identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B436B30CFFF4FFA96DB2FF1AFE7FC46F.text	B436B30CFFF4FFA96DB2FF1AFE7FC46F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora medleri Johnson 1978	<div><p>Chaetophora medleri Johnson 1978 [Figures 2‒3, 7‒8, 12‒13, 17‒18, 22A‒23C, 28]</p> <p>Type locality. Nigeria, Badeggi.</p> <p>Type material. Holotype male, deposited at MMUE “ Nigeria: Badeggi RRS NW State 19 III 1972 J. T. Medler Coll. // C-20 // Manchester Museum HOLOTYPE // Chaetophora medleri sp.n. det. C.Johnson 1977”. Three paratypes with the following label “ Nigeria: Badeggi RRS NW State 19 III 1972 J. T. Medler Coll. // C-20 // Manchester Museum PARATYPE // Chaetophora medleri sp.n. det. C.Johnson 1977”.</p> <p>Condition of holotype, male. Portion of apical end of the median lobe is broken.</p> <p>Additional material. 1 male &amp; 1 female, deposited at BMNH “Umuaha 17.XI.-15.XII.1960 // C.E. Tottenham collection. B.M. 1974-587. // NHMUK014404467” and “Umudike 10-13-iv 1960 J. L. Gregory // C.E. Tottenham collection. B.M. 1974-587. // NHMUK014385244”.</p> <p>Description. Body: Circular, dorsum convex. BL: 1.21 mm, EL: 0.87 mm, EW: 0.92 mm, PL: 0.34 mm, PW: 0.79 mm. Colour: Dorsal side reddish-brown; venter reddish-brown, tibia reddish-brown, tarsal claws light brown.</p> <p>Dorsum (Fig. 2). Head: Partially retracted into pronotum; overall surface smooth, very few punctations across surface, deep groove present from the middle of the frons and extending towards the antennal insertion. Antennae: 11 segmented, clavate; 1 st and 2 nd AS longer than wide and medium sized, 3 rd to 9 th AS short and narrow, 10 th and 11 th AS enlarged. Pronotum: Transverse, widest at posterior, narrowest at anterior, narrower than elytra; anterior margin convex, posterior margin gently convex; lateral margin sinuate with concave in the middle; surface covered with punctations, surface between punctures smooth; long and thin setae spread across surface. Scutellum: Triangular, small, longer than wide. Elytra: Slightly wider than long as wide; anterior gently concave; anterior lateral angle close to right angle; lateral margins gently curve from anterior end to anterior 2/3, increase in curvature towards the apex; smooth surface with faint elytral striae, very shallow punctuations; surface near elytral tip with small grooves; long and thin setae spread across surface.</p> <p>Venter (Fig. 7). Prosternum: Concave on anterior margin; prosternal process as long as wide, narrows towards apex, posterior margin rounded. Mesoventrite: Wider than long; anterior margin concave where prosternal process fits; lateral sub-parallel and gently concave, posterior slightly convex. Metaventrite: Wider than long, flat in the middle; relatively large and shallow punctures spread across surface. Abdomen: Five clearly visible ventrites. Intercoxal process triangular with rounded apex. Posterior margin of 1 st ventrite flat; 2 nd to 4 th ventrites concave, 5 th ventrite nearly semi-circle, posterior end flat. Surface covered with short, brown, setae on the outer margin (Fig. 17).</p> <p>Legs. Tibiae: Outer lateral margin almost straight, inner lateral margin straight. Tarsi: Simple, segments increase in length towards the apex. Tarsal claws: Simple, narrow, symmetrical.</p> <p>Male. Genitalia (Figs. 23A‒23C). Length: 0.61 mm. Parameres very much reduced. Median lobe long and thin, width constant and parallel, apical end pointed in ventral view; majority straight and obtusely angled near the apical end in lateral view; apical end pointed. Phallobase oval. (Figs. 23A‒23C). No visible variation (Figs. 2, 8, 13, 18).</p> <p>Female. No visible variation compared to male.</p> <p>Distribution. C. medleri Johnson 1978 is known only from Nigeria (Badeggi and Umuaha).</p> <p>Differential diagnosis. Combination of different key characters which can differentiate from other species are displayed in Table 1.</p></div> 	https://treatment.plazi.org/id/B436B30CFFF4FFA96DB2FF1AFE7FC46F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
B436B30CFFF4FFAC6DB2F974FBB6C07E.text	B436B30CFFF4FFAC6DB2F974FBB6C07E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora milesi Matsumoto 2021	<div><p>Chaetophora milesi sp. nov. [Figures 4, 9, 14, 19, 24A‒24C, 28]</p> <p>Type locality. Sierra Leone, Tiwai Island.</p> <p>Type material. Holotype male, deposited at BMNH “ SIERRA LEONE 120m <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-11.3525&amp;materialsCitation.latitude=7.55" title="Search Plazi for locations around (long -11.3525/lat 7.55)">Tiwai Island</a>, Moa River N07°33’00”; W11°21’09” 17-22.vi.16 // Edwards Trap leg. Takano, Miles &amp; Goff Trip Ref: SL-001 (ANHRT 18) BMNH(E) 2016-196 // NHMUK014404466”. Additional label on red paper added “ Chaetophora milesi sp. nov. K. Matsumoto det. 2020 HOLOTYPE ♂ ”.</p> <p>Description. Body: Circular, dorsum convex. BL: 1.33 mm, EL: 0.97 mm, EW: 1.06 mm, PL: 0.36 mm, PW: 0.89 mm. Colour: Dorsal side dark brown; venter brown to dark reddish brown, tibia brown, tarsal claws light brown.</p> <p>Dorsum (Fig. 4). Head: Partially retracted into pronotum; overall surface smooth, very few punctations across surface, deep groove present from the middle of the frons and extending towards the antennal insertion. Antennae: 11 segmented, clavate; 1 st and 2 nd AS longer than wide and medium sized, 3 rd to 9 th AS short and narrow, 10 th and 11 th AS enlarged. Pronotum: Transverse, widest at posterior, narrowest at anterior, narrower than elytra; anterior margin convex, posterior margin gently convex; lateral margin convex with slightly concave in the middle; surface covered with punctations, surface between punctures smooth; long and thin setae spread across surface. Scutellum: Triangular, small, longer than wide. Elytra: Slightly wider than long; anterior margin nearly straight; anterior lateral angle obtuse; lateral margins gently curve from anterior end to anterior 2/3, increase in curvature towards the apex; irregular surface with faint elytral striae punctuations; surface near elytral tip with small grooves; long and thin setae spread across surface.</p> <p>Venter (Fig. 9). Prosternum: Concave on anterior margin; prosternal process as long as wide, narrows towards apex, posterior margin rounded. Mesoventrite: Wider than long; anterior margin concave where prosternal process fits; lateral sub-parallel and gently concave, posterior slightly convex. Metaventrite: Wider than long, flat in the middle; relatively large and shallow punctures spread across surface. Abdomen: Five clearly visible ventrites. Intercoxal process triangular with rounded apex. Posterior margin of 1 st ventrite flat; 2 nd to 4 th ventrites concave, 5 th ventrite nearly semi-circle, posterior end flat. Surface covered with short, brown, setae on the outer margin (Fig. 19).</p> <p>Legs. Tibiae: Outer lateral margin almost straight, inner lateral margin straight. Tarsi: Simple, segments increase in length towards the apex. Tarsal claws: Simple, narrow, symmetrical.</p> <p>Male. Genitalia (Figs. 4‒6). Length: 0.65 mm. Parameres very much reduced. Median lobe long and thin, width constant and parallel in ventral view, apical end pointed; gently curved in lateral view; apical end pointed. Phallobase oval.</p> <p>Female. Unknown.</p> <p>Distribution. C. milesi sp. nov. is known only from the type locality.</p> <p>Etymology. This species is named after the Will Miles who collected this specimen.</p> <p>Differential diagnosis. Combination of key characters which can differentiate the new species from other species of the genus are displayed in Table 1.</p> <p>Remarks. This species is the first country record of this genus for Sierra Leone.</p></div> 	https://treatment.plazi.org/id/B436B30CFFF4FFAC6DB2F974FBB6C07E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
B436B30CFFF1FFAD6DB2F91BFBAFC7FA.text	B436B30CFFF1FFAD6DB2F91BFBAFC7FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora morettoi Matsumoto 2021	<div><p>Chaetophora morettoi sp. nov. [Figures 1A‒1I, 5, 10, 15, 20, 25A‒25C, 28]</p> <p>Type locality. Côte d’Ivoire, Gbando Village.</p> <p>Type material. Holotype male, deposited at BMNH “ IVORY COAST 417m <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.683639&amp;materialsCitation.latitude=9.571417" title="Search Plazi for locations around (long -6.683639/lat 9.571417)">Gbando Village</a> (sudanian forest with gallery forest) 09°34’17.1”N, 06°41’1.1”W 15-22.vi.2018 // MV Light Trap. Aristophanous, M., Miles, W., Moretto, P., Outtara, Y. leg. ANHRT: 2018.28, BMNH(E) 2018-153 // NMHUK014381512”. Additional label on red paper added “ Chaetophora morettoi sp. nov. K. Matsumoto det. 2020 HOLOTYPE ♂ ”. Paratypes (24 specimens. BMNH, MMUE): same locality data as holotype with additional label: “NMHUK013679876”, “NMHUK013679877”, “NMHUK014381490”, “NMHUK014381491”, “NMHUK014381492”, “NMHUK014381493”, “NMHUK014381494”, “NMHUK014381495”, “NMHUK014381496”, “NMHUK014381497”, “NMHUK014381498”, “NMHUK014381499”, “NMHUK014381500”, “NMHUK014381501”, “NMHUK014381502”, “NMHUK014381503”, “NMHUK014381504”, “NMHUK014381505”, “NMHUK014381506”, “NMHUK014381507”, “NMHUK014381508”, “NMHUK014381509”, “NMHUK014381510” and “NMHUK014381511”. All paratype specimens have been labelled with a red paratype label: “ Chaetophora morettoi sp. nov. K. Matsumoto det. 2020 PARATYPE ”.</p> <p>Description. Body: Circular, dorsum convex. BL: 1.18 mm, EL: 0.86 mm, EW: 0.90 mm, PL: 0.32 mm, PW: 0.77 mm. Colour: Dorsal side light reddish-brown; venter light reddish-brown, tibia light reddish-brown, tarsal claws light brown.</p> <p>Dorsum (Fig. 5). Head: Partially retracted into pronotum; overall surface smooth, very few punctations across surface, deep groove present from the middle of the frons and extending towards the antennal insertion. Antennae: 11 segmented, clavate; 1 st and 2 nd AS longer than wide and medium sized, 3 rd to 9 th AS short and narrow, 10 th and 11 th AS enlarged. Pronotum: Transverse, widest at posterior, narrowest at anterior, narrower than elytra; anterior margin convex, posterior margin gently convex; lateral margin convex with slightly concave in the middle; surface covered with punctations, surface between punctures smooth; long and thin setae spread across surface. Scutellum: Triangular, small, longer than wide. Elytra: Slightly wider than long; anterior margin nearly straight; anterior lateral angle nearly right angled; lateral margins gently curve from anterior end to anterior 2/3, increase in curvature towards the apex; irregular surface with faint elytral striae punctuations; surface near elytral tip with small grooves; long and thin setae spread across surface.</p> <p>Venter (Fig. 10). Prosternum: Concave on anterior margin; prosternal process as long as wide, narrows towards apex, posterior margin rounded. Mesoventrite: Wider than long; anterior margin concave where prosternal process fits; lateral sub-parallel and gently concave, posterior slightly convex. Metaventrite: Wider than long, flat in the middle; relatively large and shallow punctures spread across surface. Abdomen: Five clearly visible ventrites. Intercoxal process triangular with rounded apex. Posterior margin of 1 st ventrite flat; 2 nd to 4 th ventrites concave, 5 th ventrite nearly semi-circle, posterior end flat. Surface covered with short, brown, setae on the outer margin (Fig. 20).</p> <p>Legs. Tibiae: Outer lateral margin almost straight, inner lateral margin straight. Tarsi: Simple, segments increase in length towards the apex. Tarsal claws: Simple, narrow, symmetrical.</p> <p>Male. Genitalia (Figs. 25A‒25C). Length: 0.49 mm. Parameres very much reduced. Median lobe long and thin, width constant and parallel in ventral view, apical end pointed; strongly curved in lateral view; apical end pointed. Phallobase oval.</p> <p>Female. No external morphological difference from male.</p> <p>Distribution. C. morettoi sp. nov. is known only from the type locality.</p> <p>Etymology. This species is named after Philippe Moretto, specialist in African dung beetles, who collected these specimens.</p> <p>Differential diagnosis. Combination of key characters which can differentiate the new species from other species of the genus are displayed in Table 1.</p> <p>Remarks. This species is the first country record of this genus for Côte d’Ivoire.</p></div> 	https://treatment.plazi.org/id/B436B30CFFF1FFAD6DB2F91BFBAFC7FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
B436B30CFFF0FFA16DB2F9F5FBF0C006.text	B436B30CFFF0FFA16DB2F9F5FBF0C006.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora smithi Matsumoto 2021	<div><p>Chaetophora smithi sp. nov. [Figures 6, 11, 16, 21, 26A‒26C, 28]</p> <p>Type locality. Zambia, Nkwaji.</p> <p>Type material. Holotype male, deposited at BMNH “ ZAMBIA, 1316m, Nkwaji, open Dambo, S11°31’21”; E24°32’32”, 29.x-3.xi.13 // Yellow Pan, leg. Smith,R., Takano,H., Chmurova,L. &amp; Smith,L. BMNH(E) 2015-19 // NHMUK013996862”. Additional label on red paper added “ Chaetophora smithi sp. nov. K. Matsumoto det. 2020 HOLOTYPE ♂ ”.</p> <p>Description. Body: Elongate, oval, dorsum convex. BL: 1.59 mm, EL: 1.21 mm, EW: 1.30 mm, PL: 0.38 mm, PW: 1.03 mm. Colour: Dorsal side dark brown to reddish-brown; venter brown to reddish brown, tibia reddishbrown, tarsal claws brown.</p> <p>Dorsum (Fig. 6). Head: Partially retracted into pronotum; overall surface smooth, very few punctations across surface, deep groove present from the middle of the frons and extending towards the antennal insertion. Antennae: 11 segmented, clavate; 1 st and 2 nd AS longer than wide and medium sized, 3 rd to 9 th AS short and narrow, 10 th and 11 th AS enlarged. Pronotum: Transverse, widest at posterior, narrowest at anterior, narrower than elytra; anterior margin convex, posterior margin gently convex; lateral margin sinuate with concave in the middle; surface almost smooth with few visible punctations scattered; short and slightly thick setae spread across surface. Scutellum: Triangular, small, longer than wide. Elytra: Slightly wider than long; anterior margin concave; anterior lateral angle obtuse; lateral margins gently curved throughout towards the apex; majority of surface smooth; clearly visible elytral striae punctuations; surface near elytral tip with small grooves; short and slightly thick setae spread across surface.</p> <p>Venter (Fig. 11). Prosternum: Concave on anterior margin; prosternal process as long as wide, narrows towards apex, posterior margin rounded. Mesoventrite: Wider than long; anterior margin concave where prosternal process fits; lateral sub-parallel and gently concave, posterior slightly convex. Metaventrite: Wider than long, flat in the middle; relatively medium sized punctures spread across surface. Abdomen: Five clearly visible ventrites. Intercoxal process triangular with flat apex. Posterior margin of 1 st ventrite flat; 2 nd to 4 th ventrites strongly concave, 5 th ventrite nearly semi-circle, posterior end flat. Surface covered with short, brown, setae on the outer margin (Fig. 21). Legs. Tibiae: Outer lateral margin almost straight, inner lateral margin straight. Tarsi: Simple, segments increase in length towards the apex. Tarsal claws: Simple, narrow, symmetrical.</p> <p>Male. Genitalia (Figs. 10‒12). Length: 0.67 mm. Parameres very much reduced. Median lobe long and thin, width narrows towards the apical end, apical end pointed and pointed towards the lateral side in ventral view; thin in apical half, thicker in basal half, sinuate in lateral view; apical end pointed. Phallobase square with rounded lateral end.</p> <p>Female: Unknown.</p> <p>Distribution: C. smithi sp. nov. is known only from the type locality.</p> <p>Etymology: This species is named after Richard E.L. Smith, who is the founder of the African Natural History Research Trust (ANHRT) and collected this new species.</p> <p>Differential diagnosis: Combination of key characters which can differentiate the new species from other species of the genus are displayed in Table 1.</p> <p>Remarks. This species is the first country record of this genus for Zambia.</p></div> 	https://treatment.plazi.org/id/B436B30CFFF0FFA16DB2F9F5FBF0C006	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
B436B30CFFFFFFA36DB2F953FDC5C35E.text	B436B30CFFFFFFA36DB2F953FDC5C35E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora russelli (Fiori 1978)	<div><p>Chaetophora russelli (Fiori, 1978) [Figures 27A‒27F, 28]</p> <p>Type locality. South Africa, Rivonia.</p> <p>Type material. Holotype, deposited at BMNH “Holo-type // Holotype // S. AFRICA: TVL. Rivonia 9.xi.1968 M.I.Russell // Brit. Mus. 1972-258 // Syncalyptidae M.I.Russell det. 1974 // Syncalypta russelli g. Fiori det. G. Fiori // NHMUK011520816”. Paratype, deposited at BMNH “Para-type // S. AFRICA: TVL. Rivonia 9.xi.1968 M.I.Russell // Brit. Mus. 1972-258 // Syncalyptidae M.I.Russell det. 1974 // Syncalypta russelli g. Fiori det. G. Fiori // NHMUK011520817”.</p> <p>Remarks. The internal contents and abdomen of the holotype and paratype are missing. This species may belong to a different genus from Chaetophora, because of the following characters: 1) six short spikes on the pronotum (Fig. 6C); 2) concave area extending longitudinally along middle of the pronotum (Fig. 6B); 3) coxal cavities appear slightly different in shape from those of Chaetophora morettoi sp. nov. (Figs. 1D &amp; 6D); 4) lateral sides of the elytra parallel, rather than rounded; 5) strong punctuation and rugose surface of the lateral parts of elytra.</p> <p>Not all other genera within Syncalyptinae have been studied, therefore it is not possible at this stage to confirm the generic placement of this species.</p></div> 	https://treatment.plazi.org/id/B436B30CFFFFFFA36DB2F953FDC5C35E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
B436B30CFFFEFFA36DB2FD38FA63C735.text	B436B30CFFFEFFA36DB2FD38FA63C735.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetophora Kirby & Spence 1817	<div><p>Key to currently known species of Afrotropical Chaetophora</p> <p>1. Outer margin of elytra parallel from anterior end to anterior 2/3............................................... (2) Outer margin of elytra rounded throughout............................................................... (3)</p> <p>2. Apex of the median lobe split, individually slightly rounded................................. C. russelli (Fiori, 1978) Apex of the median lobe singular and pointed..................................... C. natalensis Jäger &amp; Pütz, 2013</p> <p>3. Length of aedeagus ≥ 0.55 mm........................................................... C. morettoi sp. nov. Length of aedeagus ≤ 0.55 mm........................................................................ (4)</p> <p>4. Basal half of aedeagus thick in lateral view............................................................... (5) Basal third or less of aedeagus thick in lateral view........................................................ (6)</p> <p>5. Aedeagus gently curved in lateral view............................................... C. fiorii Jäger &amp; Pütz, 2013 Aedeagus sinuate in lateral view............................................................ C. smithi sp. nov.</p> <p>6. Whole aedeagus consistently curved......................................................... C. milesi sp. nov. Majority of the median lobe straight in lateral view....................................... C. medleri Johnson 1978</p></div> 	https://treatment.plazi.org/id/B436B30CFFFEFFA36DB2FD38FA63C735	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsumoto, Keita	Matsumoto, Keita (2021): New species, new records and notes of Afrotropical Chaetophora Kirby & Spence 1817 (Coleoptera: Byrrhidae: Syncalptinae). Zootaxa 5067 (2): 211-223, DOI: 10.11646/zootaxa.5067.2.3
