 26 (25 – 27) long with six teeth and distinct pilus dentilis, movable digit 26 (25 – 27) long with three backwardly directed teeth. Spermatheca (Fig. 51; 96). Spermatheca bell-shaped. Calyx 16 (15 – 17) long with a distinct narrow neck at the base of atrium, minor duct and flattened major duct visible. Leg (Figs 52; 98). Macroseta on tarsus of leg IV with bulbous tip while the same on genu and tibia of leg IV are pointed and of the following lengths: genu 15 (14 – 16), tibia 20 (19 – 21) and tarsus 45 (44 – 46). Chaetotactic formula of genu II: 1 2 / 1, 2 / 0 1 and genu III 1 2 / 1, 2 / 01. Length of leg I 363 (360 – 368), leg II 240 (238 – 245), leg III 233 (230 – 236) and leg IV 388 (387 – 390).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD5FFCB2FD13189C5584543.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8385 / 20) collected from weed (Rubus elypticus), at Mawkdok: 25 ° 24 ' 30 " N, 91 ° 46 ' 44 " E, 1806 m AMSL, East Khasi Hills, Meghalaya on 1 st May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Two paratype females (Acarol. lab / BCKV / 8402 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD5FFCB2FD13189C5584543.taxon	etymology	Etymology. The specific name campana refers to the unique bell-shaped spermatheca of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD5FFCB2FD13189C5584543.taxon	discussion	Remarks. The new species Typhlodromus (Anthoseius) campana is different from all the members of subspecies group rhenanus by having bell-shaped spermatheca with conspicuous neck at the base of atrium. The new species is close to T. (A.) neorhenanus Gupta, 1977; T. (A.) sijiensis Gupta, 1986; and T. (A.) dahungensis Pramanik & Karmakar, 2016 by having similar dosum and venter. However, this new species differs from the close species by length and form of dorsal setae, shape of ventrianal shield, number of teeth on fixed and movable digit, shape of spermatheca, and number and form of leg macrosetae. All the differences between the new species and its close species are presented in Table 11.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD6FFC52FD137A4C6DF468F.taxon	description	(Figs 53 – 57; 99 – 101) Female (n = 6). Diagnosis. The species is diagnosed by having dorsum smooth, all dorsal setae short, smooth and with pointed tip, posterior margin of sternal shield wavy, setae st 3 off the sternal shield in separate platelets, fixed digit of chelicera with four teeth while movable digit with two teeth, spermatheca saccular-shaped, major duct conspicously flattened, leg IV with only one pointed macroseta on tarsus. Dorsum (Fig. 53). Dorsal shield 328 (326 – 329) long and 145 (145 – 147) wide, smooth, with six pairs of pores (gd 2, gd 4, gd 5, gd 6, gd 8 and gd 9); 18 pairs of setae on the dorsal shield and two pairs of setae on the unsclerotised cuticle: j 1 19 (17 – 20), j 3 21 (19 – 22), j 4 11 (10 – 12), j 5 11 (10 – 12), j 6 11 (10 – 12), J 2 13 (11 – 14), J 5 8 (7 – 9), z 2 13 (11 – 14), z 3 14 (12 – 14), z 4 16 (13 – 16), z 5 12 (9 – 12), Z 4 15 (14 – 16), Z 5 49 (48 – 51), s 4 15 (13 – 16), s 6 20 (18 – 21), S 2 19 (17 – 20), S 4 19 (17 – 20), S 5 22 (20 – 23), r 3 15 (14 – 16), R 1 16 (15 – 18). All setae smooth. Peritreme (Fig. 53). Extended beyond the bases of j 3. Venter (Figs 54; 99). All shields are smooth. The sternal shield square-shaped, posterior margin of sternal shield wavy, laterally with few lines. Sternal shield 65 (63 – 66) long and 82 (79 – 83) wide at level of setae st 1 - st 3 and st 3 - st 3 respectively, with two pairs of setae and two pairs of poroids; st 3 on separate shield, one pair of setae (st 4) on unsclerotised membrane 21 (21 – 23) long. Distances between st 2 - st 2 63 (62 – 64), st 5 - st 5 63 (62 – 64). Two pairs of metapodal shields present. The primary inguinal sigilla long 22 (20 – 22) and 8 (6 – 9) wide, secondary ones 11 (9 – 11) long. Genital shield smooth, broad at the base and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, pentagonal 103 (100 – 104) long, 75 (73 – 75) wide at level of ZV 2 and 60 (58 – 61) wide at anus level with four pairs of pre-anal setae, JV 1 13 (13 – 15), JV 2 13 (11 – 13), JV 3 13 (11 – 13), ZV 2 13 (11 – 13) and one pair of pre-anal pores gv 3 below seta JV 2, 16 (15 – 18) apart. Membrane surrounding ventrianal shield with four pairs of setae ZV 1 13 (11 – 13), ZV 3 10 (9 – 10), JV 4 10 (8 – 10) and JV 5 30 (28 – 31); JV 5 long and smooth. Chelicera (Figs 55; 101). Fixed digit 30 (29 – 31) long with four teeth and distinct pilus dentilis, movable digit 30 (28 – 30) long with two backwardly directed teeth. Spermatheca (Figs 56; 100). Calyx 16 (14 – 17) long, saccular with distinct atrium attached with a sclerotised structure, major duct narrow tubular, minor duct clearly visible. Legs (Fig. 57). The leg IV with only one pointed macroseta on tarsus 49 (48 – 51). Chaetotactic formula of genu II: 1 2 / 1, 2 / 0 1 and genu III 1 2 / 1, 2 / 0 1. Length of leg I 245 (242 – 245), leg II 220 (217 – 220), leg III 218 (216 – 222) and leg IV 388 (387 – 390).	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD6FFC52FD137A4C6DF468F.taxon	materials_examined	Type specimens. Holotype: female (Acarol. lab / BCKV / 8404 / 20) collected from moss associated with forest plant, at Cherrapunji 25 ° 24 ' 30 " N, 91 ° 46 ' 44 " E, 1806 m AMSL, East Khasi Hills, Meghalaya on 1 st May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Five paratype females (Acarol. lab / BCKV / 8403 – 8404 / 2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD6FFC52FD137A4C6DF468F.taxon	etymology	Etymology. The specific name cherrapunjiensis refers to the type locality of the new species.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
7D6487FFFFD6FFC52FD137A4C6DF468F.taxon	discussion	Remarks. The new species Typhlodromus (Anthoseius) cherrapunjiensis sp. nov. is different from all the species of rhenanus species group because of its unique type of spermatheca, major duct broad and flattened. This new species is very close to T. (A.) capparidis van der Merwe, 1968, T. (A.) apoxys van der Merwe, 1968 by having similar type of saccular-shaped spermatheca, sternal shield with two pairs of setae and leg IV with a single macroseta. However, this new species differs from these species by the dorsal setal form and length, number of teeth on fixed and movable digit of chelicera, posterior margin of sternal shield, shape of ventrianal shield, tip of leg macroseta and spermatheca having broad, flattened major duct. All the differences between the new species and the close species are given in Table 12.	en	Kar, Anamika, Karmakar, Krishna (2021): Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India. Zootaxa 5068 (3): 301-354, DOI: https://doi.org/10.11646/zootaxa.5068.3.1
