taxonID	type	description	language	source
4C1287DE816E8D245F6139D6FAC8FACE.taxon	type_taxon	Type species Mioscarta forcipata Breddin, 1901.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816E8D245F6139D6FAC8FACE.taxon	diagnosis	Diagnosis The genus can be identified by the following combination of characters: Habitus general shape dorso-ventrally flattened, in lateral view total length nearly 4 times height (Figs 2, 4 – 11 A). Pronotum angle not more than 25 ° (Figs 2, 4 – 11 A). Distance between ocellus and compound eye 2 times ocellus diameter (Fig. 1 A). Ocelli large, distance between eyes less than 8 times ocellus diameter (Fig. 1 A). Apical reticulation of the tegmen generally developed and reduced in few cases (Figs 4 – 11 A – C). Widest part of postclypeus in frontal view is at midheight (Fig. 1 A). Apical curve of tegmen visible in dorsal view (Figs 4 – 11 C). Widest part of habitus in dorsal view at midlength of tegmen (Figs 4 – 11 C). Male subgenital plates is at least 1.5 times longer than pygofer height. Male subgenital plates appendage always present, longer than main plate (Figs 5, 7 – 8, 10 – 11 F). Among the Rhinaulacini, Mioscarta closely resembles Trigonoschema Crispolon & Soulier-Perkins gen. nov. with respect to the distance between ocelli, postclypeus shape in frontal view, postclypeus longitudinal groove, apical cells of the tegmen, Rp posterior wing, absence of r-m crossvein, and presence of sterno-lateral plate between subgenital plate and pygofer while Peociloterpa with respect to postclypeus longitudinal groove, absence of r-m crossvein, presence of sterno-lateral plate between subgenital plate and pygofer and paramere general shape, but they differ by the following characters presented in Table 2 below.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816E8D245F6139D6FAC8FACE.taxon	distribution	Distribution India, Indonesia (Borneo, Flores and Sulawesi), Malaysia (Borneo), and Philippines.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816F8D245F653E00FCEFF833.taxon	description	Fig. 4	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816F8D245F653E00FCEFF833.taxon	materials_examined	Material examined PHILIPPINES • 3 ♀♀; “ Island of Basilan Baker ”; “ 1926 ”; “ Typus ”; “ coll. A. Jacobi ”, “ basilana Jac. ”, “ Museum für Tierkunde Dresden (MTD) ”; SMTD.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816F8D245F653E00FCEFF833.taxon	distribution	Distribution Philippines: Mindanao: Basilan.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816F8D245F653E00FCEFF833.taxon	discussion	Remarks The description provided by Jacobi (1927) was based on four females. Here we provide photographs of a female syntype and its labels. Male remains unknown.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	description	Fig. 5	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	materials_examined	Material examined PHILIPPINES • ♀; “ Leyte Mt. Pangasugan Elev. 300 m; 6 May 1952; C. R. Baltazar ”, “ Eoscarta sp. Bal 53 ” “ UPLBMNH HEM- 04043 ” • ♂; “ Mt. Maquiling Elev. 200 m.; 23 Jan. 1954; J. N. Gibe ”, “ UPLBMNH HEM- 04044 ” • ♂ ♀; “ Odell Pltn. Tagum, Davao; 16 Jan. 1956; G. B. Viado ”, “ UPLBMNH HEM- 04045, 04047 ” • ♀; “ Odell Pltn. Tagum, Davao; 17 Jan. 1956; S. M. Cendaña ”; “ UPLBMNH HEM- 04046 ” • ♀; “ Puting Lupa, Cal.; 3 Aug. 1986; Chantana B. coll. ”; “ UPLBMNH HEM- 04997 ” • ♀; “ Ex. grasses CES, Laguna; 2 Jan. 1981; M. A. Tabasa ”, “ Cercopidae ”, “ UPLBMNH HEM- 04062 ” • ♀; “ PH. MT. APO, Lake Agco, Kidapawan, Cotabato, Min; 30 Jul. 2015; E. S. Crispolon ”, “ Muséum Paris; MNHN (EH) 24853 ” • ♂ ♀; “ PH. Lake Sebu, South Cotabato, Mindanao; 20 Jul. 2016; E. S. Crispolon ”; “ Muséum Paris; MNHN (EH) 24854 to 24855 ” • 2 ♀♀; “ PH. Paniki Falls, Balabag, Kidapawan, Cotabato, Min; 18 March 2017; N. Melano ”, “ Muséum Paris, MNHN (EH) 24856 to 24857 ” • ♀; “ PH. Lapan, Kidapawan, N. C.; 5 May 2017; D. M. Cantil ”; ECpc 054 • ♂; “ Ph. Kansal, Kid. Cot.; 18 Nov. 2017; Pasion C. A. ”; ECpc 056 • ♂; “ PH. Lake AGCO; 21 Dec. 2017; ESC ”; Philippines; ECpc 057 • ♂; “ Philippines, Mt. Apo, Bongolanon, Magpet N. Cotabato ”; “ 27 to 28 Jun. 2019; JS Ramos ”; “ séquençage par Elorde Crispolon C- 00027 ”; “ Muséum Paris, MNHN (EH) 24858 ”.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	description	Description Male Terminalia In lateral view, posterior margin of pygofer slightly undulating (Fig. 5 E). Subgenital plate (Fig. 5 F) very long relative to height of pygofer with fine tapering appendage slightly longer than main plate, dorsal margin of plate largely rounded, sterno-lateral plate present, largely concave subapically on dorsal and ventral margins. Intermediate plate present, rectangular, linking internal side of lateral and subgenital plates. Paramere (Fig. 5 G) not globose, dorsal margin rounded and regularly curving, apex with a sharp spine pointing dorsally, ventral margin regularly rounded with subapical part bearing a spiniform process pointing ventrally. Aedeagus (Fig. 5 H) with basal third of dorsal margin angled and sharply bent in acute angle and last 2 ⁄ 3 vertical and roughly C-shaped, slightly humped in apical part of dorsal margin before apical extension, apical extension pointing postero-dorsally, posterior protrusion thin and hook-shaped, postero-dorsal protrusion absent.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	materials_examined	Type Locality Philippines: Visayas: Samar.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	distribution	Distribution Philippines: Luzon Island, Visayas: Samar and Leyte, Mindanao Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816C8D205F003A72FC40FADA.taxon	discussion	Remarks Walker (1851) based his description on two male specimens “ a ” and “ b ” from the “ Philippine Islands ” coming from the collections of Mr Cuming and Mr Wood. Both specimens are currently kept in the NHM. According to Walker’ description and Lallemand’ key (1949), we identified specimens, for which we provide photographs and male terminalia description and drawings.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816B8D205F5D3E05FD71F836.taxon	description	Fig. 6	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816B8D205F5D3E05FD71F836.taxon	materials_examined	Type Locality Philippines: Luzon: Baguio, Benguet province.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816B8D205F5D3E05FD71F836.taxon	distribution	Distribution Philippines: Luzon Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE816B8D205F5D3E05FD71F836.taxon	discussion	Remarks Schmidt (1925) described this species based on a single female specimen. Therefore, the description does not contain any information on the male terminalia. Here, we provide photographs of the female holotype and labels. Male remains unknown.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	description	urn: lsid: zoobank. org: act: 17979 C 2 C-D 071 - 44 EA- 9 D 16 - 974 A 742808 A 1 Fig. 7	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	diagnosis	Diagnosis General shape of M. nubisa is similar to M. obscuripennis but are distinctly different in color. M. nubisa in dorsal view presents a brownish and opaque tegmen, darker toward apex, pronotum brownish and yellowish or brown legs while M. obscuripennis has a dark brown or black tegmen, brown or orange pronotum and orange legs.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	etymology	Etymology Species name refers to the light to darker coloration forming a cloudy pattern on the tegmen and is the female superlative of the latin word “ nubis ” which means cloudy.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	materials_examined	Material examined Holotype PHILIPPINES • ♂; “ Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga ”; “ Muséum Paris; 1 May 2011; S. A. Yap, M. V. Yngente, O. L. Eusebio rec. ”; UPLBMNH HEM- 05052 ”; UPLBMNH. Paratypes PHILIPPINES • ♀; “ Trinidad; 22 June 1951; C. R. Baltazar ”; “ UPLBMNH HEM 04064 ”; UPLBMNH • ♂; “ Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga ”; “ Muséum Paris, 1 May 2011; S. A. Yap, M. V. Yngente, O. L. Eusebio rec. ”; UPLBMNH HEM- 05053 ”; UPLBMNH • 16 ♂♂; “ Philippines, Luzon, mount Isarog, Light trap ”; “ Muséum Paris; 31 May 2011; S. Yap Rec. ”; “ Muséum Paris, MNHN (EH) 23645 to 23646, 23648, 24748, 24859 to 24870 ”; MNHN • ♂; “ Philippines, Luzon isl, Laguna, Los Banos, Mount Makiling; 14 º 082 ' 29 " N, 121 º 12.242 " E; “ Muséum Paris; 7 Nov. 2008; 605 m, Primary forest, at sight, A. Soulier-Perkins, S. Yap Rec. ”; “ Muséum Paris, MNHN (EH) 23644 ”; MNHN.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	description	Description BODY. Length 10 mm (tegmina included), width 5 mm. HEAD (Figs 1 A, 7 B). In dorsal view, large ocelli, distance between eyes less than 8 times ocellus diameter, distance between ocelli equals one ocellus diameter, distance between ocellus and compound eye 2 times ocellus diameter, ocelli closer to each other than from compound eyes. Eyes not prominent, length 1.44 times than wide. Vertex and frons longitudinal median carina absent. Vertex slightly longer than wide with 3 times ocellus diameter in between two vertex grooves outside ocelli and 3.5 times ocellus diameter between anterior and posterior vertex margins. Postclypeus with longitudinal furrow, slightly swollen and ovoid shape in frontal view, widest part at mid height (Figs 1 A, 7 B), not receding and prior to anteclypeus where it bends forming obtuse angle in lateral view (Fig. 7 A). Rostrum long, reaching but not surpassing mesocoxae. Thorax (Figs 2 A, 7 A – C). In dorsal view, pronotum with anterior concavities on each side, anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margin curved, posterior margin grooved, postero-lateral margin slightly concave, longer than anterolateral margin, humeral angle rounded. In lateral view, pronotum curving not more than 25 º (Figs 2, 7 A). Scutellum as long as wide with large median dimple (Fig. 7 C). Tegmen (Fig. 7 A – C). R bifurcates on apical half, M bifurcate on basal third, apical reticulation well developed without concave apical cells. Posterior wing (Fig. 3 A), Rp separating from SC + Ra nearly at midlength, M reaches ambient vein, Cua and Cup fused at base and m-cu links M to Cua before Cua bifurcation, common base for Cup and Cua originate at base of wing, posterior wing with 7 longitudinal veins and 5 apical cells between SC + Ra and Cup, angular protrusion of costal margin near its base present. Metafemur with apical spine in inner margin, metatibiae bearing 1 lateral spine. MALE TERMINALIA. In lateral view, posterior margin of pygofer largely convex in middle with slight curved on last third (Fig. 7 E). Subgenital plates (Fig. 7 F) very long relative to height of pygofer with fine tapering appendage longer than main plate, dorsal and ventral margin of main plate straight, sterno-lateral plate present, slightly elongated. Intermediate plate present, elongated, roughly oblong shaped, linking internal sides of lateral and subgenital plates. Paramere (Fig. 7 G) not globose, dorsal margin convex and regularly curving finishing by a sharp process pointing dorsally, ventral margin convex and largely angled subapically, apex with spiniform process pointing postero-ventrally. Aedeagus (Fig. 7 H) with basal third of dorsal margin regularly bent without angle before the bent part, last 2 ⁄ 3 vertical and S-shaped, ventral margin regularly curved, apical extension pointing posteriorly, posterior protrusion axe-shaped with edge prolonged ventrally by a straight, long and thin extension, postero-dorsal protrusion absent. COLOR. Head, antennae, pronotum, abdomen, legs scutellum light brown, rostrum brownish highlighted with white. Tegmen opaque light brown to brown.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	materials_examined	Type locality Philippines: Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81688D3C5CEE3A73FE4DFAB2.taxon	distribution	Distribution Philippines: Luzon Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	description	Figs 8 – 9	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	materials_examined	Material examined PHILIPPINES • 4 ♂ ♀; “ Baguio, Phil.; 22 Jun. 1952; L. B. Uichanco ”, “ UPLBMNH-HEM 04053, 04057 to 04059, 04063 ” • ♀; “ Mt. Trail 6000 ft; 25 Jun. 1957; R. LUIS ”; “ UPLBMNH HEM 04048 ” • ♂; “ EX. grasses; 27 Jan. 1980; NBG REAL, QUEZON J. B. BALATIBAT “ UPLBMNH-HEM 04060 ” • ♂; “ MSAC; 12 Aug. 1984; A. U. MANCIA ”, “ UPLBMNH-HEM 04990 ” • ♀; “ QUEZON: NBG; 19 Aug. 1989; VP Gapud ”; “ UPLBMNH-HEM 04991 ” • ♂; “ MSAC; 12 Aug. 1984; A. U. MANCIA ”; “ UPLBMNH-HEM 04992 ” • 2 ♂♂; “ Mt. Polis, Baubo Mtn. Province 22 Dec. 2000; Shem Padua ”; “ UPLBMNH-HEM 04993 to 04994 ” • ♂; “ QUEZON REAL Nat. Bot. Garden; 1 – 2 Apr. 1977; V. P. GAPUD ”; “ UPLBMNH-HEM 04995 ” • ♀; “ Philippines, Negros volcan Canlaon, champ de coqs, 10 º 25 ' 29 " N, 123 º 05 ' 36 " E ”; “ Muséum Paris, 1050 m; 28 Oct. 2010; D. Ouvrard Rec. ”, “ Muséum Paris, MNHN (EH) 24809 ” • ♀; “ Philippines, Negros volcan Canlaon, champ de coqs, 10 º 25 ' 29 " N, 123 º 05 ' 36 " E ”; “ Muséum Paris, 1050 m; 28 Oct. 2010; A Soulier-Perkins Rec. ”; “ Muséum Paris, MNHN (EH) 23650 ” • 3 ♂♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 34 " N, 123 º 05 ' 37 " E ”, “ Muséum Paris, 1070 m; 29 Oct. 2010; D. Ouvrard Rec. ” “ Muséum Paris, MNHN (EH) 24814 to 24816 ” • 3 ♂♂; “ Philippines, Negros volcan Canlaon, forêt; 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ”; “ Muséum Paris, 1098 m; 30 Oct. 2010; D. Ouvrard Rec. ” “ Muséum Paris, MNHN (EH) 24805 to 24807 ” • 17 ♂ ♀; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ”; “ Muséum Paris, Piège lumineux 1098 m; 30 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24661, 24872 to 24886, 24808 ” • 3 ♂♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 36 " N, 123 º 05 ' 37 " E ”; “ Muséum Paris, Piège lumineux 1057 m; 31 Oct. 2010; D. Ouvrard Rec. ” “ Muséum Paris, MNHN (EH) 24900 to 24902 ” • 14 ♂ ♀; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 36 " N, 123 º 05 ' 37 " E ”; “ Muséum Paris, Piège lumineux 1057 m; 31 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 23651, 24887 to 24899 ” • 2 ♂♂; “ Philippines, Negros volcan Canlaon, forêt; 10 º 25 ' 35 " N, " 123 º 05 ' 51 " E ”; “ Muséum Paris, 1117 m; 2 Nov. 2010; D. Ouvrard Rec. ”, “ Muséum Paris, MNHN (EH) 24810 to 24811 ” • ♂; “ Philippines, Negros volcan Canlaon, forêt; 10 º 25 ' 35 " N, " 123 º 05 ' 51 " E ”; “ Muséum Paris, 1117 m; 2 Nov. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24812 ” • ♂; “ Philippines, Negros volcan Canlaon, milieu anthropisé; 10 º 25 ' 23 " N, 123 º 04 ' 46 " E ”; “ Muséum Paris, Piège lumineux 745 m; 3 Nov. 2010; D. Ouvrard Rec. ”, “ Muséum Paris, MNHN (EH) 24813 ” • 3 ♂ ♀; “ Philippines, Luzon, mount Isarog, Light trap' ”, “ Muséum Paris; 31 May 2011; S. Yap Rec. ”, “ Muséum Paris, MNHN (EH) 23649, 24803 to 24804 ”.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	description	Description MALE TERMINALIA. In lateral view, posterior margin of pygofer hump shaped in middle, last third abruptly truncated (Fig. 8 E). Subgenital plates (Fig. 8 F) very long relative to height of pygofer with fine tapering appendage longer than main plate, dorsal margin regularly convex and slightly curving on basal part, ventral margin roughly straight. Sterno-lateral plate present and roughly bean-shaped. Intermediate plate present, square shaped with a corner truncated, linking internal sides of lateral plate and subgenital plate. Paramere (Fig. 8 G) dorsal margin rounded then regularly curving up to apex. Lateral margin with spine located subapically pointing dorsally, ventral margin rounded then sharply angled subapically pointing ventrally, apex without any spines. Aedeagus (Fig. 8 H) with basal third of dorsal margin regularly bent, basal half enlarged then largely narrowed in the middle and gradually enlarging up to apex, roughly C-shaped, ventral margin regularly curved, apical extension pointing posteriorly, posterior protrusion axe-shaped with long thin extension prolonging the edge ventrally and slightly curving, postero-dorsal protrusion absent.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	materials_examined	Type Locality Philippines: Luzon: Mount Banahao, Quezon Province.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	distribution	Distribution Philippines: Luzon Island, Baguio, Quezon province, Mountain province; Visayas, Negros Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81778D385F143EEDFBB6F9CD.taxon	discussion	Remarks Schmidt (1920) described this species from a single male specimen with few details on the male terminalia. Here, we provide, a detailed illustration and description of male terminalia from identified specimens using Schmidt description (Fig. 8), along with photographs of the male holotype and its labels (Fig. 9). The coloration of the specimen we identified and the holotype are different but the specimen from which we illustrated the male terminalia (Fig. 8) has a brighter coloration and is the closest to the description given by Schmidt while the holotype has a faded coloration.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	description	Fig. 10	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	materials_examined	Material examined PHILIPPINES • ♀; “ Samar Calbayog Elev. – Sea level; 8 May 1950; R. Araneta Jr. ”, “ Eoscarta sp. Bal ’ 53 ” “ UPLBMNH HEM- 04054 ” • ♂; “ Calambog Ala Valley Elev.; 8 June 1950 S;. M Cendaña ” “ Eoscarta sp. 1 ” “ UPLBMNH HEM- 04061 ” • ♀ ♂; “ BUKIDNON Impalitaw BFD; 19 Nov. 1988; VPG ” “ UPLBMNH HEM- 04996, 04998 ” • 2 ♂♂; “ Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga ”, “ Muséum Paris; 1 May 2011; S. A. Yap, M. V. Yngente, O. L. Eusebio rec. ”, “ Muséum Paris, MNHN (EH) 24821 to 24822 ” • 3 ♂♂; “ Philippines Luzon Mount Isarog Light trap ”, “ Muséum Paris; 31 May 2011; S. Yap Rec ”, “ Muséum Paris, MNHN (EH) 23647, 24823 to 24824 ” • ♀; “ PH. MT. Makiling Los Baños, Laguna Luz; 21 Nov. 2016; WS HWANG ”, “ séquençage par Elorde Crispolon C- 00085 ”, “ Muséum Paris, MNHN (EH) 24817 ” • ♀; “ Brgy. Irosin, Sorsogon; 2 Feb. 2017; Coll. AK AMARGA ”; “ Muséum Paris, MNHN (EH) 24818 ” • ♀; “ Ph. Brgy Perez, Kidapawan City; 5 May 2017; C. S. Satohito ”, “ ECpc 057 ”, “ Ph. Mt. Akir Akir, Pigcawayan, North Cotabato; 8 – 9 Sept. 2018; G. Biacolo, K. C. Crispolon & R. Guillermo ”; “ séquençage par Elorde Crispolon C- 00017 ”, “ Muséum Paris, MNHN (EH) 24819 ”, “ Ph. Brgy Kidama, Matalam, North Cotabato; 14 Mar. 2019; K. C. Crispolon ”, “ séquençage par Elorde Crispolon C- 00022 ”, “ Muséum Paris, MNHN (EH) 24820 ”.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	description	Description MALE TERMINALIA. In lateral view, posterior margin of pygofer regularly convex last third slightly curved (Fig. 10 E). Subgenital plates (Fig. 10 F) very long relative to height of pygofer with fine tapering appendage slightly longer than main plate, dorsal margin of plate regularly rounded, ventral margin straight. Sterno-lateral plate present, slightly elongated. Intermediate plate present, cylindrical, linking internal side of lateral and subgenital plates. Paramere (Fig. 10 G) not globose, dorsal margin rounded and regularly curving, apex with a sharp spine pointing antero-dorsally, ventral margin regularly rounded then subapical part largely rounded, protruding and finishing with a sharp spine pointing ventrally. Aedeagus (Fig. 10 H) with basal third of dorsal margin angled and sharply bent in acute angle and last 2 ⁄ 3 vertical and roughly C-shaped, slightly humped in apical part of dorsal margin before apical extension, apical extension pointing posteriorly, posterior protrusion thin very short and spine-shaped, postero-dorsal protrusion absent.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	materials_examined	Type Locality Philippines: Philippine Islands.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	distribution	Distribution Philippines: Luzon, Visayas and Mindanao Islands.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81738D3A5F613D09FDA1F9D3.taxon	discussion	Remarks According to what Jacobi (1905) mentioned at the end of the description “ Mus. Berol. Nr. 9107: Semper coll., 2 Ex. ”. He examined two specimens and described this species from atleast one female without a description of the male terminalia. We illustrate and describe them here based on a specimen identified using Lallemand’s (1949) key.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	description	urn: lsid: zoobank. org: act: 7 D 16117 A- 699 D- 4 C 7 E-B 037 - E 290 B 274 EA 8 C Fig. 11	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	diagnosis	Diagnosis M. translucida is the only species with tegmen translucid without any other coloration.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	etymology	Etymology The species name refers to the translucid tegmen and is based on the latin word “ translucidus ” which means allowing light to pass through.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	materials_examined	Material examined Holotype PHILIPPINES • ♂; “ Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga ”, “ Muséum Paris; 1 May 2011; S. A. Yap, M. V. Yngente & O. L. Eusebio rec. ”; “ Muséum Paris, MNHN (EH) 24749 ”; MNHN.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	description	Description BODY. Length 10 mm (tegmina included), width 4 mm. HEAD (Figs 1 A, 11 B). In dorsal view, large ocelli, distance between eyes less than 8 times ocellus diameter, distance between ocelli equals one ocellus diameter, distance between ocellus and the compound eye 2 times ocellus diameter, ocelli closer to each other than from compound eyes. Eyes not prominent, length 1.44 times than wide. Vertex and frons longitudinal median carina absent. Vertex as long as wide with 3 times ocellus diameter in between two vertex grooves outside ocelli and 3 times ocellus diameter between anterior and posterior vertex margins. Postclypeus with longitudinal furrow, slightly swollen and ovoid shape in frontal view, widest part at mid height (Figs 1 A, 11 B), not receding and prior to anteclypeus where it bends forming right angle in lateral view (Fig. 11 A). Rostrum long, surpassing mesocoxae. Thorax (Figs 2 A, 11 A-C). In dorsal view, pronotum with anterior concavities on each side, anterior margin as wide as posterior margin of head including eyes, anterolateral margins curved, posterior margin grooved, postero-lateral margins slightly concave, longer than anterolateral margins, humeral angle rounded. In lateral view, pronotum curving not more than 25 º (Figs 2, 11 A) Scutellum as long as wide with large median dimple (Fig. 11 C). Tegmen (Fig. 11 A – C). R bifurcates on apical half, M bifurcate on basal third, apical reticulation not well developed without concave apical cells. Posterior wing (Fig. 3 A). Rp separating from SC + Ra nearly at midlength, M reaches ambient vein, Cua and Cup fused at base and m-cu links M to Cua before Cua bifurcation, common base for Cup and Cua originate at base of wing, 7 longitudinal veins and 5 apical cells between SC + Ra and Cup, angular protrusion of costal margin near its base. Metafemur with apical spine in inner margin, metatibiae bearing 1 lateral spine. MALE TERMINALIA. In lateral view, posterior margin of pygofer slightly undulating in the middle with slight curved on the last third (Fig. 11 E). Subgenital plates (Fig. 11 F) dorsal and ventral margin of main plate roughly straight, sterno-lateral plate present, slightly elongated, intermediate plate present, elongated slightly triangular shaped, linking internal sides of lateral and subgenital plates. Paramere (Fig. 11 G) not globose, dorsal margin convex and regularly curving finishing with rounded apex with very minute groove, ventral margin convex, apex with spiniform process pointing antero-ventrally. Dorsal and ventral margins of aedeagus undulating. Aedeagus (Fig. 11 H) with basal third of dorsal margin regularly bent without angle before the bent part and last 2 ⁄ 3 vertical and S-shaped, ventral margin regularly curved then slightly concaved before the base, apical extension sharp pointing postero-ventrally, posterior protrusion sharp at the apex hook-shaped, postero-dorsal protrusion absent. COLOR. Head and pronotum brown with yellow patches, rostrum yellowish white, pedicel of antenna brown, legs yellowish and abdomen light brown. Tegmen partially translucid, opaque parts being yellowish with darker patches.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	materials_examined	Type Locality Philippines: Luzon, Camarines Sur, Mount Isarog Natural Park.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	distribution	Distribution Philippines: Luzon Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81718D375F3D3D0FFE71FE97.taxon	discussion	Remarks The exact length of subgenital plate relative to height of pygofer is not provided because part of the appendage is damaged.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	description	urn: lsid: zoobank. org: act: C 8299091 - 60 D 3 - 49 B 4 - 852 F- 2145956 D 7 FE 9 Figs 12 – 16	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	type_taxon	Type species Trigonoschema manoborum sp. nov. here designated.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	diagnosis	Diagnosis In dorsal view, teardrop shaped and apex of tegmen not visible due to folding. In lateral view, very steeply declivous pronotum and crown of the head, nearly form a right angle with the rest of the dorsum in profile.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	etymology	Etymology When observed in dorsal view, the general shape of the habitus fits in a triangle even if a closer observation would lead to describe it as teardrop shaped. The name is built using two greek words “ trigonos ” for triangular and “ schema ” for shape. It is neutral.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	description	Description BODY. Length 9.5 – 12.5 mm (tegmen included), width 4 – 5 mm. GENERAL SHAPE. Not dorso-ventrally flattened, in lateral view total length around 3 times height (Figs 2 B, 12 – 16 A). Ocelli small, distance between eyes 9 – 10.5 times ocellus diameter (Figs 1, 12 – 16 B). Distance between ocellus and compound eye less than 4 times ocellus diameter (( Figs 1, 12 – 16 B). Distance between ocelli 1.5 ocellus diameter (Figs 1, 12 – 16 B). Postclypeus with longitudinal furrow, swollen laterally and slightly ovoid shape in frontal view, widest part close to frons, not receding and prior to anteclypeus where it bends rounded in lateral view, with wide longitudinal furrow (Figs 1, 12 – 16 B). Pronotum curving around 45 º angle (Figs 2 B, 12 – 16 A). Tegmen translucid without concave apical cells. Apical curve of tegmen rarely visible in dorsal view, tegmen folded in such a way that widest part of habitus is before midlength of tegmen (Figs 12 – 16 C). MALE TERMINALIA. Subgenital plates without fine appendage (Fig. 12 F), if present shorter than main plate, not forming an acute angle shape with the main plate (Figs 14, 16 F). Paramere globose, lateral margin of paramere with protrusion, on the ventral margin on the sub apex with angular protrusion ornamented with set of setae (Figs 12, 14, 16 G). Pp of aedeagus thick and beak-like shape (Figs 12, 14, 16 H).	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817C8D365CE53AC3FAC8FE89.taxon	distribution	Distribution Philippines.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	description	urn: lsid: zoobank. org: act: EDF 12 AD 9 - 920 E- 46 AA- 937 F-D 66 CE 3 ACD 6 B 1 Figs 12 – 13	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	diagnosis	Diagnosis Tegmen, white in basal 1 ⁄ 4 with orange marking on the basal half of the margin of the clavus. Scutellum orange with yellowish white coloration in middle. Most of vertex and frons bright yellow and postclypeus dull orange for male and bright orange for female.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	etymology	Etymology Manoborum, genitive plural made from Manobos. The Manobo tribe is found in Bongolanon, Magpet, North Cotabato.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	materials_examined	Material examined Holotype PHILIPPINES • ♂; “ Philippines, Mt. Apo, Bongolanon, Magpet N. Cotabato ”; “ 5 May 2018; Daniela Alcalde ”; “ holotype ”, “ séquençage par Elorde Crispolon C- 00148 ”, “ Muséum Paris, MNHN (EH) 24871 ”; MNHN. Paratype PHILIPPINES • ♀; “ PH. MT. APO, Bongolanon, Magpet, Cot, Mindanao, 17 Dec. 2015, ES Crispolon ” “ séquençage par Elorde Crispolon C- 00078 ” “ UPLBMNH HEM- 05054 ”; UPLBMNH.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	description	Description BODY LENGTH. 10.5 mm (tegmina included), width 4 mm. HEAD (Figs 1, 12 – 13 B). In dorsal view, ocelli small, distance between eyes 10.5 times ocellus diameter (Figs 1, 12 – 13 B). Distance between ocellus and compound eye less than 3.5 times ocellus diameter (( Figs 1, 12 – 13 B). Distance between ocelli 1.5 ocellus diameter (Figs 1, 12 – 13 B). Eyes not prominent, length 1.15 times than wide. Vertex with longitudinal short median carina, slightly longer than wide. Frons without any carina. Postclypeus with dimple below margin of frons and a longitudinal furrow clearly marked in male, swollen laterally and slightly ovoid shape in frontal view, widest part close to frons, not receding and prior to anteclypeus where it bends rounded in lateral view (Figs 1 B, 12 – 13 A). Rostrum long, reaching but not surpassing mesocoxae. Thorax (Figs 2 B, 12 – 13 A – C). In dorsal view, pronotum with anterior deep concavities on each side, much clearly marked in female, longitudinal median carina absent. Anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margins curved, posterior margin grooved, postero-lateral margins slightly concave, slightly longer than anterolateral margins, humeral angle rounded. In lateral view, pronotum angle around 45 º (Figs 2 B, 12 – 13 A), scutellum as long as wide with large median dimple (Figs 12 – 13 C). Tegmen (Figs 12 – 13 A – C). R bifurcates on apical half, M bifurcate on basal third, apical reticulation not well developed without concave apical cells. Posterior wing (Fig. 13 E). 2 ⁄ 3 of M length alone then fuses with Rp before reaching ambient vein, 6 longitudinal veins on posterior wing reaching apex, four apical cells between SC + Ra and Cup, angular projection near base of costal margin present. Legs. Metafemur with apical spine in inner margin, metatibiae bearing 1 lateral spine. MALE TERMINALIA. In lateral view, posterior margin of pygofer (Fig. 12 E) largely convex in middle getting concave in last third and presenting a bump in first third. Subgenital plates (Fig. 12 F) with equal length relative to height of pygofer without any appendage, regularly tapering towards apex, dorsal margin straight then curving up towards apex, ventral margin straight half its length then regularly curving dorsally, minute denticuli covering last quarter toward apex. Sterno-lateral plate present, slightly elongated. Intermediate plates present, elongated and linked with internal sides of lateral plate and subgenital plate. Paramere (Fig. 12 G) globose, dorsal margin convex then curving regularly and finishing with a sharp spine pointed antero dorsally, lateral margin with process in middle part pointing postero-dorsally, ventral margin regularly rounded with subapical part largely protruding posteriorly then finishing with a sharp spine pointing ventrally. Aedeagus (Fig. 12 H) with dorsal margin slightly humped at base then concave most its length, curving regularly half its length vertically and finishing ¼ its length convex in a C-shaped, apical extension pointing posteriorly, posterior protrusion thick, duck head shaped with minute denticuli along posterior margin, in posterior view basal half lobular (Fig. 12 I), postero-dorsal protrusion absent. COLOR. Head, and pronotum white-yellowish with orange markings, postclypeus, rostrum, abdomen and legs orange, pedicel of antennae yellowish brown, scutellum orange with yellowish white markings medially. Tegmen, translucid yellowish white, white in basal ¼ with orange marking in between white and on the basal half of the margin of the clavus, last 3 ⁄ 4 yellowish getting yellowish brown in apical fourth.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	materials_examined	Type Locality Philippines, Mt. Apo, Bongolanon, Magpet North Cotabato.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE817D8D335C5A3AD4FDA9FC0A.taxon	distribution	Distribution Philippines: Mindanao Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	description	urn: lsid: zoobank. org: act: BFF 10 DEF-C 172 - 4861 - A 9 AE- 81 AA 8 B 88657 B Fig. 14	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	diagnosis	Diagnosis In general shape, T. negrosensis is similar to T. rubercella but their coloration is distinctly different. T. negrosensis in dorsal view presents a dirty yellow pronotum followed by a light orange scutellum framed by yellow irregular patches on clavus of tegmen while T. rubercella presents a bright yellow pronotum followed by a red band running on basal parts of tegmen and across scutellum.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	etymology	Etymology The species is named after the island where it was found: Negros.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	materials_examined	Material examined Holotype PHILIPPINES • ♂; “ Philippines, Negros volcan Canlaon, champ de coqs; 10 º 25 ' 29 " N, 123 º 05.23 " E ”; “ Muséum Paris, Piège lumineux, 932 m; 28 Oct. 2010; A. Soulier-Perkins rec. ”, “ UPLBMNH HEM- 05055 ”; UPLBMNH. Paratypes PHILIPPINES • 4 ♂♂; “ Philippines, Negros volcan Canlaon, champ de coqs; 10 º 25 ' 29 " N, 123 º 05 ' 36 " E ”; “ Muséum Paris, 1050 m; 28 Oct. 2010; D. Ouvrard Rec. ”, “ Muséum Paris, MNHN (EH) 23639, 24825 to 24827 ”; MNHN • 5 ♂♂; “ Philippines, Negros volcan Canlaon, champ de coqs, 10 º 25 ' 29 " N, 123 º 05.23 " E ”; “ Muséum Paris, Piège lumineux 932 m; 28 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24828 ”, “ Muséum Paris, MNHN (EH) 24829 to 24830 ”, “ séquençage par Elorde Crispolon, C- 00076, C- 00077 ”, “ Muséum Paris, MNHN (EH) 24831 ”; MNHN. “ UPLBMNH HEM- 05056 ”; UPLBMNH • ♀; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 40 " E ”; “ Muséum Paris, Piège lumineux 1098 m; 29 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24832 ”; MNHN • ♀; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ” “ Muséum Paris, 1098 m, 30 Oct. 2010, D. Ouvrard Rec. ”, “ Muséum Paris, MNHN (EH) 24833 ”; MNHN • ♂; “ Philippines, Negros volcan Canlaon, forêt; 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ”; “ Muséum Paris, Piège lumineux 1098 m; 30 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24834 ”; MNHN • 2 ♀♀, 2 ♂♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 36 " N, 123 º 05 ' 37 " E ”; “ Muséum Paris, Piège lumineux 1057 m, 31 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24835 to 24837 ” “ séquençage par Elorde Crispolon C- 00167 ”, “ Muséum Paris, MNHN (EH) 24838 ”; MNHN • ♀; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 23 " N, 123 º 04 ' 46 " E; “ Muséum Paris, Piège lumineux 745 m; 3 Nov. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24839 ”; MNHN. • 2 ♂♂; “ Philippines, Luzon, mount Isarog, Light trap ”, “ Muséum Paris; 31 May 2011; S. Yap Rec. ” “ Muséum Paris, MNHN (EH) 24840 to 24841 ”; MNHN.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	description	Description BODY. Length 11 – 12.5 mm (tegmina included), width 4 – 5 mm. HEAD (Figs 1, 14 B). In dorsal view, ocelli small, distance between eyes 9.5 times ocellus diameter (Figs 1, 14 B). Distance between ocellus and compound eye less than 3 times ocellus diameter (Figs 1, 14 B). Distance between ocelli 1.5 ocellus diameter (Figs 1, 14 B). Eyes not prominent, length 1.33 times than wide. Vertex with longitudinal median carina, as long as wide. Frons without median carina. Postclypeus with dimple below margin of frons and a longitudinal furrow clearly marked in males, swollen laterally and slightly ovoid shape in frontal view, widest part close to frons, not receding and prior to anteclypeus where it bends rounded in lateral view (Figs 1 B, 14 A). Rostrum long, reaching but not surpassing mesocoxae. Thorax (Figs 2 B, 14 A – C). In dorsal view, pronotum with anterior deep concavities on each side, much clearly marked in female, longitudinal median carina absent. Anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margins curved, posterior margin grooved, postero-lateral margins slightly concave, slightly longer than anterolateral margins, humeral angle rounded. In lateral view, pronotum angle around 45 º (Figs 2 B, 14 A). Scutellum as long as wide with large median dimple (Fig. 14 C). Tegmen (Fig. 14 A, C). R bifurcates on apical half, M bifurcate on basal third, apical reticulation not well developed without concave apical cells. Posterior wing (Fig. 3 A). Rp separating from SC + Ra nearly at midlength, M reaches ambient vein, Cua and Cup fused at base and m-cu links M to Cua before Cua bifurcation, 7 longitudinal veins and 5 apical cells between SC + Ra and Cup, angular protrusion of costal margin near its base. Metafemur with apical spine in inner margin, metatibiae bearing 1 lateral spine. MALE TERMINALIA. In lateral view, posterior margin of pygofer (Fig. 14 E) convex in middle and clearly concave on last third. Subgenital plates (Fig. 14 F) slightly longer relative to height of pygofer with fine appendage shorter forming acute angle with main plate, dorsal and ventral margin of main plate roughly straight. Sterno-lateral plate present and slightly elongated. Intermediate plates present, boomerangshaped, linking internal sides of lateral plate and subgenital plate. Paramere (Fig. 14 G) globose, dorsal margin convex then curving regularly and finishing with a sharp spine pointed antero dorsally, lateral margin with process in middle part pointing dorsally, ventral margin regularly rounded with subapical part largely protruding posteriorly then finishing with two sharp spines pointing postero-ventrally. Aedeagus (Fig. 14 H) with basal fourth almost straight then bending vertically and slightly wave-shape up to postero-dorsal protrusion, apical extension pointing dorsally, posterior protrusion thick, beakshaped, in posterior view, very slim at the top and petal shaped and foliaceous toward the base (Fig. 14 I), postero-dorsal protrusion present. COLOR. Head including postclypeus and rostrum, legs and abdomen orange, pronotum orange white anterior half, greenish on apical part, scutellum greenish white with orange coloration, antennal scape and pedicel orange, flagellum with brown coloration. Tegmen translucid yellow, basal third with white coloration in costal margin and clavus and with orange coloration in between white coloration, last 2 ⁄ 3 yellowish brown.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	materials_examined	Type Locality Philippines: Visayas, Negros Occidental, Mount Kanlaon.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81788D0C5C8A3F54FCBDFDC5.taxon	distribution	Distribution Philippines, Visayas, Negros Island; Luzon Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0C5CE23911FE5BFB22.taxon	description	Fig. 15	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0C5CE23911FE5BFB22.taxon	distribution	Distribution Philippines: Philippine Islands.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0C5CE23911FE5BFB22.taxon	discussion	Remarks Lallemand in 1927 described this species based on the pattern of coloration and placed it in Mioscarta. He mentioned only a single male specimen (holotype), however, in the collection it is labelled as female (Fig. 15 D). This species does not conform to Breddin’s definition of Mioscarta but possesses characters of Trigonoschema gen. nov. For this reason we transfer it to this new genus. We could not borrow the holotype, however some photographs of the habitus and labels were kindly provided by Mick Webb (NHM). They are integrated here (Fig. 15) and are sufficient to see the most obvious characters of Trigonoschema gen. nov.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	description	urn: lsid: zoobank. org: act: 02 DFAF 48 - FC 8 F- 45 CE-BE 52 - FB 1 DD 05626 DF Fig. 16	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	diagnosis	Diagnosis In general shape, T. rubercella is similar to T. negrosensis but is distinctly different in color. T. rubercella presents a bright yellow pronotum followed by red band running on the basal parts of the tegmen and scutellum when T. negrosensis in dorsal view presents a dirty yellow pronotum followed by a light orange scutelum framed by yellow patches sprawling on the clavus of the tegmen.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	etymology	Etymology This species has a distinct red coloration in the network of cells at the apex of tegmen. The species name is a combination of two latin words “ ruber ” and “ cella ” respectively meaning red and cell. The name is placed in apposition.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	materials_examined	Material examined Holotype PHILIPPINES • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 29 " N, 123 º 05.36 " E ”; “ Muséum Paris, 1050 m; 28 Oct. 2010; D. Ouvrard Rec ”, “ Muséum Paris, MNHN (EH) 23638 ”, “ séquençage par Elorde Crispolon C- 00074 ”; MNHN. Paratypes PHILIPPINES • ♂; “ Philippines, Negros volcan Canlaon, champ de coqs, 10 º 25 ' 29 " N, 123 º 05 ' 23 " E ”, “ Muséum Paris, Piège lumineux 932 m; 28 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24842 ”; MNHN • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 40 " E ”, “ Muséum Paris, Piège lumineux 1098 m; 29 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24843 ”; MNHN • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ”, “ Muséum Paris, 1098 m; 30. Oct. 2010; D. Ouvrard Rec ”, “ Muséum Paris, MNHN (EH) 24844 ”; MNHN • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 31 " N, 123 º 05 ' 43 " E ”, “ Muséum Paris, Piège lumineux 1098 m; 30 Oct. 2010; A. Soulier-Perkins rec. ”, “ UPLBMNH HEM- 05058 ”; UPLBMNH. • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 36 " N, 123 º 05 ' 37 " E ”, “ Muséum Paris, Piège lumineux 1057 m; 31 Oct. 2010; D. Ouvrard Rec ”, “ UPLBMNH HEM- 05057 ”; UPLBMNH • 2 ♂♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 36 " N, 123 º 05 ' 37 " E ”, “ Muséum Paris, Piège lumineux 1057 m; 31 Oct. 2010; A. Soulier-Perkins rec. ”, “ Muséum Paris, MNHN (EH) 24845 ”, “ séquençage par Elorde Crispolon C- 00075 ”, “ Muséum Paris, MNHN (EH) 24846 ”; MNHN • ♂; “ Philippines, Negros volcan Canlaon, forêt, 10 º 25 ' 35 " N, 123 º 05 ' 51 " E ”; “ Muséum Paris, 1117 m; 2 Nov. 2010; D. Ouvrard rec. ”, “ Muséum Paris, MNHN (EH) 23640 ”; MNHN.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	description	Description BODY LENGTH. 9.5 – 10.5 mm (tegmina included), width 4.5 mm. HEAD (Figs 1, 16 B). In dorsal view, ocelli small, distance between eyes 9 times ocellus diameter (Figs 1, 16 B). Distance between ocellus and compound eye less than 4 times ocellus diameter (Figs 1, 16 B). Distance between ocelli 1.5 ocellus diameter (Figs 1, 16 B). Eyes not prominent, length 1.33 times than wide. Vertex slightly longer than wide. bearing a median longitudinal carina. Frons without carina. Postclypeus with dimple below margin of frons and a longitudinal furrow, swollen laterally and slightly ovoid shape in frontal view, widest part close to frons, not receding, prior to anteclypeus where it bends forming obtuse angle in lateral view (Figs 1 B, 16 A). Rostrum long, reaching but not surpassing mesocoxae. THORAX (Figs 2 B, 16 A – C). In dorsal view, pronotum with anterior deep concavities on each side, much clearly marked in female, longitudinal median carina absent. Anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margins curved, posterior margin grooved, postero-lateral margins slightly concave, slightly longer than anterolateral margins, humeral angle rounded. In lateral view, pronotum angle around 45 º (Figs 2 B, 16 A). Scutellum as long as wide with large median dimple (Fig. 16 C). TEGMEN (Fig. 16 A – C). R bifurcates on apical half, M bifurcate on basal third, apical reticulation not well developed without concave apical cells. Posterior wing (Fig. 3 A). Rp separating from SC + Ra nearly at midlength, M reaches ambient vein, Cua and Cup fused at base and m-cu links M to Cua before Cua bifurcation, 7 longitudinal veins and 5 apical cells between SC + Ra and Cup, angular protrusion of costal margin near its base present. Metafemur with apical spine in inner margin and metatibiae bearing 1 lateral spine. MALE TERMINALIA. In lateral view, posterior margin of pygofer (Fig. 16 E) convex in middle with slight concavity toward ventral margin. Subgenital plates (Fig. 16 F) with equal length relative to height of pygofer with fine appendage shorter than main plate directed posteriorly not forming acute angle with main plate, dorsal and ventral margin of main plate regularly curved. Sterno-lateral plates present, triangular shaped. Intermediate plates present, roughly boomerang-shaped linking internal sides of lateral plate and subgenital plate. Paramere (Fig. 16 G) globose, dorsal margin convex then curving regularly and finishing with a sharp spine pointed dorsally, lateral margin with slightly and rounded, ventral margin roughly straight with subapical part angled largely protruding posteriorly then finishing with two sharp spines pointing postero-ventrally. Aedeagus (Fig. 16 H) with dorsal margin making a right angle at its base, straight on a small portion before curving up regularly, apical part bending posteriorly and straight, apical extension pointing dorsally, posterior protrusion thick and beak-shaped, postero-dorsal protrusion absent. COLOR. Head, legs, abdomen, scutellum and anterior part of pronotum red, rest of pronotum yellow, rostrum red, antennal scape and pedicel reddish orange, flagellum yellowish. Tegmen translucid yellow, basal and apical third including network of cells red.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	materials_examined	Type Locality Philippines: Visayas, Negros Occidental, Mount Kanlaon.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	distribution	Distribution Philippines: Visayas, Negros Island.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
4C1287DE81478D0B5C983E7CFEDFFD0B.taxon	description	Molecular Phylogeny Results of the Bayesian 50 % consensus tree and ML analyses with likelihood score of − 13870.493 are shown in Figures 17 and 18 respectively. The resulting topologies are not similar with respect to the placement of Jacobsoniella and Wawi. In the Bayesian topology (Fig. 17), Jacobsoniella is recovered within a clade containing Eoscarta, Mioscarta, Poeciloterpa and Trigonoschema gen. nov. clades, each supported by a probability value (PPv) of 100 %. This clade (Eoscarta + Mioscarta + Poeciloterpa + Trigonoschema + Jacobsoniella) is supported PPv of 82.61 % and Wawi appears as sister group to it. However, the clade containing Wawi plus the five other genera is supported with a high PPv of 100 %. In the ML analysis (Fig. 18), Poeciloterpa and Trigonoschema gen. nov. are sister groups with a ML bootstrap value (MLBv) of 85.2 / 21 % (SH-aLRT support / standard bootstrap support). Then Mioscarta appears as sister group of (Eoscarta + Wawi + (Poeciloterpa + Trigonoschema )). Jacobsoniella is recovered basally as sister of the other ingroup taxa and all together the ingroup clade is supported with a MLBv of 100 / 100 % branch support. Several key aspects of both topologies are similar. Regardless of the relationships of Eoscarta, Mioscarta, Poeciloterpa, Trigonoschema, Jacobsoniella and Wawi in both topologies the monophyly of each genus is well supported with a PPv of 100 % and MLBv of 98.40 / 91 %, 97.3 / 96 %, 99.7 / 99 % and 98.8 / 77 % respectively. In both trees the ingroups are well supported with a PPv of 100 % and MLBv of 100 / 100 %.	en	Crispolon, Elorde Jr. S., Guilbert, Eric, Yap, Sheryl A., Soulier-Perkins, Adeline (2021): New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines. European Journal of Taxonomy 778: 90-135, DOI: https://doi.org/10.5852/ejt.2021.778.1571
