taxonID	type	description	language	source
F70987CA151DFFB5A5B9321FFC88FE73.taxon	type_taxon	Type and only species: Fuscopolystoechotes reticulatus Xu, Zheng, Shih & Wang, sp. nov.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151DFFB5A5B9321FFC88FE73.taxon	diagnosis	Diagnosis. Forewing elongated, slightly broad with length-width ratio about 3; membrane fuscous, with the distinct transparent patches along the anterior and posterior margins; costal space narrow at base and then expanded to the max in proximal 1 / 5 of forewing, and subsequently narrowed gradually, with a distinctly shrinking region in distal half; humeral veinlet recurrent; presence of ScA; costal crossveins mostly biforked distally; absence of scp-ra crossvein; RP with more than 10 branches, RP 1 deeply branched at the proximal half; crossveins in radial sector irregularly arranged except for the outer gradate series; M forked near the separated point of RP 1 from RP; MA and MP bearing the similarly distal pectinate branches; Cu forked before the separated point of RP; CuA distinctly longer than CuP, forming the pectinate branches in distal; CuP with 3 – 4 pectinate branches, and each forming the secondary sub-branches; A 1 dichotomously branched distally, A 2 forked close to the base, A 3 single branch, with several distal minor sub-branches.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151DFFB5A5B9321FFC88FE73.taxon	etymology	Etymology. The new generic name is a combination of the Latin prefix fusc- and polystoechotes (the representative genus of the Polystoechotes genus-group), referring to dark brown markings of forewing. The gender is masculine.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151DFFB5A5B9321FFC88FE73.taxon	discussion	Remarks. The new genus is clearly assigned to the Polystoechotes genus-group of Ithonidae for the combination of following characters: 1) forewing distinctly elongated (cf. relatively broad in Ithone genus-group); 2) absence of interlinked veinlets in between costal crossveins (cf. commonly present in Rapisma genus-group and Principiala genus-group); 3) Sc and RA fused distally (cf. generally separated in Ithone genus-groups and Rapisma genus-groups); 4) crossveins in radial sector regularly arranged or relatively sparse (cf. abundant and commonly irregularly arranged in other groups). The new genus is characterized by its special markings of forewing that is distinctly different from other genera of Polystoechotes genus-group except for Lichenipolystoechotes. However, the forewing markings in the new genus are distinctly broader and more regular comparing to Lichenipolystoechotes. In addition, their venation are also quite different, e. g., the branched costal crossveins, and the more complicated configuration of CuP, to justify setting up a new genus.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151EFFB4A5B936DBFD68FD13.taxon	description	(Fig. 1)	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151EFFB4A5B936DBFD68FD13.taxon	etymology	Etymology. The specific name reticulatus is a Latin word meaning “ reticulated ”, referring to the reticulated pattern present on forewing of this species.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151EFFB4A5B936DBFD68FD13.taxon	materials_examined	Material. Holotype. CNU−NEU−NN 2016005 (Fig. 1 A – B); Paratype. CNU−NEU−NN 2016008 P / C (Fig. 1 C), CNU−NEU−NN 2016009 (Fig. 1 D). Type locality and horizon. Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation, Callovian-Oxfordian boundary, late Middle Jurassic.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151EFFB4A5B936DBFD68FD13.taxon	diagnosis	Diagnosis. As for the genus due to monotypy.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151EFFB4A5B936DBFD68FD13.taxon	description	Description. Forewing elongated, ca. 39 mm long and 13 mm wide; membrane dark, with the distinct transparent patches along anterior and posterior margins; trichosors well-defined in most margin except for the proximal half of anterior margin; humeral vein recurrent, producing 4 – 5 short crossveins; costal crossveins with the distal forks, and absence of interlinked crossveins in between; space between Sc and RA relatively broad, sc-sa crossvein absent; ra-rp crossveins numerous; RP with 18 close-set branches, crossveins in RP area numerous and irregularly arranged, only one outer gradate series present; M forked close to the divergence of RP 1 from RP; MA with 4 distal branches, MP with 6 distal branches; Cu proximally branched; CuA with about 9 oblique pectinate branches; CuP dichotomously branched; A 1 – A 3 multi-dichotomously branched distally. Only forewing present, body features unknown.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB4A5B9343BFB2FFC7B.taxon	type_taxon	Type species. Lichenipolystoechotes angustimaculatus Fang, Zheng & Wang, 2020 Other included species. Lichenipolystoechotes ramimaculatus Fang, Ma & Wang, 2020	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	description	(Figs. 2, 3)	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	etymology	Etymology. The specific name fenestratus is a Latin word meaning “ window-like ”, referring to the diaphanous interior fenestrae in the forewing pattern of this species.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	materials_examined	Material. Holotype. CNU−NEU−NN 2020003 (Fig. 2); Paratype. CNU−NEU−NN 2020004 (Fig. 3). Type locality and horizon. Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation, Callovian-Oxfordian boundary, late Middle Jurassic.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	diagnosis	Diagnosis. Forewing broad, length-width ratio about 2.7; RP with about 36 branches; crossveins in radial sector rare except for two rows of outer and inner gradate series; RP 1 deeply branched.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	description	Description. Forewing oval shaped, about 26 mm long, and 9.4 mm wide; trichosors well-defined; membrane with the distinctive disruptive markings, forming 14 – 15 transparent fenestrae along the margin and five large fenestrae in the middle; humeral veinlet recurrent; costal crossveins mostly simple except for the distal part; ScA present; ScP and RA fused distally; the region between ScP and RA relatively narrow, no crossveins present in between; eleven crossveins preserved between RA – RP area; RP with 36 close-set branches; crossveins in RP area relatively scarce, only one outer gradate series completely present, and the inner gradate series also partly present; the first branch of RP forked deeply with several bifurcated branches; M bifurcated basally, forming relatively abundant pectinate branches distally; CuA branched in middle, forming ca. 10 oblique pectinate branches; CuP simply biforked once, with distal bifurcated branches; A 1 and A 2 well-developed, forked three times each; A 3 residually present. The paratype shows a lighter membrane with similar markings. The venations of holotype and paratype are basically the same with an exception of a plausible difference, i. e. the absence of gradate series of crossveins in paratype. However, this character is possibly not genuine because the crossveins in RP area are commonly difficultly visible owing to the overlapped wings.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
F70987CA151FFFB3A5B934D3FC1EFF27.taxon	discussion	Remarks. The new species can be easily attributed to Lichenipolystoechotes for the typical markings and highly similar venation, i. e. the length of the wing, the presence of complicated disruptive markings and diverse fenestrae in forewing, the similar configurations of M branches, the multi-branched CuA with 7 − 11 branches and the bifurcated CuP (Fang et al., 2020). Nevertheless, the new species can be differentiated from the known Lichenipolystoechotes species for the number of RP branches (36 branches in the new species vs. 18 in L. angustimaculatus and 22 branches in L. ramimaculatus). In addition, the branched pattern of the first branch of RP in the new species is also different from other species with more complicated branches.	en	Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong, Wang, Yongjie (2021): New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae). Zootaxa 5067 (2): 259-266, DOI: https://doi.org/10.11646/zootaxa.5067.2.7
