taxonID	type	description	language	source
684C87BBFFD1FFD5FF12FB64FC9AFE61.taxon	description	(Figs. 1, 2)	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD1FFD5FF12FB64FC9AFE61.taxon	materials_examined	Material examined. Southwest Atlantic. Female (USNM 1089979), Scotia Sea, near Sandwich Islands, Cruise 8, station 575 [54 ° 33 ’ S 24 ° 19 ’ W], 1867 m, no date; female (USNM 1090242), Cruise 8, station 640 [58 ° 10 ’ S 23 ° 39 ’ W], 1537 m, no date. Southeast Pacific. Female (USNM 1013495), western Drake Passage, Cruise 10, station 811 [64.767 ° S 78.1 ° W], 461 – 1263 m, 30 October 1963 (figured here). Southern Ocean, Pacific Sector. Female (USNM 1090243), Cruise 11, station 890 [59 ° 39 ’ S 114 ° 37 ’ W], 595 – 609 m, 3 January 1964. Strait of Magellan. Male (USNM 1090255), Cruise 11, station 957 [53 ° ’ S 75 ° 00 ’ W], 458 – 732 m, 5 February 1964 (Figs 1, 2). Type material. Lanceola loveni antarctica was described from seven specimens, 7 – 27 mm, collected by the Russian research vessel Ob from the following five stations between the Antarctic Continent and the Antarctic Polar Front. From the Indian Ocean Sector: station 36, off Wilkes Land [62 ° 55 ’ S 118 ° 52 ’ E], 0 – 3,700 m; station 105, near the Davis Sea [55 ° 40 ’ S 106 ° 13 ’ E], 240 – 600 m; station 111 [64 ° 25 ’ S 92 ° 52 ’ E], 0 – 2,000 m; station 285 [59 ° 29 ’ S 97 ° 08 ’ E], 0 – 4500 m. From the Pacific Sector: station 57, near the Balleny Islands [64 ° 03 ’ S 161 ° 59 ’ E], 0 – 2,000 m and 0 – 3,000 m. The syntypes from stations 36 and 57 are in the Zoological Museum of the Moscow State University (Mb – 1049 and 1050). A female is illustrated by Vinogradov (1962).	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD1FFD5FF12FB64FC9AFE61.taxon	description	Description of male 25 mm (Fig. 1). Distal corner of pereonites 6 and 7, pleonites and first urosomite produced into small, distinct, dorsal denticles. Head with relatively long, rounded rostrum. Eyes very small, reniform, without crystalline cones. A 1 slightly longer than head and first pereonite combined. A 2 length 2 x A 1, almost as long as head and first two pereonites combined. G 1; basis relatively slender, both margins fringed with long setae, about as long as remaining articles combined; carpus length 0.6 x basis, distal margin width about 1.5 x proximal margin of propodus; propodus conical-shaped, length 1.6 x proximal width; dactylus very slender, almost as long as 0.5 x propodus. G 2 with relatively long, slender articles, much longer than G 1; basis length about 1.5 x carpus; propodus gradually tapering to narrow distal margin, length about 0.8 x carpus; dactylus inserted terminally to propodus, length slightly less than 0.3 x propodus. P 3 length about 1.2 x G 2, with similar slender articles; basis length 1.5 x merus; carpus length 0.8 x merus; propodus slightly longer than carpus; dactylus inserted terminally to propodus, length marginally less than 0.2 x propodus. P 4 morphologically similar to P 3, marginally shorter, with slight differences in relative lengths of articles as follows: basis length 1.3 x merus; carpus length marginally more than 0.8 x merus; propodus and carpus sub-equal in length. P 5 – 7 all with relatively long, slender articles and fully retractile, hooded dactyls. P 5 length 1.2 x P 4; basis length almost 1.5 x merus; carpus length 0.8 x merus, 1.4 x propodus; dactylus without setae. P 6 length 1.2 x P 5, almost as long as pereon and first two pleonites combined; basis length about 1.3 x merus; carpus length almost 0.8 x merus, 0.8 x propodus; dactylus inserted in slight bulge of propodus, fringe of short setae on anterior margin. P 7 about 0.5 x P 6 length; basis length 1.6 x merus; carpus marginally shorter than merus, about 0.8 x propodus; dactylus with fringe of short setae on anterior margin. Urosome with relatively slender peduncles and rami. Rami of all uropoda sub-equal in length. Peduncle of U 1 marginally exceeds limit of peduncle of U 2. Peduncle length of U 1 = 1.8 x rami, of U 2 = 1.5 x rami, of U 3 = 1.3 x rami. Telson acute, apex rounded; length about 1.3 x width at base, reaching to about middle of peduncle of U 3. Description of female 26 mm (Fig. 2). Distal corner of pereonites 6 and 7, pleonites and first urosomite produced into small, distinct, dorsal denticles. Head with short, rounded rostrum. Eyes very small, reniform, without crystalline cones. A 1 slightly longer than head and first pereonite combined. A 2 length 2.4 x A 1, slightly longer than head and first two pereonites combined. G 1; basis relatively slender, fringed with long setae on both margins, about as long as remaining articles combined; carpus length 0.5 x basis, distal margin width about 1.4 x proximal margin of propodus; propodus conical-shaped, length 1.6 x proximal width; dactylus missing. G 2 with relatively long, slender articles, much longer than G 1; basis length about 1.8 x carpus; propodus gradually tapering to narrow distal margin, length almost 0.8 x carpus; dactylus inserted terminally to propodus, length slightly less than 0.3 x propodus. P 3 slightly longer than G 2, with similar slender articles; basis length 1.6 x merus; carpus length 0.8 x merus; propodus slightly longer than carpus; dactylus missing. P 4 morphologically similar to P 3 but marginally longer, with slight differences in relative lengths of articles as follows: basis length about 1.3 x merus; carpus slightly shorter than merus; propodus slightly longer than carpus; dactylus missing. P 5 – 7 all with relatively long, slender articles and fully retractile, hooded dactyls. P 5 length almost 1.2 x P 4; basis length 1.5 x merus; carpus length almost 0.8 x merus, 1.2 x propodus; dactylus without setae. P 6 length 1.2 x P 5, almost as long as pereon and pleon combined; basis length about 1.3 x merus; carpus length 0.8 x merus; propodus as long as merus; dactylus inserted in slight bulge of propodus, too dense to determine presence of setae. P 7 almost 0.5 x P 6; basis length 1.7 x merus; carpus marginally shorter than merus, about 0.8 x propodus; dactylus without fringe of short setae on anterior margin. Urosome with relatively slender peduncles and rami. Rami of all uropoda sub-equal in length. Peduncle of U 1 slightly exceeds limit of peduncle of U 2. Peduncle lengths of U 1 = about 2.0 x rami; of U 2 = 1.6 x rami; of U 3 = 1.4 x rami. Telson acute, with rounded apex; length about 1.4 x width at base, reaching to about the middle of peduncle of U 3. The main differences from the male, apart from minor differences in the relative lengths of articles, are as follows: i) the head has a shorter rostrum; ii) A 2 are relatively longer; iii) the basis of G 2 is relatively longer; iv) P 3 is slightly shorter than P 4; v) P 5 – 7 are relatively marginally longer compared to the body length; vi) the dactylus of P 7 is without setae (maybe also P 6, not discernable); and vii) the peduncles of U 1 – 3 are marginally longer compared to the rami.	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD1FFD5FF12FB64FC9AFE61.taxon	discussion	Remarks. The most obvious difference from the typical form is that individuals of all ages have the distal corner of the pleonites produced into small dorsal denticles, similar to L. serrata Bovallius, 1885, but that species is distinguished by the much longer telson (slightly longer or shorter than the peduncle of U 3) and the wide distal margin of the carpus of G 1. Other distinguishing characters from the typical form are as follows: i) the head has a more prominent rostrum, especially for males; ii) the carpus and propodus of G 1 are slightly more elongate; iii) the propodus of P 6 is more club-like, similar to L. loveni grossipes Shoemaker, 1945; iv) the telson is marginally more narrow; and v) there are also minor differences in the relative lengths of appendages and articles, although these may be within the confines of specific variability. The relatively long rostrum of the male, illustrated here, is particularly distinctive and may be an unusual feature of this specimen. Whether or not it is a genuine male character can only be determined with the discovery of more material. Vinogradov et al. (1982, 1996) list L. australis Stebbing, 1888 as a questionable synonym of this species, probably based on the presence of dorsal denticles on the pleonites. However, an examination of the type material in the Natural History Museum, London has confirmed that it is a synonym of L. serrata (Zeidler 2009).	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD1FFD5FF12FB64FC9AFE61.taxon	distribution	Distribution. This species seems to be restricted mainly to the Antarctic Sectors of the Southern Ocean. It has been recorded in the Atlantic Sector from the Scotia Sea to the Drake Passage and from there to the Pacific Sector as far as 114 ° 37 ’ W, and in the Indian Ocean Sector from off Wilkes Land and near the Davis Sea; from various hauls from 4,500 m to the surface, as detailed above. The male, illustrated here (USNM 1090255), from the Strait of Magellan, is a new distribution record for this species.	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD5FFD7FF12FE1CFBB3FD65.taxon	materials_examined	Material examined. Southern Ocean: immature female (USNM 1090248), Amundsen Sea, Cruise 11, station 930 [70 ° 08 ’ S 109 ° 48 ’ W], 1,391 m, 19 January 1964 (figured here). SE Pacific: immature female (USNM 1089971), off Chilean coast near Valdivia, Cruise 6, station 326 [38 ° 19 ’ S 74 ° 34 ’ W], 1,691 – 2,072 m, 25 November 1962. Type material. Lanceola sphaerica was first described as a variety of L. clausi from two damaged specimens, a mature female (8 mm) and a male (9 mm), both caught in the Kurile-Kamchatka region of the North-West Pacific. The female from Vityaz station 2076 [43 ° 39 ’ N 149 ° 24 ’ E], 4,200 – 7,800 m, 9 May 1953 and the male from Vityaz station 2208 [49 ° 23 ’ N 158 ° 45 ’ E], 0 – 6,500 m, 22 June 1953 (Vinogradov 1957). Later, Vinogradov (1970) established it as a full species with the availability of a well-preserved male (5.5 mm) from the same general area; Vityaz station 5626 [45 ° 11 ’ N 152 ° 28 ’ E], 3,000 – 4,000 m, 23 – 24 August 1966. All three specimens are in the Zoological Museum of the Moscow State University (Mb – 1051, 1052, 1053).	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD5FFD7FF12FE1CFBB3FD65.taxon	description	Description of immature female 4.5 mm (Fig. 3). Pereonites 1 – 4 inflated, almost spherical. Head without rostrum. Eyes very small, reniform, without crystalline cones. A 1 slightly longer than head and first pereonite combined. A 2 as long as A 1 with two long setae terminally. G 1; basis relatively slender, slightly shorter than remaining articles combined; carpus length 0.5 x basis, distal margin width about 1.3 x proximal margin of propodus; propodus conical-shaped, length about 2 x proximal width; dactylus length almost 0.5 x propodus. G 2 much longer and slenderer than G 1; basis length about 0.7 x remaining articles combined, about 2 x carpus length; propodus gradually tapering to narrow distal margin, length about 1.4 x carpus; dactylus inserted terminally to propodus, length about 0.4 x propodus. P 3 and P 4 similar in length and morphology; basis length slightly more than 2 x merus; merus with distinct antero-distal bulge, maximum width about 0.8 x length; carpus length 1.6 x merus for P 3, 1.8 x merus for P 4; propodus length 1.3 x carpus for P 3, 1.1 x carpus for P 4; dactylus length slightly more than 0.3 x propodus. P 5 – 7 all with relatively long, slender articles and fully retractile, hooded dactyls but propodus not club-like distally. P 5 slightly shorter than P 4; basis length almost 2 x merus; carpus length 1.3 x merus, 0.8 x propodus; dactylus in relatively broad hollow. P 6 length almost 1.3 x P 5, slightly longer than first four pereonites combined; basis length about 2 x merus; merus with very slight postero-distal bulge; carpus length 1.5 x merus; propodus length about 1.4 x merus; dactylus strongly curved, pressed against anterior margin of propodus. P 7 length 0.7 x P 6; basis length 2.5 x merus; merus with slight postero-distal bulge; carpus length 1.2 x merus, about 0.6 x propodus. Urosome with relatively slender peduncles and rami. Rami of all uropoda more or less sub-equal in length, with slightly serrated margins. Peduncles and rami of U 1 & U 2 similar in length. Peduncle length of U 3 slightly less than rami. Telson acute, apex rounded; length about 1.5 x width at base, reaching to about the middle of peduncle of U 3.	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD5FFD7FF12FE1CFBB3FD65.taxon	discussion	Remarks. Lanceola sphaerica is distinguished from all its congeners by a combination of several characters; i) the inflated pereon (especially in females); ii) the relatively long dactyls of P 3 – 7 (especially P 3 – 4 in mature males); iii) the structure of P 5 becoming sub-chelate in males; and iv) the relatively short telson. It is clearly most similar to the highly variable species, L. clausi, in the general habitus and the relative lengths of the pereopods but is readily distinguished by the slightly more inflated pereon and the relatively longer dactyls of P 3 – 7. The female specimen described here is similar to L. clausi pirloti Shoemaker, 1945 in that the antennae are of similar length and the merus of P 3 – 4 has a distinct antero-distal bulge, but differs from it in that the basis of the gnathopoda is slender; P 6 is relatively longer and the uropoda are slenderer. In the long dactyls of P 3 and P 4, it resembles L. longidactyla Vinogradov, 1964 but in that species the dactyls are relatively much longer and P 5 is not modified. It also resembles L. chelifera Vinogradov, 1970, especially in the modification of P 5 but in that species the dactylus of P 5 is not retractile or hooded and the structure of P 3 and P 4 is completely different. As noted by Vinogradov (1970), females of L. sphaerica differ from males in the following characters: i) P 3 and P 4 are more weakly armed and the dactyls are relatively shorter; ii) the articles of P 5 are more linear, with less armature and the propodus is not robust and the dactylus is not as strong, not becoming sub-chelate. The specimen described here agrees with these characters and there are only very minor differences in the relative lengths of articles that can be determined from the limited information provided for the female by Vinogradov (1957, 1970). The sexual dimorphism exhibited in the morphology of P 5 seems to become more prominent in mature males. Like L. clausi, L. sphaerica seems to occur in the colder waters of both Hemispheres.	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
684C87BBFFD5FFD7FF12FE1CFBB3FD65.taxon	distribution	Distribution. Known only from the Kurile-Kamchatka region of the Pacific Ocean, the Amundsen Sea, Antarctica and the South-East Pacific, as detailed above. The latter is a new distribution record for L. sphaerica. Vinogradov (1970) records it as “ one of the most deeply-living hyperiids, being restricted to the abyssopelagic zone (including the trench waters) and does not rise above 3,000 m into the bathypelagic ”, based on the specimens from the Kurile-Kamchatka region. It is recorded here from 1,391 m and 1,691 – 2,072 m.	en	Zeidler, Wolfgang (2021): Redescription of two rare, deep-sea species of hyperiidean amphipod, Lanceola loveni antarctica Vinogradov, 1962 and L. sphaerica Vinogradov, 1970 (Crustacea: Amphipoda: Hyperiidea: Lanceolidae). Zootaxa 5067 (1): 106-114, DOI: https://doi.org/10.11646/zootaxa.5067.1.6
