taxonID	type	description	language	source
03A8552C1B418441FF32FF78FC7A37CF.taxon	description	(Figs 1 – 3, 16, 18, 22, 23, 30, 31)	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B418441FF32FF78FC7A37CF.taxon	materials_examined	Examined specimens. 5 ♂♂, 1 ♀, Aso Commune (15.8268 N, 107.6792 E, altitude 528 m), Dong Giang District, Quang Nam Province, Vietnam, 12. v. 2014, V. T. Nguyen leg.; 12 ♂♂, 6 ♀♀, Nam Nung Nature Reserve (12.2545 N, 107.7926 E, altitude 819 m), Dak Nong Province, Vietnam, 12. v. 2020, Q. P. Ngo & Q. T. Phan leg.; 3 ♂♂, Song Thanh National Park (15.2566 N, 107.7378 E, altitude 775 m), Phuoc My Commune, Phuoc Son District, Quang Nam Province, Vietnam, 19. v. 2020, Q. P. Ngo leg.; 2 ♂♂, K Stream (11.8716 N, 107.9824 E, altitude 598 m), Ta Dung National Park, Dak Nong Province, Vietnam, 28. vi. 2020, Q. P. Ngo leg.; 1 ♂, 1 ♀, Khe Voi, Son Kim 1 Commune (18.4018 N, 105.2846 E, altitude 125 m), Huong Son District, Ha Tinh Province, Vietnam, 24. vii. 2020, Q. P. Ngo & Q. T. Phan leg.; 3 ♂♂, Khe Da Lai (18.1762 N, 105.5371 E, altitude 160 m), Phu Gia Commune, Huong Khe District, Ha Tinh Province, Vietnam, 19. vii. 2020, Q. P. Ngo & Q. T. Phan leg.; 3 ♂♂, Hoa Trung (16.0950 N, 108.0491 E, altitude 191 m), Hoa Lien Commune, Hoa Vang District, Danang City, Vietnam, 19. v. 2021, Q. T. Phan leg. Notes. This species was described from Bao Loc, Lam Dong Province, southern Vietnam, based on a male specimen (Karube 2014). The female is described based on the specimen from Nam Nung Nature Reserve, Dak Nong Province.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B418441FF32FF78FC7A37CF.taxon	description	Description of female Head. Labium greenish yellow; labrum black with a pair of yellow stripes laterally; anteclypeus greenish yellow; postclypeus black with small yellow patch each at dorso-lateral corner; frons largely greenish yellow, black medially; antennae black, ventral margin of the second segments yellowish; vertex entirely black with rounded prominence above lateral ocelli (Figs 2 – 3). Occiput with two erect horns dorsally (Figs 16, 30). Thorax (Fig. 1). Prothorax mostly black with a tiny greenish yellow spot on middle pronotal lobe. Synthorax robust, black with yellow markings; black antehumeral and humeral stripe connected, leaving only a small spot ventrally; dorsal carina yellow; interpleural stripe broad encompassing posterior half and anterior halves of mesepimeron and metepisternum respectively, this stripe interrupted medially by narrow patches of yellow; metepimeron largely yellow except for anterior and ventral margin. Wings (Fig. 1). Wings hyaline with black venation except the base tinged with yellow; anal loop 2 - celled; nodal index: Ax / Px = 17 / 21 (FW) and 14 / 23 (HW). Pt black, 4.5 – 5 mm length in both wings, covering about 8 underlying cells. Abdomen (Fig. 1). Black with greenish yellow markings as follows: S 1 with a lateral large spot; S 2 greenish yellow markings on anterior and posterior margin and part of posteroventral margin black; S 3 with a large spot on anterior margin; S 4 – 6 mostly black with a narrow basal rings; anterior half S 7 yellowish; S 8 – 10 entirely black. Anal appendages (Figs 18, 31) Cerci yellowish brown, conical, longer than S 10. Epiproct broad, shorter than S 10. Vulvar lamina on S 9, incised apically. Measurements [mm]. Abdomen (including appendages) 39, HW 32. Morphological variation in females. The variation of measements of body varies as follows: Measurements: abdomen (including appendages) 38 – 40, HW 32 – 35 mm. The basal area of wings of two specimens from Nam Nung Nature Reserve is paler than those from other localities.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B418441FF32FF78FC7A37CF.taxon	distribution	Distribution (Fig. 36). Vietnam: Ha Tinh, Thua Thien Hue, Quang Nam, Da Nang, Kon Tum, Dak Nong and Lam Dong Provinces (Karube 2014; Karube et al. 2020; this study).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	description	(Figs 4 – 11, 14, 15, 17, 19, 24, 25, 32, 33)	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	materials_examined	Holotype: ♂, Song Thanh National Park (15.5080 N, 107.4720 E, altitude 1105 m), Dak Re Commune, Nam Giang District, Quang Nam Province, central Vietnam, 28. vi. 2020, Q. P. Ngo leg. Paratypes: 1 ♀ (in copula with holotype male), same data as holotype, both in ZCDTU; 1 ♀, Song Thanh National Park (15.3113 N, 107.9097 E, altitude 725 m), Phuoc Kim Commune, Phuoc Son District, Quang Nam Province, central Vietnam, 12. vii. 2018, Alisa J. & Thomas W. Donnelly leg., Rosser W. Garrison Collection, USA.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	etymology	Etymology. The new species is named in honor of Dr. Phan Quoc Toan, a Vietnamese odonatologist, for his great contribution to the knowledge of Southeast Asian Odonata (noun in genitive case).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	description	Description of holotype male Head (Figs 4, 6, 8). Labium black, labrum black with a pair of greenish yellow markings at dorso-lateralyl; anteclypeus greenish yellow; postclypeus black with a small yellow spot at dorso-laterally; frons broadly marginated with yellow fascia lying interrupted medially with black; antennae black, yellowish basally; vertex black with two rounded prominences near lateral ocelli. Occiput black, prominent centrally. Thorax (Fig. 4). Prothorax entirely black. Synthorax mostly black with greenish yellow markings; pale antehumeral stripe and not connected with collar stripe; humeral stripe absent; dorsal carina and post-ventral part of mesinfraepisternum with yellow markings; median part of mesepimeron with pale stripe about half as wide as combined black antehumeral and humeral stripes; a small mid-dorsal yellow stripe on metepisternum; metepimeron largely greenish yellow, anterior and ventral borders black. Legs (Fig. 4). Legs stoutly built, entirely black. Femur and tibia thick and robust with closely set, gradually lengthening spurs apically. Coxa of front legs black with greenish yellow. Wings (Fig. 4). Wings hyaline, with brown patches at wing base; anal loop 2 - celled; nodal index: Ax / Px = 17 / 18 (FW) and 12 / 19 (HW); Pt dark brown, 4.5 – 5 mm length in both wings, covering about 5 or 6 - celles. Abdomen (Fig. 4). Black with greenish yellow markings; S 1 with a medio-ventral spot connecting with postventral spot; S 2 with two large lateral spots, one on auricle, the other along posterior carina; S 3 – 6 mostly black each with a narrow basal ring; S 7 with a basal spot covering about half of the segment; S 8 – 10 entirely black. Anal appendages (Figs 10, 11). Entirely black; cercus broad basally, narrower apically, gently curving ventroposteriorly, apex hooked with a dorsal small tooth near the base of cercus. Epiproct curved posterodorsally, subequal in length to cerci with a small dorsal tooth approximately at basal third. Accessory genitalia (Fig. 24). Anterior hamulus broad medially, abruptly attenuate with apical half strongly curved posteriorly. Posterior hamulus broad basally, narrower and slightly curved cephalad at apex. Vesica spermalis (Figs 14, 15, 25). Prepuce protruded ventroposteriorly, its stem slender; posterior lobe of vesica spermalis slender and short. Cornua of glans with two long, acute V-shaped flagellae (Fig. 25). Measurements (in mm). Abdomen (including appendages) 40; HW 33. Description of a paratype female Colour pattern similar to that of holotype male apart from the following characters: Head (Figs 5, 7, 9, 17, 32). The greenish yellow markings on dorsal frons much narrower (Fig. 9), (Fig. 8); occiput flattened, a projection medially on occipital crest as shown in Figs. 17 and 32, its dorsal surface minutely serrate. Thorax (Fig. 5). Prothorax entirely black. Synthorax robust, yellow making same as in holotype but lacking small mid-dorsal yellow patch on metepisternum. Wings (Fig. 5). Wings hyaline, with dark brown patches at the base; nodal index: Ax / Px = 17 / 21 (FW) and 12 / 21 (HW); Pt dark brown, 4.5 – 5 mm length in both wings, covering about 5 – 7 cells. Abdomen (Fig. 5). S 1 with lateral triangular patch; lateral S 2 laterally with a large greenish yellow marking; S 3 – 6 each with a pale basal ring; anterior fourth of S 7 yellow; S 8 – 10 entirely black. Anal appendages (Figs 19, 33). Cerci as the length of S 10, dark yellowish, slender, parallel, pointed at tip. Vulvar lamina triangular-shaped, incised apically but not forming a gap. Measurements (in mm). Abdomen (including appendages) 30 mm. HW 48 mm.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	diagnosis	Differential diagnosis. Among the known species of the genus Ophiogomphus, the new species shares the following characters with O. longihamulus, O. minimus and O. sinicus, i. e. large pincer-like appendages with cerci curving posteroventrally and epiproct curving posterodorsally, with apex notched (Figs 10, 12). Ophiogomphus phantoani can be distinguished from the other congeners by the following characters: In the male, the cornua of glans has two long, acute widely separated, V-shaped flagella (Fig. 25), whereas they are relatively short, U-shaped flagella in O. longihamulus (Fig. 21), O. minimus (Fig. 23) and O. sinicus (Fig. 27); anterior hamulus of O. phantoani with well-developed developed posterior shoulder (Fig. 24) shoulder smaller in O. longihamulus (Fig. 20), O. minimus (Fig. 22) and O. sinicus (Fig. 26). In the female, the occiput of O. phantoani has a projection with two prominent dorsally microserrated branches (Fig. 32), more widely branched than in O. longihamulus (Fig. 28), O. minimus (Fig. 30) and O. sinicus (Fig. 34); the vulvar lamina of O. phantoani differs from the other species by a pointed tip (Fig. 33), which is incised apically but not widely opened as in the other species (Figs 29, 31, 35). Habitat and ecology. Ophiogomphus phantoani was found on a small, slowly running, densely vegetated stream (about 1 – 2 m wide) within pristine forest. The stream originates from a marsh with muddy bottom. The new species shares the same habitat with Coeliccia schorri Phan & To, 2019; C. scutellum Laidlaw, 1932; Rhinocypha watsoni van Tol & Rozeendal, 1995; Burmargiolestes sp.; Mnais mneme Ris, 1916; Orthetrum glaucum (Brauer, 1865); Anotogaster chaoi Zhou, 1998; Lamelligomphus camelus Martin, 1904 and Merogomphus pavici Martin, 1904.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B428446FF32FB99FB9336EC.taxon	distribution	Distribution (Fig. 36). Vietnam: Quang Nam Province (Song Thanh National Park).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B458446FF32F95CFA9733E8.taxon	description	(Figs 20, 21, 28, 29) Notes. This species was described from Pia Oac National Park of northern Vietnam based on male and female specimens (Karube 2014). Ophiogomphus longihamulus is characterized by its long anterior hamuli (Fig. 20).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B458446FF32F95CFA9733E8.taxon	distribution	Distribution (Fig. 36). Vietnam: Cao Bang Province; China: Yunnan Province (Karube 2014; Zhang 2019).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B468445FF32FAA4FCD53337.taxon	description	(Figs 12, 13, 26, 27, 34, 35)	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B468445FF32FAA4FCD53337.taxon	materials_examined	Examined specimens. 1 ♂, Nam Toong bridge (22.7853 N, 105.8512 E, altitude 1373 m), Pia Oac National Park, Cao Bang Province, northern Vietnam, 18. v. 2015, Q. T. Phan leg. Notes. This species was described from Guangxi, China (Chao 1954) and subsequently recorded in Jiangxi, Fujian, Guangdong, and Hong Kong (Zhang 2019) and northern Vietnam on an internet blog (Kompier 2014). Ophiogomphus sinicus is the largest representative of the genus in Vietnam (measurements: Abdomen including appendages: 46 mm; Hw 37 mm). It should be noted here that, Karube (2014: p. 80) stated that the apical 2 / 3 of male cerci of Ophiogomphus sinicus from Cao Bang Province, North Vietnam are yellowish. However Wilson (per. comm.) has informed us that some specimens of O. sinicus from Hong Kong have entirely blackish-brown cerci. This may represent onto genetic or geographic variability.	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
03A8552C1B468445FF32FAA4FCD53337.taxon	distribution	Distribution. Vietnam: Cao Bang Province (Pia Oac National Park) (Fig. 36); China: Jiangxi, Fujian, Guangdong and Guangxi Provinces and Hong Kong (Zhang 2019).	en	Ngo, Quoc Phu, Nguyen, Minh Ty (2021): The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae). Zootaxa 5061 (1): 134-144, DOI: https://doi.org/10.11646/zootaxa.5061.1.6
