taxonID	type	description	language	source
03A95A4EFFC4FFF207E7FC23DF63BE8F.taxon	type_taxon	Type species. Helicomorpha scalarioides Bavay & Dautzenberg, 1904, by original designation.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC4FFF207E7FC23DF63BE8F.taxon	discussion	Remarks. Messageria was introduced by Bavay & Dautzenberg (1904) with the obliquely positioned embryonic shell and stairs-like ribs (Bavay & Dautzenberg, 1904; Thiele, 1929). Messageria was synonymized with Helicomorpha in previous studies (Egorov, 2013; Bank, 2017). In comparison with Helicomorpha (SMF 105304, H = 1.5 mm, W = 1.5 mm), the shell of Messageria is significantly larger (see measurements below). More importantly, the outer surface of the operculum in Helicomorpha is flat or with an outward bulge / inward depression in the center (possibly caused by stripped multilayer structure), while in Messageria, the operculum is concave on the outside with multispiral lamina on the outer surface. Based on this diagnostic set we consider the two as separate genera. The columellar edge of the peristome in some Helicomorpha species is strongly expanded, such a trait that has not been found in Messageria. The obliquely positioned protoconch also appears in some species of Helicomorpha, however, such trait is not diagnostic as it is common in diplommatinids and alycaeids. Affiliates of Messageria see in Discussion.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC4FFF207E7F9A5DCBFBC9A.taxon	description	(Figures 1 A – D)	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC4FFF207E7F9A5DCBFBC9A.taxon	materials_examined	Type material. Syntypes of Helicomorpha scalarioides Bavay & Dautzenberg, 1904, MNHN-IM- 2000 - 32414 / 32415, RBINS MT. 1016 / 2, Ht Tonkin [Haut Tonkin], coll. Messenger. Museum material. MNHN-IM- 2014 - 7928 (one of the labels wrote “ Messenger ” with yellow mark), Ht Tonkin [Haut Tonkin], coll. Letellier, 1949.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC4FFF207E7F9A5DCBFBC9A.taxon	description	Measurements. H = 3.20 mm, W = 2.81 mm, D A = 1.50 mm (n = 1). Vernacular name. Į 梯Ħfi ḏDzss 亚 ḋ xuán tī chóng lèi luó zhǐ míng yà zhǒng	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC4FFF207E7F9A5DCBFBC9A.taxon	distribution	Distribution. VIETNAM: Yên Bái to Long Sõn.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	description	Figures 1 E – H, 2 A	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	materials_examined	Holotype: HBUMM 10058, a big cave in mountains between Fengshan County [Hechi City] and Leye County [Baise City], Guangxi Zhuang Autonomous Region, China, ex coll. Hao Yang, 2012. Paratypes: CZY / 1, LLW / 1, same data as holotype.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	etymology	Etymology. This name comes from the local Zhuang language “ dongh ”, which means “ cave ”.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	diagnosis	Diagnosis. This subspecies can be distinguished from the nominate subspecies by the larger shell and wider lower whorls and umbilicus. Also, the aperture of M. scalarioides donghiana n. ssp. and 1 / 8 whorl behind it evidently deviate from the axis and umbilicus.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	description	Measurements. H = 3.70 mm, W = 3.37 mm, D A = 1.89 mm (holotype). Vernacular name. Į 梯Ħfi ḏ 洞Ë亚 ḋ xuán tī chóng lèi luó dòng xué yà zhǒng	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC7FFF007E7F914DC63BA75.taxon	distribution	Distribution. CHINA: Guangxi.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	description	Figures 1 I – O, 2 B	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	materials_examined	Holotype: HBUMM 10021, Botanical Garden of Shishang Forest Scenic spot, Maolan National Nature Reserve, Maolan Town, Libo County, Buyi and Miao Autonomous Prefecture of QianNan, Guizhou. 2019. VI. 16. 780 m. leg. Lu QIU. Paratype: HBUMM 10055 (one juvenile), Maolan National Nature Reserve, Maolan Town, Libo County, Buyi and Miao Autonomous Prefecture of QianNan, Guizhou. 2021. V. 21. 780 m. leg. Yan SUN.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	etymology	Etymology. This name commemorates the first discovery of this genus in China.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	description	Description. Shell small (<5 mm), dextral, roundly conical, about 5.5 whorls. Shell translucent, relatively glossy, apical part light orange. Apex slightly convex, apical whorls depressed conical. Basal whorls ovoid. Protoconch oblique compared to shell axis, 1.25 whorls with matte surface. The next 0.5 whorl of the teleoconch with regularly spaced and equally high, weak ribs. The other whorls have well-spaced, high and thin main ribs, with a distinctly lower intermediate rib in between each pair of main ribs, ca. 9 (main and intermediate) ribs per 1 mm on body whorl. The intercostal distance increases with the whorls, and the strength (height) of the two kinds of ribs becomes same at umbilical area. Umbilicus deep. Aperture rounded, slightly oblique. Peristome double, with prominent outer peristome. Inner peristome protruding, with parietal callous thick; outer peristome expanded but not reflected. Operculum rounded, corneous, concave; outer surface with multispiral lamina; inner surface simple, glossy. Measurements. H = 3.62 mm, W = 3.20 mm, D A = 1.95 mm (holotype).	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	diagnosis	Diagnosis. Compared to Messageria scalarioides, the new species is distinguished by the much more distinct lower intermediate ribs in between each pair of high ribs, particularly on the outer whorls of the shell. Additionally, M. scalarioides has moderately convex apex and nearly conical basal whorls, which are slightly convex and ovoid in the new species. The 1 / 8 whorl behind the aperture of M. sinica n. sp. grows evenly, while it is expanded in M. scalarioides (especially in M. scalarioides donghiana n. ssp.). Vernacular name. 中华Ħfi ḏ zhōng huá chóng lèi luó	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
03A95A4EFFC6FFF007E7FBD0DC63BCC6.taxon	distribution	Distribution. CHINA: Guizhou.	en	Chen, Zhe-Yu, Lin, Li-Wen (2021): Notes on the genus Messageria Bavay & Dautzenberg, 1904, with descriptions of a new speciesand anew subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea Alycaeidae). Zootaxa 5067 (1): 129-134, DOI: https://doi.org/10.11646/zootaxa.5067.1.9
