taxonID	type	description	language	source
03BA4074F366FF9DFF1689BBB06BE723.taxon	diagnosis	Diagnosis. Occipital carina absent; hind wing media not diverging before Cu-a, not separated from Cu; fore wing with marginal cell elongate, longer than stigma, closed apically; two closed submarginal cells present; one recurrent vein and two discoidal cells present; stigma large; gaster without petiole; pronotal collar with complete transverse carina.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F366FF9DFF1689BBB06BE723.taxon	biology_ecology	Biology. Members of this genus make their nests in twigs, timber, decayed wood and in buildings (Bohart & Menke 1976; Turillazzi et al. 2014). They prey on thrips, psyllids, coccids and aphids (Blüthgen 1953; Krombein 1956; de Beaumont 1964).	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F366FF9DFF1689BBB06BE723.taxon	distribution	Distribution. In all zoogeographical realms (Pulawski 2021).	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	description	(Figs 1 – 8)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	materials_examined	Type material: Holotype, ♂, India: Kerala, Kozhikode dt., Kinalur (11 ° 27 ’ 38 ’’ N & 75 ° 49 ’ 43 ’’ E, Alt. 42 m), 17. vii. 2020, Coll. C. Binoy, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17150.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	diagnosis	Diagnosis. Spilomena attenboroughi sp. nov. differs from its congeners by the following combination of characters: metasoma yellowish brown; clypeus pale ivory;.; relative length to breadth of scape = 28: 8; relative length to breadth of stigma = 45: 16; anterior carina of pronotal collar conspicuous, deeply emarginate; recurrent vein of fore wing nearly interstitial with first transverse cubital vein; propodeal dorsum transversely striate, with two longitudinal carinae medially, posterior surface with irregular transverse ridges, lateral surface finely aciculate.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	description	Description. Holotype ♂. Body length 3.5 mm; fore wing length 2.3 mm. Colour. Head and mesosoma largely black; metasoma yellowish brown; clypeus and pronotal lobe pale ivory; mandible (except apical tip brown), antenna, tegula and legs yellow, fore wing veins and stigma brown. Vestiture. T 3 – T 7 with setigerous punctures; remaining body with sparse vestiture. Head. Clypeus shiny, without appressed setae, markedly convex (Fig. 2); interantennal tubercle inconspicuous, not distinctly pointed, extending into median carina up to mid frons; mandible bidentate, inner mandibular tooth short and blunt; relative length to breadth of scape = 30: 7; lower frons imbricate, upper frons and vertex reticulate without setigerous micropunctures; inner eye margin slightly concave; gena imbricate with no micropunctures. Mesosoma. Pronotum reticulate, laterally with irregular rugae; anterior carina of pronotal collar conspicuous and deeply emarginate; mesoscutum reticulate with setigerous punctures; admedian line and parapsidal line lacking; scutellum reticulate with setigerous punctures, anterior groove of scutellum broad, crenulate; metanotum reticulate, irregularly transversely rugose laterally; mesopleuron reticulate with scattered setigerous punctures posteriorly (Fig. 4); metapleuron with irregular ridges; recurrent vein of fore wing nearly interstitial with first transverse cubital vein; stigma elongate, relative length to breadth = 45: 16; propodeal dorsum transversely striate, with two longitudinal carinae medially (Fig. 5), posterior surface with irregular transverse ridges, lateral surface finely aciculate. Metasoma. Integument shiny; T 3 – T 7 with scattered setigerous punctures on entire surface. Female. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	etymology	Etymology. The specific epithet is a commemorative, genitive noun from the patronym Attenborough after the naturalist and broadcaster, Sir David Frederick Attenborough who has made the world’s natural history accessible and understandable to millions with his outstanding documentaries.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F365FF9FFF16894EB113E744.taxon	distribution	Distribution. India: Kerala.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	description	(Figs 9 – 16)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	materials_examined	Type material: Holotype, ♂, India: Kerala, Kozhikode dt., Elathur (11 ° 20 ’ 30.48 ’’ N & 75 ° 44 ’ 24.36 ’’ E, Alt. 23 m), 21. ii. 2021, Coll. C. Binoy, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17152.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	diagnosis	Diagnosis. Spilomena fulvopleuris sp. nov. differs from its congeners by the following combination of characters: mesoscutum laterally and mesopleuron ferruginous; relative length to breadth of scape = 20: 3; pronotum coriaceous with irregular rugae; propodeum entirely transversely striated without median carina or smooth area.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	description	Description. Holotype ♀. Body length 4.6 mm; fore wing length 3.3 mm. Colour. Head and mesosoma largely black; apical margin of clypeus, antennae, femora, tibiae and tarsi pale yellow; mandible (except apical tip brown), tegula, pronotal lobe, all coxae and trochanters whitish; mesoscutum laterally and mesopleuron ferruginous; fore wing veins, stigma and metasoma brown. Vestiture. T 2 – T 6 with setigerous micropunctures posteriorly; ventral margin of T 6 with apical suberect setae; remaining body with sparse vestiture. Head. Clypeus dull, without any appressed setae, markedly convex (Fig. 10); interantennal tubercle inconspicuous, not distinctly pointed, extending into median carina up to mid frons; mandible bidentate apically, inner mandibular tooth short, relative length to breadth of scape = 20: 3; lower frons imbricate, upper frons and vertex reticulate with setigerous punctures; inner eye margin straight; gena imbricate, without any punctures. Mesosoma. Pronotum coriaceous with irregular rugae, lateral surface with longitudinal rugae; anterior carina of pronotal collar conspicuous and straight; mesoscutum reticulate, with dense micropunctures; admedian line and parapsidal line lacking; scutellum reticulate; anterior groove of scutellum crenulate; metanotum reticulate, longitudinal rugose laterally; mesopleuron reticulate, with scattered setigerous punctures posteriorly (Fig. 12); metapleuron with irregular ridges; stigma elongate, relative length to breadth = 50: 14; recurrent vein of fore wing interstitial with first transverse cubital vein; propodeum entirely transversely striate, without median carina or smooth area (Fig. 13). Metasoma. T 1 & T 2 shiny and impunctate; T 3 – T 6 with scattered setigerous punctures on entire surface; dense brush of short setae at apex of T 6. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	etymology	Etymology. The species epithet is after the yellowish brown colouration on the mesopleural region.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F364FF98FF168CE7B113E231.taxon	distribution	Distribution. India: Kerala.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F362FF99FF1688F6B761E142.taxon	description	(Figs 17 – 20)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F362FF99FF1688F6B761E142.taxon	diagnosis	Diagnosis. Female. Inner mandibular tooth broad and blunt; clypeus conspicuously convex medially; propodeum rugose. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F362FF99FF1688F6B761E142.taxon	distribution	Distribution. India: Maharashtra (Turner 1918) Note. No additional materials were available during the present study. The above diagnosis is based on the original description (Turner 1918) and digital images of the type obtained from BMNH.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF1688F6B1DDE16B.taxon	diagnosis	Diagnosis. Female (Rajan et al. 2018, figs. 1 – 10). Inner mandibular tooth pointed; clypeus weakly convex medially, anterior margin slightly incised medially; propodeum areolate with network of coarse ridges, without two longitudinal carinae near middle, with smooth area at apex; lateral face of propodeum with some strong transverse striations anteriorly and large smooth area medially. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF1688F6B1DDE16B.taxon	distribution	Distribution. India: Kerala (Rajan et al. 2018).	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	description	(Figs 21 – 28)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	materials_examined	Type material: Holotype, ♀, India: Kerala, Kozhikode dt., Elathur (11 ° 20 ’ 30.48 ’’ N & 75 ° 44 ’ 24.36 ’’ E, Alt. 23 m), 29. iii. 2021, Coll. C. Binoy, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17291.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	diagnosis	Diagnosis. Spilomena reticularis sp. nov. differs from its congeners by the following combination of characters: mesopleuron, mesoscutum and gena finely reticulate; relative length to breadth of scape = 14: 3; relative length to breadth of stigma = 22: 9; propodeal dorsum rugose, with transverse ridges laterally; propodeal enclosure with delimiting furrow; posterior surface with irregular ridges and median longitudinal carina.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	description	Description. Holotype ♀. Body length 2.9 mm; fore wing length 1.8 mm. Colour. Head and mesosoma largely black; metasoma yellowish brown; mandible (except apical tip brown), antenna, tegula and legs yellow, fore wing veins and stigma brown. Vestiture. Scattered setigerous micropunctures on mesosoma; T 5 – T 7 with setigerous punctures; remainder of body with sparse vestiture. Head. Clypeus dull, without appressed setae, slightly convex (Fig. 22); interantennal tubercle inconspicuous, not distinctly pointed, extending into short median carina at most reaching lower frons; mandible bidentate, inner mandibular tooth short and blunt; relative length to breadth of scape = 14: 3; lower frons imbricate; upper frons and vertex reticulate, without setigerous micropunctures; inner eye margin straight; gena reticulate without punctures. Mesosoma. Pronotum coriaceous, laterally with longitudinal rugae; anterior carina of pronotal collar conspicuous and slightly emarginate; mesoscutum reticulate without scattered setigerous punctures; admedian line and parapsidal line lacking; scutellum reticulate, without setigerous punctures, anterior groove of scutellum broader and crenulate; metanotum reticulate, irregularly transversely rugose laterally; mesopleuron finely reticulate, with several well defined punctures (Fig. 24); metapleuron finely aciculate; stigma elongate, relative length to breadth = 22: 9; recurrent vein of fore wing received well within first submarginal cell; propodeal dorsum rugose, with transverse ridges laterally; propodeal enclosure with delimiting furrow (Fig. 25); posterior surface with irregular ridges and median longitudinal carina. Metasoma. Integument shiny; T 3 – T 7 with scattered setigerous punctures on entire surface. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	etymology	Etymology. The species epithet is after the finely reticulate mesopleuron.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF9AFF168A0AB104E590.taxon	distribution	Distribution. India: Kerala.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	description	(Figs 29 – 36)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	materials_examined	Type material: Holotype, ♀, India: Kerala, Thiruvananthapuram dt., Neyyar Wildlife Sanctuary (8 ° 31 ’ 50 ’’ N & 77 ° 09 ’ 08 ’’ E, Alt. 93 m), 17. i. 2019, Coll. P. Girish Kumar, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17149.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	diagnosis	Diagnosis. Spilomena sahyadriensis sp. nov. differs from its congeners by the following combination of characters: lower frons imbricate; upper frons and vertex reticulate with scattered setigerous micropunctures; relative length to breadth of scape = 38: 6; relative length to breadth of stigma = 34: 15; mesoscutum reticulate with dense micropunctures; propodeal dorsum and posterior surface of propodeum rugose and rugulose, lateral surface with transverse ridges.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	description	Description. Holotype ♀. Body length 4.6 mm; fore wing length 2.6 mm. Colour. Body black with following parts yellow: mandible (except apical tip brown), labrum, palpi, antenna and legs; pronotal lobe and tegula white; fore wing veins and stigma light brown. Vestiture. Setigerous micropunctures present on frons, vertex, mesoscutum and scutellum; dense brush of short setae at apex of T 6; remainder of body with sparse vestiture. Head. Clypeus shiny, without appressed setae, markedly convex (Fig. 30); interantennal tubercle inconspicuous, not distinctly pointed, extending into short median carina at most reaching lower frons; mandible bidentate apically, inner mandibular tooth short and blunt; relative length to breadth of scape = 38: 6; lower frons imbricate; upper frons and vertex reticulate with scattered setigerous micropunctures; inner eye margin straight; gena imbricate, impunctate. Mesosoma. Pronotum laterally with longitudinal ridges; anterior carina of pronotal collar conspicuous and straight; mesoscutum reticulate, with dense micropunctures; admedian line and parapsidal line lacking; scutellum reticulate, with dense micropunctures, anterior groove of scutellum without crenulations; metanotum reticulate, without micropunctures, longitudinal rugose laterally; mesopleuron reticulate, with scattered setigerous micropunctures posteriorly (Fig. 32); metapleuron with irregular rugae; stigma elongate, relative length to breadth = 34: 15; recurrent vein of fore wing received well within first submarginal cell; propodeal dorsum and posterior surface of propodeum rugose and rugulose (Fig. 33), lateral surface with transverse ridges. Metasoma. T 1 and T 2 shiny and impunctate; T 3 – T 6 with scattered setigerous punctures on entire surface; T 6 apically with dense brush of short setae. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	etymology	Etymology. The species name is derived from “ Sahyadri ”, the vernacular name for the Western Ghat mountain ranges.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F361FF94FF168F52B104E0B9.taxon	distribution	Distribution. India: Kerala.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	description	(Figs 37 – 44)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	materials_examined	Type material: Holotype, ♂, India: Kerala, Kozhikode dt., Manipuram (11 ° 22 ’ 48 ’’ N & 75 ° 56 ’ 40 ’’ E, Alt. 99 m), 4 – 26. iv. 2019, Coll. C. Binoy, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17151.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	diagnosis	Diagnosis. Spilomena tsunekii sp. nov. differs from its congeners by the following combination of characters: frons and vertex imbricate, without setigerous micropunctures; mesopleuron finely aciculate, with illdefined punctures; relative length to breadth of scape = 20: 5; stigma not elongate, relative length to breadth = 98: 68; propodeal dorsum transversely striate, with two longitudinal carinae medially, posterior surface finely aciculate, with median longitudinal carina, lateral surface transversely striate.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	description	Description. Holotype ♂. Body length 2.6 mm; fore wing length 1.1 mm. Colour. Body black except pronotum yellowish brown; clypeus and metasoma brown; mandible (except apical tip brown), tegula, pronotal lobe, legs and antenna yellow; fore wing veins and stigma brown. Vestiture. T 5 – T 7 with scattered setigerous punctures; remaining body with sparse vestiture. Head. Clypeus dull without any appressed setae, markedly convex (Fig. 38); interantennal tubercle inconspicuous, not distinctly pointed, extending into short median carina at most reaching lower frons; mandible bidentate, inner mandibular tooth short, blunt; relative length to breadth of scape = 20: 5 frons and vertex imbricate without setigerous micropunctures; inner eye margin straight; gena imbricate, impunctate. Mesosoma. Pronotum reticulate, with irregular longitudinal rugae; anterior carina of pronotal collar conspicuous, deeply emarginate; mesoscutum reticulate, with scattered setigerous punctures; admedian line and parapsidal line lacking; scutellum dull and reticulate, with sparse setigerous punctures, anterior groove of scutellum not broadened by crenulation; metanotum dull and reticulate, irregularly transversely rugose laterally; mesopleuron finely aciculate, with ill-defined punctures (Fig. 40); metapleuron finely aciculate; stigma not elongate, relative length to breadth = 98: 68; recurrent vein of fore wing received well within first submarginal cell; propodeal dorsum transversely striate, with two longitudinal carinae medially, posterior surface finely aciculate, with median longitudinal carina (Fig. 41), lateral surface transversely striate. Metasoma. Integument shiny; T 4 – T 7 with sparse setigerous micropunctures. Female. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	etymology	Etymology. The specific epithet is a commemorative, genitive noun taken from patronym Tsuneki after late Dr. Katsuji Tsuneki, in honor of his immense contribution to aculeate hymenopteran taxonomy.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36FFF95FF168C7BB113E4B4.taxon	distribution	Distribution. India: Kerala.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	description	(Figs 45 – 52)	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	materials_examined	Type material: Holotype, ♀, India: Karnataka, Kodagu dt., Perumbadi (12 ° 08 ’ 31 ’’ N & 75 ° 47 ’ 44 ’’ E, Alt. 877 m), 03. v. 2019, Coll. P. Girish Kumar, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17141. Paratypes, all col lected by P. Girish Kumar 1 ♀, Karnataka, Kodagu dt., Perumpadi, 03. v. 2019, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17144; 1 ♀, Karnataka, Kodagu dt., Bettoli, 07. iii. 2016, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17145; Karnataka, Kodagu dt., Bettoli, Ayyappa temple, 05. v. 2019, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17146; 1 ♀, Karnataka, Kodagu dt., Madikeri, Kohinoor, 05. v. 2019, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17147; 1 ♀, Karnataka, Kodagu dt., Makutta Forest, Heggala, 08. iv. 2016, ZSIK Regd. No. ZSI / WGRC / IR / INV / 17148.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	diagnosis	Diagnosis. Spilomena tuberculata sp. nov. differs from its congeners by the following combination of characters: mesosoma brownish yellow with few black patches; relative length to breadth of scape = 23: 6; relative length to breadth of stigma = 56: 17; propodeal dorsum rugose, postereo-lateral surface with transverse ridges, posteriorly with median longitudinal carina.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	description	Description. Holotype ♀. Body length 5.6 mm; fore wing length 3.6 mm. Colour. Head black; mandible (except apical tip brown), clypeus, antenna, pronotal lobe, tegula and legs pale yellow; mesoscutum, scutellum, metanotum and metasoma yellowish brown; propodeum brown with yellow patches laterally; mesopleuron yellow; metapleuron brown; fore wing veins and stigma light brown. Vestiture. Scattered sparse setae on clypeus and mandible; T 3 – T 6 each with scattered setae; ventral margin of T 6 with apical suberect setae; remaining body with sparse vestiture. Head. Clypeus shiny, with scattered setae, markedly convex (Fig. 46); interantennal tubercle prominent, acutely pointed and extending into median carina up to mid frons; mandible bidentate apically, inner mandibular tooth short and blunt; relative length to breadth of scape = 23: 6; frons and vertex reticulate with dense setigerous micropunctures; inner eye margin straight; gena imbricate, impunctate. Mesosoma. Pronotum dull, with longitudinal rugae; anterior carina of pronotal collar conspicuous and straight; mesoscutum reticulate with dense micropunctures; admedian line and parapsidal line lacking; scutellum reticulate with dense micropunctures, anterior groove of scutellum not broadened by crenulation; metanotum reticulate, longitudinally rugose laterally; mesopleuron reticulate with scattered setigerous punctures posteriorly (Fig. 48); metapleuron with irregular ridges; stigma elongate, relative length to breadth = 56: 17; recurrent vein of fore wing received well within first submarginal cell; propodeal dorsum rugose (Fig. 49), lateral surface finely aciculate, with scattered transverse ridges, posterior surface with irregular transverse ridges and longitudinal carina medially. Metasoma. T 1 and T 2 shiny, impunctate; T 3 – T 6 with scattered setigerous punctures on entire surface; T 6 apically with dense brush of short setae. Male. Unknown.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	etymology	Etymology. The species name is derived from a distinctively conspicuous interantennal tubercle.	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
03BA4074F36DFF90FF168F53B15AE744.taxon	distribution	Distribution. India (Karnataka).	en	Rajan, Tessy, Kumar, P. Girish, Sureshan, P. M., Binoy, C. (2021): A review of the genus Spilomena Shuckard (Hymenoptera: Crabronidae) with the description of six new species from India. Zootaxa 5068 (2): 263-276, DOI: https://doi.org/10.11646/zootaxa.5068.2.6
