taxonID	type	description	language	source
03AE7920FF98FFFE5038FE3802E14A05.taxon	description	(Figs 1 - 6)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFFE5038FE3802E14A05.taxon	materials_examined	Type material. Holotype ♂, PAPUA N. G. Morobe Prov., S Aseki, 1750 m 14. IV. 1998, sifted A. Riedel (MHNG). Paratype ♀, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFFE5038FE3802E14A05.taxon	description	Description. Length 2.03 mm, width 1.27 mm. Head, thorax and elytra black, abdomen dark reddish­brown, femora and tibiae ochraceous, tarsi as tibiae or somewhat lighter, antennae light brown. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps, sulci and impressions; adsutural areas raised; sutural striae finely punctate, lateral striae impunctate; basal striae throughout deep, entire, not interrupted and joined to lateral striae. Most discal punctures about as pronotal punctures, disc with irregularly scattered, somewhat larger, and very shallow punctures. Mesoventrite convex, smooth, appearing impunctate, without impressions, with long mesal ridge extended from paxillum to posterior margin of mesocoxal process. Metaventrite not microsculptured, with elongate impression in centre, several hardly visible oblique striae converging anteriad, toward mesal impression, few fine punctures along basal margin and in middle part of mesocoxal process; submesocoxal lines margined by several coarse punctures; centre and lateral parts of metaventrite impunctate; anterior margin of mesocoxal process below level of margin of mesoventrite. Metacoxal process oblique, with margin weakly concave, notched at lateral margins. Metanepisterna in anterior halves about 0.05 mm wide. Metanepisternal suture impunctate, straight, nearly reaching margin of metepimeron. Ventrites with punctulate microsculpture absent from lateral parts of ventrite I, appearing impunctate, coarse punctures along basal margin of ventrite I excepted. Ventrite Iwith low basal hump. Protarsomeres III lacking prominent ventral lobe. ♂: Protarsomeres Ito III slightly widened, similar in width, each at apex about as wide as half of protibial apex. Protarsomere Iabout as long protarsomeres II and III combined. Protarsomere III with apicoventral pair of long, narrow setae. Aedeagus as Figs 1 - 5, 1.27 mm long. ♀: Gonocoxite as Fig. 6, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFFE5038FE3802E14A05.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFFE5038FE3802E14A05.taxon	etymology	Etymology. The species is dedicated to Luigi Maria D'Albertis (1841 - 1901), one of the early explorers of the fauna of New Guinea and Moluccas.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFFE5038FE3802E14A05.taxon	discussion	Comments. This species is similar to B. biroi in most external characters. It may be distinguished by the uninterrupted basal striae of the elytra, the metaventrite lacking coarse punctures and the mesoventrite lacking distinct striae. The shape of the parameres is unique in this species, notably by the presence of a small lobe posterior of expanded middle section.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFF95038F9D300974E1D.taxon	materials_examined	Material examined. Additional specimens were not found in the examined collections.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFF95038F9D300974E1D.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF98FFF95038F9D300974E1D.taxon	discussion	Comments. Both, B. basicolle (Pic) and B. longipes Csiki were based on specimens collected at Sattelberg, Huon Peninsula, Morobe Province. This is one of the species possessing entire basal striae joined to the lateral striae.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	description	(Figs 7  10)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	materials_examined	Type material. Holotype ♂, NEW GUINEA centr. BALIEM TAL­ 1700 m MARCH 1992 leg. Jiłí KOLIB` ¨ (NHMB). Paratypes 2 ♂, INDONESIA, Papua, Jayawijaya Prov., Nalca, 1900 - 2100 m, 8. IX. 1992, A. Riedel (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	description	Description. Length 2.23  2.40 mm, width 1.34 - 1.47 mm. Head, pronotum and legs ochraceous, elytra black, hypomera and venter of thorax reddish­brown, abdomen yellowish, legs and antennae about as light as pronotum. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps, sulci and impressions; adsutural areas slightly raised; sutural and lateral striae impunctate; basal striae shortened, broadly interrupted laterally of elytral mid­width. Elytral punctation very sparse and very fine, similar to pronotal punctation. Mesoventrite convex, smooth, appearing impunctate, with short and inconspicuous mesal ridge posterior of paxillum, without impressions. Metaventrite impressed in middle, with several hardly visible oblique striae converging toward mesal impression, not microsculptured and entirely impunctate, punctures absent from submesocoxal lines; anterior margin of mesocoxal process nearly in level with margin of mesoventrite. Metacoxal process horizontal, slightly impressed, margin weakly rounded. Metanepisterna narrow, in anterior halves about 0.02 - 0.03 mm wide. Metanepisternal suture impunctate, curved along metepimeron and nearly reaching margin of metepimeron. Ventrites with punctulate microsculpture, appearing impunctate, few distinct punctures along basal margin excepted. Ventrite Iwithout distinct basal hump. Protarsomeres III lacking prominent ventral lobe. ♂: Protarsomeres Ito III slightly widened, similar in width, each about as wide as half of protibial apex. Protarsomere Islightly shorter than protarsomeres II and III combined. Protarsomere III with apicoventral pair of long setae gradually widened, reaching about level of mid­length of protarsomere V. Aedeagus as Figs 7 - 10, 0.98 - 1.10 mm long.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	etymology	Etymology. The species epithet is a Latin adjective referring to bicolorous pattern of the elytra and pronotum.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9FFFF950DCFDD407CF4AD3.taxon	discussion	Comments. This species may be easily distinguished from its congeners by the colour pattern, teneral specimens, as the two paratypes, excepted. The species is characterized by the aedeagal characters, and in external characters by the mesally impressed metaventrite in combination with the impunctate submesocoxal lines and the smooth mesoventrite bearing a very short mesal ridge. The parameres are similar to those of B. borneense Löbl. The latter species differs conspicuously by the coarse elytral punctures forming elongate rows. The " Tal " on the label of the holotype means valley in German.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFE38071048F0.taxon	description	(Figs 11  14)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFE38071048F0.taxon	materials_examined	Material examined. PAPUA NEW GUINEA: 9, Morobe Prov., Tekadu­Kakaro, Ivinka Riv. Stat., 170 m, 3. III. 1998, sieved, A. Riedel (MHNG); 1, Morobe Prov., Wau, 1150 m, 17. V. 1992, G. Cuccodoro [# 2 C] (MHNG); 1, same data but 19. V. 1992 [# 4 A] (MHNG); 1, Morobe Prov., Wau, Kilolo Creek, 26. VIII. 1968, I. Loksa (HNMB); 1, Morobe Prov., Biaru Rd., Kaisenic, 1050 m, 28. V. 1992, G. Cuccodoro [# 11 C] (MHNG); 1, Morobe Prov., Aiewa near Poiu, S Aseki, 1750 m, 14. IV. 1998, A. Riedel (MHNG); 20, Sandaun Prov., S Mianmin, 1000 m, 19. V. 1998, sieved, A. Riedel (MHNG); 1, same data but 900 - 1200 m (MHNG); INDONESIA: 1, Papua, Merauke Prov., Asmat, Senggo, trail to Abai, ca 100 m, 15 - 17. VI. 1994, A. Riedel (MHNG); 1, Papua, Nabire Prov., rd Nabire­Ilaga km 54, 03 ° 29 ′ 517 ′′ S, 135 ° 43 ′ 913 ′′ E, 750 m, VI. 1998, M. Balke (NMEC); 2, same data but X. 1997, 250 m and 750 m respectively (NMEC); Papua, Paniai Prov., Nabire, Pusppensaat, km 54, 500 - 700 m, 13 - 16. VIII. 1991, A. Riedel (MHNG); 4, West Papua, Manokwari Prov., Testega, 1200 m, 13. IV. 1993; 3, Moluccas, Aru Isl., Wokan, 22.8.1991, D. Agosti [F 91036, leaf litter] (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFE38071048F0.taxon	distribution	Distribution. Papua New Guinea; Indonesia: Papua, West Papua, Moluccas.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFE38071048F0.taxon	discussion	Comments. This species was based on two specimens from Sattelberg, on the Huon Peninsula, Morobe Province. It differs drastically from its congeners by the parameres split to form two long lobes. The aedeagus illustrated in LÖBL (1975) has the internal sac extruded and the figures given in that paper lack detail. Therefore, new illustrations (Figs 11 - 13) are given here. The followings characters are newly given: elytra with basal striae entire, joined to lateral striae; lateral striae fairly coarsely punctate; male protarsomere I about as wide as half of protibial apex, somewhat longer than protarsomeres II and III combined; protarsomeres III slightly lobed, with long apicoventral setae not widened. Gonocoxite as Fig. 14, without gonostyle. The specimens from the island Wokan are only 1.88  1.97 mm long, smaller than specimens from New Guinea, and have the median part of the metaventrite deeper impressed and more coarsely punctate. The shape of the narrow basal sclerite of the internal sac varies from circular to oval.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFAAA07014A37.taxon	materials_examined	Material examined. Additional specimens not found in the new collections.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFAAA07014A37.taxon	distribution	Distribution. Indonesia: Moluccas, Morotai Island.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFB50DCFAAA07014A37.taxon	discussion	Comments. The followings characters are here newly given: male protarsomere Iabout as wide as half of protibial apex, shorter than protarsomeres II and III combined; protarsomeres III slightly lobed, with long ventral setae not widened.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFA50DCF9EB07544E2D.taxon	materials_examined	Material examined. Additional specimens not found in the new collections.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFA50DCF9EB07544E2D.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9DFFFA50DCF9EB07544E2D.taxon	discussion	Comments. The species was based on two specimens from Popondetta and Kokoda, both in the Oro Province. The followings characters are here newly given: male protarsomere Inearly as wide as half of protibial apex, slightly shorter than protarsomeres II and III combined; protarsomeres III not lobed, with long ventral setae not widened.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	description	(Figs 15  19)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	materials_examined	Type material. Holotype ♂, PAPUA N. G. Morobe Prov. leg. A. Riedel / Aseki, Oiwa, 1600 - 1700 m, 11 - 12. III. 1998 (MHNG). Paratype ♀, Papua New Guinea, Eastern Highlands, Kassem Pass, XI. 1957, J. Sedlacek (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	description	Description. Length 3.0 - 3.20 mm, width 1.77 - 1.92 mm. Head and body black, with apex of abdomen lighter. Femora and tibiae black, tarsi blackish­brown. Antennae blackish. Pronotum very finely punctate. Scutellum rounded. Elytra with distinct humeral humps; adsutural areas raised; sutural striae with scattered, fine punctures; basal striae shortened, broadly interrupted laterally of elytral mid­width, lateral striae finely punctate in anterior halves. Elytra lacking sulci and impressions, punctation near sutural and lateral striae, and at bases very fine, similar to pronotal punctation, on apical fourth somewhat less fine, on prevailing discal surface rather coarse and irregular. Mesoventrite hardly convex, with robust mesal ridge nearly reaching margin of mesocoxal process, and with oblique striae. Metaventrite impunctate, not microsculptured; impressed mesally; mesocoxal process below level of mesoventrite, longitudinally striate. Metacoxal process slightly oblique, with transverse margin. Metanepisterna narrow, near anterior margins about 0.04 mm wide, weakly widened apically. Metanepisternal suture impunctate, straight and nearly reaching margin of mesepimeron. Abdomen very finely punctate, except for to distinct punctures along basal margin of ventrite I. Mesal area of ventrite I lacking basal bump, with strigulate microsculpture becoming punctulate near apical margin, lateral parts of ventrite Inot microsculptured. Ventrites II to IV and base of ventrite Vwith strigulate microsculpture, prevailing surface of ventrite Vwith punctulate microsculpture. Protarsomeres III lacking prominent ventral lobe. ♂: Protarsomeres Ito III moderately widened, similar in width, each about as wide as two thirds of protibial apex. Protarsomere III longer than following three tarsomeres combined, with apicoventral pair of long setae not widened. Aedeagus as Figs 15 - 18, 1.50 mm long. ♀: Gonocoxite with gonostyle, as Fig. 19.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning large.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF9CFFF45038FD8E02BF4DCD.taxon	discussion	Comments. The species may be distinguished from its Papuan congeners by the bodysize. Only four Asian species, B. feai (Achard, 1914), B. grouvellei (Achard, 1920), B. tonkineum (Pic, 1922) and B. yunnanum Löbl, 2020, have the body 3 mm long, or more than 3 mm long, in combination with the entirely black pronotum and elytra. This new species differs also conspicuously from the Papuan congeners by the punctate ventrites and the ventrite Ihaving microsculpture to part consisting of striae. The narrow notch situated posterior of the expanded apical section of the parameres is a unique character, and the gonostyle separates this species from the remaining Papuan Bironium known in females.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF45038FDAE028B4FAC.taxon	materials_examined	Material examined. Additional specimens not found in the new collections.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF45038FDAE028B4FAC.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF45038FDAE028B4FAC.taxon	discussion	Comments. The species was based on four specimens from Wau, Morobe Prov., Papua New Guinea. The followings characters are here newly given: elytra with basal striae entire, joined to lateral striae; male protarsomere Iabout as wide as half of protibial apex, nearly as long as protarsomeres II and III combined; protarsomeres III lacking prominent ventral lobes, with long ventral setae slightly widened apically. The latter characters state is diagnostic.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF75038FC0F00B94E4D.taxon	description	(Fig. 20)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF75038FC0F00B94E4D.taxon	materials_examined	Type material. Lectotype (by inference) ♀, Mysol [handwritten] / Scaphicoma Mysol longipes m [handwritten] / longipes Reitt [handwritten] / TYP REITTER [printed] / Lectotypus [printed, red] / Bironium longipes Reitt. [handwritten] det. Löbl 1991 [printed] (NMPC). Material examined. INDONESIA: 2, Misool (NMPC, MHNG); 2, West Papua, Sorong Prov., Salawatti Island, Wayom, 0 - 610 m, 8. XI. 1996, A. Riedel (MHNG); 2, West Papua, 130 km SE Kalmana, Omba (= Yamor) river, 10 - 20 km from coast, S 4 ° 05 ′ 49 ′′, E 134 ° 54 ′ 09 ′′, 10 - 20 m, 9 - 11. II. 2011, A. Skale (NMEC); 1, West Papua, Fakfak prov., ca 20 km W Timika, SP 7, 30 m, 8 - 9. I. 1996, A. Riedel (MHNG); 5, West Papua, Manokwari Prov., Gn. Meja, 200 m, 21 - 24. VIII. 1991, A. Riedel (MHNG); 1, Papua, Jayapura, Sentani, Cyclops Mts, 300 - 1400 m, 10. VIII. 1991, A. Riedel (MHNG); 4, Papua, Jayawijaya Prov., Wamena, Angguruk­Tanggeam, 1500 - 1800 m, 28 - 29. IX. 1991, A. Riedel (MHNG); 2, Jayawijaya Prov., Emdoman, 900 - 1200 m, 29. IX. 1993, A. Riedel (MHNG); 1, Papua, Jayawijaya Prov., Samboka, upper Kollf river, ca 200 m, 10 - 14. X. 1996, A. Riedel (MHNG); PAPUA NEW GUINEA: 4, Sandaun Prov., S Mianmin, 900 - 1200 m, 19. V. 1998, A. Riedel (MHNG); 2, Sandaun Prov., N Mianmin, 700 - 1100 m, 20. V. 1998, A. Riedel (MHNG); 1, Madang Prov., Ohu Village near Gum River, S 5 ° 13 ′, E 145 ° 41 ′, 150 m, I-II. 2001, L. ¨ i-ek (NHMB).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF75038FC0F00B94E4D.taxon	distribution	Distribution. Indonesia: Misool, West Papua and Papua; Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF92FFF75038FC0F00B94E4D.taxon	discussion	Comments. REITTER (1880) based this species on an unknown number of specimens from - Mysol- (= Misool). The E. Reitter  s collection of exotic scaphidiines was puchased by R. Oberthür (HORN et al. 1990), and the bulk of the R. Oberthür collection is housed in the MHNN. Thus, Ihave assumed that also the type material of B. longipes is in the Paris collection (LÖBL 2018), although not found there while my visit. ACHARD (1924) stated to have received from R. Oberthür the type ( le type ) and transferred the species to Heteroscapha Achard, 1914 (a junior synonym of Bironium; see LÖBL 1971). As the number of original specimens is unknown, ACHARD (1924) designated the lectotype by inference, and this specimen is in the NMPC. The present concept of B. longipes (Reitter) is based on this specimen examined previously and redescribed in LÖBL (1989), though re­examined for the present study. It complies with the original Reitter  s description. The followings characters are here newly given: Elytra with basal striae interrupted at level of humeral areas; male tarsomere Iabout as wide as half of protibial apex, shorter than tarsomeres II and III combined; tarsomeres III not lobed, with long, not flattened ventral setae; gonocoxite without gonostyle (Fig. 20).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF750DCFD2E01214860.taxon	description	(Fig. 21)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF750DCFD2E01214860.taxon	materials_examined	Material examined. PAPUA NEW GUINEA: 2, Morobe Distr., Lea, H. Franz (NHMW, MHNG); 1, Morobe Distr., Wau, 1150 m, 19. V. 1992, G. Cuccodoro (MHNG); 1, Morobe Distr., above Wau, 1400 m, 16. V. 1992, G. Cuccodoro (MHNG); 2, Morobe Distr., Mt. Kaindi, 1350 m, 24. V. 1992, G. Cuccodoro (MHNG); 1, Morobe Distr., Lakekamu Basin, Tekadu, 200 - 400 m, 28. II. - 1. III. 1998, A. Riedel (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF750DCFD2E01214860.taxon	distribution	Distribution. Papua New Guinea.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF750DCFD2E01214860.taxon	discussion	Comments. This species was based on two specimens collected in 1957 at Popondetta and Kokoda (Oro Province), respectively. The followings characters are here added: ventrite Iwith small basomedian hump, male protarsomere Iabout as wide as three fourth of protibial apex and about as long as protarsomeres II and III combined; protarsomeres III weakly lobed, with long, not widened apicoventral setae; gonocoxite as Fig. 21, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	description	(Figs 22 - 25)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	materials_examined	Type material. Holotype ♂: WEST PAPUA: Foja Mountains mosses and leaves, forest stream, 200 m 2 ° 27 ′ 32.37 ′′ S 138 ° 46 ′ 30.19 ′′ E leg. Tobias Mainda 28.05. (MHNG). Paratype ♂, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	description	Description. Length 1.47 - 1.60 mm, width 0.83 - 0.90 mm. Head and body black, legs ochraceous, tibiae and tarsi lighter than femora, antennae light brown. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas flat; sutural and lateral striae impunctate; basal striae deep, shortly interrupted at level of humeral areas, discal punctation about as fine as pronotal punctation, hardly visible at 100 times magnification. Mesoventrite convex, appearing impunctate, not microsculptured, with short ridge present only on paxillum, few very shallow striae between mesocoxal cavities. Median part of metaventrite hardly convex, with anterior margin conspicuously impressed below margin of mesoventrite, lacking obvious longitudinal striae, not microsculptured, with pair of coarse punctures in centre; metacoxal process horizontal, with margin shallowly notched in middle, lateral notches deeper and narrower than median notch. Lateral parts of metaventrite lacking coarse punctures. Submesocoxal lines with a few coarse punctures. Metanepisterna evenly, about 0.03 mm wide. Metanepisternal suture impunctate, curved posteriad, nearly reaching margin of metepimeron. Ventrites lacking microsculpture. Ventrite Iwithout basomedian hump, with few rather fine punctures along basal margin. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III hardly widened, similar in width, protarsomere Islightly wider than half of protibial apex. Protarsomere III about as long as following two tarsomeres combined, with apicoventral pair of long setae of not widened. Aedeagus as Figs 22 - 25, 0.59 mm long.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	etymology	Etymology. The species is named in honour of its collector, Tobias Mainda (Nauen, Germany).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF91FFF650DCFB3A01724F7B.taxon	discussion	Comments. This species may be readily distinguished from the remaining species by its small body­size. It is also characterized by the reduced ridge of the mesoventrite in combination with the metaventrite lacking striae. The shape of the parameres, as seen in ventral view, with angulate inner margin posterior of narrowed basal section, is also diagnostic for the species.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	description	(Figs 26 - 30)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	materials_examined	Type material. Holotype ♂, INDO: Irian Jaya, Monokwari Distr. Maibri vill. Arfak Mts., 1570 m 6 - 19. xii. J. HorAEk leg. (MHNG). Paratype ♀, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	description	Description. Length 2.50 - 2.56 mm, width 1.40 - 1.50 mm. Head and thorax black. Elytra black, each with two ochraceous spots. Anterior spot transverse, expanded near sutural stria; interval between anterior spot and sutural stria somewhat shorter than interval to anterior elytral margin and about as large as interval to lateral stria. Abdomen blackish with dark brown apical segments. Antennae brown, legs blackish­brown, darker than antennae. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas raised; sutural striae impunctate, lateral striae with few punctures; basal striae entire, very shallow and hardly visible in level of humeral areas, discal punctation about as fine as pronotal punctation on prevailing elytral surface, distinctly coarser punctures scattered on middle third of disc. Mesoventrite conspicuously convex, appearing impunctate, with sharply delimited mesal ridge extended from paxillum to apical third of mesal length, lacking striae. Median part of metaventrite flat between mesocoxae, with anterior margin well below margin of mesoventrite, bearing few longitudinal striae, not microsculptured; apical stripe and centre impressed, flattened; metacoxal process horizontal, with bisinuate margin. Lateral parts of metaventrite lacking coarse punctures. Submesocoxal lines punctate. Metanepisterna evenly about 0.08 mm wide. Metanepisternal suture impunctate, slightly sinuate, not curved along metepimeron, nearly reaching margin of metepimeron. Ventrite Iwithout basomedian hump, with strigulate microsculpture, distinctly punctate along basal margin. Following ventrites with punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III distinctly widened, protarsomere Iabout as wide wo thirds of protibial apex, following two tarsomeres narrower, protarsomere Ias long as following two tarsomeres. Apicoventral pair of long setae of protarsomere III not widened apically. Aedeagus as Figs 26 - 29, 0.98 mm long. ♀: Gonocoxite as Fig. 30, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	distribution	Distribution. Indonesia: West Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning provided and referring to the presence of elytral spots.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF90FFF15038FC1501594EFB.taxon	discussion	Comments. This species may be easily distinguished from the congeners by the elytral spots, in combination with the pattern of elytral punctation. It is also characterized by the mesoventrite strongly convex and lacking striae, and the inner margin of the parameres notched posterior of bases.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	description	(Figs 31 - 36)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	materials_examined	Type material. Holotype ♂: IRIAN JAYA Prov. Jayawijaya, Samboka Upper Kollf River, ca 200 m, 10 - 14. X. 96, Riedel (MHNG). Paratypes 2 ♂, 2 ♀, with the same data as the holotype; 1 ♀, Papua, Jayawijaya Prov., Lereh, 300 - 550 m, 25. I. 1996, A. Riedel (MHNG); 1 ♀, Papua, Jayawijaya Prov., Sumo - Uarn, 100 - 1000 m, 26 - 27. VI. 1994, A. Riedel (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	description	Description. Length 1.95 - 2.15 mm, width 1.20 - 1.34 mm. Head, thorax and elytra black or blackish­brown, abdomen very dark brown, light brown to yellowish apical segments excepted. Legs reddish­brown, with tarsi lighter than tibiae, antennae light brown. Pronotum very finely punctate. Scutellum rounded. Elytra lacking humeral humps; adsutural areas hardly raised; sutural and lateral striae punctate; basal striae entire, very shallow and hardly visible at humeral areas. Elytral disc impunctate along base and lateral striae, with very fine, hardly visible punctation on apical third; prevailing discal surface coarsely punctate. Coarse punctures forming two longitudinal rows, outer row impressed, interval between rows somewhat raised. Mesoventrite weakly convex, with well delimited mesal ridge extended from paxillum nearly to apical margin and with longitudinal striae evanescent anteriorly; area near lateral margins of mesocoxal process densely punctate and uneven. Metaventrite not microsculptured. Median part of metaventrite with minute hump between mesocoxae, flat anterior of hump, impressed posterior of hump, with anterior margin below margin of mesoventrite, bearing few longitudinal striae at each side of mesocoxal process; hump and area posterior of hump irregularly punctate; apical stripe flat, not impressed; metacoxal process horizontal, with rounded margin. Lateral parts of metaventrite coarsely punctate, except near apical margin. Submesocoxal lines coarsely punctate. Metanepisterna narrow, about 0.03 mm wide. Metanepisternal suture impunctate, straight, not curved along metepimeron, reaching or nearly reaching margin of metepimeron. Ventrite Iwithout basomedian hump, with punctulate microsculpture, distinctly punctate along basal margin. Following ventrites with punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III distinctly widened, protarsomere Iabout as wide as two thirds of protibial apex, somewhat shorter than following two tarsomeres combined; protarsomeres II and III slightly narrower that protarsomere I. Apicoventral pair of long setae of protarsomere III not flattened apically. Aedeagus as Figs 31 - 35, 0.86 mm long. ♀: Gonocoxite as Fig. 36, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning punctate.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF97FFED50DCFC9D07144EFB.taxon	discussion	Comments. This species may be readily distinguished from its congeners having similar body­colour by the pattern of the elytral punctation combined with the coarsely punctate lateral parts of the metaventrite, and the presence of a hump situated between mesocoxae. The comparatively narrow parameres lacking lobes are similar to those in B. flavapex while the hook­like shape of the apex of the median lobe is unique.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	description	(Figs 37 - 41)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	materials_examined	Type material. Holotype ♂, Irian Jaya Prov. Jayawijaya, Borme 1500 - 2000 m, 12 - 15.8.1992, Riedel [24] (MHNG). Paratypes 2 ♂, 1 ♀, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	description	Description. Length 1.92 - 1.98 mm, width 1.17 mm. Head and body uniformly ochraceous, appendages slightly lighter. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas not or slightly raised; sutural striae with few fine punctures or impunctate, lateral striae impunctate; basal striae widely interrupted at level of humeral areas; discal punctation irregular, to part similar to pronotal punctation, very shallow and hardly visible at 100 times magnification. Mesoventrite convex, with well delimited mesal ridge extended from paxillum nearly to apical margin, impunctate, lacking microsculpture, with hardly visible longitudinal striae on mesocoxal process. Metaventrite not microsculptured, entirely very finely punctate. Median part of metaventrite flattened, with anterior margin below level of margin of mesoventrite, with hardly visible longitudinal striae; apical stripe flat, not impressed; metacoxal process inflexed, with margin notched near coxae. Submesocoxal lines with few coarse punctures. Metanepisterna narrow, about 0.04 - 0.05 mm wide. Metanepisternal suture impunctate, mostly straight, curved apically, reaching near to margin of metepimeron. Ventrite Iwithout basomedian hump, not microsculptured, coarsely punctate along basal margin. Following ventrites with punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III hardly widened, protarsomere Islightly wider than half of protibial apex, following two tarsomeres still somewhat narrower. Protarsomere I somewhat longer than following two tarsomeres combined. Apicoventral pair of long setae of protarsomere III not flattened apically. Aedeagus as Figs 37 - 40, 0.64 - 0.67 mm long. ♀: Gonocoxite as Fig. 41, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	etymology	Etymology. The species is dedicated to Alexander Riedel (Karlsruhe, Germany) who significantly contributed to the knowledge of New Guinean beetles and collected most of the scaphidiines dealt in the present paper.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8BFFEC50DCFCA701E14DCD.taxon	discussion	Comments. This species may be distinguished by the shape of the parameres of the sclerotized structures of the internal sac. In external characters it is similar to B. basicolle, B. loksai and B. glabrum (see the key).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	description	(Figs 42 - 45)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	materials_examined	Type material. Holotype ♂, INDO: West Papua Manokwari Prov. Testega, 1200 m 13. IV. 93, A. Riedel (MHNG). Paratypes 5 ♀, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	description	Description. Length 1.80 - 2.0 mm, width 1.02 - 1.80 mm. Head, thorax, elytra and legs uniformly rufous, abdomen lighter, with apical segments yellowish, antennae yellowish. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas hardly raised; sutural and lateral striae impunctate; basal striae interrupted at level of humeral areas; discal punctation as fine as pronotal punctation. Mesoventrite convex, lacking mesal ridge, impunctate, not microsculptured and not striate. Metaventrite not microsculptured, impunctate. Median part of metaventrite flattened between mesocoxae, with anterior margin below level of margin of mesoventrite, distinct longitudinal striae extended to mid­length; centre with large, shallow impression situated posterior of mid­length; apical stripe flat, not impressed; metacoxal process horizontal, margin notched in middle and near coxae. Submesocoxal lines with few coarse punctures. Metanepisterna to large part concealed, exposed posterior section about 0.01 - 0.02 mm wide. Metanepisternal suture impunctate, curved apically, ending well anterior margin of metepimeron. Ventrite Iwithout basomedian hump, not microsculptured, lacking puncture row along basal margin. Following ventrites with punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III hardly widened, protarsomere Iabout as wide as half of protibial apex, following two tarsomeres hardly narrower. Protarsomere Inearly as long as following two tarsomeres combined. Apicoventral pair of long setae of protarsomere III not flattened apically. Aedeagus as Figs 42 - 44, 0.70 mm long. ♀: Gonocoxite as Fig. 45, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	distribution	Distribution. Indonesia: West Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning rufous.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEC5038FDBC07084A41.taxon	discussion	Comments. The species may be distinguished from most Papuan congener by the mesoventrite lacking, or having a reduced mesal ridge, as it is in B. basicolle, B. bicolor, B. flavapex, and B. maindai. These four species have elytra with entire basal striae and wider exposed parts of the metanepisterna, and two of them, B. maindai and B. bicolor, differ drastically by their body size and colour pattern, respectively. The habitus and the aedegus of B. rufulum are similar to those of B. riedeli, suggesting relationships. However, the mesoventrite with a robust mesal ridge, and the subquadrate lobes and the broad apical section of the parameres of the latter species are diagnostic.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	description	(Figs 46 - 49)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	materials_examined	Type material. Holotype ♂, IRIAN JAYA Jayapura Prov., leg. A. Riedel / Lereh 300 - 550 m, 25. I. 1996 / from base of dead palm leaves (MHNG). Paratypes 6 ♀, with the same data as the holotype (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	description	Description. Length 2.20 - 2.35 mm, width 1.32 - 1.35 mm. Head, thorax and elytra dark reddish­brown to blackish, abdomen lighter than thorax, appendages ochraceous to yellowish. Pronotum very finely punctate. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas flat or slightly raised; sutural and lateral striae punctate; basal striae entire, joined to lateral striae; discal punctation sparse, irregular, to part similar to pronotal punctation, very shallow and hardly visible at 100 times magnification, most punctures much larger than pronotal punctures, well visible at 15 times magnification. Mesoventrite slightly convex, with well delimited mesal ridge extended from paxillum to apical margin, impunctate, lacking microsculpture, with conspicuous longitudinal to oblique striae on mesocoxal process. Metaventrite not microsculptured, entirely very finely punctate. Median part of metaventrite slightly convex, with anterior margin below level of margin of mesoventrite, short longitudinal striae between mesocoxae and oblique mesal striae, shallowly impressed posterior centre; apical stripe somewhat convex, not impressed; metacoxal process inflexed, with margin sinuate, notched near coxae. Submesocoxal lines margined by fairly fine punctures and elongate striae. Metanepisterna about 0.03 - 0.05 mm wide. Metanepisternal suture impunctate, straight, curved along margin of metepimeron. Ventrite Iwithout basomedian hump, with punctulate microsculpture, distinctly punctate along basal margin. Following ventrites with well visible punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III slightly widened, protarsomere Iabout as wide as half of protibial apex, following two tarsomeres hardly narrower. Protarsomere Iabout as long as following three tarsomeres combined. Apicoventral pair of long setae of protarsomere III not flattened apically. Aedeagus as Figs 46 - 48, 0.93 mm long. ♀: Gonocoxite as Fig. 48, without gonostyle.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning striate.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF8AFFEF5038F92D0786495B.taxon	discussion	Comments. This species may be distinguished from its Papuan congeners by the striae margining submesocoxal lines. Its aedeagal characters are similar to those of B. riedeli and B. rufulum, the middle section of the parameres is significantly narrower than that of these two species, and the internal sac has a single, narrow semi­circular sclerite.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	description	(Figs 50 - 53)	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	materials_examined	Type material. Holotype ♂, IRIAN JAYA Jayawijaya Prov., Borme. 1000 - 1300 m, 13 - 18. VIII. 92, leg. A. Riedel (MHNG).	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	description	Description. Length 2.45 mm, width 1.48 mm. Head and thorax blackish, elytra, abdomen and appendages brown or reddish­brown. Pronotal punctation extremely fine and scattered, hardly visible at 100 times magnification. Scutellum triangulate. Elytra lacking humeral humps; adsutural areas distinctly raised; sutural and lateral striae sparsely punctate; basal striae entire, joined to lateral striae; prevailing discal punctation similar to pronotal punctation, several larger punctures scattered on disc. Mesoventrite convex, impunctate, lacking microsculpture, with well delimited mesal ridge extended from paxillum nearly to apical margin and becoming lower and broader apically, and with striae on mesocoxal process. Metaventrite not microsculptured, entirely very finely punctate. Median part of metaventrite impressed, with anterior margin below level of margin of mesoventrite, longitudinally striate between mesocoxae, deeply impressed at centre of anterior margin; with two coarse admesal punctures; apical stripe somewhat flattened, not impressed; metacoxal process inflexed, with slightly concave margin. Submesocoxal lines margined by coarse punctures. Metanepisterna about 0.05 mm wide. Metanepisternal suture impunctate, straight, ending comparatively far from margin of metepimeron. Ventrite Iwithout basomedian hump, with punctulate microsculpture evanescent on lateral areas, impunctate along basal margin. Following ventrites with well visible punctulate microsculpture. Protarsomeres lacking prominent ventral lobe. ♂: Protarsomeres Ito III widened, protarsomere Iabout as wide two thirds of protibial apex, following two tarsomeres narrower. Protarsomere Iabout as long as following two tarsomeres combined. Apicoventral pair of long setae of protarsomere III not flattened apically. Aedeagus as Figs 50 - 53, 1.20 mm long.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	distribution	Distribution. Indonesia: Papua.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	etymology	Etymology. The species epithet is a Latin noun meaning bull. It refers to the two prominent processes of the median lobe of the aedeagus.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
03AE7920FF89FFE950DCFA0707BB4FBB.taxon	discussion	Comments. This species is unique in having robust processes on the ventral side of the aedeagus, near bases of the parameres. It may be also distinguished from most congeners by the conspicuously elongate widened part of the parameres. In external characters it is similar to B. albertisi from which it may be readily distinguished by the impressed median part of the metaventrite and the impunctate basal margin of ventrite I.	en	Löbl, Ivan (2021): A review of the Bironium Csiki, 1909 (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea and the Moluccas. Acta Musei Moraviae 106 (2): 227-248, DOI: http://doi.org/10.5281/zenodo.5706173
