identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A05F87AD171DFD76D6C9F9F5905BFDA3.text	A05F87AD171DFD76D6C9F9F5905BFDA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leistus (Pogonophorus) spinibarbis subsp. zagrosensis Muilwijk & Seiedy & Wrase 2021	<div><p>Leistus (Pogonophorus) spinibarbis zagrosensis ssp. n.</p> <p>urn:lsid:zoobank.org:act: AEF2F2E3-B739-4A34-8D34-36C96041C008</p> <p>(Fig. 1.1, 1.2)</p> <p>Type material. Holotype ♂, labelled: “ Iran, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.16639&amp;materialsCitation.latitude=32.180836" title="Search Plazi for locations around (long 50.16639/lat 32.180836)">Chahar Mahaal</a> and Bakhtiari / Chery pass, 31.v.2019, 3260 m / Muilwijk leg. / 32°10ˊ51´´ N, 50°09ˊ59´´ E ” // “ Holotype / Leistus (Pogonophorus) spinibarbis / zagrosensis ssp. n. / Muilwijk, Seiedy &amp; Wrase des. 2021” (black print on red label). Deposited in HMIM.</p> <p>Description. Upper surface of head and pronotum dark with bright blue-violet lustre, elytra more brownish with blue lustre; lateral margins of pronotum translucent reddish; propleura yellow, epipleura brown, appendages entirely yellow, underside dark except brownish apical sternites (Fig. 1.1). BL: 8.4 mm, EW: 3.3 mm.</p> <p>1.1 habitus holotype, scale: 5 mm.</p> <p>1.2 median lobe holotype, lateral view, scale: 1 mm.</p> <p>Head punctate, longitudinally superficial rugulose; eyes hemispherical; antennae long and slender.</p> <p>Pronotum transverse (PW/PL:1.37); punctuation on the base, lateral margins and near the apex strong, punctation on the disc rather fine; lateral furrow broad, sides narrowed to the rectangular basal angles. Middle line fine and only visible on disc.</p> <p>Elytra subparallel (EL/EW:1.7); weakly rounded laterally, their maximum width apical of middle; basal margin concave, humeral denticle small and sharp; striae rather deep, superficial apically, evidently punctate, intervals moderately convex, flattened near the apex with fine punctuation.</p> <p>Ventral side with dense and rather strong punctuation on prosternum, pro-, mes- and metepisterna, mesosternum, lateral sides of metasternum, lateral sides of abdominal sternites 1 and 2, lateral sides of sternite 3 with finer punctation, middle of sternite 3 and sternites 4–7 with fine scarce punctation; prosternal lobe pointed and not bordered at the tip with brown colour.</p> <p>Microsculpture superficially, isodiametric on head, and pronotal disk; elytra with less superficially, isodiametric meshes.</p> <p>Median lobe (Fig. 1.2) with a regularly concave apex.</p> <p>Comparative notes. L. (P.) spinibarbis zagrosensis ssp. n. differs from the other subspecies, Leistus (P.) spinibarbis rufipes Chaudoir 1843, L. (P.) spinibarbis ponticus Kryzhanovskij &amp; Shilenkov, 1999 and L. (P.) spinibarbis abdominalis Reiche &amp; Saulcy, 1855, in the Middle East by the concave apex of the median lobe, which is clearly asymmetrical in the other subspecies. In addition, L. (P.) spinibarbis rufipes differs from L. (P.) spinibarbis zagrosensis ssp. n. by shorter antennae; antennomere 3 shorter and thicker; more transverse pronotum; elytral sides more rounded; prosternal lobe apically rounded and completely bordered and black coloured. L. (P.) spinibarbis ponticus differs from L. (P.) spinibarbis zagrosensis ssp. n. by more coarsely and densely punctation of head, head and pronotum brownish black, elytra less parallel, widest point in the middle. L. (P.) spinibarbis abdominalis differs from L. (P.) spinibarbis zagrosensis ssp. n. by blue-green elytra, elytral striae more finely and densely punctured, the intervals completely flat and ventral side completely brown-red.</p> <p>Distribution. Iran (Chaharmahal and Bakhtiari). So far only known from collecting place.</p> <p>Habitat. The specimen was collected under a stone at high altitude, near a small snow field.</p> <p>Etymology. The name of the subspecies refers to the Zagros Mountains, where the specimen was collected.</p></div> 	http://treatment.plazi.org/id/A05F87AD171DFD76D6C9F9F5905BFDA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD171BFD74D6C9FD419102FE33.text	A05F87AD171BFD74D6C9FD419102FE33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trechus kouroshi Muilwijk & Seiedy & Wrase 2021	<div><p>Trechus kouroshi sp. n.</p> <p>urn:lsid:zoobank.org:act: 83977913-FB4E-48E7-AC91-5C77E066A015</p> <p>(Figs. 2.1, 2.2)</p> <p>Type material. Holotype, ♂ labeled: “ Iran, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.427197&amp;materialsCitation.latitude=30.823862" title="Search Plazi for locations around (long 50.427197/lat 30.823862)">Kohgiluyeh</a> and Boyer-Ahmad / Suq cave, 812 m / 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E, 5.vi.2019 / S. Rahideh, Y. Fatemii &amp; J. Muilwijk leg.” // “ Holotype / Trechus kouroshi sp. n. / Muilwijk, Seiedy &amp; Wrase des. 2021” (black print on red label), deposited in HMIM. Paratype, 1 ♀ with same data but “ Paratype ”, in CJM.</p> <p>Comparing material: Trechus saulcyanus Csiki, 1928: 1 ♂, Israel, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.040554&amp;materialsCitation.latitude=33.039165" title="Search Plazi for locations around (long 35.040554/lat 33.039165)">Upper Galilee Ya'ar Bar'am</a>, 683 m, 23.iii.2011, 33°02´21´´ N, 35°2´26´´ E; Meybohm leg.; W. Ziegler det. 2011. CJM.</p> <p>Description. A small to middle sized and bicoloured Trechus species (3.5 mm) with piceous brown head and pronotum and yellowish elytra; appendages yellow; pronotum slightly shiny; micropterous (Fig. 2.1).</p> <p>Head eyes slightly convex, much longer than tempora (tempora/eye: 0.33–0.36). Frontal furrows moderately deep, evenly impressed, semi-circular. Clypeus with two pairs of setae, located in a slight deepening of the clypeus. Two pairs of supraorbital setae, the anterior setae at level of the middle of the eye, the posterior setae behind the eye. Antennae short (1.95 mm), reaching to one quarter of the elytra.</p> <p>Pronotum cordate (PW/PL: 1.28–1.37); slightly convex. Lateral sides of pronotum sinuate towards posterior angles, posterior angles distinct, nearly rectangular, apically upwards pointed. Anterior angles rounded. Posterior margin slightly emarginate laterally. Basal fovea deep. Middle line deep on disc, not reaching the anterior edge; lateral margin with 2 pairs of setae, one pair in the posterior angles, the other pair at the widest point.</p> <p>Elytra oblong-ovate (EL/EW: 1.3–1.4); humeri prominent, though rounded; maximum width around the middle; the marginal border reaching anteriorly interval 5. Three setiferous punctures in intervals, the anterior 2 pores situated along striae 3, the preapical pore connects stria 2 with 3; all striae punctated; striae 1–2 deep, ending at apical setiferous punctures, striae 3–4 superficial, towards apex shortened, striae 5–6 consists of a row of punctures, strongly shortened, stria 7 indistinct, consisting of a few punctures in the middle. Parascutellar striole short, superficially; parascutellar pore small surrounded through a keel formed by the thickened suture. The deep recurrent striole not reaching stria 5; series umbilicata with 8 setiferous punctures, 4 punctures at the base, 2 punctures just behind the middle, and 2 punctures apically.</p> <p>Microsculpture of head with isodiametric meshes, pronotum with more transverse meshes and elytra with superficial, transverse meshes.</p> <p>Legs slender; protarsomeres 1–2 dilated in male, each with a massive inner and obtuse outer tooth. Front tibiae grooved on exterior surface.</p> <p>Ventral side underside colour yellow except the dark meso- and metasternum and mes- and metepisternum.</p> <p>Median lobe (Fig. 2.2) small, 0.5 mm long, regularly bent, apex in vertical view asymmetric. Sclerites of endophallus in lateral view (l) triangular plate shaped, in dorsal view (d) and ventral view (v) arrow shaped, pointed to the apex. Left paramere with three setae, right paramere with four setae.</p> <p>2.1 habitus holotype, scale: 2 mm.</p> <p>2.2 median lobe holotype, l = lateral, v =ventral, d= dorsal view, scale: 0.5 mm.</p> <p>2.3 Trechus saulcyanus, median lobe, l = lateral, v =ventral view, scale: 0.5 mm; Israel.</p> <p>Comparative notes. So far only three Trechus species are known from Zagros: T. liopleurus Chaudoir, 1850, T. melanocephalus Kolenati, 1845 and T. quadristriatus (Schrank, 1781) (Pawłowski, 1979). T. kouroshi sp. n. differs from these species by the shape of the pronotum, shape of median lobe and sclerites in the endophallus. T. quadristriatus and T. liopleurus have slightly convex lateral sides of the pronotum. T. melanocephalus has a dark brown dorsal and ventral side and elongated elytra.</p> <p>Probably Trechus kouroshi sp. n. is more related to T. saulcyanus Csiki, 1928 by the combination of punctated striae, small body size, cordiform pronotum and lamelliform sclerite in the median lobe (Fig. 2.3). However, in T. saulcyanus the sclerite has in horizontal view an arch shape and in lateral view a plate shape. The median lobe is more bent than in Trechus kouroshi sp. n.</p> <p>Distribution. Iran (Kohgiluyeh and Boyer-Ahmad). Only known from the collecting site, a small cave.</p> <p>Habitat. The cave is located in a gorge where a small brook flow through. The cave is approximately 100 m long, almost horizontal, with a streamlet between stones and small rocks. At the far end of the cave streams a larger brook, too narrow and too low to follow without diving equipment. There is hardly any debris or mud between the stones. The specimens were found not far from the cave entrance. The cave is surrounded by an extensively agricultural area with very high temperatures (more than 35 ° C) and very low humidity in summer.</p> <p>Etymology. The species’ epithet refers to Koerosh (Cyrus the Great), the founder of the Achaemenid Empire.</p> <p>Identification references: Morvan &amp; Pawłowski (1977); Pawłowski (1979).</p></div> 	http://treatment.plazi.org/id/A05F87AD171BFD74D6C9FD419102FE33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1719FD75D6C9FDB697E2FD87.text	A05F87AD1719FD75D6C9FDB697E2FD87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atranus dariushi Muilwijk & Seiedy & Wrase 2021	<div><p>Atranus dariushi sp. n.</p> <p>urn:lsid:zoobank.org:act: D65213E8-E48D-4639-A60E-1BA6A5E3FC7A</p> <p>(Figs. 3.1, 3.2)</p> <p>Type material. Holotype, ♂, labelled: “ Iran, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.427197&amp;materialsCitation.latitude=30.823862" title="Search Plazi for locations around (long 50.427197/lat 30.823862)">Kohgiluyeh</a> and Boyer-Ahmad / Suq cave, 812 m / 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E / 16.vi.2018 / S. Rahideh &amp; J. Muilwijk leg.” // Atranus dariushi sp. nov. / Muilwijk, Seiedy &amp; Wrase des. 2021“ (black print on red label). Holotype deposited in HMIM.</p> <p>Comparing material: Atranus ruficollis (Gautier des Cottes, 1858): 1 ♂, Iran, Alborz, Mazandaran, near Siah Bisheh, ca 2000 m, 14.vi.2017; Muilwijk leg.; CJM.</p> <p>Description. Elytra and appendices yellow, head and pronotum slightly shiny yellowish red; dorsum pubescent; eyes flat; pronotum slender; elytra elongate elliptical; micropterous. (Fig. 3.1). Body length 6.4 mm.</p> <p>Head temples long; eyes flat; frons lateral flat impressed, wrinkled in the middle; finely pubescent on basal part of vertex; with anterior margin weakly emarginate, clypeus with two setae; mandibles long and pointed; palpi long and slender; 2 pairs of supraorbital setae, the posterior setae far behind the eye; antennae from antennomere 2 onwards pubescent; reaching to one third of the elytra length.</p> <p>Pronotum slender, cordate (PW/PL 1.02), lateral sides slightly sinuated and constricted to the base; greatest width at about the anterior fourth; anterior angles protruding, posterior angles subrounded, apically slightly upwards pointed; basal fovea moderately deep and round extending along the lateral side; lateral margin with 2 pairs of setae, one pair basolateral, the other pair at the widest point; middle line from apex to base. Pubescence less dense and finer on the disc.</p> <p>Elytra elongate-elliptical (EL/EW: 1.55); shoulders strongly rounded, without teeth; striae at the disc coarsely punctated, finer towards the apex, striae 1–4 at the base impressed, superficially near the apex; intervals slightly convex, with 2–3 rows of hairs; scutellar striole at base with a scutellar setiferous pore; three setiferous punctures in intervals 3; series umbilicata with 14 setiferous punctures.</p> <p>Microsculpture head with isodiametric meshes, pronotum and elytra with dense transverse meshes.</p> <p>Legs rather short and stout; tarsi with dorsal dense pubescence, ventral with hairbrush; protarsomeres 1–3 dilated (male), protarsomere 4 slightly bilobed.</p> <p>Ventral side colour underside yellow except the yellowish-red head; mentum with median tooth, slightly emarginate in the middle; sterna with pubescence; ratio length/width at anterior margin metepisterna: 1.67.</p> <p>Median lobe (Fig. 3.2) in lateral view comma-shaped, 1.44 mm long; the large erected hook at apex spatulashaped, directed at the apical lamella at an acute angle (seen laterally, in A. ruficollis the hook is located at an approximate right angle on the apical lamella, Fig.3.4).</p> <p>Comparative notes. Atranus dariushi sp. n. is compared with numerous specimens of A. ruficollis (Gautier des Cottes, 1858) from almost the entire range (Fig. 3.3). The new species differs from A. ruficollis by colour, the slenderer habitus, the pronotum less sinuate before hind angles, slightly weaker shoulders, reduced hind wings, metepisterna: ratio length/width at anterior margin (1.67) shorter as in A. ruficollis (2.11), and by the apex of the median lobe which is curved (as in P. ruficollis, Fig. 3.4) but the apex is spatula-shaped and the hook is located at an acute angle on the apical lamella (seen laterally). A. ruficollis is lighter or darker red-brown, often pronotum somewhat lighter, the pronotum and elytra a little shorter (PW/PL about 1.03-1.10, EL/EW 1.47-1.50, but measuring of more specimens of A. dariushi sp. n. in future should show the variability of these characters), pronotum as a rule somewhat more strongly sinuate before hind angles, shoulders less rounded, hind wings fully developed (distinctly longer than elytra, at the end folded twice, hence the species is capable of flight (some specimens from Greece were collected by car-catcher).</p> <p>Distribution. Iran (Kohgiluyeh and Boyer-Ahmad). Only known from the same small cave where Trechus kouroshi sp. n. was found.</p> <p>Habitat. This species was collected about 50 m from the entrance.</p> <p>Remarks: Atranus dariushi sp. n. is likely a cave-adapted beetle, lacking of functional hind wings and without pigmentation, and differs considerably in its ecology from the other species of this genus. Both A. ruficollis and the North American species A. pubescens Dejean, 1828 are epigean species from river banks, vegetal debris and floodplain forests (Larochelle &amp; Larivière 2003, own observations) and are capable of flight.</p> <p>Etymology. The species epithet refers to Dariush, king of Persia, of the house of the Achaemenids.</p> <p>Identification reference. Arndt et al. (2011).</p></div> 	http://treatment.plazi.org/id/A05F87AD1719FD75D6C9FDB697E2FD87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1718FD7BD6C9FD38940EF974.text	A05F87AD1718FD7BD6C9FD38940EF974.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philorhizus flavocorpus Muilwijk & Seiedy & Wrase 2021	<div><p>Philorhizus flavocorpus sp. n.</p> <p>urn:lsid:zoobank.org:act: 3F40F6E0-963D-47B3-9311-33F0A1DBE036</p> <p>(Figs. 4.1, 4.2)</p> <p>Type material. Holotype ♂ labeled: “ Iran, Kerman / Kuh-e Bahr Aseman / 22.iv.2007 / Muilwijk leg.“ // “ Holotype / Philorhizus flavocorpus sp. n. / Muilwijk, Seiedy &amp; Wrase des. 2021“ (black print on red label). Paratypes with same data but “ Paratype “. 1 ♀ same data as holotype and red label // “ Paratype “. 1 ♀: “ Iran, Kerman / Kuh-e Lalezar / 3400–3600 m / 04.vi.2014, 29°27´52´´ N 56°45´65´´ E / A. Weigel leg. IR 10a HF” // Philorhizus flavocorpus sp. n. // “ Paratype “ red label. Holotype deposited in HMIM, 1 paratype in CJM and 1 paratype in CAW.</p> <p>Description. A small sized Philorhizus (3.1–3.2 mm.) with elytra strongly widened to apex; unicolorous, slightly shining pale yellowish including appendages (Fig. 4.1); microsculpture superficially in male, more evident in female; brachypterous.</p> <p>Head longer than pronotum (HL/PL: 1.16); eyes slightly convex; temples oblique to neck. Labium convex, with 5 setae; antennomeres 1–3 without hairs except normal setae, 4–11 with fine, long, suberect hairs. Microsculpture consisting of large isodiametric superficial meshes.</p> <p>Pronotum slightly transverse (PW/PL: 1.3), widest at insertion of lateral seta. Anterior angles not protruding. Lateral sides regularly sinuate to rounded hind angles. Lateral furrow narrow, with 2 setae, one at anterior fourth, the other at the hind angle. Base medially more or less rectilinear, laterally toward hind angles oblique. Middle line well-developed. Basal impression prominent. Microsculpture consisting of superficially transverse, somewhat irregular meshes.</p> <p>Elytra long-oval (EL/EW: 1.35); humeri strongly rounded; strongly widened to apex, with maximum width at about 1/6 of the apex. Striae 1–3 superficially, other striae evanescent. Scutellar striole present. Marginal umbilicate series consist of 14 setigerous pores. Microsculpture consisting of superficial isodiametric meshes.</p> <p>Legs metatibia in females slightly bent, in males straight; meso- and metatarsi with long claws with 2–3 small dents; meso- and metatarsomere 1 about the same length as tarsomere 5.</p> <p>Ventral side with wide ligula, slightly rounded at apex; mentum without tooth. Anal segment with 2 setae on each side, segment 4–5 with 1 seta in middle on each side.</p> <p>Microsculpture consists of superficially transversal meshes; more evident on anal segment.</p> <p>Median lobe (Fig. 4.2) relatively stout, 0.84 mm long, apex pointed in lateral view. Endophallus with two twisted bands of scales and spines, the middle band more extensive than the apical one.</p> <p>Comparative notes. This wingless species resembles the mono-coloured species such as Philorhizus tinauti Anichtchenko, 2005 or P. alpinus Meschnigg, 1934 from the West Palearctic due to its yellow colour. However, the combination of features and especially the structure of the endophallus are different from all other species of this genus.</p> <p>Distribution. Iran (Kerman / Kuh-e Bahr Aseman).</p> <p>Habitat. The type specimens were collected in sub-alpine grasslands interspersed at lower elevations with shrubberies. This species is uncapable of flight, hence its distribution area is certainly restricted.</p> <p>3.1 habitus holotype, scale: 5 mm.</p> <p>3.2 median lobe holotype, lateral view, scale: 1 mm.</p> <p>3.3 Atranus ruficollis habitus, scale: 5 mm. Alborz.</p> <p>3.4 Atranus ruficollis median lobe, lateral view, scale: 1 mm. Alborz.</p> <p>4.1 habitus holotype, scale: 2 mm.</p> <p>4.2 median lobe holotype, rl = right lateral, v = ventral, ll = left lateral, scale 0.2 mm.</p> <p>Etymology. The name refers to its body colour.</p> <p>Identification references: Sciaky (1991); Wrase (2005); Wrase &amp; Assmann (2008); Arndt et al. (2011); Allegro et al. (2015).</p></div> 	http://treatment.plazi.org/id/A05F87AD1718FD7BD6C9FD38940EF974	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1716FD79D6C9F8B19656FC1F.text	A05F87AD1716FD79D6C9F8B19656FC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philorhizus kermanus Muilwijk & Seiedy & Wrase 2021	<div><p>Philorhizus kermanus sp. n.</p> <p>urn:lsid:zoobank.org:act: 0FB2556C-8447-4ADD-8539-461A1940C888</p> <p>(Figs. 5.1)</p> <p>Type material. Holotype ♀ labeled: “C. Iran, 2000 m / 12 km NW Dowlat-abad / 21.v.1977 // loc. no 341/ Exped. Nat. Mus Praha “// “ Holotype / Philorhizus kermanus sp. nov. / Muilwijk, Seiedy &amp; Wrase des. 2021“ (black print on red label). Deposited in MNP.</p> <p>Additional comparison species material: Philorhizus adoxus (Andrewes, 1923): photo of specimen with label “ NHMUK 0105804362017-03-24 16-56-05 (C)-. 1500x 5792”.</p> <p>5.1 habitus holotype, scale: 2 mm.</p> <p>5.2 Philorhizus adoxus habitus, scale: 2 mm.</p> <p>Description. A large Philorhizus (length 3.8 mm). Head with distinctly narrowed neck; upper side dull yellowish including appendages, except a dark brown pattern on elytra, consisting of broad transverse median band, broad dark brown bands along lateral sides directed to apex in striae 5–8, and an arrow-shaped darkening along the suture directed to the base; wings developed.</p> <p>Head about as long as pronotum (HL/PL: 1.01); nearly as wide as pronotum (HW/PW: 0.9). Eyes convex, twice longer than tempora; tempora with an evident blunt angle with neck. Anterior margin of labrum convex with 5 long setae. Mandibles long and sharply pointed. Antennomeres 1–3 without hairs except the normal setae; Antennomeres 4–11 with sparse thick decumbent hairs.</p> <p>Pronotum transverse (PW/PL: 1.24); front angles slightly protruding, lateral sides regularly sinuate to the blunt hind angles. Lateral sides slightly bent upwards. Obtuse hind angles slightly elevated; Lateral furrow narrow with 2 setae, one at anterior fourth, the other in the hind angle. Base medially more or less rectilinear, laterally toward hind angles oblique. Middle line distinct. Basal impression superficial. Microsculpture consists of slightly transversal meshes.</p> <p>Elytra long-oval (PL/PW: 1.4); humeri rounded; slightly widened to apex, with maximum width at about 1/4 of the apex. Striae 1–6 superficial. Scutellar striole present. umbilicate series consisting of 14 setigerous pores, interrupted in the middle. Microsculpture consisting of isodiametric meshes.</p> <p>Legs metatibia slightly bent, claws stout, without denticles; metatarsomere 1 about the same length as tarsomere 5.</p> <p>Ventral side with wide ligula, slightly rounded at apex, mentum without tooth. Anal segment with 2 setae on each side, segment 4–5 with 1 seta in middle on each side.</p> <p>Microsculpture consists of transversal meshes.</p> <p>Comparative notes. The combined characteristics of yellow colour of forebody and blunt angle between temples and neck of Philorhizus kermanus sp. n. are different from all other Palaearctic species of this genus. P. adoxus (Fig. 5.2) from Pakistan, India and adjacent countries also has a pronounced angle between the temples and neck, but differs from P. kermanus sp. n. by the squarer pronotum, piceous head and dark elytral pattern.</p> <p>Etymology. The name of the species refers to the Iranian province of Kerman.</p> <p>Distribution. The species is only known from the type locality.</p> <p>Habitat. Occurring in degraded mountain steppe forest with sparse growth of Amygdalus scoparia, Amygdalus eburnea, Pistacia khinjuk, Acantholimon sp. and Astragalus sp. and were collected from the vegetation (Hoberlandt 1983).</p> <p>Identification references: Andrewes (1923); Sciaky (1991).</p></div> 	http://treatment.plazi.org/id/A05F87AD1716FD79D6C9F8B19656FC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1714FD79D6C9FBE8918BF8D9.text	A05F87AD1714FD79D6C9FBE8918BF8D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poecilus (Derus) advena (Quensel 1806)	<div><p>Poecilus (Derus) advena (Quensel, 1806)</p> <p>(Fig. 6)</p> <p>Poecilus (Ancholeus) iranicus (Morvan, 1974) syn. n.</p> <p>Examined type. 1 ♂, Paratype of Pseudopedius iranicus Morvan, 1974 with labels: “Varamine / 19–28.v.1947 / Mirz” // “ Pseudopedius iranicus Morv ” // red label “ Paratype ”. HMIM.</p> <p>Additional examined material: Iran: 2 ♂♂, 2 ♀♀, East Azerbaijan, Moghan, 17.viii.1967, Daman leg., HMIM; 2 ♀♀, Khorasan, Sarakhs, Tajan, 250 m, 17.viii.1993, Ebra &amp; Badii leg., HMIM. Ukraine: 1 ♂, without data. CJM. Georgia: 2 ♂♂, 2 ♀♀, Transcaucasus, Kumisi nr. Tbilisi, 3–24.vi.1987, Wrase &amp; Schülke leg., CDW. Kazakhstan: 1 ♀, Aksai, Oblast Uralsk, 3.vii.–11.ix.1989, H. Wandsleb leg., CDW.</p> <p>Comparative notes. Morvan described Pseudopedius iranicus in 1974 (Fig. 6). Subsequently, Pseudopedius Seidlitz, 1887 was synonymized with Ancholeus Dejean, 1828 and P. iranicus was listed by Bousquet, 2017 as Poecilus (Ancholeus) iranicus. Morvan mentioned in the description of this species that antennomere 3 is not keeled. Species of the subgenus Ancholeus are characterized by the presence of a keel on antennomere 3, while species of the related subgenus Derus Motschulsky, 1850 are characterized by the absence of this keel. A comparison of a paratype of P. iranicus with P. (Derus) advena showed no difference between these taxa, hence P. iranicus is synonymous with P. advena.</p> <p>Distribution. Transcaucasia, the Southern Russian Plain, Ciscaucasia, the uplands of Armenia, the plains of Kazakhstan, and the plains of Middle Asia to the Altais-Sayan Mt. Land. (Kryzhanovskij et al. 1995). The species also occurs in Ukraine and Georgia, both countries were not mentioned in Löbl &amp; Löbl 2017.</p></div> 	http://treatment.plazi.org/id/A05F87AD1714FD79D6C9FBE8918BF8D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1712FD7DD6C9FF1D9175FD87.text	A05F87AD1712FD7DD6C9FF1D9175FD87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poecilus (Ancholeus) puncticollis (Dejean 1828)	<div><p>Poecilus (Ancholeus) puncticollis (Dejean, 1828)</p> <p>(Figs. 7.1–7.7)</p> <p>Poecilus (Ancholeus) aeneolus (Chaudoir, 1868) syn. n.</p> <p>Poecilus (Ancholeus) hafezi (Morvan, 1975) syn. n.</p> <p>Examined types. P. (A.) aeneolus (Chaudoir, 1868): Holotype ♂ with label “ Holotypus ” (red, printed). (MNHN). Without further labels. Obviously, the label “ Holotypus ” is added subsequently, Chaudoir never used the word “ Holotypus ”.</p> <p>Not examined type. P. (A.) hafezi (Morvan, 1975), described from Iran, East Azerbaijan, Mt. Sabalan, South of Tabriz, 2600 m, vi.1973. The holotype in collection Morvan was not available for study. The description was based on one female specimen.</p> <p>Additional examined material of P. (A.) puncticollis (Dejean, 1828): France: 2 ♂♂, 1 ♀, Herault, Palavas, i.1955, L. Schaefer leg., CDW. Austria: 1 ♀, Burgenland, Illmitz, Albensee, 3.v.1991; M. Egger leg., CDW. Italy: 1 ♂, Lazio, Frosinone, Lago di Cantern, 9.viii.1983, Gigli leg., CJM. Greece: 1 ♀, Préveza, Ammoudia, 40 km N Préveza (swamp area), v.2002; T. Lebenbauer leg., CDW; 1 ♀, Peloponnes, W. Nemea, Stymphalia lake, 616 m, 37°51´03.8´´ N, 22°26´47.3´´ E (cultural landscape, reed banks), 29.iv.2007; P. Schnitter leg., CDW; 3 ♂♂, 3 ♀♀, Peloponnes, Tripolis env., ii 1991; P. Pokornı leg., CDW. Bulgaria: 1 ♂, Stara Planina, Dolno Sakhrane, reservoir Georgi Dimitroff, 18.v.1984; D.W. Wrase leg., CDW; 1 ♀, Pomorie, 25.v.1984; D.W. Wrase leg., CDW; 1 ♀, Pomorie, saline, 9–13.v.1985; D.W. Wrase leg.; CDW. Ukraine: Crimea, Krasnogvardejs region, Kalinino, steppe, 24.iv.1989, Lobko leg., CDW. 2 ♂♂, 1 female. Georgia: Transcaucasus, Kumisi near Tbilisi (Salt Lake), 3– 24.vi.1987; D.W. Wrase / M. Schülke leg.; CDW. 3 ♂♂, 3 ♀♀. Azerbaijan: 1 ♀, Gobustan, 19–25.vi.1981, Scholz leg., CDW, Turkey: 2 ♂♂, 1 ♀, Bingöl, Gönük, 1800 m, 25–29.iv.1989, W. Heinz leg., SMNS; 1 ♂, 2 ♀♀, Bingöl, Solhan, 1200 m, 20.iv.1992, W. Heinz leg., SMNS; 1 ♂, 1 ♀, Bingöl, Yolcati, 1300 m, 18.iv.1981, W. Heinz leg., SMNS; 1 ♂, Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg., SMNS; 1 ♂, Şanlıurfa, Hilvan Urfa, 700 m, 14.iv.1980, Heinz leg., SMNS; 1 ♂, 1 ♀, Van, Gürpinar, Pass S.W. Van, 2200 m, 3.vii.1987, W. Heinz leg., SMNS; 2 ♀♀, Van, Muradiye, 1800 m, 10.v.1989, W. Heinz leg., SMNS; 1 ♂, Van, ca. 25 km S Başkale, 1950–2100 m, 7.v.1989, W. Heinz leg., CDW; 1 ♂, 1 ♀, Van, ca. 10 km S Kisiltas, S. Gevaş, 2100 m, 8.v.1989, W. Heinz leg., CDW; 1 ♂, Hakkâri, Esendere, 1900–2000 m, 5.v.1989, W. Heinz leg., SMNS; 1 ♀, Erzurum, Kuruca, 1600 m, 19.iv.1992, W. Heinz leg., SMNS; 1 ♂, 1 ♀, Kahramanmaraş, Pören geçidi, nr. Göksun, 1500 m, 24.iv.1989, W. Heinz leg., CDW; 1 ♂, Kars, Sarıkamış env., 1800 m, 8.vi.1992; S. Kadlec leg.; CDW. Iran: 1 ♂, 1 ♀, East Azerbaijan, ca. 20 km NW Kivi, 14.iv.96, Heinz leg., SMNS; 1 ♂, Mazandaran, Dasht, 900 m, 2.iv.96, W. Heinz leg., SMNS; 5 ♂♂, 7 ♀♀, Chaharmahal and Bakhtiari, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.824165&amp;materialsCitation.latitude=31.965834" title="Search Plazi for locations around (long 50.824165/lat 31.965834)">Sulegan</a> lagoon, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.824165&amp;materialsCitation.latitude=31.965834" title="Search Plazi for locations around (long 50.824165/lat 31.965834)">Cheshmeye Gholamhossein</a>, 28.v.2019, 31° 57ˊ57 N, 50° 49ˊ27 E, J. Muilwijk leg., HMIM &amp; CJM; 5 ♂♂, 6 ♀♀, Chaharmahal and Bakhtiari, Zagros Mts., Choghakhor lake nr. Boldaghi vill., 2270 m, 31°55´13´´ N 50°52´58 E (muddy shore/under stones, or running), 22.iv.2018, D.W. Wrase &amp; B. Laser leg., CDW; 2 ♂♂, 5 ♀♀, same data but: M. Hartmann leg., NME; 1 ♂, 2 ♀♀, same data but: A. Weigel leg., NME. Israel: 1 ♀, Upper Galilea, Alma plateau, 630 m, D.W. Wrase leg., CDW; 7 ♂♂, 3 ♀♀, North district, Bental Reservoir, W Merom Golan, ca 1000 m (stony wetland near shore), 30.iv.2006, D.W. Wrase leg., CDW; 1 ♂, 2 ♀♀, North district, Mt. Bental, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.7842&amp;materialsCitation.latitude=33.146885" title="Search Plazi for locations around (long 35.7842/lat 33.146885)">Merom Golan</a>, 33°08.813 N, 035°47.052 E, 30.iv.2018, W. Grosser leg., CDW. Syria: 1 ♂, as-Suwaida, Djebel Drouz, between Soueida and Salkhad, ca. 1400 m, 8.iv.1982, W. Heinz leg., SMNS.</p> <p>Comparative notes. P. (A.) puncticollis (Dejean, 1828) is a black coloured Poecilus with roughly punctate head and pronotum with or without light metallic lustre (Fig. 7.1). The species is distributed in Southern Europe, Levant, Transcaucasus, Iran and Turkmenistan (Bousquet, 2017).</p> <p>Later, Chaudoir described P. (A.) aeneolus (Chaudoir, 1868) (Fig. 7.2). Unfortunately, Chaudoir did not give any information on the locality in the description of his new species. The consultation of Chaudoir’s paper of 1876 indicated P. aeneolus originated from Syria and was collected by Kindermann. At the time of Chaudoir Syria encompassed several countries, therefore, it can mean modern Israel, Lebanon, Syria, and also south-eastern Turkey. This species also has punctured head and pronotum, but more metallic lustre.</p> <p>Morvan described P. (A.) hafezi (Morvan, 1975a) from East Azerbaijan. This species is characterised by head and pronotum covered with punctures and bronze lustre on head and elytra. Morvan compared P. (A.) hafezi with P. (A.) puncticollis and P. (A.) lissoderus Chaudoir, 1876 (= P. (A.) laevicollis Chaudoir, 1842) but not with P. (A.) aeneolus. P. (A.) laevicollis has no punctures on the head and along the midline of the pronotum, and is therefore easy to distinguish from P. (A.) hafezi. According to Morvan, the differences to P. (A.) puncticollis are mainly based on features of body shape such as a less sinuate pronotum and a shorter body length.</p> <p>We examined a huge material of the Ancholeus species with punctured forebody and more or less metallic lustre from Turkey and Iran (Fig. 7.3), including regions surrounding the type locality of P. (A.) hafezi (see Distribution map, Fig. 7.7). All characters indicated by the authors to distinguish these three species are indeed considerably variable. No differences in the structure of the median lobe (Figs. 7.4, 7.5) (including endophallus shape and copulatory piece, Figure 7.6) were found either. We can conclude, that only one species with punctured forebody inhabit Iran and adjacent regions. Therefore, the following new synonymies are proposed: Poecilus (Ancholeus) puncticollis (Dejean, 1828) = Poecilus (Ancholeus) aeneolus (Chaudoir, 1868) syn. n. = Poecilus (Ancholeus) hafezi (Morvan, 1975) syn. n. The gradual change of colour of the upper side from dark or slightly metallic from the western range to a more vivid metallic colouration in the eastern or south-eastern region is interpreted as a phenomenon of clinal variability.</p> <p>Identification references: Chaudoir (1868); Morvan (1975); Arndt et al. (2011).</p></div> 	http://treatment.plazi.org/id/A05F87AD1712FD7DD6C9FF1D9175FD87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1710FD7DD6C9FD7097E2FA93.text	A05F87AD1710FD7DD6C9FD7097E2FA93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porotachys ottomanus subsp. ottomanus ssp. ottomanus Schweiger 1968	<div><p>Porotachys ottomanus ottomanus Schweiger, 1968</p> <p>(Fig. 8)</p> <p>Examined material: 1 ♂, Iran, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.427197&amp;materialsCitation.latitude=30.823862" title="Search Plazi for locations around (long 50.427197/lat 30.823862)">Kohgiluyeh</a> and Boyer-Ahmad, Suq cave, 812 m, 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E, 16.vi.2018, S. Rahideh &amp; J. Muilwijk leg., CJM; 2 ♂♂, 2 females, same place but: 5.vi.2019, S. Rahideh, Y. Fatemi &amp; J. Muilwijk leg., HMIM, CJM.</p> <p>Remarks: The difference between Porotachys and Tachyura is based on the presence of labial pores in Porotachys while these pores are absent in Tachyura and in Porotachys the 8th stria is only present at the apex, while in Tachyura the 8 th stria is present at the base and apex.</p> <p>Distribution. Iran, South-East Europe, Turkey and eastern Mediterranean region, the new locality is the most southeastern finding of the species.</p> <p>Habitat. The same cave where Trechus kouroshi sp. n. was found.</p> <p>Remark. Porotachys bisulcatus (Nicolai, 1822), a sister species of P. ottomanus, was also regularly collected in caves (Bellés 1992).</p> <p>Identification reference. Arndt et al. (2011).</p></div> 	http://treatment.plazi.org/id/A05F87AD1710FD7DD6C9FD7097E2FA93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD1710FD7DD6C9FAD4960CF8D3.text	A05F87AD1710FD7DD6C9FAD4960CF8D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tachyura (Tachyura) shahinei (Schatzmayr & Koch 1934)	<div><p>Tachyura (Tachyura) shahinei (Schatzmayr &amp; Koch, 1934)</p> <p>(Fig. 9)</p> <p>Examined material: Iran: 1 ♂, 2 ♀♀, Sistan and Baluchestan, 13 km SSE Nikshahr riv., 8–9.iv.1973, Loc. no. 152, Exp. Nat. Mus. Praha. Muilwijk det., MNP, CJM; 1 ♂, Sistan and Baluchestan, Mask, valley of river Sarbaz, 3–4.iv.1973, Loc. no. 146, Exp. Nat. Mus. Praha. Muilwijk det., MNP; 1 ♂, Sistan and Baluchestan, env. Sarbaz, valley of river Sarbaz, 1–2.iv.1973, Loc. no. 145, Exp. Nat. Mus. Praha. Muilwijk det., MNP.</p> <p>Remark. The species can be recognized by its rather big size, and pale colour. Along the suture the elytra are darker orange coloured. The 8 th stria is parallel to the lateral margin. Striae 1–4 are well visible and punctated; pronotum cordiform.</p> <p>Distribution. Iran and Egypt.</p> <p>Remark. This species was listed in the genus Tachys by Azadbakhsh &amp; Nozari (2015) but was not listed for Iran by Löbl &amp; Löbl (2017).</p> <p>Identification reference. Schatzmayr &amp; Koch (1934).</p></div> 	http://treatment.plazi.org/id/A05F87AD1710FD7DD6C9FAD4960CF8D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170EFD63D6C9FD4B912FFBA9.text	A05F87AD170EFD63D6C9FD4B912FFBA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyderis algiricus (Lucas 1846)	<div><p>Polyderis algiricus (Lucas, 1846)</p> <p>(Fig. 11)</p> <p>Examined material: Iran: 1 ♂, 1 ♀, Chaharmahal and Bakhtiari, 10 km W Naghan, 3.iv.2007; Muilwijk leg., confirmed by J. Coulon. CJM; 2 ♂♂, 1 ♀, Khuzestan, Lake Izeh, 1.iv.2007, Muilwijk leg., CJM.</p> <p>Remarks: A very small (1.80 mm), brownish Tachyina with a very short recurrent stria, not reaching posterior puncture; elytra with 3–4 visible striae, and isodiametric microsculpture. Pronotum is cordiform, with distinct nearly rectangular posterior angles.</p> <p>Distribution. Iran, Mediterranean region, South East Europe and Levant. The new locality is the most southeastern finding of the species.</p> <p>Identification reference. Schatzmayr &amp; Koch (1934); Arndt et al. (2011).</p></div> 	http://treatment.plazi.org/id/A05F87AD170EFD63D6C9FD4B912FFBA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170EFD63D6C9FB0E9790FA64.text	A05F87AD170EFD63D6C9FB0E9790FA64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyderis impressipennis (Motschulsky 1860)	<div><p>Polyderis impressipennis (Motschulsky 1860)</p> <p>(Fig. 12)</p> <p>Examined material: 2 ♂♂, 1 ♀, Iran, Hormozgan, Minab, 19–20.v.1973, Loc. no 203; Exp. Nat. Mus. Praha. Confirmed by J. Coulon. MNP, CJM.</p> <p>Remarks: A very small (1.81 mm), palish Tachyina with a very short recurrent stria, not reaching posterior puncture. Front discal elytral pore on or immediately adjoining stria 3; border of elytra dentate, angulate at shoulder. Elytra with striae 1 and 2 slightly impressed and vaguely crenulate, the rest obsolete though often visible; preapical pore just in front of the apical striole.</p> <p>Distribution. Iran, Japan, India, Yemen, East Africa (Réunion), Oriental and Australian Region.</p> <p>Identification reference. Andrewes (1935).</p></div> 	http://treatment.plazi.org/id/A05F87AD170EFD63D6C9FB0E9790FA64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170EFD63D6C9FF1D960CFDEA.text	A05F87AD170EFD63D6C9FF1D960CFDEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tachyura (Tachyura) conspicua (Schaum 1863)	<div><p>Tachyura (Tachyura) conspicua (Schaum, 1863)</p> <p>(Fig. 10)</p> <p>Examined material: Iran: 1 ♂, 1 ♀, Hormozgan, Kuh-e- Geno, 600–1000 m, 27–28.iv.1977, Loc. no. 318, Exp. Nat. Mus. Praha. det. Muilwijk; confirmed by J. Coulon, MNP, CJM; 1 ♂, Hormozgan, Kuh-e Geno, Bagh Tang, 410 m, 7–8.v.1977, Pazuki leg., HMIM; 1 ♀, Hormozgan, Angorhan, SE Minab, 24.iv.2002, M. Rejzek leg., identified by T. Kopeckı, 2010 as sudanensis Schatzmayr &amp; Koch, 1934 (this name is synonymous with conspicua), CDW; 1 ♀, Yemen, Ta’izz, 10.01.2001, A.van Harten &amp; A.R. Yarimi leg., Felix det., CJM.</p> <p>Remarks: This species is even bigger and more convex than the preceding one; the colour is similar but the brown spot along the elytral suture is more developed. The distance between the 8 th stria and interval 9 is wider after the second humeral seta. Head and pronotum are dull with microsculpture. The lateral sides of the cordiform pronotum are clearly sinuated. 1 st –3 rd striae of elytra are deepened and punctated.</p> <p>Distribution. Iran, and widely from North Afrika to the Middle East.</p> <p>Identification reference. Schatzmayr &amp; Koch (1934).</p></div> 	http://treatment.plazi.org/id/A05F87AD170EFD63D6C9FF1D960CFDEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170EFD60D6C9F9069764FA3D.text	A05F87AD170EFD60D6C9F9069764FA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pogonopsis pallida Bedel 1898	<div><p>Pogonopsis pallida Bedel, 1898</p> <p>(Figs. 13.1-13.4)</p> <p>Olegius turkmenicus Komarov, 1996 syn. n.</p> <p>Examined types. Olegius turkmenicus Komarov, 1996, holotype ♀ with labels: “Turkmenia, 40 km E / of Nebit-Dag, SE / Chilmamedkum Sands / 25.05.1993 Yu. Arzanow ”; “Kyzyl-Takyr / at light” (printed on white label), and: “ Holotypus Olegius turkmenicus sp.n. / Komarov det., 1996.” (“ Holotypus ” printed, the rest handwritten by E.V. Komarov on red label). ZIN.</p> <p>Additional examined material: 1 ♀ with label “ Tunisie / 7.1906 / Kébilli ” and “ Pogonopsis pallida Bed. / Comparé au TYPE ”; 1 ♂, 1 ♀: “ Tunisie / 7.1906 / Kébilli” “ Tunisie, Kébilli, 7.1906”, “ Pogonopsis pallida Bedel ” and “Muséum Paris / Ex M. Maindron / G. Babault 1939” (black print on blue label). MNHN (The species was described from Kébilli, so these specimens can be regarded as topotypes); 3 ♀♀, Iran, Kerman, Lut desert, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.973694&amp;materialsCitation.latitude=29.580696" title="Search Plazi for locations around (long 58.973694/lat 29.580696)">Ab-e Yalan</a>, 396 m, 29°34´50.5 N, 58°58´25.3 E, 29.iii.2017; H. Rajaei leg.; SMNS, CJM.</p> <p>Remark. Azadbakhsh (2019) synonymized Olegius Komarov, 1996 with Pogonopsis Bedel, 1898, but made no statement at the species level. After study of the holotype of Olegius turkmenicus we synonymize O. turkmenicus with P. pallida.</p> <p>Distribution. So far, the species is known from the deserts in Tunisia and Turkmenistan (Löbl &amp; Löbl 2017), and now also from the Lut desert in Iran.</p> <p>Identification reference. Bedel (1898).</p> <p>13.1 Pogonopsis pallida habitus, Lut desert, scale: 5 mm.</p> <p>13.2 Olegius turkmenicus holotype habitus.</p> <p>13.3 Olegius turkmenicus holotype head ventral.</p> <p>13.4 Olegius turkmenicus holotype labels.</p> </div>	http://treatment.plazi.org/id/A05F87AD170EFD60D6C9F9069764FA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170DFD66D6C9F9BC94CEFB4A.text	A05F87AD170DFD66D6C9F9BC94CEFB4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Trichochlaenius) albissoni Reitter 1908	<div><p>Chlaenius (Trichochlaenius) albissoni Reitter 1908</p> <p>(Fig. 14)</p> <p>Examined material: Iran: 5 ♂♂, 4 ♀♀, Khuzestan, Mahshar, 9.v.2006; Muilwijk leg. Confirmed by E. Kirschenhofer. CJM; 1 ♂, 3 ♀♀, Khuzestan, lake Iseh, 1.iv.2007; Muilwijk leg., CJM, CDW; 1 ♂, Busher, Dorahak vill. env., 10 m, 26.iii.2018, I. Pljushtch leg., CDW.</p> <p>Remark. A species of green-bluish lustre, appendages light; body narrow and elongate, with dense punctation. Antennae pubescent from antennomere 2. Antennomere 3 longer than 4. Tarsi pubescent dorsally. Easy to separate from the two other Iranian species of the subgenus Trichochlaenius Seidlitz, 1887: C. gotschii Chaudoir, 1846, which has a bluish upperside and dark legs and antennae and C. aeneocephalus aeneocephalus Dejean, 1826 which has a shiny golden head.</p> <p>Distribution. Iran and Syria.</p> <p>Identification reference. Reitter (1908).</p> <p>15.1 habitus, W Bazargan, scale: 1 mm.</p> <p>15.2 median lobe, scale: 0.2 mm.</p></div> 	http://treatment.plazi.org/id/A05F87AD170DFD66D6C9F9BC94CEFB4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170BFD66D6C9F9399624F870.text	A05F87AD170BFD66D6C9F9399624F870.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Demetrias monostigma Samouelle 1819	<div><p>Demetrias monostigma Samouelle, 1819</p> <p>(Fig. 16)</p> <p>Examined material: Iran: 1 ♂, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.668335&amp;materialsCitation.latitude=32.07028" title="Search Plazi for locations around (long 50.668335/lat 32.07028)">Chaharmahal</a> and Bakhtiari, S Shahr-e Kord, Ardal, 1960 m, 32°04´13 N, 50°40´06 E, 21.x.2011, J. Frisch leg., MNB.</p> <p>Distribution. This species is known from adjacent countries of Iran, such as Armenia and Azerbaijan. The distribution area is from Western and Southern Europe to Central Asia and Western Siberia (Löbl &amp; Löbl 2017). The new locality is the southernmost finding of the species.</p> <p>Identification reference. Müller-Motzfeld (2006).</p></div> 	http://treatment.plazi.org/id/A05F87AD170BFD66D6C9F9399624F870	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
A05F87AD170BFD66D6C9FAC09652F9FF.text	A05F87AD170BFD66D6C9FAC09652F9FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philorhizus crucifer subsp. crucifer (Lucas 1846) ssp. crucifer (Lucas 1846	<div><p>Philorhizus crucifer crucifer (Lucas, 1846)</p> <p>(Figs. 15.1, 15.2)</p> <p>Examined material: Iran: 1 ♂, West Azerbaijan, W. Bazargan, 2157 m., 30°21´02.5´´ N, 44°07´08.4´´ E, 27.viii.2008, Serri &amp; Frisch leg., MNB.</p> <p>Remark. The sclerites in the endophallus are characteristic for this species (Fig. 15.2).</p> <p>Distribution. Iran, and from West Mediterranean Europe, East Europe, Middle East to Middle Asia.</p> <p>Identification references: Lucas (1846); Arndt et al. (2011).</p></div> 	http://treatment.plazi.org/id/A05F87AD170BFD66D6C9FAC09652F9FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Muilwijk, Jan;Seiedy, Marjan;Wrase, David W.	Muilwijk, Jan, Seiedy, Marjan, Wrase, David W. (2021): Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera). Zootaxa 5067 (1): 55-81, DOI: https://doi.org/10.11646/zootaxa.5067.1.3
