taxonID	type	description	language	source
534F336D7707BD1AFDFFFB8EBD1C51C3.taxon	diagnosis	Diagnosis (according to Gruber & Martens 1968, slightly modified) Genus of Nemastomatidae Simon, 1872, with body length 1.5 – 2.5 mm, with scutum compositum; uniformly black or with silver or golden lateral spots. Pe with base perpendicular to dorso-ventrally flattened truncus, glans bilaterally symmetric or subsymmetric, without or with 1 – 8 pairs of lateral spines or denticles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7707BD1BFE42FAB0BD4956E3.taxon	diagnosis	Diagnosis Uniformly black subspecies of Nemastoma bidentatum and closely related species, with glans with two pairs of lateral lancet-like, articulated spines, and saw-like series of denticles on Fe IV.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7707BD1BFE42FAB0BD4956E3.taxon	description	Morphology The Pe is characterized by a bulbous, muscular, posteriorly medially split basis, a thin truncus and triangular glans, eventually with a shoulder-like truncated tip, either slightly narrower, evenly wide or slightly wider than the truncus, with two pairs of lateral lancet-like, articulated spines. The distal part of the glans is characteristic of some taxa. The shape and size of the Ch basal article and especially of Ch- Apo, and the shape, size and armament of Pa-Fe, and in some taxa Pa-Ti, are further diagnostic characters. Other characters, such as the Pa and leg lengths, may support identification, but are not definitively distinctive. The identification key includes both sexes in the nine pure taxa known to date, except the unknown female of N. pluridentatum stat. nov., and typical male hybrids.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7707BD1BFE42FAB0BD4956E3.taxon	discussion	Remarks To date, four additional taxa of the N. bidentatum complex have been discovered in Slovenia: N. b. gruberi Novak, Slana Novak, Kozel & Raspotnig ssp. nov., N. b. martensi Novak, Slana Novak & Raspotnig ssp. nov., N. b. schmidti Novak, Raspotnig & Slana Novak ssp. nov. and N. b. sneznikensis Novak, Komposch, Slana Novak & Raspotnig ssp. nov., and one in Bosnia and Herzegovina: N. kozari Novak, Kozel, Podlesnik & Raspotnig sp. nov. Within these new taxa, the relatively consistent external morphology of the male Ch-Apo, Pa and glans allows reliable assignment of most individuals. However, N. b. gruberi ssp. nov. shows considerable variation in morphology, especially in the Pa-Fe armament, which in some particular cases still raises doubts about the reliability of the assignment (see the description of the taxon for more details). This is because we are still missing a broad review of potentially syntopic hybrids that should be based on data sets independent from morphology, e. g., chemical characters from highly specific scent gland secretions. Such individuals arise from some marginal localities of the area, e. g., from the very south-west of Slovenia, where we still have not been able to exclude the eventual existence of hybrids with N. b. schmidti ssp. nov. Morphological differences between females based on the Ch, Pa and Rec sem characters are less conspicuous, but allow correct identification of over 90 % of individuals belonging to pure subspecies. Besides, a pairwise vicariance in most neighboring subspecies reveals their different ecological requirements, resulting in niche partitioning. On the other hand, hybrids in the contact zones of adjacent populations of some taxa reveal active gene exchange, supporting the subspecies status of these taxa. Nemastoma bidentatum in Austria deserves a separate treatise, while scarce records from the Balkan countries do not allow any further conclusion for the time being. Thus, to date, there are four species in the Nemastoma b. complex in total: Nemastoma bidentatum Roewer, 1914, N. relictum Gruber & Martens, 1968 stat. nov. and N. pluridentatum (Hadži, 1973) stat. nov., both raised to the species level, and the newly discovered N. kozari Novak, Kozel, Podlesnik & Raspotnig sp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD1BFD89FB90B945539D.taxon	diagnosis	Diagnosis (according to Gruber & Martens 1968, slightly modified) Species of the Nemastoma b. complex with male Pa-Fe having either at least two conspicuous spines or long, rigid setae instead of some spines, or 6 – 7 irregular tubercles, or 5 – 11 denticles, and Pa-Fe min: max width 1: 2.5 – 3.5, Rec sem with 1 or none tubular and 12 – 18 balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD1BFD89FB90B945539D.taxon	discussion	Remarks To date, the species includes six subspecies: N. b. bidentatum Roewer, 1914, N. b. sparsum Gruber & Martens, 1968, and the newly discovered N. b. gruberi Novak, Slana Novak, Kozel & Raspotnig ssp. nov., N. b. martensi Novak, Slana Novak & Raspotnig ssp. nov., N. b. schmidti Novak, Raspotnig & Slana Novak ssp. nov. and N. b. sneznikensis Novak, Komposch, Slana Novak & Raspotnig ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	description	Figs 2 – 3, 4 A, 5 A, 6 A, 7 B, 8 B, 9 A, 10 A, 11 B, 12 A, 13 A; Tables 1 – 2	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	diagnosis	Diagnosis (according to Gruber & Martens 1968, slightly modified) Subspecies of N. bidentatum with large, arched dorsal hump on male Ch basal article, and large, apically, sagittally shallowly bifid, quarter moon-like Ch-Apo, ~ 2.4 times as high as wide. Pa-Fe distinctively club-shaped (Pa-Fe min: max width ~ 1: 3.5), with 3 stout spines in distal Pa-Fe quarter, spine 2 the largest and longest. Pa-Pt Apo equilateral triangular with rounded apex. Pa-Ti with conspicuous proximal dorsomedial hump. Glans isosceles triangular, terminally rounded-truncated. Rec sem of 1 tubular and 12 elongated balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	materials_examined	Material examined AUSTRIA – VM 44 • 1 ♀; Bärental; 23 Apr. 2018; L. Slana Novak and T. Novak leg. (6 / 2018); PMSL. SLOVENIA – VM 44 • 2 ♀♀; Ljubelj; 1 Jun. 1993; S. Brelih leg. (799 / 1998, rev. 2018); PMSL • 1 ♂, 1 ♀; Ljubeljski prelaz; 20 Jun. 2018; T. Novak leg. (23 / 2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 23 Jul. 2021; L. Slana Novak and T. Novak leg. (298 / 2021); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	description	Description (according to Gruber & Martens 1968, slightly modified) Male (from Ljubelj) BODY. Length 1.5 − 1.7. CHELICERAE. Ch basal article with large, arched hump in front of dorsal indentation, and large ventral triangular hump; Ch-Apo conspicuously quarter moon-like, Apo apex sagittally bilobed, medial lobe with inconspicuous apical pinnacle (Figs 4 A, 6 A, 9 A). Secretion field apically medially. Proximal article length 0.57, distal article length 0.54, max width 0.17, movable finger length 0.25. PEDIPALPS. Pa-Tr with high, subequilateral dorsal hump. Pa-Fe distinctively club-shaped (Pa-Fe min: max width ~ 1: 3.5), ventrally bent; with 3, rarely 4 stout spines in distal Pa-Fe quarter; spine 2 the largest, with 2 – 3 pinnacles. Pa-Pt distinctively ventrally bent, with large, stout, apically rounded medio-ventral Pt-Apo. Pa-Ti straight or slightly bent dorsally, with small, but distinct proximal dorso-medial hump. Pa-Ta terminally narrowing, conically oval (Figs 5 A, 6 A, 10 A). Pa article lengths in Table 1. PENIS. Pe length 1.7, Pe basis ⅓ Pe length, glans isosceles triangular, terminally rounded-truncated (Figs 7 B, 8 B). LEGS. Leg article lengths in Tables 1 – 2. Female (from Ljubelj) BODY. Length 1.6 – 1.8. CHELICERAE. Ch with subparallel dorsal and ventral margins, slightly arched dorsally, Ch length: width ~ 2.6: 1 (Fig. 13 A). Proximal article length 0.57, distal article length 0.63, max width 0.21, movable finger length 0.32. PEDIPALPS. Pa-Tr long, Pa-Tr length: width ~ 2.5: 1, Pa-Tr dorsal margin with low hump with long, straight posterior portion and stair-like front indentation in distal quarter; Pa-Fe max: min width ~ 1.9: 1; Pa-Ti straight, simply rod-shaped; Pa-Ta narrow long-oval (Fig. 13 A). Pa article lengths in Table 1. OVIPOSITOR. Ovipositor length 1.05, Rec sem with one tubular and 12 elongated balloon-like vesicles (Fig. 11 B). LEGS. Leg article lengths in Tables 1 – 2.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	discussion	Remarks The type locality Comana Vlasca in Romania (Roewer 1914) is erroneous, since besides N. b. sparsum (sic! not described by Roewer 1914), the type series also contains N. b. bidentatum and N. triste, both endemic to the Eastern Alps (Gruber & Martens 1968: 143). Later, Roewer (1951) designated a type specimen from the type series from an unknown locality, but the mixed composition of the type series remained unresolved (Gruber & Martens 1968). Thus, there was a need to clarify the type locality (ICZN 1999: Art. 75.3.1.). Respecting both Roewerʼs drawing of the male Ch characteristic of individuals in the Karawanken / Karavane Mts and the syntopy of the Roewerʼs type specimens with N. triste, Gruber & Martens (1968) redescribed the nominal subspecies N. b. bidentatum upon a neotype (Gruber & Martens 1968: figs 4 − 6) from Bärental, Carinthia, Austria (not mapped in Gruber & Martens 1968: fig. 24). Bärental is considered the type locality of the type subspecies. However, according to our findings the figured Pe tip from Leutschach / Lučine, Styria, Austria (Gruber & Martens 1968: fig. 3), considered under the type subspecies, belonged to N. b. schmidti ssp. nov. All so far published evidence on N. b. bidentatum in Slovenia, Croatia and NE Italy (Hadži 1927, 1928; Gruber & Martens 1968; Martens 1978; Novak & Gruber 2000; Komposch & Gruber 2004; Novak 2004 a, 2004 b, 2005 b, 2005 c; Novak et al. 1984, 1995, 1996, 2002, 2006, 2017) refer to other taxa. The type series deposited in Naturhistorisches Museum Wien (Gruber & Martens 1968) also includes N. b. schmidti ssp. nov. See remarks under N. pluridentatum stat. nov., N. b. schmidti ssp. nov., N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. and N. b. martensi ssp. nov. × N. b. schmidti ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	distribution	Distribution The South-Eastern Alps: Austria, Slovenia (Gruber & Martens 1968; Martens 1978), perhaps endemic to the Karawanken / Karavanke Mts. Vertical distribution (incomplete data): 980 − 1407 m a. s. l. (partly Gruber & Martens 1968 and Martens 1978; this work). Type locality: Bärental / Zavrh−Rute (46.47 ° N, 14.15 ° E, 980 m a. s. l.) near Feistritz im Rosental / Bistrica v Rožu, Carinthia, Austria.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7706BD16FE50F961B95E574A.taxon	biology_ecology	Ecology Montane subspecies of N. bidentatum inhabiting submontane and montane neutrophile fir forests within the area of distribution of the montane beech forests, dominated by Abies alba Mill., with a great admixture of Picea abies (L.) H. Karst. and Fagus sylvatica L. Phenology: adults probably eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	description	Figs 2 – 3, 4 B, 5 B, 6 B, 7 C, 8 C, 9 B, 10 B, 11 A, C, 12 B, 13 B; Tables 3 – 4	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	diagnosis	Diagnosis (according to Gruber & Martens 1968, slightly modified) Male Ch basal article with nearly straight dorsal margin in front of dorsal indentation, and short, roundly oblong Ch-Apo, with small apical pinnacle. Pa-Fe moderately club-shaped (Pa-Fe min: max width ~ 1: 2.5), with 2 distal, pointed spines, spine 2 the largest. Pa-Pt Apo conical. Glans equilaterally triangular. Rec sem with 14 balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	materials_examined	Material examined CROATIA – XM 05 • 1 ♀; Jurovčak; 3 Apr. 1983; T. Novak and L. Slana Novak leg. (LSN 7 / 1983, TN rev. 2008); PMSL. SLOVENIA – WM 06 • 1 ♂; Cvitrško sedlo, Otiški vrh; 28 May 2008; T. Novak leg. (160 / 2011); PMSL. – WM 42 • 1 ♂, 2 ♀♀, 1 juv.; Mestinje; 4 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 43 / 1983); PMSL • 1 ♂, 1 ♀; ibid.; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1138 / 2002); PMSL • 1 ♀; Šmarje pri Jelšah; 6 Apr. 1986 (1169 / 1997, rev. 2015); PMSL • 1 ♀; Zgornje Poljčane; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1160 / 2002); PMSL. – WM 45 • 1 ♂; Ledine; 25 Jul. 1987; F. Janžekovič leg. (7 / 1999); PMSL • 1 ♂; Maribor; 24 Aug. 1986; Malavašić and Koren leg. (265 / 2002); PMSL. – WM 47 • 1 ♀; Kresnica; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (162 / 2011); PMSL. – WM 52 • 1 ♂; Maršečka vas; 11 Oct. 1997; L. Slana Novak and T. Novak leg. (1893 / 1998); PMSL. – WM 53 • 2 ♂♂, 1 ♀; Pečke; 24 Jun. 2007; T. Novak leg. (45 / 2007); PMSL • 1 ♂, 2 ♀♀; Šega, Makole; 9 Apr. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 70 / 1983); PMSL. – WM 56 • 1 ♀; Kaniža; 20 May 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 80 / 1983); PMSL • 1 ♀; Mt. Piramida, Maribor, 1 − 30 Nov. 1978; (909 / 1997, rev. 2015); PMSL • 1 ♀; Spodnje Dobrenje; 23 Apr. 2008; L. Slana Novak and T. Novak leg. (165 / 2011); PMSL • Ranca, Pesnica. – WM 57 • 1 ♂; Ceršak; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (113 / 2011); PMSL • 1 ♂; Kozjak pri Ceršaku; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (116 / 2011); PMSL. – WM 66 • 2 ♀♀; Črnetina; 23 Apr. 2008; L. Slana Novak and T. Novak leg. (163 / 2011); PMSL • 1 ♂; Kremberg; 4 Jun. 1984; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 343 / 1985); PMSL • 2 ♂♂; ibid.; 23 Apr. 2008; L. Slana Novak and T. Novak leg. (114 / 2011); PMSL • 1 ♀; Spodnji Žerjavci; 23 Apr. 2008; L. Slana Novak and T. Novak leg. (115 / 2011); PMSL. – WM 67 • 1 ♂; Sv. Ana; 3 Jul. 2002; T. Novak leg. (799 / 2002); PMSL • 1 ♀; Nasova; 3 Jun. 1984; L. Slana Novak and T. Novak leg. (314 / 2002, rev. 2015); PMSL • 1 ♂; Stogovci; 13 Apr. 1984; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 323 / 1985); PMSL • 1 ♂, 1 ♀; Zgornja Velka; 20 Mar. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 8 / 1983); PMSL. – WM 73 • 1 ♀; Repišče; 24 Jun. 2007; T. Novak leg. (31 / 2007, rev. 2015); PMSL. – WM 74 • 1 ♀; Podvinci; 26 Oct. 1994; M. Varžič leg. (21 / 1999); PMSL • 1 ♂; Žamenci; 22 Oct. 1994; M. Varžič leg. (18 / 1999); PMSL • 1 ♂; ibid.; (20 / 1999); PMSL • 1 ♂; ibid.; 23 Oct. 1994; M. Varžič leg. (19 / 1999, rev. 2015); PMSL. – WM 75 • 1 ♂; Trnovce; 2 Jun. 2002; L. Slana Novak and T. Novak leg. (760 / 2002); PMSL • 7 ♂♂, 4 ♀♀; Velika Slavšina; 13 Oct. 2005; L. Slana Novak and T. Novak leg. (158 / 2005, rev. 2018); PMSL • 5 ♂♂, 3 ♀♀; Vitomarci; 30 Jun. 2002; T. Novak leg. (787 / 2002); PMSL • 1 ♀; ibid.; 3 Jul. 2002; T. Novak leg. (792 / 2002); PMSL • 1 ♂; ibid.; (797 / 2002); PMSL • 1 ♀; ibid.; 3 Jul. 2002; T. Novak leg. (794 / 2002, rev. 2008); PMSL • 1 ♀; ibid.; (796 / 2002); PMSL • 1 ♂; ibid.; (797 / 2002); PMSL • 1 ♂; ibid.; (800 / 2002); PMSL • 1 ♀; ibid.; 2 Oct. 2002; T. Novak leg. (1280 / 2002); PMSL • 1 ♂; ibid.; (1279 / 2002); PMSL. – WM 77 • 1 ♀; Črešnjevci; 24 Apr. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 79 / 1983, TN rev. 2008); PMSL • 1 ♂, 1 ♀; Korovci; 7 Apr. 1984; L. Slana Novak and T. Novak leg. (LSN 191 / 1985); PMSL • 1 ♂, 1 ♀; Lomanoše; 1 Oct. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 110 / 1983); PMSL. – WM 78 • 1 ♂; Gerlinci; 11 May 1986; T. Novak and L. Slana Novak leg. (LSN 139 / 1986); PMSL. – WM 86 • 2 ♂♂, 6 ♀♀; Hrašenski vrh; 9 Sep. 1983; T. Novak and L. Slana Novak leg. (LSN 115 / 1983); PMSL • 4 ♂♂, 3 ♀♀; Kapelski vrh; 17 Sep. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 112 / 1983); PMSL • 1 ♂; Murski vrh; 19 May 1984; L. Slana Novak and T. Novak leg. (LSN 240 / 1984); PMSL • 1 ♀; Radenci; 25 Jul. 1983; M. Slana Novak and L. Slana Novak leg. (LSN 111 / 1983, TN rev. 2007); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 20 Sep. 1983; L. Slana Novak leg. (LSN 119 / 1983); PMSL • 1 ♂; Rihtarovci; 26 Feb. 1997; B. Drovenik leg. (94 / 2001); PMSL. – WM 87 • 3 ♂♂, 3 ♀♀; Cankova; 15 Sep. 1983; T. Novak and L. Slana Novak leg. (LSN 114 / 1983); PMSL • 3 ♂♂, 3 ♀♀; Skakovci; 27 Mar. 1983; T. Novak and L. Slana Novak leg. (LSN 4 / 1983); PMSL • 1 ♂; Sodišinci, Radenci; 19 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 68 / 1983); PMSL. – WM 88 • 1 ♂; Gerlinci; 11 Aug. 1983; L. Slana Novak, M. Slana Novak and T. Novak leg. (1322 / 1998); PMSL. – WM 89 • 2 ♂♂, 4 ♀♀, 1 juv.; Trdkova; 30 Jul. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 49 / 1983); PMSL. – WM 94 • 1 ♀; Pavlovci; 15 Apr. 2000; L. Slana Novak and T. Novak leg. (744 / 2002); PMSL. – WM 95 • 1 ♀; Podgradje, Ljutomer; 27 May 2001; S. Brelih leg. (125 / 2001); PMSL. – WM 96 • 1 juv.; Bratonci; 9 Aug. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 335 / 1985, TN rev. 2008); PMSL. – WM 97 • 1 ♂; Andrejci; 28 Jul. 1981; T. Novak leg. (1188 / 1981, JG det.); PMSL • 1 ♂; Moravci; 16 Mar. 1984; T. Novak and L. Slana Novak leg. (LSN 243 / 1984); PMSL • 4 ♂♂; Murska Sobota; 24 Nov. 1982; T. Novak leg. (603 / 1982); PMSL • 3 ♀♀; Rakičan – Noršinci; 23 Sep. 1984; L. Slana Novak leg. (322 / 1985); PMSL • 1 ♂, 2 ♀♀; ibid.; 13 Sep. 1981; T. Novak leg. (371 / 1982, JG det.); PMSL. – WM 98 • 2 ♂♂, 3 ♀♀; Košarovci; 24 Jul. 1982; L. Slana Novak, T. Novak and M. Slana Novak leg. (529 / 1982, JG det.); PMSL • 1 ♀; ibid.; 1 Sep. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 318 / 1985); PMSL • 1 ♂, 3 ♀♀; Mačkovci; 31 Jul. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 67 / 1983); PMSL. – WM 99 • 2 ♂♂; Lucova; 2 Jun. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 239 / 1984); PMSL. – XM 05 • 1 ♀; Bobri; 24 Aug. 1988; F. Velkovrh leg. (13 / 1995); PMSL • 2 ♂♂, 1 juv.; Dolnja Bistrica; 17 Aug. 1996; I. Kovač leg. (SL 2 / 1997); PMSL • 2 ♀♀; ibid.; (SL 3 / 1997); PMSL. – XM 06 • 2 ♂♂, 3 ♀♀; Banuta / Bánuta; 10 Aug. 1996; I. Kovač leg. (SL 5 / 1997); PMSL • 2 ♂♂, 1 ♀; ibid.; (SL 22 / 1997); PMSL • 2 ♂♂, 2 ♀♀; ibid.; (SL 21 / 1997); PMSL • 13 ♂♂, 15 ♀♀; ibid.; 13 Aug. 2012; T. Novak leg. (GR 3837 − 3849, 3792 − 3797, TN det.); PMSL • 4 ♂♂, 3 ♀♀; ibid.; (GR 3770 − 3776); PMSL • several specs; Polanski log, Mala Polana; 17 Jul. 2000; T. Novak leg.; PMSL • 1 ♂, 1 ♀; Radmožanci / Radamos; 17 Mar. 1984; T. Novak and L. Slana Novak leg. (LSN 109 / 1984); PMSL. – XM 07 • 4 ♂♂, 3 ♀♀; Dobrovnik / Dobronak; 8 Aug. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 57 / 1983); PMSL • 1 ♀, 1 juv.; ibid.; 18 Aug. 1996; I. Kovač leg. (SL 1 / 1997); PMSL. – XM 08 • 3 ♂♂, 1 ♀; Prosenjakovci / Pártosfalva; 8 Aug. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 50 / 1983); PMSL • 2 ♀♀; Središče / Szerdahely; 12 Aug. 1996; I. Kovač leg. (SL 6 / 1997); PMSL • 4 ♂♂; ibid.; 24 Aug. 1996; I. Kovač leg. (SL 7 / 1997); PMSL • 2 ♀♀; ibid.; (SL 8 / 1997); PMSL • 1 ♂, 3 ♀♀; ibid.; (SL 20 / 1997); PMSL • 4 ♂♂; ibid.; (SL 23 / 1997); PMSL • 4 ♂♂; ibid.; (SL 61 / 1997); PMSL • 3 ♂♂; ibid.; (SL 108 / 1997); PMSL • 12 ♀♀; ibid.; (SL 105 / 1997); PMSL. – XM 15 • 3 ♀♀; Benica / Benice; 1 Aug. 1996; I. Kovač leg. (SL 11 / 1997); PMSL • 3 ♀♀; ibid.; (SL 12 / 1997); PMSL • 2 ♂♂; ibid.; 19 Aug. 1996; I. Kovač leg. (SL 25 / 1997); PMSL • 2 ♀♀; ibid.; (SL 26 / 1997); PMSL • 5 ♀♀; ibid.; (SL 28 / 1997); PMSL • 3 ♂♂; ibid.; (SL 29 / 1997); PMSL • 2 ♂♂; ibid.; (SL 39 / 1997); PMSL • 2 ♀♀; ibid.; (SL 64 / 1997); PMSL • 1 ♂, 1 ♀; ibid.; (SL 65 / 1997); PMSL • 1 ♂, 3 ♀♀; Gaberje / Gyetyános; 22 Aug. 1996; I. Kovač leg. (SL 9 / 1997); PMSL • 1 ♀; ibid.; (SL 10 / 1997); PMSL • 1 ♂; ibid.; (SL 15 / 1997); PMSL • 3 ♀♀; ibid.; (SL 16 / 1997); PMSL • 1 ♀; ibid.; 12 Oct. 1996; I. Kovač leg. (SL 30 / 1997); PMSL • 2 ♀♀; ibid.; (SL 55 / 1997); PMSL • 5 ♂♂; ibid.; 19 Oct. 1996 (SL 17 / 1997); PMSL • 4 ♂♂, 4 ♀♀; ibid.; (SL 24 / 1997); PMSL • 4 ♀♀; ibid.; (SL 103 / 1997); PMSL • 8 ♀♀; ibid.; (SL 104 / 1997); PMSL • 2 ♀♀; ibid.; (SL 106 / 1997); PMSL • 7 ♂♂; ibid.; (SL 107 / 1997); PMSL • 2 ♂♂; ibid.; (SL 109 / 1997); PMSL • 14 ♀♀; ibid.; 22 Aug. 1997; I. Kovač leg. (SL 27 / 1997); PMSL • 1 ♂, 1 juv.; ibid.; (SL 32 / 1997); PMSL • 1 ♂; ibid.; (SL 68 / 1997); PMSL • 4 ♂♂; ibid.; 24 Aug. 1997; I. Kovač leg. (SL 19 / 1997); PMSL • 2 ♀♀; ibid.; 28 Aug. 1997; I. Kovač leg. (SL 13 / 1997); PMSL • 2 ♂♂; ibid.; (SL 14 / 1997); PMSL • 1 ♂; ibid.; 12 Oct. 1997; I. Kovač leg. (SL 4 / 1997); PMSL • 4 ♂♂; ibid.; (SL 20 / 1997); PMSL • 1 ♀; ibid.; (SL 56 / 1997); PMSL • 1 ♂; ibid.; (SL 100 / 1997); PMSL • 4 ♀♀; ibid.; (SL 101 / 1997); PMSL. – XM 16 • 2 ♂♂; Murska šuma; 29 Aug. 1998; T. Novak leg. (1538 / 1998); PMSL • several specs; ibid.; 18 Jun. 2001; T. Novak leg.; PMSL • 1 ♂, 1 ♀; Dolgovaške Gorice / Hossúfaluhegy; 31 Mar. 1984; T. Novak, D. Kuhar and L. Slana Novak leg. (LSN 141 / 1984); PMSL • 4 ♂♂, 1 ♀; Lendavske gorice / Lendvahegy; 17 Aug. 1996; I. Kovač leg. (SL 31 / 1997); PMSL. – XM 25 • 1 ♂; Muriša, Murska šuma; B. Drovenik and A. Pirnat leg. (26 / 2005); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	description	Description (according to Gruber & Martens 1968, slightly modified) Male (from Prosenjakovci) BODY. Length 1.5 − 2.0. Males from Slovenia larger than those from the type locality. Length 1.96, width 1.44. CHELICERAE. Ch basal article with nearly straight dorsal margin in front of dorsal indentation and small ventral hump, Ch-Apo short, roundly oblong, with convex anterior margin and small apical pinnacle (Figs 4 B, 6 B, 9 B). Secretion field medially. Proximal article length 0.53, distal article length 0.59, max width 0.18, movable finger length 0.25. PEDIPALPS. Pa-Tr with highly arched dorsal hump. Pa-Fe moderately club-shaped (Pa-Fe min: max width ~ 1: 2.5), slightly ventrally bent, with 2 stout, pointed spines in distal Pa-Fe quarter; spine 2 the largest. Pa-Pt Apo triangular-conical. Pa-Ti simple, straight. Pa-Ta conically tapering (Figs 5 B, 6 B). Pa article lengths in Table 3. PENIS. Pe length 1.6, Pe basis ¼ Pe length, glans equilaterally triangular (Figs 7 C, 8 C). LEGS. Leg article lengths in Tables 3 – 4. Female (from Prosenjakovci) BODY. Body length 1.6 − 2.4. Females from Slovenia larger than those from the type locality. Body length 2.40, body width 1.97. CHELICERAE. Ch basal article with low-arched dorsal margin in front of dorsal indentation, and straight ventral margin with triangular hump, Ch basal article length: width> 2.6: 1. Proximal article length 0.57, distal article length 0.63, max width 0.21, movable finger length 0.32 (Fig. 13 B). PEDIPALPS. Pa-Tr with moderately high, arched dorsal margin; Pa-Fe with distal bonce, Pa-Fe max: min width ~ 1.7: 1; Pa-Ti elongated-oval (13 B). Pa article lengths in Table 3. OVIPOSITOR. Ovipositor length 1.0, Rec sem with 14 balloon-like vesicles (Fig. 11 A, C). LEGS. Leg article lengths in Tables 3 – 4.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	discussion	Remarks In Slovenia, N. b. sparsum lives in the northeast, at the margins of the Pannonian basin (Martens & Gruber 1968; Martens 1978; Kovač 1997). The type series deposited in Naturhistorisches Museum Wien (Gruber & Martens 1968) also includes N. b. gruberi ssp. nov. distributed south of the Sava River in Slovenia and Croatia. All prior identifications of N. b. sparsum south of the Sava River (Novak et al. 1995, 2002; Novak 2005 b) refer to N. b. gruberi ssp. nov. Martens (1978: 112) reported a sparsum - like population living in the mountain pine association (Hyperico grisebachii-Pinetum mugo Horvat 1938) on Mt. Veliki Snežnik (the peak at 1796 m) at least up to the altitude of 1650 m a. s. l. These individuals may refer to N. b. gruberi ssp. nov., N. b. martensi ssp. nov. or N. b. sneznikensis ssp. nov. Individuals from Bosnia and Herzegovina (Mt. Plasa, Jajce) and Dalmatia, Croatia (Split) (Gruber & Martens 1968) should be revised for identification. See remarks under N. b. gruberi ssp. nov. and N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov., N. b. martensi ssp. nov. and N. b. sneznikensis ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	distribution	Distribution Central and eastern Europe: Austria, Bulgaria, Bosnia and Herzegovina, Czech Republic, Denmark, Hungary, Croatia, Montenegro, Romania, Russia, Slovakia, Slovenia, Serbia and Turkey (Gruber & Martens 1968; Staręga 1976; Martens 1978, 2006; Mučalica 1988; Klimeš 2000; Komposch 2002; Komposch & Gruber 2004; Schönhofer & Holle 2007). Vertical distribution: 120 − 1650 m a. s. l. (Komposch 1999; Komposch & Gruber 2004), in Slovenia 155 – 644 m a. s. l. Type locality: Sonnenberg (47.88 ° N, 16.31 ° E, 480 m a. s. l.), Leitha Mts, Burgenland, Austria.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770BBD13FE02FD2ABE6E51BE.taxon	biology_ecology	Ecology Nemastoma b. sparsum is a lowland and hilly land subspecies inhabiting various types of acidophilous and neutrophilous forests of Fagus sylvatica, Carpinus betulus L. and Quercus petraea (Matt.) Liebl., riverine forests, and thermophilous scrubs in north-eastern Slovenia, mostly on non-calcareous soils. Phenology: adults eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	description	Figs 1 B, 2, 4 C, 5 C, 6 C, 7 D, 8 D, 9 C, 10 C, 11 D, 12 C, 13 C; Tables 5 – 6	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	diagnosis	Diagnosis Species within the N. b. complex with button-shaped Ch-Apo, and secretion field oriented frontally. Pa- Fe without spines, slightly club-shaped (Pa-Fe min: max width ~ 1: 2.3). Rec sem with 5 lobate, tubularsaccular vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	materials_examined	Material examined AUSTRIA • 2 ♂♂, 3 ♀♀; Grosses Kar, the Koralpe Mts; 25 Oct. 2005; C. Komposch leg. (CK det.); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	description	Description (according to Gruber & Martens 1968, slightly modified) Male (from Grosses Kar, Salzburg) BODY. Length 1.55 − 1.60. CHELICERAE. Ch with flat triangular dorsal margin in front of dorsal indentation; ventral margin straight, with minute triangular hump. Ch-Apo button-like, inferior and frontal margins straight, embracing a 90 ° angle, Ch-Apo dorsally round, secretion field with dense setae (brush) frontally (Figs 4 C, 6 C, 9 C). Proximal article length 0.54, distal article length 0.53, max width 0.18, movable finger length 0.19. PEDIPALPS. Pa-Tr dorsal margin high, oval, leaned distally; Pa-Fe distended distally (max: min width ~ 1.7: 1), without spines. Pa-Pt with conspicuously convex ventral margin and finger-like medial Apo. Pa-Ti and Pa-Ta straight (Figs 5 C, 6 C). Pa article lengths in Table 5. PENIS. Pe length 1.6, Pe basis ¼ Pe length, glans isosceles triangular (Figs 7 C, 8 C). LEGS. Leg article lengths in Tables 5 – 6. Female (from Grosses Kar, Salzburg) BODY. Length 1.70. CHELICERAE. Ch basal article with low-arched dorsal margin in front of indentation, and ventral lowarched concavity, Ch basal article length: width ~ 2.3: 1 (Fig. 13 C). Proximal article length 0.56, distal article length 0.55, max width 0.19, movable finger length 0.20. PEDIPALPS. Pa short, ~ 2 mm long, Pa-Tr with high, isosceles triangular dorsal margin arched on the top, Pa-Fe distended distally (max: min width ~ 1.6: 1), Pa-Ta short, Pa-Ta length: Pa-Ti length ~ 1: 1.6 (Fig. 13 C). Pa article lengths in Table 5. OVIPOSITOR. Rec sem with 5 lobate, tubular-saccular vesicles (Fig. 11 D). LEGS. Leg article lengths in Tables 5 – 6.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	discussion	Remarks We did not examine the original type material; for the current description, we used other specimens from the type location. Rec sem were found much different from the other taxa within the N. b. complex.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	distribution	Distribution Austria, endemite of eastern Central Alps (Gruber & Martens 1968; Martens 1978; Komposch 1999, 2009). Vertical distribution: 1325 − 2240 m a. s. l. (Komposch 1999, 2009). Type locality: Grosses Kar, near the Tappenkarsee Lake, Kleinartal, Salzburg (47.19 ° N, 13.32 ° E, 1760 m a. s. l.), Radstädter Tauern Mts, Austria.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D770EBD0DFE30FB41BD8557F6.taxon	biology_ecology	Ecology Nemastoma relictum stat. nov. is a montane to alpine species within the N. b. complex inhabiting forests below the krummholz zone, alpine meadows, spring-areas, and gravel and block heaps in the eastern Central Alps (Gruber & Martens 1968; Martens 1978; Komposch 1999, 2009) in 1325 – 2240 m a. s. l. (Komposch 1999). Phenology: evidence of adults from the end of May till the end of October, but probably eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	description	Figs 2, 4 D, 5 D, 6 D, 9 D, 10 D, 12 D; Table 7	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	diagnosis	Diagnosis (partly according to Hadži 1973 a) Robust, short-legged member of the Nemastoma b. complex, with extremely stout Ch and Pa. Ch-Apo small, slightly trapezoid button-like, wider than high, with secretion field in frontal third. Pa-Tr nearly round, Pa-Fe and Pa-Pt very stout, Pa-Fe club-shaped, with 4 large, wide humps.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	materials_examined	Material examined Holotype BOSNIA AND HERZEGOVINA – YK 10 • 1 ♂; Mt. Paljenik, Vlašić planina Mts; 1933 m a. s. l.; 18 Jun. 1962; E. Pretner leg. (4 / 2005, JH’s microscopic preparation); PMSL. (Note: UTM code in Novak (2005 a) was erroneously cited as YJ 19.)	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	description	Partial redescription Note: male body and penis missing; female unknown. Male (holotype) CHELICERAE. Ch basal article length (without Apo) 0.57, stout, 1.8 times as long as wide at Ch max width at dorsal hump, distally rounded. Ch-Apo rounded low trapezoid, with longest margin distally, secretion field frontally. Distal article length 0.57, max width 0.26, movable finger length 0.28 (Figs 4 D, 6 D, 9 D). PEDIPALPS. Pa stout, Pa-Fe extremely stout, with 4 large, bulky humps (instead of spines), club-shaped (Pa-Fe min: max width ~ 1: 3.6), Pa-Pt extremely stout (length: max width ~ 1: 2.8) (Figs 5 D, 6 D). Pa article lengths in Table 7. LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 2 − 1 − 2. Tarsomere formula I−IV: 8 − 13 − 5 − 8. Leg article lengths in Table 7.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	discussion	Remarks Under the name Nemastoma (Lugubrostoma) triste pluridentatum, Hadži (1973 a) described four taxa: a) N. pluridentatum (Hadži, 1973) stat. nov. (Hadži 1973 a: fig. 33 a) from Mt. Paljenik (1943 m a. s. l.) in the Vlašić Mts, Bosnia, b) a hybrid N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. (Hadži 1973 a: fig. 33 b, č) from the surroundings of Novo mesto, Slovenia, c) a hybrid N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. (Hadži 1973 a: fig. 33 c) from the surroundings of Novo mesto, which Hadži (ibid.) erroneously cited for Mt. Vlašić, and d) Nemastoma b. schmidti ssp. nov. from Mt. Šmarna gora, Slovenia (Hadži 1973 a: fig. 34 a), from the surroundings of Ljubljana (Hadži 1973 a: fig. 34 b), and from the cave Jama Sv. Janeza pri Prestranku near Postojna, Slovenia (Hadži 1973 a: fig. 34 c). Figs 33 and 34 were drawn freehand; consequently, shapes and dimensions only partly correspond to the real measurements. Hadži’s original microscopic preparation labeled “ Nemastoma bidentatum Roewer pulli, A- 1065 (Novo Mesto), legit Karaman ” includes two males and a female, but no juveniles. Hadži (ibid.) drew some details of the males, including damaged legs; see remarks on Nemastoma (Stridulostoma) seliskari Hadži, 1973 under N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. Given that some important parts, e. g., Pa of one male, are missing, Hadži probably made this incomplete preparation after observation of the specimens under a microscope. In his original paper (Hadži 1973 a), the type specimen and the type locality of N. pluridentatum stat. nov. were not designated, but he did that in the catalogue (Hadži 1973 b: 13) under a mistyped name: “ N. (L.) triste pleuridentatum Hadži, 1973 ”. There he wrote: “ Loc. typ.: Novo mesto ”. Since the specimens from this locality belong to the above-mentioned hybrids, Novo mesto cannot be the type locality of N. pluridentatum stat. nov.; that is the first-mentioned location in Bosnia, as fig. 33 a (Hadži 1973 a) is the only drawing that refers to this taxon. Unfortunately, only the preparation of one Ch, one Pa and four legs of the male holotype are preserved, while the body, as well as a further male and female mentioned by Hadži (1973 a: 44) are missing. Because of the lack of any information on the Pe, the saw on Fe IV in preserved legs is the only available character allowing us to place this taxon into the N. b. complex, interpreted as N.? bidentatum pluridentatum (Novak 2005 a: fig. 34) or N. b. pluridentatum (Schönhofer 2013). In 2019, we visited the type locality Mt. Paljenik, but we did not find further specimens of N. pluridentatum. Instead we found a new, much different N. kozari sp. nov. there, which unlikely hybridize with N. pluridentatum stat. nov. Consequently, we ascribed the species status to N. pluridentatum. See remarks under N. relictum stat. nov., N. kozari sp. nov. and hybrids N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. and N. b. martensi ssp. nov. × N. b. schmidti ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	distribution	Distribution Bosnia and Herzegovina, endemic to central Bosnia, probably endemic to the Vlašić Mts. Type locality: Mt. Paljenik in the Vlašić planina Mts, Bosnia, (44.29 ° N, 17.64 ° E, 1933 m a. s. l.), Bosnia and Herzegovina.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7710BD0FFE7CFC8BB910544E.taxon	biology_ecology	Ecology Nemastoma pluridentatum stat. nov. inhabit the highest peak areas with the Mastgrass Gentianello crispatae-Nardetum strictae Redžić 1990 communities. Phenology: adults probably eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	description	urn: lsid: zoobank. org: act: 17 EA 28 A 1 - 5 F 36 - 4 DFC- 9191 - 9 C 8680496 FAC Figs 1 A, C, 2 – 3, 4 E, 5 E, 6 E, 7 E, 8 E, 9 E, 10 E, 11 E, 12 E, 13 D; Table 8	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	diagnosis	Diagnosis Relatively long-legged subspecies of Nemastoma bidentatum, with strait Ch ventral margin, and Pa-Fe with either 4 subequidistantly positioned spines or pointed tubercles or long setae on low protrusions, or their combination in the distal Pa-Fe half, or alternatively, with 2 – 3 distal stout thorns. Rec sem with one tubular and 12 elongated balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	etymology	Etymology The subspecies name ʻ gruberi ʼ is dedicated to Jürgen Gruber (Vienna), our teacher, colleague and friend, who generously introduced us to the taxonomy of harvestmen and helped in identifying problematic specimens.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	materials_examined	Material examined Holotype SLOVENIA – WL 18 • 1 ♂; Dolenje Laknice, Mokronog; 45.93 ° N, 15.21 ° E; 237 m a. s. l.; 26 Oct. 2013; L. Slana Novak and T. Novak leg.; mixed forest litter sift; PMSL-Opiliones-GR 4661. Paratypes SLOVENIA – WL 18 • 1 ♀; same collection data as for holotype; PMSL-Opiliones-GR 4664 • 5 ♂♂, 4 ♀♀; same collection data as for holotype; PMSL-Opiliones-GR 4662, GR 4666, GR 4667, GR 4669, GR 4673, GR 4676 to GR 4679. Other material ITALY – VL 05 • 1 ♀; Chiusa / Ricmanje; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (275 / 1999, rev. 2015); PMSL. – VL 15 • 1 ♀; Basovizza / Bazovica; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (271 / 1999, rev. 2015); PMSL. CROATIA – UL 92 • 3 ♂♂, 2 ♀♀ (paratypes of N. b. sparsum); vicinity of Buje, and VL 02, Vižinada; Sep. 1963; R. Sturany leg.; NHMW 4700. – VL 01 • 1 ♂; Karojba; 28 Feb. 1990; F. Potočnik leg. (1307 / 1998, rev. 2018); PMSL. – VL 13 • 1 ♀; Mlini; 30 Sep. 1990; L. Slana Novak and T. Novak leg. (32 / 1990, rev. 2015); PMSL. – XL 90 • 1 ♂, 2 ♀♀; Ponor Sušik, Drežnik; 8 Sep. 2009; A. Schönhofer leg. (Coll. ASc 334, ASc det., TN rev. 5 / 2012); PMSL. SLOVENIA – UL 93 • 4 ♀♀, 1 juv.; Abrami; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (148 / 1999, rev. 2018); PMSL • 2 ♀♀; ibid.; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (188 / 1999, rev. 2015); PMSL • 1 ♀; Dragonja; 14 Sep. 2000; L. Slana Novak and T. Novak leg. (154 / 2001, rev. 2015); PMSL • 4 ♂♂, 4 ♀♀; Dragonja valley; 26 Jun. 2015; T. Novak leg. (78 / 2015 = GR 5123 − 5130); PMSL • 2 ♂♂, 3 ♀♀; Ivankovec; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1791 / 1998, rev. 2018); PMSL • 1 ♀; Lucija – Seča – Debernardo; 22 Sep. 1998; L. Slana Novak and T. Novak leg. (1817 / 1998, rev. 2019); PMSL • 1 ♂; Korte; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1821 / 1998, rev. 2018); PMSL • 33 ♂♂, 30 ♀♀; Medljan, Korte; 21 Aug. 1999; T. Novak leg. (239 / 1999, rev. 2018); PMSL • 1 juv.; ibid.; 22 Jun. 2000; B. Bertoncelj, B. Ogrizek and T. Novak leg. (217 / 2000, rev. 2018); PMSL • 1 ♀; ibid.; (218 / 2000, TN rev. 2019); PMSL • 1 ♂; Padna; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (168 / 1999, rev. 2017); PMSL • 2 ♀♀; Parecag; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1788 / 1998, rev. 2018); PMSL • 1 ♀; Seča; 10 Oct. 1998; L. Slana Novak and T. Novak leg. (1844 / 1998, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; 24 Oct. 1998, L. Slana Novak and T. Novak leg. (1881 / 1998, rev. 2007); PMSL • 3 ♀♀; ibid.; 21 Aug. 1999; M. Štangelj and T. Novak leg. (236 / 1999, rev. 2008); PMSL • 15 ♂♂, 21 ♀♀; ibid.; 21 Aug. 1999; M. Štangelj and T. Novak leg. (231 / 1999, rev. 2015); PMSL • 1 ♂, 6 ♀♀; Sečovlje; 1 Mar. 2009; L. Slana Novak and T. Novak leg. (1 / 2015); PMSL • 1 ♂; Sv. Peter; 11 Oct. 1998; L. Slana Novak and T. Novak leg. (1850 / 1998, rev. 2015); PMSL • 4 ♂♂, 2 ♀♀; Sv. Sab, Padna; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (151 / 1999, rev. 2009); PMSL • 1 ♀; Škrline, Dragonja; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (194 / 1999, rev. 2015); PMSL. – VL 03 • 2 ♂♂, 1 ♀; Boršt; 14 Apr. 2011; T. Novak leg. (76 / 2011, rev. 2015); PMSL • 2 ♀♀; ibid.; (81 / 2011, rev. 2015); PMSL. – VL 04 • 1 ♀; Ankaran; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (276 / 1999, rev. 2015); PMSL • 1 ♂, 1 ♀; Kolombini, Spodnje Škofije; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (94 / 2007, rev. 2018); PMSL • 5 ♂♂; Mlinarji – Osp; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (97 / 2007, rev. 2017); PMSL • 1 ♂, 1 ♀; Rižana; 24 Sep. 2016; L. Slana Novak and T. Novak leg. (18 / 2018); PMSL • 1 ♀; Škocjanski zatok, Koper; 15 Oct. 2009; S. Polak leg. (PK 1 / 2015); NM • 4 ♂♂, 3 ♀♀; Zgornje Škofije – Tinjan; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (88 / 2007); PMSL. – VL 07 • 4 ♀♀; Hrastje, Šmarje; 17 Aug. 2000 (674 / 2000, rev. 2015); PMSL. – VL 13 • 1 ♂; Poletiči, Kubed; 4 Sep. 2016; T. Marinko leg. (27 / 2016); PMSL • 1 ♂, 1 ♀; Sočerga; 28 Sep. 1990; L. Slana Novak and T. Novak leg. (31 / 1990, rev. 2015); PMSL. – VL 14 • 4 ♂♂, 4 ♀♀; Bezovica; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (LSN 1 / 1990, TN rev. 2018); PMSL • 1 ♂; Hrpelje, Kozina; 2 Apr. 1993; S. Brelih leg. (651 / 1998, rev. 2018); PMSL • 2 ♂♂; Predloka; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (35 / 1990, rev. 2015); PMSL • 2 ♂♂, 1 ♀; Rižana; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (33 / 1990, rev. 2015); PMSL. – VL 15 • 1 ♀; Botač; 22 Mar. 1990; M. Černila, B. Horvat and I. Sivec leg. (1246 / 1997); PMSL. – VL 16 • 1 ♂; Divača; 2 Apr. 1993; S. Brelih leg. (174 / 2001, rev. 2015); PMSL. – VL 17 • 1 ♂; Vrabče, Podnanos; 10 Mar. 1994; S. Brelih leg. (632 / 1998, rev. 2017); PMSL. – VL 18 • 1 ♂; Uhanje; 3 Sep. 2001; T. Novak leg. (329 / 2001, rev. 2018); PMSL. – VL 23 • 1 ♂, 1 ♀; Poljane pri Podgradu; 18 Aug. 2011; L. Slana Novak and T. Novak leg. (166 / 2012, rev. 2019); PMSL. – VL 25 • 3 ♀♀; Buje; 10 Mar. 1994; S. Brelih leg. (163 / 2001, rev. 2018); PMSL • 1 ♂, 2 ♀♀; Goriče pri Framljah; L. Slana Novak and T. Novak leg. (343 / 2011); PMSL • 1 ♀; ibid.; (344 / 2011); PMSL. – VL 28 • 1 ♂, 1 ♀; Podvrh, Raka; 18 Jul. 2012; P. Kozel, T. Kozel and T. Novak leg. (55 / 2012); PMSL • 9 ♂♂, 7 ♀♀; ibid.; (GR 3734 – 3746); PMSL • 3 ♀♀; ibid.; (GR 3791); PMSL. – VL 29 • 6 ♂♂, 12 ♀♀; Dolnje Brezovo, Sevnica; 30 Jun. 2012; (GR 3670 – 3683); PMSL. – VL 36 • 1 ♂, 1 ♀; Palčje – Mt. Javorniki; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (361 / 2011, rev. 2019); PMSL. – VL 45 • 1 ♀; Koritnice − Mašun; 2 Oct. 2011; L. Slana Novak and T. Novak leg. (329 / 2011, rev. 2014); PMSL. – VL 47 • 1 ♀; Suha reber, Koritnice; 21 Jan. 2007; S. Polak leg. (107 / 2007); NM. – VL 54 • 1 ♀; Sviščaki; 20 May 1989; T. Trilar leg. (271 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; 25 May 1989; T. Trilar leg. (275 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; 21 Jul. 2001; B. Bertoncelj, U. Bertoncelj and T. Novak leg. (29 / 2001, rev. 2018); PMSL • 1 ♀; ibid.; (34 / 2001, rev. 2011); PMSL • 1 ♂, 3 ♀♀; Travni Dolci, Mt. Snežnik; 12 Aug. 2001; L. Slana Novak and T. Novak leg. (195 a / 2001, rev. 2020); PMSL • 2 ♂♂, 4 ♀♀; Mt. Veliki Snežnik; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (185 a / 1999, rev. 2020); PMSL • 1 ♂; ibid.; 12 Sep. 2018; L. Slana Novak and T. Novak leg. (116 / 2018); PMSL. – VL 36 • 1 ♂, 1 ♀; Palčje − Javorniki; L. Slana Novak and T. Novak leg. (361 / 2011, rev. 2015); PMSL. – VL 66 • 1 ♀; Benete, Runarsko; 31 May 2004; L. Slana Novak and T. Novak leg. (33 / 2004, rev. 2015); PMSL. – VL 67 • 5 ♂♂, 3 ♀♀; Nova vas – Volčje; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (25 / 2006, rev. 2019); PMSL • 1 ♂; Runarsko; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (38 / 2006, rev. 2015); PMSL • 1 ♂, 1 ♀; Veliki vrh, Runarsko; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (36 / 2006, rev. 2015); PMSL • 7 ♂♂, 5 ♀♀; Volčje; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (28 / 2006, rev. 2015); PMSL. – VL 69 • 1 ♂; Veliki vrh pri Šmarju; 10 May 2008; L. Slana Novak and T. Novak leg. (156 / 2011); PMSL. – VL 74 • 1 ♂, 1 ♀; Mt. Goteniški Snežnik; 28 May 2000; L. Slana Novak and T. Novak leg. (247 / 2000, rev. 2007); PMSL • 1 ♀; Strma reber, Zgornji Čačiči; 27 Sep. 1996; L. Slana Novak and T. Novak leg. (LSN 126 / 1996, TN rev. 2008); PMSL. – VL 75 • 2 ♂♂; Draga; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (94 / 2011); PMSL • 1 ♂, 2 ♀♀; Mt. Goteniški Snežnik; 28 May 2000; L. Slana Novak and T. Novak leg. (253 / 2000, rev. 2007); PMSL. – VL 76 • 1 ♂; Bukovica, Ribnica; Jul. 1994; B. Oven leg. (LSN 25 / 1996, TN rev. 2009); PMSL • 2 ♂♂, 1 ♀; Lipovšica, Sodražica; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (100 / 2011, rev. 2018); PMSL • 1 ♂; Nova Štifta; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (105 / 2011); PMSL • 2 ♂♂, 3 ♀♀; Ribnica; 6 Sep. 2000; M. Mlakar leg. (828 / 2000, rev. 2008); PMSL • 1 ♂, 2 ♀♀; ibid.; (831 / 2000, rev. 2019); PMSL • 1 ♀; Mt. Turn; 27 Jun. 1980; B. Drovenik leg. (383 / 2002, rev. 2018); PMSL • 1 ♂; ibid.; 28 Aug. 1980; B. Drovenik leg. (394 / 2002, rev. 2009); PMSL. – VL 77 • 1 ♂; Dolnje Retje; 3 May 2008; L. Slana Novak and T. Novak leg. (161 / 2011); PMSL • 1 ♀; Rašica, Velike Lašče; 16 Jun. 1986 (261 / 2002, rev. 2015); PMSL • 1 ♂, 1 ♀; Velike Poljane, Ortnek; 3 May 2008; L. Slana Novak and T. Novak leg. (142 / 2011, rev. 2019); PMSL. – VL 78 • 1 ♂; Laporje; 3 May 2008; L. Slana Novak and T. Novak leg. (151 / 2011); PMSL • 1 ♀; Zagradec pri Grosupljem; 7 Jul. 1986; L. Slana Novak and T. Novak leg. (500 / 2002, rev. 2015); PMSL. – VL 79 • 1 ♀; Veliki Lipoglav − Dobrinje; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (467 / 2002, rev. 2015); PMSL. – VL 83 • 2 ♂♂; Vršič, Banjaloka; 24 Apr. 2001; S. Brelih leg. (121 / 2001, rev. 2019); PMSL. – VL 84 • 1 ♂; Jelenja vas; 3 May 2008; L. Slana Novak and T. Novak leg. (137 a / 2011); PMSL • 1 ♂; ibid.; (153 / 2011); PMSL • 1 ♂; Pleš, Borovec pri Kočevski Reki; 19 Sep. 2013; P. Kozel and T. Novak leg. (71 / 2013, rev. 2018); PMSL. – VL 85 • 1 ♀; Grčarice; 18 Oct. 2013; T. Novak leg. (63 / 2013, rev. 2015); PMSL • 1 ♂, 2 ♀♀; Grčarske Ravne; 18 Oct. 2013; T. Novak leg. (66 / 2013, TN rev. 2019); PMSL • 3 ♀♀; Kočevje; 6 Sep. 2000; M. Mlakar leg. (822 / 2000, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; (825 / 2000, rev. 2007); PMSL • 2 ♂♂, 1 ♀; Mt. Stojna, Kočevje; 25 Oct. 1996; S. Brelih leg. (92 / 2001, rev. 2008); PMSL. – VL 86 • 1 ♀; Dolenja vas; 3 May 2008; L. Slana Novak and T. Novak leg. (110 / 2011); PMSL • 1 ♂, 1 ♀; ibid.; (154 / 2012, rev. 2018); PMSL. – VL 87 • 1 ♂; Breg pri Zagradcu; 4 Nov. 1993; S. Brelih leg. (805 / 1998, rev. 2007); PMSL. – VL 89 • 1 ♀; Potok, Stična; 4 Mar. 1994; S. Brelih leg. (658 / 1998, rev. 2015); PMSL. – VL 93 • 1 ♂; Brsnik, Fara; 3 May 2008; L. Slana Novak and T. Novak leg. (157 / 2011, rev. 2018); PMSL. – VL 94 • 1 ♂, 1 ♀; Dolenja Žaga; 29 Apr. 2001; S. Brelih leg. (131 / 2001, rev. 2015); PMSL • 1 ♀; Dolenja Žaga – Podstene; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (90 / 1999, rev. 2008); PMSL • 1 ♂, 1 ♀; Gorenja Žaga; 3 May 2008; L. Slana Novak and T. Novak leg. (111 a / 2011); PMSL • 1 ♂, 1 ♀; ibid.; (159 / 2011); PMSL • 1 ♀; Ograja; 3 May 2008; L. Slana Novak and T. Novak leg. (109 a / 2011); PMSL • 1 ♀; ibid.; (121 a / 2011); PMSL • 1 ♀; ibid.; (144 / 2011); PMSL • 1 ♀; ibid.; (148 / 2011); PMSL • 1 ♂, 1 ♀; Ograja – Suhor − Morava; 3 May 2008; L. Slana Novak and T. Novak leg. (120 a / 2011); PMSL • 1 ♂, 1 ♀; ibid.; (138 a / 2011); PMSL • 1 ♂, 1 ♀; ibid.; (152 / 2011); PMSL • 1 ♂; Ograja – Zdihovo; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (76 / 1999, rev. 2008); PMSL • 3 juvs; ibid.; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (32 / 1999, rev. 2008); PMSL • 2 juvs; Podstene pri Kostelu; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (24 / 1999, rev. 2008); PMSL • 1 ♂; Rajndol; 21 Mar. 1994; S. Brelih leg. (821 / 1998, rev. 2008); PMSL • 5 ♂♂, 3 ♀♀; Srednji Potok, Banja Loka; 8 Oct. 2012; P. Kozel and T. Novak leg. (68 / 2015); PMSL • 1 ♂, 1 ♀, 2 juvs; Štalcerski preval; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (33 / 1999, rev. 2008); PMSL. – VL 95 • 1 ♂, 2 ♀♀; Dolga vas; 17 Aug. 1985; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 97 / 1986, TN rev. 2007); PMSL • 1 ♂, 1 ♀; Livold; 3 May 2008; L. Slana Novak and T. Novak leg. (123 a / 2011); PMSL • 1 ♂, 1 ♀; ibid.; (158 / 2011); PMSL. – VL 97 • 1 ♀; Dvor, Lašče; 10 May 2008; L. Slana Novak and T. Novak leg. (112 / 2011); PMSL • 1 ♂, 2 ♀♀; Železno, Dobrnič; 4 Nov. 1993; S. Brelih leg. (812 / 1998, rev. 2007); PMSL. – VL 99 • 3 ♀♀; Nova gora, Čatež; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (61 / 2007, rev. 2015); PMSL. – VM 80 • 1 ♂; Litija; 10 Sep. 2000; M. Mlakar leg. (809 / 2000, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; (834 / 2000, rev. 2015); PMSL. – WL 03 • 2 ♂♂; Dolenje Kozice; 18 Apr. − 13 May 2002; B. Drovenik and A. Gergeli leg. (509 / 2018); PMSL • 1 ♂; ibid.; 18 Apr. − 13 May 2002; B. Drovenik and A. Gergeli leg. (550 / 2018); PMSL • 1 ♂; ibid.; 5 Jul. 2002; B. Drovenik and A. Pirnat leg. (1135 / 2018); PMSL • 1 ♀; ibid.; 6 Aug. 2002; B. Drovenik and A. Pirnat leg. (1090 / 2018); PMSL • 1 ♀; ibid.; 30 Sep. – 28 Oct. 2002; B. Drovenik and A. Gergeli leg. (PK 64 / 2012); PMSL • 1 ♀; ibid.; 25 Nov. 2002; B. Drovenik and A. Pirnat leg. (270 / 2018); PMSL • 2 ♀♀; ibid.; 25 Nov. − 23 Dec. 2002; B. Drovenik and A. Gergeli leg. (1027 / 2018); PMSL • 4 ♂♂, 5 ♀♀; Srednji Radenci; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 51 / 1986, rev. 2008); PMSL • 3 ♂♂, 4 ♀♀; Stari trg ob Kolpi; 21 Mar. 1994; S. Brelih leg. (851 / 1998, rev. 2008); PMSL • 5 ♂♂, 5 ♀♀; Dolenji Radenci – Breg pri Sinjem vrhu; 3 May 2008; L. Slana Novak and T. Novak leg. (135 a / 2011); PMSL • 5 ♂♂, 2 ♀♀; ibid.; (149 / 2011); PMSL. – WL 05 • 1 ♂, 3 ♀♀, 1 juv.; Črmošnjice; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (160 / 1997); PMSL • 1 ♀; Koprivnik, Kočevski Rog; 21 Mar. 1994; S. Brelih leg. (867 / 1998, rev. 2008); PMSL • 1 ♂, 2 ♀♀; Mt. Mirna gora; 8 Oct. 2012; P. Kozel and T. Novak leg. (59 / 2015); PMSL • 4 ♂♂, 4 ♀♀; ibid.; 8 Oct. 2012; P. Kozel and T. Novak leg. (66 / 2015); PMSL. – WL 06 • 1 ♂; Cerovec − Selišče, Podturn pri Dolenjskih toplicah; 10 May 2008; L. Slana Novak and T. Novak leg. (145 / 2011, rev. 2017); PMSL. – WL 07 • 1 ♂; Boršt pri Dvoru; 10 May 2008; L. Slana Novak and T. Novak leg. (113 / 2012); PMSL • 1 ♀; Prečna, Mirna; Jun. 2000; V. Jaćimović leg. (PK and TN 100 / 2020); PMSL. – WL 08 • 1 ♂; Rdeči kal, Dobrnič; 4 Nov. 1993; S. Brelih leg. (872 / 1998, rev. 2007); PMSL • 1 ♂, 1 ♀; Trebnje; 29 Aug. 2000; M. Mlakar leg. (787 / 2000, rev. 2018); PMSL • 1 ♂; ibid.; (787 a / 2000, rev. 2018); PMSL • 1 ♀; ibid.; (788 / 2000, rev. 2008); PMSL. – WL 13 • 4 ♀♀; Balkovci, Vinica; 15 Aug. 1999; M. Štangelj and T. Novak leg. (212 / 1999, rev. 2008); PMSL • 2 ♂♂, 2 ♀♀; Dolenji Radenci – Breg pri Sinjem vrhu; 3 May 2008; L. Slana Novak and T. Novak leg. (146 / 2011); PMSL • 4 ♂♂, 1 ♀; Kot pri Dramlju; 16 Aug. 1999; M. Štangelj and T. Novak leg. (207 / 1999, rev. 2015); PMSL • 3 ♂♂, 2 ♀♀; Špeharji, Vinica; 16 Aug. 1999; M. Štangelj and T. Novak leg. (201 / 1999, rev. 2006); PMSL • 6 ♂♂, 5 ♀♀; Vinica – Sinji vrh; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 59 / 1986, TN rev. 2007); PMSL • 2 ♂♂; Vukovci; 18 Apr. 1996; S. Brelih leg. (764 / 1998, rev. 2007); PMSL. – WL 14 • 1 ♂, 1 ♀; Bistrica, Črnomelj; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (65 / 1999, rev. 2017); PMSL • 1 ♀; Črnomelj; 15 Aug. 1999; M. Štangelj and T. Novak leg. (221 / 1999, rev. 2008); PMSL • 3 ♂♂, 3 ♀♀, 2 juvs; Desinec – Črnomelj; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (385 / 1998, rev. 2007); PMSL • 2 ♂♂, 2 ♀♀; Tanča gora; 21 Mar. 1994; S. Brelih leg. (861 / 1998, rev. 2015); PMSL • 5 ♂♂, 2 ♀♀; Sinji vrh, Vinica; 16 Aug. 1999; M. Štangelj and T. Novak leg. (205 / 1999, rev. 2007); PMSL. – WL 15 • 1 ♀; Krupa spring; 10 Mar. 1994; S. Brelih leg. (808 / 1998, rev. 2007); PMSL • 6 ♂♂, 3 ♀♀, 3 juvs; Lokve, Črnomelj; 2 Jul. 1995; L. Slana Novak and T. Novak leg. (105 / 1997, rev. 2009); PMSL • 3 ♂♂, 2 ♀♀; Mihelja vas; 18 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 55 / 1986, TN rev. 2017); PMSL • 1 ♀; Praprot; 6 Aug. 1997; T. Novak leg. (418 / 1998, rev. 2008); PMSL • 1 ♀; ibid.; (432 / 1998, rev. 2008); PMSL • 1 ♀; ibid.; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (37 / 1999, rev. 2008); PMSL • 1 ♀; ibid.; (110 / 1999, rev. 2008); PMSL • 2 ♂♂, 2 ♀♀; Stranska vas; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (108 / 1999, rev. 2007); PMSL • 2 ♂♂, 1 ♀; Vrčice; 10 May 2008; L. Slana Novak and T. Novak leg. (110 / 2012); PMSL • 1 ♂, 3 ♀♀; Vrčice – Semič; 8 Oct. 2012; P. Kozel and T. Novak leg. (64 / 2015); PMSL. – WL 16 • 2 ♀♀; Laze, Uršna Sela; 10 May 2008; L. Slana Novak and T. Novak leg. (103 / 2012, rev. 2015); PMSL • 1 ♀; ibid.; (108 / 2012, rev. 2018); PMSL. – WL 17 • 1 ♂, 1 ♀; Brod, Novo Mesto; 31 Aug. 2000; M. Mlakar leg. (811 / 2000, rev. 2007); PMSL • 1 ♂; Novo mesto; Karaman leg. (87 / 1983 microscopic preparation, JH det., TN rev. 2011); PMSL • 1 ♂, 1 ♀; ibid.; Karaman leg. (108 / 1983 microscopic preparation, JH det., TN rev. 2011); PMSL • 1 ♀; ibid.; (814 / 2000); PMSL • 1 ♂; ibid.; (153 / 2018 microscopic preparation, JH det., TN rev. 2018); PMSL • 1 ♀, 1 ♀; ibid.; 31 Aug. 2000; M. Mlakar leg. (811 / 2000, rev. 2007); PMSL • 1 ♂, 2 ♀♀; Ratež; 29 Apr. 1995; S. Brelih leg. (750 / 1998, rev. 2007); PMSL. – WL 18 • 6 ♂♂, 5 ♀♀; Dolenje Laknice; 26 Oct. 2013; T. Novak leg. (GR 4661, 4662, 4664, 4666, 4667, 4669, 4673, 4676, 4677 − 4679); PMSL. – WL 19 • 1 ♀; Boštanj – Mokronog – Križ; L. Slana Novak and T. Novak leg. (70 / 2007, rev. 2011); PMSL • 1 ♂; Jelovec; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (56 / 2007); PMSL. – WL 23 • 1 ♂; Zilje; 18 Apr. 1996; S. Brelih leg. (767 / 1998, rev. 2007); PMSL • 1 ♂, 2 ♀♀; Zilje – Vinica; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (364 / 1998, rev. 2007); PMSL. – WL 24 • 5 ♂♂, 1 ♀, 1 juv.; Gorenjci pri Adlešičih; 15 Aug. 1999; M. Štangelj and T. Novak leg. (218 / 1999, rev. 2007); PMSL • 1 ♂; Dragoši, Griblje; 18 Apr. 1994; S. Brelih leg. (733 / 1998, rev. 2007); PMSL • 4 ♂♂, 2 ♀♀; Griblje − Desinec; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (161 / 1997, rev. 2009); PMSL. – WL 25 • 1 ♂; Bečka jama cave, Cad. No. 6275, Boldraž; 9 Jun. 2004; V. Germovšek, T. Tkavc, A. Kocuvan and T. Novak leg. (21 / 2004, rev. 2007); PMSL • 1 ♂; Božakovo; 3 May 2004; L. Slana Novak and T. Novak leg. (48 / 2004, rev. 2007); PMSL • 2 ♂♂, 3 ♀♀; Bušinja vas; 18 Apr. 1996; S. Brelih leg. (743 / 1998, rev. 2007); PMSL • 1 ♂, 4 ♀♀; Kapljišče; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 113 / 1986, TN rev. 2007); PMSL • 2 ♂♂, 1 juv.; ibid. (LSN 119 / 1986, TN rev. 2007); PMSL • 3 ♀♀; Metlika; 31 Aug. 2000; M. Mlakar leg. (816 / 2000, rev. 2007); PMSL • 1 ♂; ibid.; (818 / 2000, rev. 2015); PMSL • 7 ♂♂, 9 ♀♀; Trnovec, Metlika; 23 Jul. 1999; L. Slana Novak and T. Novak leg. (141 / 1999, rev. 2008); PMSL • 1 ♂, 2 ♀♀; Sv. Urban, Metlika; 18 Apr. 1996; S. Brelih leg. (784 / 1998, rev. 2007); PMSL. – WL 26 • 1 ♂; Krašnji vrh; 9 Jun. 2004; V. Germovšek, T. Tkavc, A. Kocuvan and T. Novak leg. (37 / 2004, rev. 2007); PMSL • 1 ♂; Markučev Mlin, Krašnji vrh; 1 Jul. 2007; L. Slana Novak and T. Novak leg. (21 / 2007); PMSL • 4 ♀♀; ibid.; (17 / 2007, rev. 2011); PMSL. – WL 27 • 1 ♂; Gabrje, Novo Mesto; 11 Apr. 1984; I. Sivec leg. (875 / 1997, rev. 2018); PMSL • 1 ♀; Gabrje – Sv. Miklavž – Krvavi kamen, Gorjanci Mts; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (123 / 1999, rev. 2019); PMSL • 5 ♂♂, 6 ♀♀; Glažarica, Mt. Gorjanci; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (118 / 1999, rev. 2015); PMSL • 1 ♂, 1 ♀; ibid.; (134 / 1999, rev. 2018); PMSL • 1 ♀; Golo, Šentjernej; 29 Apr. 1995; S. Brelih leg. (757 / 1998, rev. 2007); PMSL • 1 ♀; Sv. Miklavž, Gorjanci Mts; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (128 / 1999, rev. 2019); PMSL. – WL 28 • 3 ♂♂, 7 ♀♀; Brezje pri Raki; 27 Jul. 2012; S. Samu, P. Kozel and T. Novak leg. (115 / 2012); PMSL • 1 ♂, 1 ♀; Podvrh, Raka; 18 Jul. 2012; P. Kozel, T. Kozel and T. Novak leg. (55 / 2012); PMSL • 9 ♂♂, 10 ♀♀; ibid.; (GR 3734 − 3746, 3791, TN det.); PMSL • 4 ♂♂, 11 ♀♀; Škocjan; 26 Oct. 2013; T. Novak leg. (GR 4680 − 4694, TN det.); PMSL. – WL 29 • 3 ♂♂, 1 juv.; Boštanj; 5 Jul. 1987; L. Slana Novak, M. Slana Novak and T. Novak leg. (433 / 1998, rev. 2007); PMSL • 6 ♂♂, 12 ♀♀; Dolnje Brezovo; 30 Jun. 2012; T. Novak leg. (GR 3670 − 3683, TN det.); PMSL • 1 ♂, 1 ♀; Sevnica; 29 Aug. 2000; M. Mlakar leg. (785 / 2000, rev. 2009); PMSL. – WL 37 • 1 ♂, 1 ♀; Grič; 29 Apr. 1995, S. Brelih leg. (763 / 1998, rev. 2008); PMSL • 2 ♂♂, 2 ♀♀; Velike Vodenice − Opatova gora; 15 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (144 / 1997, rev. 2015); PMSL. – WL 38 • 3 ♀♀; Krakovski gozd, Sajevce; 5 May 1993; S. Brelih leg. (792 / 1998, rev. 2015); PMSL • 68 ♂♂, 59 ♀♀; Dolenje Radulje – Raški vrh; 5 Sep. 2012; P. Kozel and T. Novak leg. (PK 98 / 2012); PMSL • 84 ♂♂, 78 ♀♀; ibid.; 17 Sep. 2012; P. Kozel and T. Novak leg. (PK 99 / 2012); PMSL • 73 ♂♂, 83 ♀♀; ibid.; 4 Oct. 2012; P. Kozel and T. Novak leg. (PK 100 / 2012); PMSL • 44 ♂♂, 38 ♀♀; ibid.; 29 Apr. 2013; P. Kozel and T. Novak leg. (PK 43 / 2013); PMSL • 74 ♂♂, 71 ♀; Podulce – Zabukovje pri Raki; 20 Aug. 2012; P. Kozel and T. Novak leg. (PK 96 / 2012); PMSL. – WL 39 • 1 ♂, 1 ♀; Arto, Brestanica; 18 Nov. 1994; S. Brelih leg. (189 / 2001, rev. 2017); PMSL • 3 ♂♂, 3 ♀♀; ibid.; 1 Jul. 2007; L. Slana Novak and T. Novak leg. (28 / 2007); PMSL • 5 ♂♂, 3 ♀♀; Kremen; 12 Oct. 2005; L. Slana Novak and T. Novak leg. (141 / 2005, rev. 2008); PMSL • 1 ♀; Krško; 11 Apr. 1978 (42 / 2019); PMSL • 1 ♂, 5 ♀♀; ibid.; (47 / 2019); PMSL • 2 ♀♀; ibid.; (63 / 2019); PMSL • 1 ♀; ibid.; M. Mlakar leg. (704 / 2000, rev. 2008); PMSL • 1 ♂; Stara vas; 24 Aug. 2000; M. Mlakar leg. (709 / 2000, rev. 2007); PMSL • 2 ♂♂; Stolovnik – Anže; 9 Oct. 2005; L. Slana Novak and T. Novak leg. (134 / 2005, rev. 2008); PMSL • 5 ♂♂, 7 ♀♀; ibid.; 14 Oct. 2005; L. Slana Novak and T. Novak leg. (154 / 2005, rev. 2007); PMSL. – WL 48 • 1 ♂; Brežice; 30 Aug. 2000; M. Mlakar leg. (794 / 2000, rev. 2008); PMSL • 1 ♂; ibid.; 30 Aug. 2000; M. Mlakar leg. (792 / 2000, rev. 2007); PMSL • 1 ♀; Vitovec, Velike Malence; Jul. 1994; M. Potokar leg. (289 / 1995, rev. 2008); PMSL. – WL 49 • 6 ♂♂, 1 ♀; Podsreda castle; 17 Jun. 2001; L. Slana Novak and T. Novak leg. (145 / 2001, rev. 2008); PMSL. – WL 59 • 3 ♂♂, 4 ♀♀; Stara vas; 14 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (141 / 1997, rev. 2007); PMSL • 1 ♂; ibid.; 24 Aug. 2000; M. Mlakar leg. (709 / 2000, rev. 2007); PMSL • 1 ♂; ibid.; (714 / 2000, rev. 2007); PMSL • 1 ♂; ibid.; (716 / 2000, rev. 2007); PMSL. – WM 01 • 3 ♀♀; Hrastnik; 30 Aug. 2000; M. Mlakar leg. (802 / 2000, rev. 2018); PMSL • 2 ♀♀; ibid.; (806 / 2000, rev. 2018); PMSL. – WM 10 • 1 ♂, 1 ♀; Lokavec, Zidani most; 26 May 2004; A. Kapla leg. (56 / 2004, rev. 2007); PMSL • 2 ♀♀; Radeče; 29 Aug. 2000; M. Mlakar leg. (796 / 2000, rev. 2015); PMSL • 1 ♀; ibid.; (801 / 2000, rev. 2015); PMSL. – WM 20 • 1 ♂; Mt. Lisca; 26 May 2004; A. Kapla leg. (54 / 2004, rev. 2007); PMSL • 1 ♂; ibid.; (62 / 2004, rev. 2007); PMSL. – WM 30 • 7 ♂♂, 8 ♀♀, 1 juv.; Šentvid pri Planini; 14 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 52 / 1983, TN rev. 2007); PMSL • 10 ♂♂, 4 ♀♀; Mt. Bohor; 20 Aug. 1994, L. Slana Novak and T. Novak leg. (66 / 1997, rev. 2008); PMSL • 3 ♂♂, 2 ♀♀; Tovornik, Mt. Bohor; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1134 / 2002, TN rev. 2007); PMSL • 1 ♂, 1 ♀; ibid.; (1149 / 2002, rev. 2007); PMSL • 4 ♂♂, 1 ♀; Zgornji Reštanj, Senovo; 20 May 2012; T. Novak leg. (53 / 2012); PMSL. – WM 40 • 1 ♂; Bistri graben, Kozje; 11 Jul. 2003; T. Novak leg. (160 / 2003, rev. 2007); PMSL • 2 ♀♀; ibid.; (162 / 2003, rev. 2018); PMSL • 1 ♂; ibid.; (170 / 2003, rev. 2007); PMSL • 1 ♂; ibid.; (165 / 2003, rev. 2007); PMSL • 1 ♂; ibid.; (170 / 2003, rev. 2007); PMSL • 3 ♂♂, 4 ♀♀, 1 juv.; ibid.; 16 Jun. 2019; L. Slana Novak and T. Novak leg. (6 / 2019); PMSL • 1 ♂, 1 juv.; ibid.; (11 / 2019); PMSL • 1 ♀; ibid.; (12 / 2019); PMSL • 1 ♂, 1 ♀, 1 juv.; ibid.; (13 / 2019); PMSL • 2 ♂♂, 2 ♀♀; Kozje; 14 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 72 / 1983, TN rev. 2011); PMSL • 1 ♀; ibid.; 31 Jul. 1995; S. Polak leg. (177 / 2001, rev. 2015); NM • 1 ♀; ibid.; 1 Aug. 1995; S. Polak leg. (168 / 2001); NM • 3 ♂♂, 2 ♀♀; ibid.; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1150 / 2002, rev. 2007); PMSL • 3 ♂♂, 5 ♀♀; Podsreda; 14 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 71 / 1983, TN rev. 2007); PMSL • 2 ♂♂, 3 ♀♀; Stara Glažuta; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1145 / 2002, rev. 2007); PMSL. – WM 41 • 1 ♂, 1 ♀; Podčetrtek; 10 Aug. 1996; L. Slana Novak and T. Novak leg. (64 / 1997, rev. 2018); PMSL. – WM 50 • 4 ♂♂, 1 ♀; Bistrica ob Sotli; 13 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 46 / 1983, TN rev. 2007); PMSL • 1 ♂, 1 ♀; Svete gore; 17 May 1997; L. Slana Novak and T. Novak leg. (2 / 1997, rev. 2008); PMSL • 1 ♂; ibid.; 29 Aug. 2009; C. Komposch leg.; PMSL • 11 ♂♂, 9 ♀♀; Orešje na Bizeljskem; 10 Aug. 1996; L. Slana Novak and T. Novak leg. (82 / 1997, rev. 2009); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	description	Description Male (holotype) BODY. Length 1.99, body width 1.48. CHELICERAE. Ch basal article (without Apo) length 0.60, relatively long, 2.4 times as long as wide at Ch max width at dorsal hump. Ch-Apo small, 0.3 times as high as basal article long, rounded-trapezoid, widest distally and highest posteriorly; Ch-Apo 0.18 high, max width 0.17, min width 0.10 (at the base), ventral margin straight, with 0.70 long and 0.02 deep indentation with 10 – 20 μm high tipped tubercles and denticles constituting a saw. Secretion field fronto-medially, apically. Distal article length 0.67, max width 0.24, movable finger length 0.29 (Figs 4 E, 6 E, 9 E). PEDIPALPS. Pa-Fe relatively thin (Pa-Fe min: max width ~ 1: 2.8), with variable armament: either (the main population) with 4 subequidistant tubercles, spines or long setae, or their combination in the distal Pa-Fe half, or alternatively (marginal population in SW Slovenia), with 2 – 3 distal stout thorns. Pa-Pt relatively thin (length: max width ~ 1: 4.2) (Figs 5 E, 6 E, 10 E). Pa article lengths in Table 8. PENIS. Pe length 1.57, base 0.57, glans 0.18, stylus 0.07 (Figs 7 E, 8 E). LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 1 − 1 − 2. Tarsomere leg-Fe formula I−IV: 10 − 16 − 9 − 10. Leg article lengths in Table 8. Female BODY. Body length 2.49, body width 1.76. CHELICERAE. Ch basal article length 0.65, 2.6 times as long as wide at dorsal hump, with low-arched dorsal margin in front of dorsal indentation, and low ventral round-triangular hump (Fig. 13 D). Distal article length 0.67, max width 0.20, movable finger length 0.31. PEDIPALPS. Pa-Tr with relatively high, symmetrically, slightly knee-like arched dorsal margin, Pa-Fe with conspicuous distal bonce, Pa-Ti elongated ellipsoid (Fig. 13 D). Pa article lengths in Table 8. OVIPOSITOR. Ovipositor length 0.81, Rec sem with one tubular and 12 elongated balloon-like vesicles (Fig. 11 E). LEGS. Leg article lengths in Table 8. Variability In eastern Slovenia, males mostly have four thin medio-ventral spines of similar size on the Pa-Fe, the proximal one placed approximately in the middle of the Pa-Fe. In south-eastern Slovenia, some spines can be very small or diminished to a tubercle with a seta. In contrast, in south-western Slovenia, males have three or rarely two large, stout thorns, sometimes resembling those in N. b. bidentatum. Further investigation in Croatian Istria is required to elucidate the whole range of morphological variation and the expected transition between the tender eastern and robust western armament of the Pa-Fe.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	discussion	Remarks Nemastoma b. gruberi ssp. nov. is the most variable subspecies within N. bidentatum. Prior to this study, a few collected individuals had been considered within the variability range of N. b. sparsum, e. g., individuals from Buje and Vižinada considered as paratypes of N. b. sparsum (NHMW 4700). Accordingly, Novak et al. (2002) and Novak (2004 b, 2005 b) misidentified specimens, and those from SW Slovenia were erroneously reported (Novak 2005 b) as a disjunctive population of N. b. sparsum in the Sub Mediterranean region in Slovenia. Near the border of its distribution area in SW Slovenia, N. b. gruberi ssp. nov. has relatively uniform, robust Pa-Fe, with 3 subequidistant, rarely 2, large, stout thorns, much different from the armament in the main population.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	distribution	Distribution Croatia, Italy, Slovenia. Vertical distribution in Slovenia: 4 − 1619 m a. s. l. Type locality: Dolenje Laknice, Mokronog (45.93 ° N, 15.21 ° E, 237 m a. s. l.), Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7712BD05FE91FE31B92652D3.taxon	biology_ecology	Ecology Nemastoma b. gruberi ssp. nov. inhabits various habitat types in lowland and hilly land, from thermophile shrub and Quercetalia pubescenti – petraeae Klika 1928 communities in southwestern regions to riverine, floodplain and submontane woods in southern and south-eastern Slovenia. Phenology: adults eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	description	urn: lsid: zoobank. org: act: 7 F 207 FF 7 - 9241 - 4854 - A 4 C 3 - B 7 DED 507 F 740 Figs 2, 4 F, 5 F, 6 F, 7 F, 8 F, 9 F, 10 F, 11 F, 12 F, 13 E; Table 9	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	diagnosis	Diagnosis Species of the N. b. complex with highly arched, symmetrical Pa-Tr dorsal margin, concave dorsal margin of Pa-Ti and equilateral triangular glans, and Rec sem of 5 elongated vesicles attached to a bladder.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	etymology	Etymology The species name ʻ kozari ʼ is dedicated to Senad Kozar (Travnik), whose friendship and hospitality helped us in a searching expedition on the Vlašić planina Mts.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	materials_examined	Material examined Holotype BOSNIA AND HERZEGOVINA – YK 10 • 1 ♂; Mrazišče, the Vlašić planina Mts, Travnik; 44.28 ° E, 17.63 ° N; 1550 m a. s. l.; 1 Oct. 2019; S. Kozar, J. Podlesnik, P. Kozel and T. Novak leg.; light submontane beech and fir forest litter; PMSL-Opiliones-TN 137 / 2019. Paratypes BOSNIA AND HERZEGOVINA – YK 10 • 2 ♂♂, 1 ♀; same collection data as for holotype; PMSL- Opiliones-TN 137 / 2019. Other material BOSNIA AND HERZEGOVINA – YK 10 • 4 ♂♂, 10 ♀♀; Oštrike, Vlašić Planina Mts; 1 Oct. 2019, S. Kozar, J. Podlesnik, P. Kozel and T. Novak leg. (GR 5534 – 5543, GR 5547 – 5550); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	description	Description Male (holotype) BODY. Length 1.90, body width: 1.41. CHELICERAE. Ch basal article (without Apo) length 0.61, 2.0 times as long as wide at Ch max width at dorsal hump. Ch-Apo small, 0.24 times as high as basal article length, rounded-trapezoid, widest distally; Ch-Apo height 0.14, max width 0.12, min width 0.09 (at the base), ventral margin slightly concave. Secretion field apically. Distal article length 0.60, max width 0.19, movable finger length 0.23 (Figs 4 F, 6 F, 9 F). PEDIPALPS. Pa-Tr relatively very high (Pa-Tr height: length ~ 0.7: 1), Pa-Fe moderately robust (Pa-Fe min: max width ~ 1: 2.7), with 4 subequidistant stout spines in the distal Pa-Fe third. Pa-Pt relatively thin (length: max width ~ 1: 4.5), Pa-Ti dorsally conspicuously concave (Figs 5 F, 6 F). Pa article lengths in Table 9. PENIS. Pe length 1.55, base 0.60, glans 0.13, stylus 0.04 (Figs 7 F, 8 F). LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 1 − 1 − 2. Tarsomere leg formula I−IV: 8 − 14 − 8 − 9. Leg article lengths in Table 9. Female BODY. Length 2.03, body width 1.67. CHELICERAE. Ch basal article length 0.63, 2.4 times as long as wide at dorsal hump, with slightly arched dorsal margin in front of indentation, and slightly concave ventral margin (Fig. 13 E). Distal article length 0.66, max width 0.23, movable finger length 0.28. PEDIPALPS. Pa-Tr with moderately high, arched dorsal margin, Pa-Fe with conspicuous distal bonce, Pa-Pt and Pa-Ti elongated (Fig. 13 E). Pa article lengths in Table 9. OVIPOSITOR. Ovipositor length 0.93, Rec sem consisted of bladder and 5 elongated sac-like vesicles (Fig. 11 F). LEGS. Leg lengths in Table 9.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	distribution	Distribution Bosnia and Herzegovina, probably endemic to the Vlašić planina Mts. Vertical distribution: 1550 − 1630 m a. s. l. Type locality: Mrazišče, Mt. Paljenik in the Vlašić planina Mts, Travnik (44.28 ° N, 17.63 ° E, 1550 m a. s. l.), Bosnia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771BBD07FE28FE8CB8DB500F.taxon	biology_ecology	Ecology Nemastoma kozari sp. nov. individuals have been found in light submontane beech and fir forests, but may inhabit also other vegetation types. Phenology: adults probably eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	description	urn: lsid: zoobank. org: act: CCDC 54 EF-D 41 A- 4547 - A 4 D 4 - 50370212 C 243 Figs 2 – 3, 4 G, 5 G, 6 G, 7 G, 8 G, 9 G, 10 G, 11 G, 12 G, 13 F; Table 10	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	diagnosis	Diagnosis Subspecies of Nemastoma bidentatum with Ch basal article with a saw-like series of 1 – 3 μm high denticles on anterior margin of ventral hump, and a row of 5 – 11 denticles and tubercles in the distal half of Pa-Fe. Rec sem of 12 – 14 slightly elongated balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	etymology	Etymology The subspecies name ʻ martensi ʼ is dedicated to Jochen Martens (Mainz), our teacher, colleague and friend, who provided the first modern revision of harvestmen in Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	materials_examined	Material examined Holotype SLOVENIA – VL 23 • 1 ♂; Poljane pri Podgradu; 45.50 ° N, 14.10 ° E; 597 m a. s. l.; 25 Sep. 2011; L. Slana Novak and T. Novak leg.; thermophile scrub and mixed forest litter sift; PMSL-Opiliones-TN 287 / 2011. Paratypes SLOVENIA – VL 23 • 8 ♂♂, 5 ♀♀; same collection data as for holotype; PMSL-Opiliones-TN 287 / 2011. Other material SLOVENIA – VL 15 • 3 ♂♂, 8 ♀♀; Buje; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (316 / 2011, rev. 2015); PMSL • 2 ♂♂; ibid.; (316 a / 2011); PMSL • 1 ♀; Mt. Ostrič; 29 Aug. 2014; L. Slana Novak and T. Novak leg. (32 / 2014); PMSL. – VL 23 • 4 ♂♂, 1 ♀; Poljane pri Podgradu; 15 May 2011; L. Slana Novak and T. Novak leg. (40 / 2012); PMSL • 1 ♂; ibid.; 18 Aug. 2011; (165 / 2012); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 20 May 2012; L. Slana Novak and T. Novak leg. (GR 3597 − 3601, TN det.); PMSL • 1 ♂, 2 ♀♀; ibid.; 24 Oct. 2013; L. Slana Novak and T. Novak leg. (GR 6017, 6018, 6020, TN det.); PMSL • 2 ♂♂, 4 ♀♀; ibid.; 28 Oct. 2013; L. Slana Novak and T. Novak leg. (GR 4651, 4653, 4654, 4656, 4657, 4659, TN det.); PMSL • 1 ♂, 1 ♀; Mt. Velika Medvižica; 18 Jul. 2014; L. Slana Novak and T. Novak leg. (15 / 2014); PMSL. – VL 24 • 1 ♂; Mala Plešivica, Golac; 20 Sep. 2012; T. Novak leg. (169 / 2012); PMSL • 1 ♂; ibid.; 5 Oct. 2012; L. Slana Novak and T. Novak leg. (PK 20 / 2012, TN rev. 2018); PMSL • 1 ♀; ibid.; 21 May 2014; P. Kozel and T. Novak leg. (PK 18 / 2014); PMSL • 1 ♀; ibid.; (PK 18 / 2014); PMSL • 1 ♂; Mt. Mala Pleševica – Mt. Lipica; 19 Oct. 2014; T. Novak leg. (26 / 2015); PMSL • 1 ♂; Obrov – Golac; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (253 / 2011); PMSL • 1 ♂; Mt. Velika Pleševica; 19 Oct. 2014; T. Novak leg. (14 / 2015); PMSL • 1 ♂, 1 ♀; ibid.; (16 / 2015); PMSL • 1 ♂; Velika vrata, Mt. Velika Pleševica; 6 Oct. 2012; L. Slana Novak and T. Novak leg. (186 / 2012); PMSL • 4 ♂♂, 2 ♀♀; ibid.; (PK 16 / 2012, TN rev. 2018); PMSL. – VL 25 • 1 ♀; Buje – Suhorje; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (298 / 2011); PMSL. – VL 26 • 3 ♂♂; Fameljska loza, Senožeče; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (351 / 2011); PMSL. – VL 33; PMSL • 3 ♂♂; Starod; 25 Sep. 2011; L. Slana Novak and T. Novak leg. (269 / 2011); PMSL. – VL 34 • 1 ♂, 1 ♀; Harije; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (224 a / 2011); PMSL. – VL 43 • 2 ♀♀; Novokrajska jama cave, Cad. No. 810, Novokračine; 21 Jul. 2006; P. Trontelj and M. Zagmajster leg. (PK and TN 96 / 2020); PMSL • Novokračine; 2 ♂♂, 2 ♀♀; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (307 / 2011, rev. 2015); PMSL. – VL 44 • 1 ♂, 1 ♀; Jablanica – Trpčane; 9 Sep. 2011; L. Slana Novak and T. Novak leg. (252 / 2011, rev. 2015); PMSL. – VL 53 • 1 ♂, 1 ♀; Mt. Šešnovica; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (300 / 2011); PMSL. – VL 54 • 1 ♀; Bezgovec, Zabiče; 13 Nov. – 10 Dec. 2002; B. Drovenik and A. Gergeli leg. (PK 50 / 2012; TN rev. 2018); PMSL • 1 ♀; ibid.; 16 Oct. – 13 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 58 / 2012; TN rev. 2018); PMSL • 1 ♂; Mt. Snežnik; 12 Sep. 2018; L. Slana Novak and T. Novak leg. (116 b / 2018); PMSL. – VL 84 • 1 ♂, 1 ♀; Pleš, Borovec pri Kočevski Reki; 19 Sep. 2013; T. Novak leg. (60 / 2013, rev. 2015); PMSL. – VL 87 • 1 ♂; Veliki Grintovec, Zagradec, Žužemberk; 10 May 2008; L. Slana Novak and T. Novak leg. (117 / 2011, rev. 2015); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	description	Description Male (holotype) BODY. Length 1.63, width 1.26. CHELICERAE. Ch basal article (without Apo) length 0.57, 2.5 times as long as wide at Ch max width at dorsal hump, and with a saw-like series of 1 – 3 μm high denticles on anterior margin of ventral hump (not visible on Figs). Ch-Apo small, 0.31 times as high as basal article length, rounded-trapezoid, widest distally, highest posteriorly; Ch-Apo height 0.18, max width 0.15, min width 0.09 (at the base), frontal margin quarter moon-like. Secretion field medially. Distal article length 0.58, max width 0.19, movable finger length 0.23 (Figs 4 G, 5 G, 6 G). PEDIPALPS. Pa-Tr oval-trapezoid, moderately high (Pa-Tr height: length ~ 0.6: 1), Pa-Fe moderately robust (Pa-Fe min: max width ~ 1: 3.0), with 7 subequidistant low, stout, irregularly shaped tubercles and denticles in the distal Pa-Fe half. Pa-Pt moderately robust (length: max width ~ 1: 4.0) (Figs 5 G, 6 G). Pa article lengths in Table 10. PENIS. Pe length 1.44, base 0.45, glans 0.12, stylus 0.07 (Figs 7 G, 8 G). LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 1 − 1 − 2. Tarsomere leg-Fe formula I−IV: 8 − 17 − 8 − 9. Leg article lengths in Table 10. Female BODY. Length 2.23, width 1.60. CHELICERAE. Ch basal article length 0.60, 2.9 times as long as wide at dorsal hump, with slightly, evenly arched dorsal margin in front dorsal indentation, and medial straight portion in middle of slightly concave ventral margin (Fig. 13 F). Distal article length 0.63, max width 0.20, movable finger length 0.23. PEDIPALPS. Pa-Tr with moderately high, arched, with straight posterior portion of dorsal margin, Pa-Fe slightly widened distally; Pa-Pt, Pa-Ti and Pa-Ta elongated (Fig. 13 F). Pa article lengths in Table 10. OVIPOSITOR. Ovipositor length 0.94, Rec sem of 12 – 14 slightly elongated vesicles (Fig. 11 G). LEGS. Leg article lengths in Table 10.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	distribution	Distribution Croatia, Slovenia. Vertical distribution in Slovenia: 320 − 1030 m a. s. l. Type locality: Poljane pri Podgradu (45.50 ° N, 14.10 ° E, 597 m a. s. l.), Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771ABD01FEF3FAF1BD85535F.taxon	biology_ecology	Ecology Nemastoma b. martensi ssp. nov. is native to lowland to submontane habitats with thermophile scrub and Quercetalia pubescenti – petraeae and Tilia platyphyllos Scop. forest communities, with partly substantial admixture of Fagus sylvatica and Corylus avellana L. in southwestern Slovenia. Phenology: adults probably eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	description	urn: lsid: zoobank. org: act: 389 E 4 C 27 - 24 B 9 - 44 F 9 - A 724 - 11475 CB 9 E 12 B Figs 2 – 3, 4 H, 5 H, 6 H, 7 A, H, 8 A, H, 9 H, 10 H, 11 H, 12 H, 13 G; Table 11	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	diagnosis	Diagnosis Subspecies of Nemastoma bidentatum with elongated oval-trapezoid Ch-Apo, sagittally shallowly bilobe apically, wider distally, with a prominent apical pinnacle posteriorly, and conspicuously club-shaped Pa- Fe, with dorsal margin bent in the middle and nearly straight anterior and posterior portions, and evenly arched ventral margin, with 3 stout spines in distal fourth, and low, inconspicuous Pa-Ti hump. Rec sem of one tubular and 12 slightly elongated vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	etymology	Etymology The subspecies name ʻ schmidti ʼ is dedicated to Ferdinand Joseph Schmidt (1791 – 1878; Ljubljana), who collected the first specimens of this taxon preserved in his arthropod collection in the Slovenian Museum of Natural History, Ljubljana.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	materials_examined	Material examined Holotype SLOVENIA – VM 31 • 1 ♂; Dolenja Sevnica, Poljane nad Škofjo Loko; 46.12 ° N, 14.15 ° E; 517 m a. s. l.; 27 Mar. 2018; T. Novak leg.; mixed forest litter; PMSL-Opiliones-TN 1145 / 2018. Paratypes SLOVENIA – VM 31 • 2 ♀♀; same collection data as for holotype; PMSL-Opiliones-GR 3809. Other material AUSTRIA – VM 74 • 1 ♂; Eisenkappel / Železna Kapla – Ebriach / Obirsko, Carinthia; 7 May 2018; T. Novak leg. (38 / 2018); PMSL • 1 ♀; Seilach / Sele – Ebriach / Obirsko, Carinthia; 7 May 2018; T. Novak leg. (41 / 2018); PMSL. CROATIA – WL 67 • 1 ♂, 1 ♀; Mt. Medvednica; A. Schönhofer leg. (6 / 2012, rev. 2019); PMSL. ITALY – UM 52 • 7 ♂♂, 7 ♀♀; Artegna; 4 Mar. − 11 Aug. 1992; F. Gasparo leg. (64 / 2002, rev. 2017); PMSL. – UM 62 • 1 ♂; Alta Val Torre; 5 Aug. 1992; G. Governatori leg. (1062 / 2003, rev. 2017); PMSL • 1 ♂, 1 ♀; Villanova, Lusevera; 23 Oct. 1997; G. Governatori leg. (244 / 2003); PMSL • 3 ♂♂, 4 ♀♀; Tarcento / Čenta; 19 Aug. 2001; L. Slana Novak and T. Novak leg. (259 / 2001, rev. 2017); PMSL • 1 ♂, 5 ♀♀; ibid.; (257 / 2001, rev. 2018); PMSL • 1 ♂; Valle del Torre Vedronca valley; 5 Aug. 1992; G. Governatori leg. (1062 / 2003, rev. 2017); PMSL. – UM 65 • 2 ♂♂, 1 ♀; Pontebba; 19 Aug. 2001; L. Slana Novak and T. Novak leg. (240 / 2001, rev. 2017); PMSL. – UM 80 • 4 ♂♂; Altana / Utana; 26 Aug. 2001; L. Slana Novak and T. Novak leg. (283 / 2001, rev. 2017); PMSL. SLOVENIA • 1 ♂, 1 ♀ (Schmidt’s collection); (84 / 1985 microscopic preparation, JH det. sub N. triste, TN rev.); PMSL • 1 ♂, 1 ♀ (Schmidt’s collection); (89 / 1985 microscopic preparation, JH det. sub N. triste, TN rev.); PMSL. – UL 99 • 1 ♂, 2 ♀♀; Panovec; 17 Aug. 2000; T. Novak leg. (668 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; 18 Mar. 2001; S. Brelih leg. (63 / 2001, TN rev. 2019); PMSL • 9 ♂♂, 5 ♀♀; ibid.; 11 Nov. 2006; L. Slana Novak and T. Novak leg. (152 / 2006, rev. 2017); PMSL • 1 ♂; Replje; 8 Apr. 1992; S. Brelih leg. (649 / 1998, rev. 2017); PMSL. – UM 72 • 1 ♂; Mt. Kobariški Stol; 24 May 2003; L. Slana Novak and T. Novak leg. (720 a / 2003, TN and SL rev. 2017); PMSL. – UM 73 • 1 ♂; Mt. Kobariški Stol; 8 Jun. 2017; T. Novak leg. (6 / 2017); PMSL. – UL 82 • 1 ♀; Tavenica, Bela; 10 Oct. 1995; S. Brelih leg. (789 / 1998, rev. 2018); PMSL • 1 ♂; Sedlo – Mt. Kobariški Stol; 1 Aug. 1998; L. Slana Novak and T. Novak leg. (52 / 1999, rev. 2018); PMSL. – UM 90 • 2 ♀♀; Avče; 2 Aug. 2000; T. Novak leg. (298 / 2000, rev. 2019); PMSL • 2 ♂♂; Doblar; 2 Aug. 2000; leg. T. Novak (285 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; 17 Jun. 2001; L. Slana Novak and T. Novak leg. (151 / 2001, rev. 2017); PMSL • 1 ♂; Lepenka valley, Doblar; 25 Sep. 1998; L. Slana Novak and T. Novak leg. (1795 / 1998, rev. 2017); PMSL • 1 ♂; Levpa – Seniški breg; 2 Aug. 2000; T. Novak leg. (319 / 2000, rev. 2017); PMSL • 1 ♀; Marija Snežna, Avče; 2 Aug. 2000; T. Novak leg. (293 / 2000; rev. 2019); PMSL. – UL 91 • 1 ♂; Kamno; 31 Jul. 1998; L. Slana Novak and T. Novak leg. (1337 / 1998, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid.; 25 Sep. 1998; L. Slana Novak and T. Novak leg. (1809 / 1998, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Solarji, Livške Ravne; 8 Aug. 2000; T. Novak leg. (562 / 2000, TN and SL rev. 2017); PMSL. – UM 92 • 6 ♂♂, 1 ♀; Ladrski vrh; 14 May− 11 Jun. 2002; B. Drovenik and A. Gergeli leg. (482 / 2018); PMSL • 1 ♀; ibid.; 11 Jun. – 7 Aug. 2002; B. Drovenik and A. Pirnat leg. (792 / 2018); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 8 Jul. − 7 Aug. 2002; B. Drovenik and A. Pirnat leg. (PK 9 / 2013, TN rev. 2019); PMSL • 1 ♀; ibid.; (PK 10 / 2013, TN rev. 2019); PMSL • 1 ♀; ibid.; (585 / 2018); PMSL • 1 ♀; ibid.; 7 Aug. − 4 Sep. 2002; B. Drovenik and A. Pirnat leg. (780 / 2018); PMSL • 1 ♂; ibid.; (787 / 2018); PMSL • 2 ♂♂, 6 ♀♀; ibid.; 4 Sep. − 1 Oct. 2002; B. Drovenik and A. Pirnat leg. (827 / 2018); PMSL • 2 ♂♂; ibid.; (910 / 2018); PMSL • 4 ♀♀; ibid.; (915 / 2018); PMSL • 3 ♀♀; ibid.; (923 / 2018); PMSL • 2 ♀♀; ibid.; (928 / 2018); PMSL • 1 ♂; ibid.; (933 / 2018); PMSL • 3 ♂♂, 5 ♀♀; ibid.; 1 − 29 Oct. 2002; B. Drovenik and A. Pirnat leg. (959 / 2018); PMSL • 1 ♂, 3 ♀♀; ibid.; (961 / 2018); PMSL • 10 ♀♀; ibid.; (965 / 2018); PMSL • 4 ♀♀, 1 juv.; ibid.; (971 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (974 / 2018); PMSL • 3 ♂♂, 6 ♀♀; ibid.; (981 / 2018); PMSL • 8 ♀♀; ibid.; (1003 / 2018); PMSL • 1 ♂; ibid.; 29 Oct. − 7 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 36 / 2013, TN rev. 2019); PMSL • 3 ♀♀; ibid.; (PK 37 / 2013, TN rev. 2019); PMSL • 2 ♂♂, 13 ♀♀; ibid.; (871 / 2018); PMSL • 2 ♀♀; ibid.; (877 / 2018); PMSL • 1 ♂, 3 ♀♀; ibid.; (880 / 2018); PMSL • 2 ♀♀; ibid.; (881 / 2018); PMSL • 3 ♂♂, 4 ♀♀; ibid.; (897 / 2018); PMSL • 4 ♂♂, 6 ♀♀; ibid.; (900 / 2018); PMSL • 3 ♂♂, 1 ♀; ibid.; (939 / 2018); PMSL • 2 ♀♀; ibid.; 27 Nov. − 21 Dec. 2002; B. Drovenik and A. Gergeli leg. (948 / 2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; (950 / 2018); PMSL • 1 ♂; ibid.; (1010 / 2018); PMSL • 3 ♀♀; ibid.; (1014 / 2018); PMSL • 2 ♀♀; ibid.; (1019 / 2018); PMSL • 1 ♂; ibid.; (1026 / 2018); PMSL • 1 ♂, 2 ♀♀; ibid.; 21 Dec. 2002 − 28 Jan. 2003; B. Drovenik and A. Pirnat leg. (718 / 2018); PMSL • 1 ♂, 2 ♀♀; ibid.; (742 / 2018); PMSL • 2 ♂♂, 1 ♀; ibid.; (772 / 2018); PMSL • 3 ♀♀; ibid.; (778 / 2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; (813 / 2018); PMSL • 2 ♂♂, 3 ♀♀; ibid.; (1007 / 2018); PMSL • 1 ♂; ibid.; (1013 / 2018); PMSL. – UL 92 • 1 ♂, 1 ♀; near Kamno; Sep. 1915; Andreini leg.; PMSL • 1 ♂, 2 ♀♀; Selce; 10 Oct. 1995; S. Brelih leg. (786 / 1998, rev. 2017); PMSL • 1 ♂; Šturmi; 8 Aug. 2000; T. Novak leg. (545 / 2000, rev. 2017); PMSL. – UL 93 • 1 ♀; Bavšica; 9 Aug. 1996; S. Brelih leg. (744 / 1998, rev. 2018); PMSL • 3 ♀♀; Brvce; 11 Oct. 1995; S. Brelih leg. (745 / 1998, rev. 2018); PMSL. – VL 07 • 1 ♀; Velike Žablje; 17 Aug. 2000; T. Novak leg. (677 / 2000, rev. 2017); PMSL. – VL 09 • 1 ♂, 2 ♀♀; Čepovanska reber; 2 Aug. 2000; T. Novak leg. (344 / 2000, rev. 2017); PMSL • 1 ♂; Lijak spring, Ozeljan; 17 Apr. 2002; B. Drovenik and A. Gergeli leg. (219 / 2011, rev. 2017); PMSL • 1 ♂; ibid.; 17 Apr. – 14 May 2002; B. Drovenik and A. Pirnat leg. (540 / 2018); PMSL • 2 ♂♂; ibid.; (553 / 2018); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 14 May− 11 Jun. 2002; B. Drovenik and A. Gergeli leg. (503 / 2018); PMSL • 1 ♀; ibid.; (518 / 2018); PMSL • 1 ♂; ibid.; 11 Jun. 2002; B. Drovenik and A. Pirnat leg. (212 / 2011, rev. 2017); PMSL • 1 ♂; ibid.; 8 Jul. 2002; B. Drovenik and A. Pirnat leg. (1093 / 2018); PMSL • 1 ♀; ibid.; (1097 / 2018); PMSL • 1 ♀; ibid.; 7 Aug. − 4 Sep. 2002; B. Drovenik and A. Gergeli leg. (567 / 2018); PMSL • 1 ♂; ibid.; 1 Oct. 2002; B. Drovenik and A. Pirnat leg. (313 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; 18 Feb. 2003; B. Drovenik and A. Gergeli leg. (200 / 2011, TN rev. 2017); PMSL • 1 ♂; Lokve; 2 Aug. 2000; T. Novak leg. (350 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; (364 / 2000, rev. 2017); PMSL • 1 ♀; Smrečje, Trnovski gozd; 18 Aug. 1973; I. Sivec leg. (519 / 1981, JG det. 1982, TN rev. 2018); PMSL. – VL 16 • Divača – VL 19 • 1 ♂; Gorenja Kanomlja; 1 Aug. 2000; T. Novak leg. (401 / 2000, rev. 2017); PMSL • 1 ♀; Predmeja; 21 Aug. 2006; L. Slana Novak and T. Novak leg. (4 / 2006, rev. 2017); PMSL • 2 ♀♀; Srednja Kanomlja; 1 Aug. 2000; T. Novak leg. (391 / 2000, rev. 2017); PMSL. – VL 26 • 3 ♂♂, 3 ♀♀; Senožeče; 20 Sep. 2011; L. Slana Novak and T. Novak leg. (262 / 2011, rev. 2017); PMSL • 1 ♀; Mt. Vremščica; 10 Mar. 1994; S. Brelih leg. (680 / 1998, rev. 2017); PMSL. – VL 28 • 2 ♂♂, 1 ♀; Črni vrh – Col; L. Slana Novak and T. Novak leg. (239 / 2011, rev. 2017); PMSL. – VL 29 • 3 ♂♂, 3 ♀♀; Godovič – Črni vrh; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (241 / 2011, rev. 2017); PMSL • 2 ♂♂, 1 ♀; Gore; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (155 / 2000, rev. 2017); PMSL • 1 ♀; Idrija; Jun. 1997; M. Vončina leg. (1702 / 1998, rev. 2017); PMSL • 1 ♂; ibid.; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (137 / 2000, rev. 2017); PMSL • 8 ♂♂, 1 ♀; ibid.; (229 / 2000, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 1 Aug. 2000; T. Novak leg. (382 / 2000, rev. 2017); PMSL • 2 ♂♂; Razpotje; 1 Aug. 2000; T. Novak leg. (408 / 2000, rev. 2017); PMSL • 1 ♂; Spodnja Idrija; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (143 / 2000, rev. 2017); PMSL • 2 ♂♂; Srednja Kanomlja; 1 Aug. 2000; T. Novak leg. (389 / 2000, rev. 2017); PMSL • 1 ♀; ibid.; (397 / 2000, rev. 2017); PMSL • 1 ♂; Urbanovec; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (149 / 2000, rev. 2017); PMSL. – VL 36 • 1 ♂; Jama Sv. Janeza cave, Cad. No. 897, Prestranek; 11 Aug. 1952; E. Pretner leg. (56 / 1983, JH det., TN rev. 2017); PMSL • 1 ♀; ibid.; 29 Aug. 1954; E. Pretner leg. (174 / 1984, JH det., TN rev. 2018); PMSL • 1 ♂; ibid.; 11 Aug. 1957; E. Pretner leg. (221 / 1984, TN rev. 2017); PMSL • 3 ♀♀; Jernejeva jama cave, Cad. No. 929, Koče; 22 Apr. 2001; S. Vidmar, S. Polak, T. Novak leg. (157 / 2001, rev. 2008); PMSL • Postojna – VL 37 • 1 ♂; Belsko; 14 Apr. 2011; L. Slana Novak and T. Novak leg. (86 / 2011); PMSL • 1 ♂; Landol; 9 May 1994; S. Brelih leg. (888 / 1998); PMSL. – VL 38 • 2 ♂♂, 3 ♀♀; Dolenji Novi Svet; L. Slana Novak and T. Novak leg. (230 / 2011); PMSL • 1 ♀; Novi svet, Hotederšica; 22 Mar. 1993; S. Brelih leg. (855 / 1998, rev. 2019); PMSL. – VL 39 • 3 ♂♂, 3 ♀♀; Kugu, Rovte; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (244 / 2011); PMSL. – VL 46 • 1 ♂, 1 ♀; Mt. Javorniki; 9 May 1994; S. Brelih leg. (836 / 1998); PMSL • 2 ♂♂, 8 ♀♀; Mali Kožljek, Mt. Javorniki; 2 Oct. 2011; L. Slana Novak and T. Novak leg. (336 / 2011); PMSL. – VL 47 • 1 ♂, 2 ♀♀; Planina pri Rakeku; 24 Mar. 1992; F. Velkovrh leg. (221 / 1995, rev. 2017); PMSL • 1 ♂; Rakov Škocjan; Jul. 1994; K. Musar leg. (79 / 1995, rev. 2017); PMSL • 1 ♂; ibid.; 12. – 15 May 1994; S. Polak leg. (640 / 1998, rev. 2018); NM • 7 ♂♂, 17 ♀♀; ibid.; 10 Sep. 2009; A. Schönhofer leg. (3 / 2012, rev. 2017); PMSL. – VL 48 • 1 ♀; Bistra; 29 Mar. 1993; S. Brelih leg. (641 / 1998, rev. 2017); PMSL • 1 ♂; Laze; 16 Oct. − 21 Nov. 1994; M. Kuntner leg. (176 / 1995, rev. 2017); PMSL • 1 ♀; ibid.; Jul. – Aug. 1994; M. Kuntner leg. (20 / 1997, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; Aug. 1994; M. Kuntner leg. (183 / 1995, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; Dec. 1994 − Mar. 1995; M. Kuntner leg. (170 / 1995, rev. 2017); PMSL • 1 ♀; Vranja jama cave, Cad. No. 88, Laze pri Planini; 7 Jun. 1994; stud. biol. leg. (101 / 2019); PMSL. – VL 49 • 1 ♀; Retovje, Vrhnika; Jul. 1994, A. Valenčak leg. (17 / 1997, rev. 2017); PMSL • 1 ♂; ibid.; Jun. 1998; M. Zagmajster leg. (1493 / 1998, rev. 2017); PMSL • 1 ♀; Vrhnika; Jul. 1994; A. Valenčak leg. (17 / 1997, rev. 2017); PMSL. – VL 54 • 1 ♂; Mt. Snežnik; 21 Jul. 2001; B. Bertoncelj, U. Bertoncelj and T. Novak leg. (150 / 2001, rev. 2017); PMSL • 1 ♀; ibid.; 24 Nov. 2003; S. Polak leg. (120 / 2006, rev. 2018); NM; PMSL. – VL 56 • 1 ♂, 3 ♀♀; Cerkniško jezero; 19 Oct. 1994; S. Polak leg. (721 / 1998, rev. 2017); NM • 1 ♀; ibid.; 20 Nov. 1994; S. Polak leg. (687 / 1998, rev. 2017); NM • 2 ♂♂, 1 ♀; ibid.; 24 Apr. 2001; S. Brelih leg. (72 / 2001, rev. 2017); PMSL. – VL 57 • 2 ♂♂, 4 ♀♀; Dolenje Otave – Selšček; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (36 / 2011, rev. 2017); PMSL. – VL 58 • 1 ♂; Mt. Krim; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (42 / 2011, rev. 2017); PMSL • 2 ♀♀; Rakitna; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (40 / 2011, rev. 2017); PMSL. – VL 59 • 2 ♂♂, 1 ♀; Bevke; 20 Oct. 2018; L. Slana Novak and T. Novak leg. (136 / 2018); PMSL • 1 ♂; Brezovica pri Ljubljani; 25 May 1997; L. Slana Novak and T. Novak leg. (14 / 1997, rev. 2018); PMSL • 1 ♀; Draga pri Igu; 13 Apr. 1995; S. Brelih leg. (732 / 1998, rev. 2018); PMSL • 3 ♂♂, 10 ♀♀; Žabnica; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (33 / 2011, rev. 2017); PMSL. – VL 68 • 1 ♂, 3 ♀♀; Iška vas – Mt. Krim; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (48 / 2011, rev. 2017); PMSL • 1 ♂; ibid.; 31 Mar. 1997; S. Brelih leg. (53 / 2001, rev. 2017); PMSL • 1 ♀; Kremenica, Ig; 29 Jun. 1980 (1155 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 2 Mar. 1996; S. Brelih leg. (45 / 2001, rev. 2017); PMSL • 1 ♂; ibid.; 1 May 1997; S. Brelih leg. (101 / 2001, rev. 2017); PMSL • 1 ♂; ibid.; 7 Jun. 1997; S. Brelih leg. (117 / 2001, rev. 2017); PMSL • 1 ♂; ibid.; 10 Jul. 1997; S. Brelih leg. (139 / 2001, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; 25 Aug. 1997; S. Brelih leg. (113 / 2001, rev. 2017); PMSL • 1 ♀; Mt. Mokrec; 18 Jul. 1980 (LSN 151 / 1984, TN rev. 2017); PMSL • 1 ♀; Ščurki, Turjak; 3 May 2008; L. Slana Novak and T. Novak leg. (155 / 2011, rev. 2017); PMSL • 1 ♀; ibid.; 20 Jun. 1980; (1149 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; Škrilje; 20 Jun. 1980; (1159 / 1981, JG det., TN rev. 2018); PMSL. – VL 69 • 2 ♀♀; Šmarje Sap; 10 May 2008; L. Slana Novak and T. Novak leg. (119 a / 2011, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; (147 / 2011, rev. 2018); PMSL. – VL 79 • 1 ♂, 4 ♀♀; Brinjski hrib; L. Slana Novak and T. Novak leg. (99 / 2012, TN and SL rev. 2017); PMSL • 1 ♂; Stara vas; 16 Mar. 1993; S. Brelih leg. (665 / 1998, rev. 2017); PMSL. – VL 85 • 1 ♀; Mt. Stojna; 21 Oct. 1996; S. Brelih leg. (90 / 2001, rev. 2017); PMSL. – VL 89 • 1 ♂, 4 ♀♀; Vintarjevec; T. Novak leg. (173 / 2011, rev. 2017); PMSL • 2 ♂♂, 4 ♀♀; ibid.; (178 / 2011, rev. 2017); PMSL. – VL 93 • 1 ♀; Kostel, Dolenja Žaga; 16 Apr. 1996; S. Brelih leg. (737 / 1998, rev. 2018); PMSL. – VL 99 • 1 ♀; Cerovec, Gabrovka; L. Slana Novak and T. Novak leg. (74 / 2007, rev. 2017); PMSL. – VM 00 • 1 ♂, 2 ♀♀; Cvetrež, Voglarji; 2 Aug. 2000; T. Novak leg. (324 / 2000, TN and SL rev. 2017); PMSL • 5 ♂♂, 4 ♀♀; Čepovan; 1 Aug. 2000; T. Novak leg. (361 / 2000, rev. 2017); PMSL • 1 ♂; Čepovan – Most na Soči; 10 Jun. 1997; S. Brelih leg. (110 / 2001, rev. 2017); PMSL • 1 ♂; Preval, Lokovec, Čepovan; 2 Aug. 2000; T. Novak leg. (329 / 2000, rev. 2017); PMSL. – VM 05 • several specs; Podkoren; J. Hadži leg.; PMSL. – VM 09 • 1 ♀; Smrečje; 18 Aug. 1973; I. Sivec leg. (519 / 1981, JG det., TN rev. 2018); PMSL. – VM 10 • 1 ♂; Bukovo; 8 Aug. 2000; T. Novak leg. (501 / 2000, rev. 2017); PMSL • 3 ♂♂, 5 ♀♀; Oblakov vrh; 1 Aug. 2000; T. Novak leg. (377 / 2000, rev. 2017); PMSL • 1 ♂; Stopnik; 8 Aug. 2000; T. Novak leg. (492 / 2000, rev. 2018); PMSL. – VM 12 • 1 ♀; Šavnica, Bohinjska Bistrica; 26 Jul. 2002; B. Drovenik and A. Pirnat leg. (1094 / 2018); PMSL. – VM 13 • 1 ♀; Suha; 25 Oct. 1993; S. Brelih leg. (868 / 1998, rev. 2018); PMSL. – VM 14 • 1 ♀; Peričnik waterfall, Vrata valley; 14 Aug. 1928; J. Hadži leg.; PMSL. – VM 20 • 1 ♀; Fužine; 8 Aug. 2000; T. Novak leg. (484 / 2000, rev. 2017); PMSL • 1 ♂, 1 ♀; Govejk; 8 Aug. 2000; T. Novak leg. (486 / 2000, rev. 2017); PMSL • 1 ♀; Osojnica; 8 Aug. 2000; T. Novak leg. (498 / 2000, rev. 2017); PMSL • 2 ♂♂; Otalež; 8 Aug. 2000; T. Novak leg. (479 / 2000, rev. 2017); PMSL. – VM 21 • 3 ♂♂, 1 ♀; Davča; 27 Jul. 2003; L. Slana Novak and T. Novak leg. (703 / 2003, rev. 2017); PMSL • 1 ♀; ibid.; (706 / 2003, rev. 2017); PMSL • 1 ♂; Velbnik, Mt. Porezen; 7 Jul. 2003; L. Slana Novak and T. Novak leg. (700 / 2003); PMSL. – VM 22 • 1 ♀; Nomenj; 25 Oct. 1993; S. Brelih leg. (849 / 1998, rev. 2018); PMSL. – VM 23 • 2 ♀♀; the Pokljuka Mts; 19 Jul. 1940; J. Hadži leg. (323 / 1983, rev. 2017); PMSL. – VM 30 • 1 ♂, 1 ♀; Gorenja vas – Lučine; 15 May 1994; S. Brelih leg. (862 / 1998, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Srednja vas – Poljane; 8 Aug. 2000; T. Novak leg. (483 / 2000, rev. 2017); PMSL. – VM 31 • 15 ♂♂, 14 ♀♀; Dolenja Sevnica, Poljane; 15 Aug. 2012; L. Slana Novak and T. Novak leg. (GR 3777 − 3783, 3805 − 3811, 3819 − 3825, 3833 − 3835); PMSL. – VM 32 • 1 ♂, 2 ♀♀; Selca; 15 Aug. 2012; L. Slana Novak and T. Novak leg. (139 / 2012, TN and SL rev. 2017); PMSL. – VM 33 • 1 ♂; Begunje; 12 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (166 / 1997, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; (374 / 1998, rev. 2017); PMSL • Bled. – VM 40 • 5 ♂♂, 5 ♀♀; Polhov Gradec – Črni vrh; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (235 / 2011, rev. 2017); PMSL. – VM 41 • 2 ♀♀; Log nad Škofjo Loko; 8 Aug. 2000; T. Novak leg. (475 / 2000, rev. 2017); PMSL • 1 ♂; Mt. Lubnik; Jul. 1998; R. Potočnik leg. (1666 / 1998, rev. 2017); PMSL • 2 ♂♂, 6 ♀♀; Sopotniška grapa; 8 Aug. 2000; T. Novak leg. (505 / 2000, rev. 2017); PMSL • 1 ♂; Štinetova jama cave, Cad. No. 240, Crngrob, Škofja Loka; 25 Jul. 1980; T. Novak, I. Sivec leg. (LSN 214 / 1984, TN rev. 2019); PMSL. – VM 42 • 1 ♂; Bukovščica; 15 May 1994; S. Brelih leg. (881 / 1998, rev. 2017); PMSL • 1 ♂; Mt. Šmarjetna gora; 27 Apr. − 4 May 1987; A. Ziherl, T. Trilar leg. (492 / 1998, rev. 2017); PMSL • 1 ♀; ibid.; 18 − 24 Oct. 1987; A. Ziherl, T. Trilar leg. (483 / 1998, rev. 2017); PMSL • 1 ♂; ibid.; (513 / 1998, TN and SL rev. 2017); PMSL. – VM 43 • 1 ♀; Kriška planina; May 1968; (1218 / 1981, JG det., TN rev. 2017); PMSL • 1 ♂; Križe, Tržič; 6 Aug. 2015; L. Slana Novak and T. Novak leg. (101 / 2015, rev. 2017); PMSL. – VM 50 • 1 ♀; Ljubljana; Nov. 1986; (631 / 2002, rev. 2017); PMSL • 1 ♀; ibid.; Nov. 1987; (125 / 1997, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid.; Dec. 1987; F. Velkovrh leg. (106 / 1997, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; Mar. 1990; F. Velkovrh leg. (285 / 1995, rev. 2017); PMSL • 1 ♀; Podutik; Jun. 1998; (1648 / 1998, rev. 2017); PMSL • 1 ♀; ibid.; Oct. 1928; J. Staudacher leg. (16 / 1983 microscopic preparation, rev. 2017); PMSL • 1 ♀; ibid.; May – Jun. 2000; A. Pokrivač leg. (PK and TN 74 / 2020); PMSL • 1 ♀; Preval, Ljubljana; 18 Mar. 1993; S. Brelih leg. (664 / 1998, rev. 2017); PMSL. – VM 51 • 1 ♂; Mt. Grmada, Medno; 28 Jul. 1969; (531 / 1981, rev. 2018); PMSL • 2 ♂♂; ibid.; 3 Jun. 1986; (242 / 2002, rev. 2017); PMSL • 1 ♂; ibid.; 28 Jul. 1969; (531 / 1981, rev. 2018); PMSL • 1 ♂; ibid.; 15 May 1970; (526 / 1981, rev. 2018); PMSL • 1 ♀; ibid.; 19 Apr. − 15 May 2002; B. Drovenik and A. Gergeli leg. (188 / 2011, rev. 2017); PMSL • 1 ♀; ibid.; (446 / 2018); PMSL • 1 ♀; ibid.; 15 May 2002; B. Drovenik and A. Pirnat leg. (216 / 2011, rev. 2017); PMSL • 1 ♂; ibid.; 15 May− 12 Jun. 2002; B. Drovenik and A. Gergeli leg. (427 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (523 / 2018); PMSL • 1 ♂; ibid.; (558 / 2018); PMSL • 1 ♀; ibid.; 12 Jun. – 9 Jul. 2002; B. Drovenik and A. Gergeli leg. (602 / 2018); PMSL • 1 ♀; ibid.; (PK 34 / 2014, TN rev. 2019); PMSL • 2 ♂♂, 1 ♀; ibid.; 8 Aug. – 6 Sep. 2002; B. Drovenik and A. Gergeli leg. (PK 86 / 2012; TN rev. 2019); PMSL • 1 ♂; ibid.; 6 Sep. 2002; B. Drovenik and A. Pirnat leg. (354 / 2018); PMSL • 1 ♀; ibid.; 2 Oct. 2002 (173 / 2003, rev. 2017); PMSL • 1 ♀; ibid.; (1100 / 2018); PMSL • 2 ♀♀; ibid.; (1124 / 2018); PMSL • 1 ♀; ibid.; 28 Oct. – 25 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 28 / 2012, TN rev. 2019); PMSL • 1 ♂; ibid.; (264 / 2018); PMSL • 1 ♀; ibid.; 25 Nov. 2002; B. Drovenik and A. Pirnat leg. (277 / 2018); PMSL • 4 ♀♀; ibid.; (279 / 2018); PMSL • 1 ♂; ibid.; 19 Mar. 2003; B. Drovenik and A. Pirnat leg. (330 / 2018); PMSL • 1 ♀; ibid.; (375 / 2018); PMSL • 3 ♂♂, 2 ♀♀; Mavčiče; Jun. 1998; A. Žunič leg. (1568 / 1998, rev. 2017); PMSL. – VM 52 • 14 ♂♂, 12 ♀♀; Predoslje; 29 Jan. 1954; A. Polenec leg. (263 / 1984, rev. 2017); PMSL • 11 ♂♂, 23 ♀♀; ibid.; 31 Mar. 1954; A. Polenec leg. (337 / 1984, rev. 2017); PMSL • 8 ♂♂, 7 ♀♀; ibid.; 10 Oct. 1954; A. Polenec leg. (106 / 1984, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 15 Oct. 1954; A. Polenec leg. (351 / 1984, TN and SL rev. 2017); PMSL • 6 ♂♂, 14 ♀♀; ibid.; 24 Oct. 1954, A. Polenec leg. (63 / 1984, rev. 2017); PMSL • 4 ♂♂, 7 ♀♀; ibid.; 30 Oct. 1954; A. Polenec leg. (416 / 1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 4 ♀♀; ibid.; 6 Nov. 1954; A. Polenec leg. (556 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 11 ♀♀; ibid.; (319 / 1984, rev. 2017); PMSL • 2 ♂♂; ibid.; 14 Nov. 1954; A. Polenec leg. (345 / 1984, TN and SL rev. 2017); PMSL • 3 ♂♂, 9 ♀♀; ibid.; (393 / 1984, TN and SL rev. 2017); PMSL • 3 ♂♂, 1 ♀; ibid.; 28 Nov. 1954; A. Polenec leg. (364 / 1984, TN and SL rev. 2017); PMSL • 1 ♂, 4 ♀♀; ibid.; (410 / 1984, TN and SL rev. 2017); PMSL • 28 ♂♂, 32 ♀♀; ibid.; 5 Dec. 1954; A. Polenec leg. (382 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 18 Dec. 1954; A. Polenec leg. (20 / 1984, rev. 2017); PMSL • 32 ♂♂, 34 ♀♀; ibid.; (58 / 1984, rev. 2017); PMSL • 1 ♀; ibid.; 31 Dec. 1954; A. Polenec leg. (24 / 1984, rev. 2017); PMSL • 9 ♂♂, 21 ♀; ibid.; 9 Jan. 1955; A. Polenec leg. (324 / 1984, TN and SL rev. 2017); PMSL • 12 ♂♂, 8 ♀♀; ibid.; 15 Jan. 1955; A. Polenec leg. (209 / 1984, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; (210 / 1984, TN and SL rev. 2017); PMSL • 6 ♂♂, 17 ♀♀; ibid.; 22 Jan. 1955; A. Polenec leg. (342 / 1984, TN and SL rev. 2017); PMSL • 18 ♂♂, 15 ♀♀; ibid.; 7 Feb. 1955; A. Polenec leg. (103 / 1984, TN and SL rev. 2017); PMSL • 20 ♂, 27 ♀♀; ibid.; 12 Feb. 1955; A. Polenec leg. (242 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 15 Feb. 1955; A. Polenec leg. (328 / 1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 14 ♀♀; ibid.; 19 Feb. 1955; A. Polenec leg. (390 / 1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid.; 25 Mar. 1955; A. Polenec leg. (29 / 1984, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; (33 / 1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 7 ♀♀; ibid.; 27 Mar. 1955; A. Polenec leg. (285 / 1984, TN and SL rev. 2017); PMSL • 8 ♂♂, 12 ♀♀; ibid.; 29 Mar. 1955; A. Polenec leg. (99 / 1984, TN and SL rev. 2017); PMSL • 9 ♂♂, 10 ♀♀; ibid.; (277 / 1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 6 Apr. 1955; A. Polenec leg. (90 / 1984); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 9 Apr. 1955; A. Polenec leg. (229 / 1984, TN and SL rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; 8 May 1955; A. Polenec leg. (69 / 1984, TN and SL rev. 2017); PMSL • 4 ♂♂, 6 ♀♀; ibid.; (247 / 1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 5 ♀♀; ibid.; 21 May 1955; A. Polenec leg. (279 / 1984, TN and SL rev. 2017); PMSL • 9 ♂♂, 3 ♀♀; ibid.; 4 Jun. 1955; A. Polenec leg. (298 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; (333 / 1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid.; 23 Apr. 1955; A. Polenec leg. (313 / 1984, TN and SL rev. 2017); PMSL • 13 ♂♂, 11 ♀♀; ibid.; 21 Jun. 1955; A. Polenec leg. (50 / 1984, TN and SL rev. 2017); PMSL • 3 ♀♀; ibid.; 2 Jul. 1955 (379 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 27 Aug. 1955; A. Polenec leg. (253 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; (305 / 1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid.; 8 Sep. 1955; A. Polenec leg. (258 / 1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 12 ♀♀; ibid.; 8 Oct. 1955; A. Polenec leg. (362 / 1983, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid.; 22 Oct. 1955; A. Polenec leg. (383 / 1983, TN and SL rev. 2017); PMSL • 3 ♂♂, 4 ♀♀; ibid.; (389 / 1983, rev. 2017); PMSL • 11 ♂♂, 9 ♀♀; ibid.; 9 Nov. 1955; A. Polenec leg. (91 / 1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 16 Apr. 1956; A. Polenec leg. (482 / 1983, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid.; 22 Apr. 1956; A. Polenec leg. (280 / 1983, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 26 May 1957; A. Polenec leg. (441 / 1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 4 Jun. 1956; A. Polenec leg. (289 / 1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 22 Jun. 1956; A. Polenec leg. (472 / 1983); PMSL • 1 ♂; ibid.; 8 Jul. 1956; A. Polenec leg. (208 / 1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 22 Jul. 1956; A. Polenec leg. (LSN 9 / 1983, TN rev. 2018); PMSL • 1 ♂; ibid.; (532 / 1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 10 Aug. 1956; A. Polenec leg. (256 / 1983, TN and SL rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; 24 Oct. 1956; A. Polenec leg. (537 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 11 ♀♀; ibid.; 29 Oct. 1956; A. Polenec leg. (508 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 1 ♀; ibid.; 30 Nov. 1956; A. Polenec leg. (423 / 1983, TN and SL rev. 2017); PMSL • 1 ♂, 5 ♀♀; ibid.; 16 Dec. 1956; A. Polenec leg. (196 / 1983, TN and SL rev. 2017); PMSL • 7 ♂♂, 11 ♀♀; ibid.; (544 / 1983, TN and SL rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 30 Dec. 1956; A. Polenec leg. (214 / 1983, TN and SL rev. 2017); PMSL • 4 ♀♀; ibid.; (295 / 1984, TN and SL rev. 2017); PMSL • 8 ♂♂, 10 ♀♀; ibid.; 20 Feb. 1957; A. Polenec leg. (449 / 1983, TN and SL rev. 2017); PMSL • 4 ♂♂, 10 ♀♀; ibid.; 23 Feb. 1957; A. Polenec leg. (418 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 5 ♀♀; ibid.; (215 / 1984, JH det., TN and SL rev. 2017); PMSL • 4 ♂♂, 10 ♀♀; ibid.; 8 Mar. 1957; A. Polenec leg. (247 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 12 ♀♀; ibid.; 6 Apr. 1957; A. Polenec leg. (491 / 1983, TN and SL rev. 2017); PMSL • 10 ♂♂, 11 ♀♀; ibid.; 9 Mar. 1957; A. Polenec leg. (465 / 1983, TN and SL rev. 2017); PMSL • 12 ♂♂, 4 ♀♀; ibid.; 3 May 1957; A. Polenec leg. (506 / 1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 5 ♀♀; ibid.; 18 May 1957; A. Polenec leg. (551 / 1983, TN and SL rev. 2017); PMSL. – VM 60 • 12 ♂♂, 29 ♀♀; Ljubljana; Jun. 1972; (518 / 1981, JG det., TN rev. 2020); PMSL • 1 ♂; ibid.; 26 May 1986; Hribernik, Koren and Mencinger leg. (255 / 2002, rev. 2017); PMSL. – VM 61 • 4 ♂♂, 1 ♀; Mt. Gobavica, Mengeš; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (97 / 2011, rev. 2017); PMSL • 1 ♀; Rašica; Jun. 1994; H. Urbas leg. (241 / 1995, TN rev. 2017); PMSL. – VM 62 • 1 ♂, 1 ♀; Stahovica – Ambrož; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (61 / 2011, rev. 2017); PMSL. – VM 63 • 1 ♀; Mt. Kalški greben; 12 Aug. 2009; L. Slana Novak and T. Novak leg. (28 / 2009, rev. 2017); PMSL • 1 juv.; ibid.; (31 / 2009, rev. 2017); PMSL • 1 ♂, 2 ♀♀; Kamnik; 29 Oct. 2000; N. Žmaher leg. (885 / 2000, rev. 2017); PMSL • 1 ♀; Mt. Koren; 8 Sep. 2009; L. Slana Novak and T. Novak leg. (50 / 2009, rev. 2017); PMSL • 1 ♂; Mt. Košutna; 22 Sep. 2009; L. Slana Novak and T. Novak leg. (68 / 2009, rev. 2017); PMSL • 1 ♀; Mt. Mokrica; 22 Sep. 2009; L. Slana Novak and T. Novak leg. (80 / 2009, rev. 2017); PMSL. – VM 70 • 1 ♂, 3 ♀♀; Jevnica; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (538 / 2002, rev. 2017); PMSL • 5 ♂♂, 3 ♀♀; Senožeti – Velika vas; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (55 / 2011, rev. 2017); PMSL • 2 ♂♂, 3 ♀♀; Spodnja Javoršica; 16 Aug. 2000; T. Novak leg. (603 / 2000, rev. 2017); PMSL • 1 ♂; Vinje; 16 Aug. 2000; T. Novak leg. (693 / 2000, rev. 2017); PMSL. – VM 71 • 1 ♂; Brezje pri Dobu; 16 Aug. 2000; T. Novak leg. (605 / 2000, rev. 2017); PMSL • 5 ♂♂, 1 juv.; Lukovica pri Domžalah; L. Slana Novak and T. Novak leg. (64 / 2011, rev. 2017); PMSL • 1 ♀; Prevoje pri Šentvidu, Šentvid pri Lukovici; 30 Mar. 1993; S. Brelih leg. (637 / 1998, rev. 2018); PMSL. – VM 72 • 7 ♂♂; Smrečje v Črni; L. Slana Novak and T. Novak leg. (73 / 2011, rev. 2017); PMSL. – VM 73 • 1 ♂, 1 ♀; Robanov kot; 4 Aug. 2015; T. Novak leg. (85 / 2015, rev. 2017); PMSL. – VM 74 • 2 ♀♀; Podolševa; 4 Aug. 2015; T. Novak leg. (83 / 2015, rev. 2017); PMSL. – VM 80 • 3 ♂♂, 3 ♀♀; Kresnice − Litija; 9 Jul. 1986; L. Slana Novak and T. Novak leg. (496 / 2002, rev. 2017); PMSL • 1 ♀; Ribče; 20 Nov. 1992; S. Brelih leg. (724 / 1998, rev. 2017); PMSL. – VM 81 • 6 ♂♂, 4 ♀♀; Mt. Limbarska gora; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (68 / 2011, rev. 2017); PMSL • 1 ♀; Negastern; 16 Aug. 2000; T. Novak leg. (593 / 2000, rev. 2017); PMSL • 1 ♀; Podmilj; 10 May 2008; L. Slana Novak and T. Novak leg. (106 / 2012, rev. 2017); PMSL. – VM 82 • 3 ♂♂, 1 ♀; Mt. Gradišče, Bočna, Gornji Grad; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (75 / 2011, rev. 2017); PMSL. – VM 83 • 1 ♂; Primož pri Ljubnem; 31 Mar. 1994; S. Brelih leg. (875 / 1998, rev. 2017); PMSL. – VM 84 • 1 ♀; Bistra; 4 Aug. 2015; T. Novak leg. (89 / 2015, rev. 2017); PMSL. – VM 85 • 1 ♂, 1 ♀; Najbrževo, Mt. Peca; 10 Aug. 1999; F. Janžekovič leg. (82 / 2002, JG det., TN rev. 2018); PMSL • 1 ♂; ibid.; 21 Oct. 1999; F. Janžekovič leg. (45 / 2003, rev. 2018); PMSL • 1 ♂, 2 ♀♀; Topla valley; 12 Jul. 2003; L. Slana Novak and T. Novak leg. (187 / 2003, rev. 2017); PMSL. – VM 90 • 1 ♀; Cerovec, Gabrovka; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (74 / 2007, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Renke; 9 Jul. 1986; L. Slana Novak and T. Novak leg. (518 / 2002, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; Sava; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (55 / 2007, rev. 2017); PMSL. – VM 91 • 1 ♂, 4 ♀♀; Mt. Čemšenik; 1 Jul. 2002; L. Slana Novak and T. Novak leg. (1202 / 2002, rev. 2018); PMSL • 5 ♂♂, 1 ♀; Mt. Črni vrh, Mt. Čemšeniška planina; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1171 / 2002, rev. 2018); PMSL • 1 ♀; Sv. Primož in Felicijan, Mt. Čemšeniška planina; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1197 / 2002, rev. 2017); PMSL. – VM 93 • 1 ♂, 1 ♀; Mt. Golte; 14 Dec. 1984; A. Polenec leg. (LSN 57 / 1985, TN rev. 2018); PMSL • 1 ♂; Žlabor; 31 Mar. 1994; S. Brelih leg. (886 / 1998, rev. 2017); PMSL. – VM 94 • 1 ♂, 1 ♀; Florjan; 6 Jul. 2003; L. Slana Novak and T. Novak leg. (196 / 2003, rev. 2018); PMSL • 1 ♂; Mt. Smrekovec; 18 Sep. 2000; (579 / 2002, rev. 2017); PMSL. – VM 95 • 3 ♂♂; Tolsti vrh, Ravne na Koroškem; 11 Oct. 2006; T. Novak leg. (55 / 2006, rev. 2019); PMSL • 4 ♂♂, 3 ♀♀; Mt. Uršlja gora; L. Slana Novak and T. Novak leg. (65 / 2006, TN and SL rev. 2017); PMSL. – VM 96 • 2 ♂♂, 3 ♀♀; Jamnica; 10 Oct. 2006; T. Novak leg. (11 / 2006, rev. 2019); PMSL • 1 ♂, 1 ♀; Zgornja Jamnica; Ravne na Koroškem; 10 Oct. 2006; T. Novak leg. (13 / 2006, rev. 2018); PMSL. – WL 07 • 1 ♂; Mali Lipovec; 10 May 2008; L. Slana Novak and T. Novak leg. (107 / 2011, rev. 2017); PMSL. – WL 19 • 1 ♂; Krmelj – Tržišče; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (66 / 2007, rev. 2017); PMSL. – WL 39 • 1 ♂; Arto; 18 Nov. 1994; S. Brelih leg. (188 / 2001, rev. 2017); PMSL. – WL 48 • 1 ♂, 4 ♀♀; Prilipe, Brežice; 26 May 1984; L. Slana Novak and T. Novak leg. (LSN 228 / 1984, TN rev. 2018); PMSL. – WM 00 • 1 ♀; Sopota valley, Podkum; 1 Apr. − 6 May 2002; B. Drovenik and A. Pirnat leg. (642 / 2018); PMSL • 1 ♀; ibid.; (648 / 2018); PMSL • 1 ♂; ibid.; 27 May 2002; B. Drovenik and A. Pirnat leg. (214 / 2011, ver. 2017); PMSL • 1 ♂; ibid.; 27 May− 1 Jul. 2002; B. Drovenik and A. Gergeli leg. (185 / 2011, rev. 2017); PMSL • 3 ♀♀; ibid.; (187 / 2011, rev. 2017); PMSL • 1 ♀; ibid.; (189 / 2011, rev. 2017); PMSL • 2 ♀♀; ibid.; 20 Aug. – 19 Sep. 2002; B. Drovenik and A. Gergeli leg. (PK 24 / 2012, TN rev. 2018); PMSL • 1 ♀; ibid.; (858 / 2018); PMSL • 1 ♂, 2 ♀♀; ibid.; (888 / 2018); PMSL • 1 ♂; ibid.; 19. Sep. 2002; B. Drovenik and A. Pirnat leg. (256 / 2018); PMSL • 1 ♀; ibid.; 19 Sep. − 15 Oct. 2002; B. Drovenik and A. Pirnat leg. (943 / 2018); PMSL • 1 ♀; ibid.; (985 / 2018); PMSL • 1 ♀; ibid.; 15 Oct. 2002; B. Drovenik and A. Pirnat leg. (1030 / 2018); PMSL • 1 ♀; ibid.; (1078 / 2018); PMSL • 2 ♀♀; ibid.; 15 Oct. 2002 – 12 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 81 / 2012, TN rev. 2018); PMSL • 1 ♂; ibid.; (898 / 2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 12 Nov. 2002 − 6 Mar. 2003; B. Drovenik and A. Pirnat leg. (685 / 2018); PMSL • 1 ♂; ibid.; (713 / 2018); PMSL • 1 ♀, 1 juv.; ibid.; (736 / 2018); PMSL • 1 ♂, 3 ♀♀; ibid.; (743 / 2018); PMSL • 1 juv.; ibid.; 6 Mar. 2003; B. Drovenik and A. Pirnat leg. (212 / 2018); PMSL • 1 ♀; ibid.; (238 / 2018); PMSL • 1 ♂; Šklendrovec, Mt. Kum; 22 Apr. 1992; S. Brelih leg. (727 / 1998, rev. 2017); PMSL. – WM 02 • 3 ♂♂, 1 ♀; Braslovško jezero, Braslovče; 26 Oct. 2013; T. Novak leg. (35 / 2013, rev. 2017); PMSL. – WM 03 • 3 ♂♂, 3 ♀♀; Andraž nad Polzelo; 26 Oct. 2013; T. Novak leg. (47 / 2013, rev. 2017); PMSL. – WM 04 • 6 ♂♂; Mislinjska Dobrava; 26 Oct. 2013; T. Novak leg. (37 / 2013, rev. 2018); PMSL • 1 ♀; Rdečka jama cave, Cad. No. 3488, Mt. Rdečki vrh; 10 Jul. 1977; V. Kuštor, T. Novak leg. (1119 / 1981, JG det., TN rev. 2018); PMSL. – WM 06 • 1 ♂, 1 ♀; Dom na Košenjaku; 14 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 103 / 1996, TN rev. 2004); PMSL • 1 ♂, 1 ♀; Mt. Košenjak; 14 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 103 / 1996, TN rev. 2019); PMSL • 2 ♂♂; Pernice; 18 Aug. 2005; T. Novak leg. (57 / 2005, rev. 2018); PMSL • 1 ♂; ibid.; 18 Aug. 2005; T. Novak leg. (66 a / 2005, rev. 2017); PMSL • 1 ♂; ibid.; 1 Aug. 2005; T. Novak leg. (69 / 2005, rev. 2019); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (86 / 2005, rev. 2017); PMSL • 1 ♂; ibid.; 22 Aug. 2005; L. Slana Novak and T. Novak leg. (108 / 2005, rev. 2017); PMSL • 1 ♂; ibid.; (110 / 2005, rev. 2017); PMSL • 1 ♀; ibid.; (112 / 2005, rev. 2017); PMSL • 2 ♂♂, 1 ♀; ibid.; (114 / 2005, rev. 2018); PMSL. – WM 13 • 1 ♂, 1 ♀; Lipje, Vinska Gora; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (741 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; Vodosteč, Črnova; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (737 / 2002, rev. 2019); PMSL • 1 ♂, 1 ♀; Zavrh nad Dobrno; 22 Apr. 1995; S. Brelih leg. (770 / 1998, rev. 2018); PMSL. – WM 14 • 2 ♀♀; Paka pri Velenju; 9 Jun. 2015; T. Novak leg. (50 / 2015, rev. 2018); PMSL. – WM 15 • 3 ♂♂, 1 ♀; Sv. Primož; 22 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 104 / 1996, TN rev. 2018); PMSL • 1 ♂, 1 ♀; Mt. Kremžarjev vrh; 26 May 2005; L. Slana Novak and T. Novak leg. (187 / 2005, rev. 2017); PMSL. – WM 16 • 2 ♂♂; Mt. Kapunar; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (60 / 2005, rev. 2018); PMSL • 1 ♂; Pernice; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (75 / 2005, rev. 2018); PMSL • 1 ♀; ibid.; 27 Apr. 2008; L. Slana Novak and T. Novak leg. (167 / 2011, rev. 2017); PMSL • 1 ♀; Prevalov graben, Muta; 27 Apr. 2008; L. Slana Novak and T. Novak leg. (150 / 2011, rev. 2017); PMSL • 1 ♀; Radlje ob Dravi; 2 Jul. 1939; (334 / 1983, rev. 2017); PMSL • 1 ♀; ibid.; 15 May 1966; I. Leitinger leg. (510 / 1981, rev. 2018); PMSL • 1 ♀; ibid.; (513 / 1981, JG det., TN rev. 2018); PMSL • 1 ♂; ibid.; 12 Jun. 1966; I. Leitinger leg. (514 / 1981, rev. 2018); PMSL • 1 ♂; ibid.; (517 / 1981, rev. 2018); PMSL • 1 ♀; ibid.; 10 Jul. 1966; I. Leitinger leg. (525 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 8 Aug. 1966; I. Leitinger leg. (1222 / 1981, JG det., TN rev. 2018); PMSL • 3 ♂♂; ibid.; 26 Aug. 1966; I. Leitinger leg. (511 / 1981, rev. 2018); PMSL • 1 ♀; ibid.; (512 / 1981, JG det., TN rev. 2018); PMSL • 2 ♀♀; ibid.; (515 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; (524 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 9 Sep. 1966; I. Leitinger leg. (516 / 1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; (671 / 1981, rev. 2017); PMSL • 1 ♀; ibid.; (152 / 1982, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 8 Oct. 1973; T. Novak leg. (994 / 1981, rev. 2018); PMSL • 3 ♂♂, 1 ♀; Sv. Trije Kralji; 19 Aug. 2005; T. Novak leg. (97 / 2005, rev. 2017); PMSL • 1 ♀; ibid.; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (82 / 2005, rev. 2017); PMSL • 2 ♂♂; ibid.; (73 / 2005, rev. 2018); PMSL. – WM 23 • 2 ♂♂, 1 ♀; Mt. Stenica; 25 Apr. 1982; T. Novak; G. Kolar leg. (267 / 1982, JG det., TN rev. 2018); PMSL. – WM 32 • 1 ♂; Razbor, Ponikva; 9 Jun. 2015; T. Novak leg. (54 / 2015, rev. 2017); PMSL. – WM 33 • 6 ♂♂, 7 ♀♀; Preloge; 17 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 73 / 1983, TN rev. 2018); PMSL. – WM 34 • 1 ♀; Sv. Trije Kralji, the Pohorje Mts; 10 Aug. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 250 / 1984, TN rev. 2018); PMSL. – WM 35 • 1 ♀; Smolnik; 8 Aug. 2015; L. Slana Novak and T. Novak leg. (108 / 2015, TN rev. 2017); PMSL. – WM 36 • 2 ♂♂, 2 ♀♀; Ožbalt; 25 Oct. 2013; T. Novak leg. (45 / 2013, rev. 2017); PMSL. – WM 43 • 1 ♀; Brezje pri Poljčanah; 10 Apr. 1995; S. Brelih leg. (714 / 1998, rev. 2017); PMSL • 1 ♀; Zbelovo; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1158 / 2002, rev. 2018); PMSL. – WM 45 • 1 ♀; Sv. Bolfenk, the Pohorje Mts; 27 Jul. 1987; F. Janžekovič leg. (5 / 1999, TN rev. 2018); PMSL • 1 ♂; Limbuš; 11 Oct. 2001; D. Kos leg. (217 / 2004, rev. 2017); PMSL • 16 ♂♂, 36 ♀♀; ibid.; (224 / 2004, rev. 2017); PMSL • 1 ♂; ibid.; (218 / 2004, rev. 2018); PMSL • 7 ♂♂; ibid.; (225 / 2004, rev. 2018); PMSL. – WM 46 • 1 ♀; Gaj nad Mariborom; 16 Apr. 1983; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 64 / 1983, TN rev. 2018); PMSL • 1 ♀; ibid.; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (164 / 2011, rev. 2018); PMSL • 1 juv.; Gaj nad Mariborom − Žavcarjev vrh; 1 Aug. 1981; T. Novak leg. (1084 / 1981, rev. 2019); PMSL • 1 ♀; Jurski vrh; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (166 / 2011, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Kamniška graba; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (129 / 2011, rev. 2018); PMSL • 5 ♂♂, 1 ♀; Zgornji Slemen; 16 Apr. 1983; T. Novak, M. Slana Novak, L. Slana Novak and M. Štangelj leg. (LSN 10 / 1983, TN rev. 2018); PMSL. – WM 54 • 1 ♂, 1 ♀; Marjeta na Dravskem polju; 8 Jun. 2015; P. Kozel leg. (57 / 2015, rev. 2017); PMSL. – WM 55 • 1 ♀; Maribor – Ruperče; 20 Apr. 1993; S. Brelih leg. (852 / 1998, rev 2018); PMSL. – WM 64 • 1 ♂; Krčevina pri Vurbergu; 15 Sep. 2015; T. Marinko leg. (134 / 2015, TN rev. 2017); PMSL • 1 ♂; Placar; 16 Oct. 1994; M. Varžič leg. (17 / 1999, tev. 2018); PMSL. – WM 65 • 1 ♂; Hum; 23 Apr. 2008; L. Slana Novak and T. Novak leg (130 / 2011, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Zgornja Voličina – Hrastovec; 2 Oct. 2002; T. Novak leg. (1284 / 2002, rev. 2018); PMSL. – WM 74 • 1 ♂; Hlaponci; 23 Apr. 1998; S. Brelih leg. (73 / 2001, rev. 2017); PMSL • 1 ♂; ibid.; (74 / 2001, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; Pacinje; 13 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 60 / 1983, T. Novak rev. 2018); PMSL. – WM 75 • 2 ♀♀; Drbetinci; 28 Apr. 2002; L. Slana Novak and T. Novak leg. (731 / 2002, rev. 2017); PMSL • 1 ♂, 1 ♀; Kokolanjšček; 26 Sep. 2005; L. Slana Novak and T. Novak leg. (121 / 2005, rev. 2017); PMSL • 1 ♂, 1 ♀; Velika Slavšina; 13 Oct. 2005; L. Slana Novak and T. Novak leg. (149 / 2005, rev. 2017); PMSL • 3 ♂♂; Vitomarci; 2 Jun. 2002; L. Slana Novak and T. Novak leg. (756 / 2002, rev. 2018); PMSL. – WM 76 • 1 ♂; Kočki vrh; 30 Jun. 2002; T. Novak leg. (784 / 2002, rev. 2017); PMSL. – WM 95 • 1 ♂; Gresovščak; L. Slana Novak and T. Novak leg. (162 / 2012, TN and SL rev. 2017); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	description	Description Male (holotype) BODY. Length 1.94, width 1.43. CHELICERAE. Ch basal article (without Apo) length 0.58, 2.0 times as long as wide at Ch max width at dorsal hump. Ch-Apo elongated oval-trapezoid, shallowly bilobe apically, wider distally, with a prominent apical pinnacle posteriory. Ch-Apo height 0.21, min width 0.08 (at the base) and max width 0.15. Secretion field frontally. Distal article length 0.60, max width 0.25, movable finger length 0.28 (Figs 4 H, 6 H, 9 H). PEDIPALPS. Pa-Tr with dorsal margin conspicuously inclined distally. Pa-Fe conspicuously club-shaped (Pa-Fe min: max width ~ 1: 2.8) with dorsal margin bent in the middle and nearly straight anterior and posterior portions, and evenly arched ventral margin, with 3 stout spines in distal fourth. Pa-Ti hump low, inconspicuous (Figs 5 H, 6 H). Pa-Pt length: max width ~ 1: 4.1. Pa article lengths in Table 11. PENIS. Pe length 1.47, base 0.48, glans 0.09, stylus 0.07 (Figs. 7 A, H, 8 H). LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 1 − 1 − 2. Tarsomere leg-Fe formula I−IV: 10 − 17 − 10 − 10. Leg article lengths in Table 11. Female BODY. Length 2.11, width 1.65. CHELICERAE. Ch basal article width 0.63, 2.7 times as long as wide at dorsal hump, with nearly straight dorsal margin in front of indentation and ventral margin (Fig. 13 G). Distal article length 0.64, max width 0.23, movable finger length 0.27. PEDIPALPS. Pa-Tr with low dorsal margin, highest in the first third and straight posteriorly. Pa-Fe with conspicuous distal bonce, Pa-Pt, Pa-Ti and Pa-Ta elongated (Fig. 13 G). Pa article lengths in Table 11. OVIPOSITOR. Ovipositor length 0.84, Rec sem of one tubular and 12 slightly elongated vesicles (Fig. 11 G). LEGS. Leg article lengths in Table 11.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	discussion	Remarks Hadži (1927, sub N. bidentatum) made the first detailed drawings of N. b. schmidti ssp. nov. from Schmidt’s collection. Prior to this study, all published evidence citing either N. bidentatum or N. b. bidentatum in Slovenia (all localities), Croatia (Zagreb) and NE Italy (all localities) refers to N. b. schmidti ssp. nov. Previously, all authors considered this subspecies to be within the range of variability of N. b. bidentatum (Roewer 1917, 1923; Hadži 1927, 1928, 1931; Gruber & Martens 1968; Marcellino 1973; Martens 1978; Novak et al. 1984, 1995, 1996, 2002, 2017; Novak & Gruber 2000; Komposch & Gruber 2004; Novak 2004 a, 2005 b, 2005 c). However, Roewer (1951) noticed that Hadžiʼs individuals designated as N. bidentatum probably belonged to another species, while Novak & Gruber (2000) considered N. b. schmidti ssp. nov. a possible hybrid N. b. bidentatum × N. b. sparsum. The glans tip of a specimen from Leutschach / Lučane, (46.67 ° N, 15.47 ° E, 350 m a. s. l.; Gruber & Martens 1968: fig. 24; Martens 1978: fig. 142), Austria, is the first detailed drawing of the glans in N. b. schmidti ssp. nov. See remarks under N. pluridentatum stat. nov., N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. and N. b. martensi ssp. nov. × N. b. schmidti ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	distribution	Distribution Austria, Italy, Croatia, Slovenia. Vertical distribution in Slovenia: 77 − 2108 m a. s. l. Type locality: Dolenja Sevnica, Poljane nad Škofjo Loko (46.12 ° N, 14.15 ° E, 517 m a. s. l.), Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D771CBD3BFEEDF920BD34567F.taxon	biology_ecology	Ecology Nemastoma b. schmidti ssp. nov. is a mesophilous colline to montane subspecies occupying most regions of Slovenia, except the southern and eastern territory, inhabiting various forest and scrub communities, from mixed hardwood riverine forests in the lowland to subalpine Nardion strictae Br. - Bl. 1926 with isolated patches of Pinus mugo Turra and low-growing or krummholz communities. Phenology: adults eurychronous.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	description	urn: lsid: zoobank. org: act: 7690 FA 07 - 5 D 60 - 4545 - 8660 - 282 ADED 3 F 576 Figs 2 – 3, 4 I, 5 I, 6 I, 7 I, 8 I, 9 I, 10 I, 11 I, 12 I, 13 H; Table 11	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	etymology	Etymology The subspecies name ʻ sneznikensis ʼ references Mt. Snežnik, southern Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	diagnosis	Diagnosis Short-legged (leg II <6 mm) species of the N. b. complex, with slightly elongated-trapezoidal Ch-Apo, with row of low tubercles on Pa-Fe and with equilateral triangular glans. Rec sem of 18 balloon-like vesicles.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	materials_examined	Material examined Holotype SLOVENIA – VL 54 • 1 ♂; Mt. Snežnik; 45.59 ° N, 14.44 ° E; 1553 m a. s. l.; 21 Aug. 1998; Ljuba Slana Novak and Tone Novak leg.; deep-humus in Dinaric Pinus mugo scrub in contact with the Dinaric Fagus sylvatica – Abies alba treeline litter sift; PMSL-Opiliones-TN 1494 / 1998. Paratypes SLOVENIA – VL 54 • 2 ♂♂, 1 ♀; same collection data as for holotype; PMSL-Opiliones-TN 1494 / 1998. Other material SLOVENIA – VL 54 • 3 ♂♂, 4 ♀♀; Mt. Snežnik; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (185 / 1999, rev. 2009); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	description	Description Male (holotype) BODY. Length 1.81, width 1.18. CHELICERAE. Ch basal article (without Apo) length 0.53, 2.7 times as long as wide at Ch max width at dorsal hump. Ch-Apo oval-trapezoid, wider distally, without apical pinnacle. Ch-Apo height 0.13, max width 0.13, min width 0.08 (at the base). Secretion field frontally. Distal article length 0.51, max width 0.17, movable finger length 0.22 (Figs 4 I, 6 I, 9 I). PEDIPALPS. Pa-Tr with dorsal margin slightly inclined distally. Pa-Fe slightly club-shaped (Pa-Fe min: max width ~ 1: 2.6) with terminal bonce, and 7 irregular granula in distal third. Pa-Pt length: max width ~ 1: 4.3. Pa-Ti slender (Figs 5 I, 6 I, 10 I). Pa article lengths in Table 12. PENIS. Pe length 1.22, base 0.46, glans 0.09, stylus 0.04 (Figs 7 I, 8 I). LEGS. Pseudoarticle leg-Fe formula I−IV: 0 − 1 − 1 − 2. Tarsomere leg formula I−IV: 9 − 14 − 6 − 8. Leg article lengths in Table 12. Female BODY. Length 2.11, width 1.43. CHELICERAE. Ch basal article length 0.59, 2.7 times as long as wide at dorsal hump, with low-arched dorsal margin in front of indentation and straight ventral margin (Fig. 13 H). Distal article length 0.66, max width 0.20, movable finger length 0.30. PEDIPALPS. Pa-Tr low, with roof-like broken dorsal margin, highest in the first two fifths. Pa-Fe distally with low ventral hump, Pa-Pt, Pa-Ti and Pa-Ta elongated (Fig. 13 H). Pa article lengths in Table 12. OVIPOSITOR. Ovipositor length 0.62, Rec sem of 18 balloon-like vesicles (Fig. 11 I). LEGS. Leg article lengths in Table 12.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	discussion	Remarks Regarding external morphology, N. b. sneznikensis ssp. nov. is very similar to N. relictum, including the habitus, the similarly shaped Pa and relatively short legs. No individuals belonging to the subspecies were found in the last decade. See remarks on taxa of Nemastoma on Mt. Snežnik under N. b. gruberi ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	distribution	Distribution Despite over 30 collections, this taxon has been recorded only in a limited area on Mt. Snežnik. Endemic to Mt. Snežnik, southern Slovenia. Type locality: Mt. Snežnik (45.59 ° N, 14.44 ° E, 1553 m a. s. l.), Slovenia.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	biology_ecology	Ecology Nemastoma b. sneznikensis ssp. nov. is a psychrophilous subspecies. To date it has been recorded only in a cold, well-drained, deep mull humus Dinaric Pinus mugo scrub in contact with the Dinaric Fagus sylvatica – Abies alba treeline on calcareous soils. Likely, it inhabits other cold habitats, like scree and stone heap edges in contact with humus accumulations, as is known for N. relictum (Komposch 1999; Komposch & Gruber 2004), and perhaps a humus-rich shallow subterranean habitat (SSH), i. e., Milieu Souterrain Superficiel (MSS) (Mammola et al. 2016; Halse 2018). Since this habitat is restricted to an area of less than 1 km 2 on Mt. Snežnik, this subspecies is considered highly endangered. Phenology: probably eurychronous. Hybrids among the subspecies of Nemastoma bidentatum	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7726BD35FF4DFC04B8BC536F.taxon	discussion	Remarks In Austria, a few localities are known where N. b. bidentatum lives in syntopy with N. relictum (Gruber & Martens 1968; Martens 1978; in both papers sub N. b. relictum), and with N. b. sparsum (Komposch & Gruber 2004; Komposch 2009). However, individuals of these three taxa do not interbreed and have been considered as potential species (Gruber & Martens 1968; Martens 1978; Komposch & Gruber 2004; Komposch 2009). In Slovenia, N. b. schmidti ssp. nov., with its central geographic position, interbreeds with all other adjacent subspecies: N. b. bidentatum in the North, and N. b. martensi ssp. nov., N. b. sneznikensis ssp. nov., N. b. gruberi ssp. nov. and N. b. sparsum from the Southwest to the Northeast of the country. Besides, N. b. gruberi ssp. nov. interbreed with N. b. martensi ssp. nov., and with N. b. sparsum. In Bosnia, N. pluridentatum stat. nov. has not been found; consequently, the genitalia have not been examined. Despite this, major differences from all the other known taxa within the N. b. complex allow it to be considered a separate species. This is because interbreeding of N. pluridentatum stat. nov., which might be a subalpine species, with the montane N. kozari sp. nov. from the same mountain is quite unlikely, given their great dissimilarity. However, data on taxa of Nemastoma in Bosnia and Herzegovina are nearly completely missing, allowing no further conclusions. Although being morphologically variable, hybrids possess larger Ch-Apo than pure subspecies and are well distinguishable from these. In interbreeding subspecies, hybrids between the two subspecies frequently occur syntopically with individuals belonging to pure subspecies. Consequently, such a collection consists of either one or two pure subspecies and a series of hybrids with varying Ch-Apo shapes. An overview of interbreeding relations among the taxa of the N. b. complex is presented in Table 13. In Slovenia, N. b. bidentatum and N. b. sparsum do not interbreed, since the areas are several tens of kilometres distant from each other. In contrast, we have found hybrids between all the adjacent subspecies (Fig. 3), except N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. × N. b. sparsum, which is expected in the meeting area of the three taxa. Besides, data on hybridization of N. b. sneznikensis ssp. nov. are too scarce to be properly evaluated in full. The zone of hybridization between N. b. schmidti ssp. nov. and N. b. sparsum is the widest, measuring up to 100 km in north-eastern Slovenia, while these zones are narrow to virtually non-identifiable among the others, and hybrids are accordingly scarce. Hybrids N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. were found in a zone up to 10 km wide, while N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. and N. b. gruberi ssp. nov. × N. b. martensi ssp. nov. are limited to zones a few kilometers wide. In Slovenia, there are no hybrids between allopatric N. b. martensi ssp. nov. and N. b. sparsum, while these are expected in Croatia. Hybridization patterns were not simple clines in the transition zones between two subspecies. Instead, in some cases, morphologically various hybrids between the neighboring subspecies, sharing various ratios of the hybridizing subspecies characters (cf. Gruber & Martens 1968; Martens 1978), appeared scattered across the whole width of the transition zone. In some places in the hybridization zone, individuals of both subspecies, together with hybrids of various intermittent stages, occurred syntopically.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7728BD36FEA5F915B8325100.taxon	diagnosis	Diagnosis Typically, males with narrow quarter moon-like, apically bifid Ch-Apo, ~ 2.4 times as high as wide, with wide, inconspicuous apical pinnacle posteriorly, and Pa-Fe ~ intermediate between both subspecies.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D7728BD36FEA5F915B8325100.taxon	materials_examined	Material examined SLOVENIA – VM 44 • 1 ♂, 1 ♀; Gromov rob, Jelendol, Tržič; 23 Aug. 2017; T. Novak leg. (GR 5339, GR 5338); PMSL. – WM 06 • 1 ♂, 1 ♀; Mt. Košenjak; 27 Sep. 1981; T. Novak leg. (520 / 1981, rev. 2018); PMSL • 4 ♂♂, 2 ♀♀; ibid.; 14 Aug. 2012; T. Novak leg. (GR 3784, GR 3799, GR 3800, GR 3826, GR 3830, GR 3832); PMSL • 2 ♀♀; ibid.; 24 Aug. 2012; T. Novak leg. (153 / 2012, rev. 2018); PMSL • 1 ♂; Sv. Simon, Pernice; 18 Aug. 2005; T. Novak leg. (66 b / 2005, rev. 2018); PMSL • 7 ♂♂, 5 ♀♀; Sv. Urban, Kozji vrh nad Dravogradom; 24 Sep. 2005; L. Slana Novak and T. Novak leg. (127 / 2005, rev. 2019); PMSL • 3 ♂♂, 7 ♀♀; ibid.; 9 Aug. 2017; S. Lipovšek and T. Novak leg. (GR M 1021, GR M 1024, GR M 1025, GR M 1027, GR M 1033, GR M 1035 − GR M 1039); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772BBD36FF7DFBFFBD2D504B.taxon	discussion	Remarks Every allegation of hybrids N. b. bidentatum × N. b. sparsum in Slovenia and Croatia (Gruber & Martens 1968; Martens 1978; Novak & Gruber 2000; Novak et al. 2002; Novak 2004 b) refer either to N. b. schmidti ssp. nov. × N. b. sparsum or N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov.; see these references for further remarks.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772BBD37FEF2FA2ABE0054C0.taxon	description	Figs 3, 9 K, 10 K	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772BBD37FEF2FA2ABE0054C0.taxon	diagnosis	Diagnosis Typical male hybrids are characterized by Ch basal article with large hump in front of dorsal indentation, and broad, low quarter moon-like Ch-Apo, ~ 1.6 times as high as wide, and slightly club-shaped, relatively thin Pa-Fe, and five, subequidistant, simple ventral Pa-Fe denticles in the frontal half of the Pa-Fe.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772BBD37FEF2FA2ABE0054C0.taxon	materials_examined	Material examined CROATIA – WL 00 • 8 ♂♂, 11 ♀♀; Debeli Lug, Jasenak; 8 Sep. 2009; (2 / 2012); PMSL. – WL 20 • 7 ♂♂, 13 ♀♀; Ambarac sinkhole, Ogulin; 7 Sep. 2009; R. Ozimec and A. Schönhofer leg. (Coll. ASc 324, 1 / 2012); PMSL. SLOVENIA – VL 26 • 3 ♂♂, 1 ♀; Senožeče – Laže; 20 May 2012; L. Slana Novak and T. Novak leg. (50 / 2012, rev. 2019); PMSL. – VL 33 • 2 ♂♂, 3 ♀♀; Plazine, Starod; 25 Oct. 2012; L. Slana Novak and T. Novak leg. (194 / 2012, rev. 2018); PMSL. – VL 54 • 1 ♂; Travni Dolci, Mt. Snežnik; 12 Aug. 2001; L. Slana Novak and T. Novak leg. (195 b / 2001, rev. 2020); PMSL • 1 ♂; Mt. Snežnik; 12 Sep. 2018; L. Slana Novak and T. Novak leg. (116 a / 2018); PMSL • 1 ♂; ibid.; 25 Oct. 2019; T. Novak. leg. (138 / 2019); PMSL. – VL 95 • 1 ♂, 2 ♀♀; Mt. Kočevska Mala gora; 17 Aug. 1985; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 98 / 1986, TN rev. 2019); PMSL. – VL 96 • 1 ♂; Pečke − Mali vrh, Kočevski Rog; 20 Sep. 2001; B. Drovenik and A. Pirnat leg. (10 / 2005, rev. 2015); PMSL • 1 ♀; ibid.; (16 / 2005, rev. 2015); PMSL. – WL 37 • 3 ♂♂, 2 ♀♀; Mirčev grič, Mt. Gorjanci; 29 Apr. 1995; S. Brelih leg. (780 / 1998, rev. 2019); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD37FEF2FDBFBE4F5135.taxon	description	Figs 3, 9 L, 10 L	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD37FEF2FDBFBE4F5135.taxon	diagnosis	Diagnosis Typical male hybrids are characterized by Ch-Apo with nearly symetrically convex ventro-anterior margin and straight posterior margin, and club-shaped, robust Pa-Fe with five simple, more or less equidistant ventral Pa-Fe spines in the frontal half, and Pa-Ti with small dorsal hump.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD37FEF2FDBFBE4F5135.taxon	materials_examined	Material examined SLOVENIA – VL 09 • 2 ♂♂; Ozeljan; 17 Apr. 2002; B. Drovenik and A. Pirnat leg. (218 / 2011, rev. 2017); PMSL. – VL 26 • 3 ♂♂, 4 ♀♀; Laže, Razdrto; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (358 / 2011, rev. 2017); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD30FEF3FBC8B8615181.taxon	description	Figs 3, 9 M, 10 M	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD30FEF3FBC8B8615181.taxon	diagnosis	Diagnosis In typical male hybrids, Ch-Apo is boomerang-shaped, Pa-Fe club-shaped, but relatively thin, with four simple, equidistant ventral Pa-Fe spines in the frontal half, Pa-Ti without proximal dorso-medial hump.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772ABD30FEF3FBC8B8615181.taxon	materials_examined	Material examined CROATIA – WL 97 • 1 ♂, 1 ♀; Zagreb − Dugo selo; J. Martens et al. leg. (JM 837); PMSL • 16 ♂♂, 13 ♀♀; ibid.; 9 Sep. 2009; A. Schönhofer leg. (ASc 342, 4 / 2012, ASc det., TN rev. 2018); PMSL. – XL 06 • 3 ♂♂, 2 ♀♀; Prevlaka; (826 / 2002, rev. 2018); PMSL • 4 ♂♂, 6 ♀♀; ibid.; Oct. 1984; (236 / 2002, rev. 2018); PMSL • 2 ♂♂, 2 ♀♀; ibid.; (278 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid.; (280 / 2002, rev. 2018) • 1 ♂, 1 ♀; ibid.; 15 Aug. 1985; (LSN 28 / 1996, TN rev. 2018); PMSL • 1 ♀; ibid.; (891 / 2002, rev. 2018); PMSL • 1 ♂; ibid.; 30 Aug. 1985; (609 / 2002, rev. 2019); PMSL • 2 ♂♂; ibid.; (861 / 2002, rev. 2018); PMSL • 2 ♂♂, 1 ♀; ibid.; Oct. 1985; (896 / 2002); PMSL • 1 ♀; ibid.; 7 Apr. 1986; I. Kos leg. (328 / 2002, rev. 2018); PMSL • 1 ♂; ibid.; (843 / 2002, rev. 2018); PMSL. SLOVENIA – VL 13 • 3 ♂♂, 3 ♀♀; Rakitovec; 21 Sep. 2011; L. Slana Novak and T. Novak leg. (263 / 2011, rev. 2017); PMSL. – VL 26 • 5 ♂♂, 2 ♀♀; Fameljska loza, Senožeče; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (350 / 2011, rev. 2017); PMSL • 1 ♂; Senožeče – Vremščica; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (355 / 2011, rev. 2017); PMSL. – VL 27 • 3 ♂♂, 1 ♀; Mt. Nanos; 1 Sep. 2011; T. Novak leg. (198 / 2011, rev. 2018); PMSL. – VL 34 • 1 ♂, 3 ♀♀; Harije; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (224 / 2011, rev. 2015); PMSL • 1 ♀; Petelinje; 27 Apr. 1994; S. Brelih leg. (815 / 1998, rev. 2015); PMSL. – VL 58 • 2 ♂♂; Brezovica pri Borovnici; 12 May 1997; S. Brelih leg. (83 / 2001, rev. 2015); PMSL. – VL 67 • 1 ♂, 2 ♀♀; Lužarji; 25 Aug. 2011; T. Novak leg. (168 / 2011, rev. 2018); PMSL. – VL 78 • 1 ♀; Mt. Limberk; Jun. 1998; I. Bertoncelj leg. (1544 / 1998, rev. 2009); PMSL • 1 ♂; Veliko Mlačevo; 13 Mar. 1992; S. Brelih leg. (813 / 1998, rev. 2018); PMSL. – VL 79 • 1 ♂; Veliki Lipoglav; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (472 / 2002, rev. 2018); PMSL. – VL 84 • 1 ♂, 2 ♀♀; Pleš, Borovec pri Kočevski Reki; 19 Sep. 2013; P. Kozel and T. Novak leg. (27 / 2013, rev. 2015); PMSL. – VL 88 • 1 ♂, 1 ♀; Krka, Krška vas pri Muljavi; 7 Jul. 1986; L. Slana Novak and T. Novak leg. (474 / 2002, rev. 2018); PMSL • 1 ♀; ibid.; Jun. 1998; V. Kovačič leg. (1444 / 1998, rev. 2017); PMSL. – VL 93 • 1 ♂; Slavski Laz; 29 Apr. 2001; S. Brelih leg. (79 / 2001, rev. 2018); PMSL. – VL 99 • 5 ♂♂, 1 ♀; Cirnik; 27 May 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 249 / 1984, TN rev. 2015); PMSL • 1 ♀; Temenica; 4 Mar. 1994; S. Brelih leg. (680 a / 1998, rev. 2018); PMSL. – WL 04 • 5 ♂♂; Brezovica pri Predgradu; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (147 / 1997, rev. 2018); PMSL. – WL 05 • 5 ♂♂; Črmošnjice; 18 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (152 / 1997, rev. 2018); PMSL. – WL 06 • 1 ♂, 1 ♀; Baza 20, Kočevske Poljane; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 76 / 1986, TN rev. 2018); PMSL • 2 ♂♂; Podstene, Kočevski Rog; 19 Jun. 1986 (672 / 2002, rev. 2017); PMSL. – WL 08 • 1 ♀; Velika jama nad Trebnjem cave, Cad. No. 104, Mokronog; 28 Jul. 2009; M. Zagmajster leg. (PK and TN 6 / 2020); PMSL. – WL 09 • 1 ♂; Šentrupert; 28 Feb. 1994; S. Brelih leg. (825 / 1998, rev. 2018); PMSL. – WL 19 • 3 ♂♂; Puščava, Mokronog; 28 Feb. 1994; S. Brelih leg. (801 / 1998, rev. 2018); PMSL. – VL 28 • 1 ♂; Mt. Homski hrib, Sela pri Zburah; Jun. 1997; D. Vrček leg. (1709 / 1998); PMSL. – WL 29 • 1 ♂, 2 ♀♀; Dolenji Boštanj; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (73 / 2007, rev. 2015); PMSL. – WM 01 • 1 ♂, 6 ♀♀; Mt. Javor, Znojile; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1194 / 2002, rev. 2018); PMSL. – WM 40 • 1 ♂; Lesično; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1130 / 2002, rev. 2018); PMSL • 5 ♂♂; ibid.; (1132 / 2002, rev. 2018); PMSL. – WM 42 • 1 ♂; Mestinje; 4 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 44 / 1983, TN rev. 2015); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD30FF4DFB7DB82E53BB.taxon	description	Figs 3, 9 N, 10 N	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD30FF4DFB7DB82E53BB.taxon	diagnosis	Diagnosis Typical male hybrids with elongated Ch-Apo with knee-like flexion of frontal margin, and equidistant triangular upper portion, and club-shaped, but relatively thin Pa-Fe, with tubercle tightly in front of the first spine of four simple, equidistant ventral spines in distal Pa-Fe half, either pointed or almost without the tip.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD30FF4DFB7DB82E53BB.taxon	materials_examined	Material examined CROATIA – XL 03 • 1 ♂; Jezero Posavsko, Suša; 3 Dec. 1985; (61 / 2019); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD31FEFBF93BBF945301.taxon	description	Figs 3, 9 O	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD31FEFBF93BBF945301.taxon	diagnosis	Diagnosis Typical male hybrids with Ch basal article with large hump in front of dorsal indentation, Ch-Apo with knee-like frontal flexion and narrowly drawn out upper portion, Pa-Fe club-shaped, but thinner than in N. b. schmidti ssp. nov., with four simple, equidistant ventral spines in distal Pa-Fe half, and Pa-Ti with minute hump.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD31FEFBF93BBF945301.taxon	materials_examined	Material examined SLOVENIA – VL 15 • 1 ♂; Prelože pri Lokvi; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (310 / 2011); PMSL. – VL 35 • 3 ♂♂, 5 ♀♀; Ribnica – Ostrožno Brdo; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (324 / 2011, rev. 2015); PMSL. – VL 53 • 1 ♂; Zabiče; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (301 / 2011, rev. 2015); PMSL. – WL 05 • 1 ♂; Mt. Mirna gora; 28 Jul. 1948; A. Seliškar leg; PMSL. – WL 17 • 1 ♂; Novo Mesto; Karaman leg. (107 / 1983 microscopic preparation, JH det., TN rev. 2017); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772DBD31FEFBF93BBF945301.taxon	discussion	Remarks In Hadži’s original preparation, labelled “ Nemastoma bidentatum Roewer pulli ”, with two cover glasses on the same microscope slide, there are two males and a female. One male corresponds to the type specimen of Nemastoma (Stridulostoma) seliskari Hadži, 1973 (Hadži 1973 a). For this species, Hadži (ibid.) founded a new subgenus Stridulostoma based on the existence of an alleged “ stridulatory organ ” on the Fe III (Hadži 1973 a: figs 37 č – d). Gruber (1976: 797) explained that the stridulatory organ alone does not justify establishing of a new subgenus, especially because this new species differs from N. (Lugubrostoma) triste pluridentatum Hadži, 1973 exclusively by possession of this organ, and accordingly, synonymized Stridulostoma with Nemastoma. Following these remarks, Schönhofer (2013: 36) synonymized N. seliskari with N. b. pluridentatum. We found that the portion of the Fe III, identified by Hadži (1973 a) as a stridulatory organ in the type specimen, is in fact a depression-damage, caused by desiccation. Moreover, such damage is present in nearly all (sic!) the other femora of this freshly moulted specimen, but nothing resembles the fine lamellated structure shown in Hadži’s drawing (1973 a: figs 37 č – d). No similar structure was found in the specimens of Nemastoma collected later on Mt. Mirna gora. One might imagine that a careless inspection under a microscope at low magnification could result in the illusion of a ʻstridulatory organʼ. However, the drawing of its lamellae cannot be considered anything but a pure fabrication. Moreover, Pa-Fe in Hadži (1973 a: fig. 33 c), being under the same cover glass in the preparation, belongs to the same specimen, although Hadži (ibid.) ascribed it to N. (Lugubrostoma) triste pluridentatum. See remark under N. pluridentatum stat. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772CBD32FE8CF9FFBED8546A.taxon	diagnosis	Diagnosis In typical male hybrids, Ch basal article oblong, sneznikensis - like, with Ch-Apo with small apical pinnacle posteriorly, Pa-Fe club-shaped, thinner than in N. b. schmidti ssp. nov., with three simple ventral spines in distal Pa-Fe half, Pa-Pt schmidti - like, Pa-Ti with inconspicuous tubercle.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772CBD32FE8CF9FFBED8546A.taxon	materials_examined	Material examined SLOVENIA – VL 54 • 1 ♂; Mt. Snežnik; 14 Sep. 2019; L. Slana Novak and T. Novak leg. (GR 5468); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772CBD32FE8CF9FFBED8546A.taxon	discussion	Remarks To date, only one male has been found.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772FBD2CFF50FE15BF37505F.taxon	description	Figs 3, 9 P, 10 O	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772FBD2CFF50FE15BF37505F.taxon	diagnosis	Diagnosis Hybrids exhibiting varying Ch-Apo morphology, but with knee-like flexion of anterior margin, and either being relatively tiny with more or less elongated superior portion or more massive with superior portion drawn out in tiny apical portion, and club-shaped Pa-Fe with intermediate form with respect to both subspecies.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772FBD2CFF50FE15BF37505F.taxon	materials_examined	Material examined SLOVENIA – VM 91 • 2 ♂♂, 4 ♀♀; Čemšemik; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1183 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; Jesenovo – Znojile; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1187 / 2002, TN rev. 2018); PMSL • 6 ♂♂, 5 ♀♀; Mt. Čemšeniška planina; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1191 / 2002, rev. 2018); PMSL • 2 ♀♀; Zaplanina, Ločica pri Vranskem; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1210 / 2002, rev. 2018); PMSL • 1 ♂, 5 ♀♀; ibid.; (1179 / 2002, rev. 2018); PMSL. – VM 92 • 4 ♂♂, 9 ♀♀; Ločica pri Vranskem; 10 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 101 / 1986, TN rev. 2018); PMSL. – WM 01 • 7 ♂♂, 5 ♀♀; Marija Reka; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1186 / 2002, rev. 2018); PMSL. – WM 05 • 4 ♂♂, 7 ♀♀; Slovenj Gradec; 9 Oct. 1983; T. Novak and L. Slana Novak leg. (LSN 113 / 1983, TN rev. 2018); 3 ♂♂, 4 ♀♀; ibid.; 6 Jul. 2003; L. Slana Novak and T. Novak leg. (190 / 2003, rev. 2018); PMSL. – WM 06 • 1 ♂, 2 ♀♀; Dravograd; 25 Oct. 1995; S. Hudrap and M. Pavlin leg. (11 / 1996, rev. 2018); PMSL. – WM 10 • 4 ♂♂, 1 ♀; Lože – Globoko; 14 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 45 / 1983, TN rev. 2018); PMSL. – WM 11 • 1 ♂, 2 ♀♀; Brdce; 22 Apr. 1995; S. Brelih leg. (758 / 1998, rev. 2018); PMSL • 1 ♂, 3 ♀♀; Udmat; 15 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 47 / 1983, TN rev. 2018); PMSL. – WM 12 • 7 ♂♂, 3 ♀♀; Arja vas; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (513 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; Velika Pirešica; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (452 / 2002, rev. 2018); PMSL • 1 ♂, 4 juvs; ibid.; (470 / 2002, rev. 2018); PMSL. – WM 13 • 4 ♂♂, 3 ♀♀; Vodosteč, Črnova; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (453 / 2002, rev. 2018); PMSL • 2 ♂♂; ibid.; 22 Apr. 1995; S. Brelih leg. (772 / 1998, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid.; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (737 / 2002, rev. 2019); PMSL. – WM 16 • 2 ♂♂, 3 ♀♀; Sv. Trije Kralji, Muta; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (79 / 2005, rev. 20018); PMSL. – WM 20 • 2 ♂♂, 3 ♀♀; Jurklošter; 15 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 53 / 1983, TN rev. 2018); PMSL. – WM 21 • 1 ♂, 2 ♀♀; Lahovna; 22 Apr. 1995; S. Brelih leg. (740 / 1998, TN rev. 2018); PMSL • 2 ♂♂, 3 ♀♀, 1 juv.; Trobni dol; 16 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 48 / 1983, TN rev. 2018); PMSL. – WM 22 • Celje • 1 ♂; ibid.; 30 Oct. 2000; N. Žmaher leg. (881 / 2000, rev. 2018); PMSL • 7 ♂♂, 2 ♀♀; Šmartinsko jezero; 25 Mar. 1995; S. Brelih leg. (704 / 1998, rev. 2018); PMSL • 1 ♂, 4 ♀♀; Vojnik; 17 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 58 / 1983, TN rev. 2018); PMSL. – WM 23 • 1 ♂, 1 ♀; Črešnjice; 25 Mar. 1995; S. Brelih leg. (709 / 1998, rev. 2018); PMSL • 3 ♂♂, 3 ♀♀, l juv.; Frankolovo; 17 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 76 / 1983, TN rev. 2018); PMSL • 1 ♂; Mt. Konjiška gora; 14 Jun. 1981; V. Kuštor, T. Novak leg. (1120 / 1981, rev. 2018); PMSL • 1 ♂, 2 ♀♀; Mt. Stenica; 30 May 1981; T. Novak, V. Kuštor leg. (228 / 1981, rev. 2018); PMSL • 1 ♀; Stenica; 24 Jul. 2002; B. Drovenik and A. Gergeli leg. (1138 / 2018); PMSL • 1 ♀; ibid.; 24 Jul. – 21 Aug. 2002; B. Drovenik and A. Gergeli leg. (PK 73 / 2012, TN rev. 2018); PMSL • 1 ♀; ibid.; (PK 12 / 2013, TN rev. 2018); PMSL • 1 ♂; ibid.; 21 Aug. 2002; B. Drovenik and A. Gergeli leg. (1071 / 2018); PMSL • 1 ♀; ibid.; 21 Aug. − 16 Sep. 2002; B. Drovenik and A. Gergeli leg. (782 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (819 / 2018); PMSL • 1 ♀; ibid.; (856 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (804 / 2018); PMSL • 1 ♀; ibid.; 16 Sep. − 14 Oct. 2002; B. Drovenik and A. Gergeli leg. (817 / 2018); PMSL • 1 ♀; ibid.; 14 Oct. 2002; B. Drovenik and A. Gergeli leg. (361 / 2018); PMSL • 1 ♀; ibid.; (783 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (794 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (814 / 2018); PMSL • 1 ♀; ibid.; (824 / 2018); PMSL • 1 ♀; ibid.; (831 / 2018); PMSL • 1 ♀; ibid.; (846 / 2018); PMSL • 1 ♂; ibid.; 11 Nov. 2002; B. Drovenik and A. Gergeli leg. (208 / 2018); PMSL • 1 juv.; ibid.; (266 / 2018); PMSL • 1 ♂; ibid.; 11 Nov. 2002 − 4 Mar. 2003; B. Drovenik and A. Gergeli leg. (694 / 2018); PMSL • 1 ♀; ibid.; (698 / 2018); PMSL • 2 ♀♀; ibid.; (701 / 2018); PMSL • 1 ♂, 1 ♀; ibid.; (710 / 2018); PMSL • 1 ♀; ibid.; (716 / 2018); PMSL • 1 ♀; ibid.; (721 / 2018); PMSL • 1 ♂; ibid.; (724 / 2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; (729 / 2018); PMSL • 1 ♂; ibid.; 8 Apr. 2003; B. Drovenik and A. Gergeli leg. (267 / 2018); PMSL • 3 juvs; ibid.; (271 / 2018); PMSL • 1 ♂; ibid.; 8 Apr. 2003; B. Drovenik and A. Gergeli leg. (189 / 2018); PMSL • 1 juv.; ibid.; (266 / 2018); PMSL. – WM 24 • 1 ♂, 1 ♀; Jurgovo; 24 Jul. – 3 Aug. 2005; A. Kapla leg. (178 / 2005, rev. 2018); PMSL • 3 ♂♂; Resnik; 12 Aug. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 252 / 1984, TN rev. 2018); PMSL. – WM 25 • 3 ♂♂; Glažuta, Mislinjski jarek; 9 Sep. 2017; M. Delakorda, S. Lipovšek, L. Slana Novak and T. Novak leg. (107 / 2018); PMSL • 1 ♀; ibid.; 19 Sep. 2018; S. Lipovšek, P. Kozel and T. Novak leg. (132 / 2018); PMSL • 1 ♂; Ribnica na Pohorju; 4 Jul. 1939; J. Hadži leg. (145 / 1984, rev. 2018); PMSL. – WM 30 • 1 ♂, 1 ♀; Loke – Lažiše; 11 Jul. 2003; T. Novak leg. (154 / 2003, rev. 2018); PMSL. – WM 31 • 3 ♂♂, 2 ♀♀; Bezovje pri Šentjurju; 16 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 56 / 1983, TN rev. 2018); PMSL • 1 ♂; Slivniško jezero; 7 Apr. 1995; S. Brelih leg. (701 / 1998, rev. 2018); PMSL. – WM 33 • 3 ♂♂, 2 ♀♀, 1 juv.; Tepanje; 17 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 62 / 1983, TN rev. 2018); PMSL • 2 ♂♂; Žiče monastery; 25 Mar. 1995; S. Brelih leg. (706 / 1998, rev. 2018); PMSL. – WM 40 • 1 ♂; Lesično; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1130 / 2002, rev. 2018); PMSL. – WM 41 • 1 ♂, 5 ♀♀; Podčetrtek; 13 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 51 / 1983, TN rev. 2018); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 7 Jul. 2012; T. Novak leg. (51 / 2012, rev. 2018); PMSL • 2 ♂♂, 3 ♀♀; Virštajn; 7 Apr. 1995; S. Brelih leg. (699 / 1998, rev. 2018); PMSL. – WM 42 • 1 ♀; Mt. Boč; 17 Apr. 1982; T. Novak and G. Kolar leg. (125 / 1982, rev. 2018); PMSL • 2 ♂♂, 1 ♀; ibid.; 16 Oct. 1983; L. Slana Novak and T. Novak leg. (LSN 118 / 1983, TN rev. 2018); PMSL • 2 ♂♂; Mestinje; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1165 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid.; (1168 / 2002, rev. 2018); PMSL • 1 ♂, 1 ♀; Zgornji Gabrnik; 11 Oct. 1997; L. Slana Novak and T. Novak leg. (412 / 1998, rev. 2018); PMSL. – WM 43 • 4 ♂♂, 1 ♀; Ložnica; 17 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 75 / 1983, TN rev. 2018); PMSL. – WM 44 • 2 ♂♂, 4 ♀♀; Kočno pri Polskavi; 8 Aug. 2015; L. Slana Novak and T. Novak leg. (110 / 2018); PMSL. – WM 45 • 1 ♂; Limbuš; 11 Oct. 2001; D. Kos leg. (226 / 2004, rev. 2018); PMSL. – WM 52 • 1 ♀; Sv. Donat, Donačka gora; 16 Sep. 2002; B. Drovenik and A. Pirnat leg. (247 / 2018); PMSL • 1 ♀; ibid.; (248 / 2018); PMSL • 1 ♀; ibid.; 14 Oct. 2002; B. Drovenik and A. Pirnat leg. (1032 / 2018); PMSL • 6 ♂♂, 2 ♀♀, 2 juvs; Tuncovec; 4 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 66 / 1983, TN rev. 20018); PMSL. – WM 53 • 4 ♂♂, 1 ♀; Majšperk; 24 Jun. 2007; T. Novak leg. (43 / 2018); PMSL • 4 ♂♂, 1 ♀; Pečke; 10 Apr. 1995; S. Brelih leg. (713 / 1998, rev. 2018); PMSL • 1 ♂; Stogovci; 24 Jun. 2007; T. Novak leg. (50 / 2007, rev. 2018); PMSL. – WM 62 • 1 ♂, 2 ♀♀; Dobovec pri Rogatcu; 7 Oct. 2001; T. Novak leg. (605 / 2002, rev. 2007); PMSL • 2 ♂♂; ibid.; 16 Aug. 2003; J. Ravljen and T. Novak leg. (807 / 2003, rev. 2018); PMSL • 4 ♂♂; ibid.; 16 Aug. 2003; J. Ravljen, L. Slana Novak and T. Novak leg. (812 / 2003, rev. 2007); PMSL • 1 ♂, 2 ♀♀; ibid.; 30 Sep. 2012; T. Novak leg. (63 / 2015); PMSL • 7 ♂♂, 4 ♀♀, 1 juv.; Kozminci; 6 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 61 / 1983, TN rev. 2018); PMSL • 1 ♀; Mt. Macelj; Jun. 1980; V. Kuštor and T. Novak leg. (LSN 5 / 1983, TN rev. 2018); PMSL • 2 ♀♀; Žale, Dobrina; 24 Apr. 1998; S. Brelih leg. (85 / 2001, rev. 2018); PMSL. – WM 63 • 4 ♂♂, 1 ♀; Majski vrh − Tržec; 6 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 65 / 1983, TN rev. 2018); PMSL. – WM 64 • 3 ♂♂, 3 ♀♀; Ptuj; 6 Aug. 1983, T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 77 / 1983, TN rev. 2018); PMSL. – WM 65 • 2 ♂♂; Selca; 14 Jun. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 238 / 1984, TN rev. 2018); PMSL • 1 ♂, 2 ♀♀; Zgornja Voličina; 23 Apr. 1998; S. Brelih leg. (124 / 2001, rev. 2018); PMSL. – WM 73 • 1 ♂, 4 ♀♀; Berinjak; 24 Jun. 2007; T. Novak leg. (36 / 2007, rev. 2018); PMSL • 1 ♂, 8 ♀♀; Borl castle, Dolane; 7 Aug. 1983; L. Slana Novak, M. Štangelj and T. Novak leg. (LSN 59 / 1983, TN rev. 2018); PMSL • 1 ♂, 2 ♀♀; Ljubslava, Podlehnik; 24 Jun. 2007; T. Novak leg. (49 / 2007, TN rev. 2018); PMSL • 5 ♂♂; Spodje Gruškovje, Podlehnik; 24 Jun. 2007; T. Novak leg. (13 / 2007, rev. 2018); PMSL • 4 ♂♂, 1 ♀; Vareja; 6 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 78 / 1983, TN rev. 2018); PMSL. – WM 76 • 4 ♂♂, 3 ♀♀; Brengova; 26 Jul. 1983; T. Novak and L. Slana Novak leg. (LSN 63 / 1983, TN rev. 2018); PMSL • 2 ♂♂, 1 ♀; Kočki vrh; 30 Jun. 2002; T. Novak leg. (783 / 2002, rev. 2018); PMSL • 1 ♂, 2 ♀♀; ibid.; (789 / 2002, rev. 2018); PMSL. – WM 83 • 3 ♂♂, 5 ♀♀; Korenjak – Turški vrh; 7 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. (LSN 55 / 1983, TN rev. 2018); PMSL. – WM 84 • 4 ♂♂, 3 ♀♀; Vudina, Velika Nedelja; 29 May 2001; P. Igerc and T. Novak leg. (22 / 2001, rev. 2018); PMSL. – WM 85 • 2 ♀♀; Radoslavci; 10 Sep. 1983; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 117 / 1983, TN rev. 2018); PMSL. – WM 94 • 2 ♂♂, 3 ♀♀; Pavlovci; 2 Apr. 1983; T. Novak and L. Slana Novak leg. (LSN 6 / 1983, TN rev. 2018); PMSL. – WM 95 • Noršinci pri Ljutomeru; 27 Aug. 1983; T. Novak and L. Slana Novak leg. 2 ♂♂, 2 ♀♀ (LSN 109 / 1983, TN rev. 2018); PMSL. – WM 95 • 3 ♂♂, 3 ♀♀; Gresovščak; L. Slana Novak and T. Novak leg. (163 / 2012, rev. 2018); PMSL • 2 ♂♂, 2 ♀♀; Slamnjak, Ljutomer; 10 Jun. 1984; L. Slana Novak and T. Novak leg. (LSN 241 / 1984, rev. 2018); PMSL.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
534F336D772FBD2CFF50FE15BF37505F.taxon	discussion	Remarks To date, N. b. schmidti ssp. nov. has been reported under N. b. bidentatum, and N. b. gruberi ssp. nov. under N. b. sparsum. Since hybrids N. b. bidentatum × N. b. sparsum have not been identified in any locality, all these published data likely refer either to N. b. schmidti ssp. nov. × N. b. sparsum or N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. Nemastoma b. bidentatum referred from Germany (Schönhofer & Holle 2007) is a hybrid, probably N. b. schmidti ssp. nov. × N. b. sparsum. Since both N. b. sparsum and N. b. gruberi ssp. nov. inhabit the lowland in Croatia, hybrids south of the Sava River or nearby probably belong to N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov.	en	Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor, Raspotnig, Günther (2021): Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence. European Journal of Taxonomy 777: 1-67, DOI: https://doi.org/10.5852/ejt.2021.777.1561
