taxonID	type	description	language	source
5E4E87DAFD652E58FC803BDBFC6157E7.taxon	materials_examined	Material examined. 2 ♂ & 4 ♀ PCAS & MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD652E58FC803BDBFC6157E7.taxon	distribution	Distribution: This species has yet been reported from the Philippine islands Leyte, Luzon, Mindanao and Siargao.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD652E58FC8038B0FB335694.taxon	discussion	Currently, 20 species of Baeocera are known to occur in the Philippines, only four of them have been described or reported from Mindanao. Three of them are in the new collections, together with additional three new species.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E59FF3D3DA5FC5555C7.taxon	description	(Figs 1 – 4)	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E59FF3D3DA5FC5555C7.taxon	materials_examined	Holotype ♂ MHNG: Philippines Mindanao. Araibo. Pantukan. Compostela Valley. 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E59FF3D3DA5FC5555C7.taxon	etymology	Derivatio nominis: The species epithet is an adjective derived from the name of the mountains where it was found.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E59FF3D3DA5FC5555C7.taxon	description	Description: Length 1.48 mm, width 1.10 mm. Frons and body very dark brown, almost blackish, apex of abdomen ochraceous, femora and tibiae reddish-brown, tarsi and antennomeres I to VI ochraceous, following antennomeres light brown. Body strongly convex dorsally. Eyes in dorsal view slightly larger than half of interval between them. Length / width rations of antennomeres as: III 25 / 8: IV 25 / 8: V 30 / 8: VI 26 / 8: VII 45 / 11: VIII 44 / 11: IX 45 / 14: X 44 / 14: XI 51 / 14. Pronotum and elytra not microsculptured, with lateral contours separately arcuate. Pronotum very finely punctate, with lateral margins rounded, lateral margin carinae concealed in dorsal view. Apex of scutellum exposed. Elytra not overlapping abdominal apex, moderately narrowed apically, lateral margin carinae not visible in dorsal view, sutural striae entire, parallel in basal third of sutural length, converging posteriad, bent along pronotal lobe and extended along bases to form complete basal striae joined with lateral striae. Sutural striae distinctly punctate, except along apical third, punctures along lateral striae almost indistinct. Basal striae deep and well visible near lateral margins. Punctation on basal 0.15 to 0,20 mm, on 0.15 mm broad stripe along sutural striae, on narrow stripe along lateral margins and on apical halves of elytra fine, very shallow and not well delimited, with punctures distinctly larger than pronotal punctures. Conspicuous patch of coarse and sharply delimited punctures situated in anterior half of each elytron; intervals between coarse punctures about 1.5 to two times puncture diameters. Hind wings not reduced. Hypomera extremely finely punctate. Mesepimera about four times as long as wide and four times as long as intervals to mesocoxae. Metaventrite convex and impunctate in middle, with several coarse punctures delimiting smooth centre; lateral parts of metaventrite extremely finely punctate. Mesocoxal lines parallel, with coarse puncture row extended laterally almost to mid-length of mesepimera; mesocoxal areas about 0.03 mm, as fifth of shortest intervals to metacoxae. Exposed part of metanepisterna large, near anterior angles about 0.10 mm wide, narrowed posteriad, with distinct, impunctate suture. Metepimera each with longitudinal stria. Protibiae straight, mesotibiae and metatibiae slightly bent. Propygidium and ventrite I not microsculptured, pygidium and ventrites II to VI with hardly visible punctulate microsculpture. Basal half of propygidium densely and rather coarsely punctate, with punctures well delimited, smaller than coarse elytral punctures, to part about as large as puncture intervals. Apical part of propygidium and pygidium very finely and sparsely punctate. Ventrite I without wrinkles, with basal row of coarse elongate punctures not interrupted in middle; punctation extremely fine posterior of basal row, as on following ventrites. Male protarsomere I strongly widened, somewhat narrower than apices of protibiae, Protarsomere II and III slightly widened. Mesotarsomeres I slightly widened. Ventrite VI shallowly emarginate at each side of triangular, about 0.10 mm long and acute median process. Aedeagus (Figs 1 – 4) 0.87 mm long.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E59FF3D3DA5FC5555C7.taxon	diagnosis	Differential diagnosis: This new species falls in the key to Philippine species (LÖbl, 2012) under the couplet 5, to B. alticola LÖbl, 2012 and B. fortis LÖbl, 2012, its aedeagal characters suggest relationships with B. fortis. The new species is readily distinguishable from B. fortis by its smaller size, the darker body colour, the antennomeres III as long as the antennomeres IV and the antennomeres V clearly longer than antennomeres IV or VI, the shape of the apical margin of the male ventrite VI, the parameres widened apically and lacking lobes or processes, the blunt and in dorsal view not incurved tip of the median lobe, and the shape of the sclerotized pieces of the internal sac. Diagnostic for the new species are also tufts of elongate spine-like and lamellar structures within the internal sac.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E5EFC8038EFFC2F582F.taxon	description	Figs 5 – 7)	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E5EFC8038EFFC2F582F.taxon	materials_examined	Holotype ♂ MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. Paratypes 3 ♂ PCAS & MHNG: data as the holotype.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E5EFC8038EFFC2F582F.taxon	etymology	Derivatio nominis: The species epithet is a Latin adjective meaning glabrous.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E5EFC8038EFFC2F582F.taxon	description	Description: Length 0.88 – 0.91 mm, width 0.63 – 0.65 mm. Frons and most of body blackish, apex of abdomen, femora and tibiae reddish-brown, tarsi and antennae ochraceous to yellowish. Body strongly convex dorsally. Bode strongly convex dorsally. Eyes in dorsal view slightly smaller than half of interval between them. Length / width rations of antennomeres as: III 16 / 5: IV 17 / 5: V 18 / 6: VI 17 / 5: VII 24 / 8: VIII 18 / 5: IX 25 / 10: X 25 / 10: XI 34 / 10. Pronotum and elytra not microsculptured, with lateral contours continuously arcuate. Pronotum very finely punctate, with lateral margins rounded, lateral margin carinae concealed in dorsal view. Scutellum concealed. Elytra not overlapping abdominal apex, moderately narrowed apically, lateral margin carinae not visible in dorsal view, sutural striae entire, parallel, bent along pronotal lobe and extended along bases to form complete basal striae joined to lateral striae. Adsutural areas and sutural striae with few rather distinct fine puncture, discal punctation very fine, as that on pronotum, hardly visible at 100 times magnification. Hind wings fully developed. Hypomera appearing impunctate. Mesepimera about three times as long as wide and somewhat more than two times as long as intervals to mesocoxae. Metaventrite convex and impunctate in middle, with coarse punctures delimiting smooth centre; lateral parts of metaventrite smooth near metacoxae, coarsely and irregularly punctate on remaining surface. Mesocoxal lines slightly convex, with coarse puncture row extended laterally almost to mesepimera; mesocoxal areas about 0.02 mm, as third of shortest intervals to metacoxae. Exposed part of metanepisterna narrow, parallel, about 0.04 mm wide, with distinct, punctate suture. Metepimera each with longitudinal stria. Tibiae straight. Ventrite I without wrinkles, with basal row of coarse, to part slightly elongate punctures not interrupted in middle, punctation posterior of basal row extremely fine, hardly visible at 100 times magnification, as that on following ventrites. Male protarsomere I to III slightly widened. Aedeagus (Figs 5 – 7) 0.21 – 0.24 mm long.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD642E5EFC8038EFFC2F582F.taxon	diagnosis	Differential diagnosis: This new species is a member of the B. lenta species-group and is similar to B. danielae LÖbl, 2012. It may be distinguished by the elytra entirely very finely punctate, in combination with the minute body-size and the internal sac lacking membranous denticles and bearing a proximally widened sclerite (seen in lateral view). [scale bar 0.1 mm].	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3DA5FE045193.taxon	materials_examined	Material examined. 3 ♂ & 3 ♀ PCAS & MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3DA5FE045193.taxon	discussion	Comments. This species is known only from Luzon and Mindanao.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3CD0FC095467.taxon	description	Figs 8 – 10)	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3CD0FC095467.taxon	materials_examined	Holotype ♂ MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg. Paratypes 8 ♂ & 6 ♀ PCAS & MHNG: same data as in holotype.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3CD0FC095467.taxon	etymology	Derivatio nominis: The species epithet is a Latin adjective meaning difficult.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3CD0FC095467.taxon	description	Description: Length 1.08 – 1.18 mm, width 0.71 – 0.81 mm. Frons and body blackish, with apical abdominal segments light, ochraceous to yellowish. Femora and tibiae dark reddish-brown, tarsi and antennae light, nearly yellowish. Body strongly convex dorsally. Eyes in dorsal view as large as half of interval between them. Length / width rations of antennomeres as: III 28 / 6: IV 21 / 6: V 30 / 6: VI 28 / 6: VII 34 / 8: VIII 33 / 7: IX 35 / 10: X 35 / 13: XI 40 / 13. Pronotum and elytra not microsculptured, with lateral contours nearly continuously arcuate. Pronotum very finely punctate, with lateral margins rounded, lateral margin carinae concealed or hardly visible in dorsal view. Scutellum concealed. Elytra not overlapping abdominal apex, strongly narrowed apically, lateral margin carinae visible only near bases in dorsal view, sutural striae entire, parallel in basal half of sutural length, converging posteriad mid-length, bent along pronotal lobe and extended along bases to form complete basal striae joined with lateral striae. Anterior third of sutural and lateral striae distinctly punctate. Basal striae very shallow and hardly visible near lateral margins. Punctation coarse and sparse on basal third of elytral disc, except on narrow band along base and near sutural striae, intervals between coarse punctures mostly about 1.5 to 2 times as large as puncture diameters. Punctation on remaining surface of elytra very fine and sparse. Hind wings not reduced. Hypomera appearing impunctate. Mesepimera about four times as long as wide and three times as long as intervals to mesocoxae. Metaventrite flattened in middle, with impunctate mesal stripe, remaining mesal and entire lateral surfaces coarsely and very densely punctate, punctures mostly larger than puncture intervals. Mesocoxal lines parallel, mesocoxal areas about 0.02 mm, as fourth of shortest intervals to metacoxae. Exposed part of metanepisterna very narrow, with suture indicated by outer coarse punctures. Metepimera without longitudinal striae. Tibiae straight. Ventrite I without wrinkles, with coarse row of basal punctures narrowly interrupted in middle and several scattered rather large punctures on sides; remaining abdominal punctation very fine. Male protarsomere I to III slightly widened. Aedeagus (Figs 8 – 10) 0.33 – 0.35 mm long.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFF3D3CD0FC095467.taxon	diagnosis	Differential diagnosis: This new species is also a member of the B. lenta species-group and falls under the couplet 13 in the key to Philippine Baeocera (LÖbl 2012). Its aedeagal characters, especially the complex sclerotized parts of the internal sac, differ drastically from both species under this couplet, B. louisi LÖbl, 2012 and B. danielae LÖbl, 2012. Baeocera onerosa sp. nov. may be readily distinguished from these two species by the much dense punctation on lateral parts of the metaventrite and by several rather large punctures on lateral parts of the abdominal ventrite I, situated posterior of basal puncture row. The internal sac of B. onerosa sp. nov. is similar to that of B. boettcheri (LÖbl, 1972) while the shape of the parameres and the elytral punctation differ drastically.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFC80398FFC5355F7.taxon	materials_examined	Material examined. 2 ♂ PCAS & MHNG: Mindanao, Davao Prov., Mt. Talomo (Mt. Apo) Catigan, / 800 - 1000 m, 7 ° 01 ’ 21.0 ’’ N, 125 ° 22 ^ 30.5 ’’ E, 29.4. - 1.5. 2019, A. V. Shavrin leg.; 6 ♂ & 4 ♀ PCAS, MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019, Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFC80398FFC5355F7.taxon	distribution	Distribution: The species was known only from the island Leyte.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5FFC803B3FFAD35734.taxon	synonymic_list	This genus comprises three species, with B. derougemonti LÖbl, 1983 appearing restricted to Sulawesi (Greater Sunda Islands, Indonesia), B. punctatissima (Reitter, 1880) known both from the Philippines and Sulawesi, and B. basicollis Löbl, 2011 described from the Philippine specimens.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5CFC803A7AFD0E50F7.taxon	materials_examined	Material examined. 1 ♀ PCAS: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD622E5CFC803A7AFD0E50F7.taxon	distribution	Distribution: This species is currently known from Luzon and Mindanao, the Philippines.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD612E5CFF3D3C3FFAD750F7.taxon	discussion	This genus is with 31 species widely distributed in the Oriental region and in Melanesia. The four Philippine species of Bironium were keyed in LÖbl (2011 a). Only one of them, B. rufescens LÖbl, 1972, is known to occur in Mindanao, the Philippines.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD612E5DFC803C3FFF45514E.taxon	materials_examined	Material examined. 1 ♂ & 1 ♀ PCAS & MHNG: Mindanao, Davao Prov., Mt. Talomo (Mt. Apo) Catigan, 800 - 1000 m, 7 ° 01 ’ 21.0 ’’ N, 125 ° 22 ’ 30.5 ’’ E, 29.4. - 1.5. 2019, A. V. Shavrin leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD612E5DFC803C3FFF45514E.taxon	distribution	Distribution: This species is reported from Leyte and Mindanao in LÖbl et al. (2020). Unlike other examined specimens of B. rufescens, both are black.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3F7BFD2C53C4.taxon	materials_examined	Material examined. 1 ♂ & 2 ♀ PCAS & MHNG: Mindanao, Davao Prov., Mt. Talomo (Mt. Apo) Catigan, / 800 - 1000 m, 7 ° 01 ’ 21.0 ’’ N, 125 ° 22 ’ 30.5 ’’ E, 29.4. - 1.5. 2019, A. V. Shavrin leg.; 3 ♂ PCAS & MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3F7BFD2C53C4.taxon	distribution	Distribution: The species has been reported from Leyte, Luzon, Mindanao and Panay.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3C75FD515234.taxon	discussion	Scaphisoma comprises 112 species reported from the Philippines (LÖbl & Ogawa 2016) and is the far most species-rich genus of the scaphidiines. A single species is present in the recent material.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3EEBFDD554EE.taxon	discussion	The Philippine species were dealt with and keyed in LÖbl (2011 b). From the 18 currently recognized Philippine species, only four are known to occur on Mindanao. An additional, new species is present in the examined collections.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3815FC4355E3.taxon	description	Figs 11 – 12)	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3815FC4355E3.taxon	materials_examined	Holotype ♂ MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg. Paratype ♂ PCAS: Mindanao, Davao Prov., Mt. Talomo (Mt. Apo) Catigan, / 800 - 1000 m, 7 ° 01 ’ 21.0 ’’ N, 125 ° 22 ^ 30.5 ’’ E, 29.4. - 1.5. 2019, A. V. Shavrin leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3815FC4355E3.taxon	etymology	Derivatio nominis. The species epithet is a Latin adjective meaning wide, referring to the wide parameres.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3815FC4355E3.taxon	description	Description: Length 1.35 mm, width 0.69 mm, dorsoventral diameter 0.71 mm. Head and body dark brown with somewhat reddish shine, apical abdominal segments and appendages lighter, reddish-brown. Length / width rations of antennomeres as: III 17 / 7: IV 22 / 5: V 27 / 5: VI 22 / 7: VII 33 / 9: VIII 18 / 8: IX 30 / 19: X 33 / 14: XI 40 / 14. Thorax not microsculptured, pronotum very finely punctate. Tip of scutellum exposed. Elytra with strigulate microsculpture hardly visible at 100 times magnification, not iridescent, with punctation similar with that of pronotum; sutural striae starting near pronotal lobe, parasutural striae hardly visible. Hypomeron without longitudinal stria. Middle part of metaventrite impressed, lacking stria, densely and finely punctate and bearing short pubescence on posterior half, impunctate in middle of anterior half. Sides of metaventrite sparsely, extremely fine punctate, with fairly long pubescence. Mesocoxal lines with row of fine punctures not exceeded laterally along mesepimera; mesocoxal areas about 0.04 mm long, almost as fourth of shortest intervals to metacoxae. Metanepisterna flat, 0.06 mm wide, narrowed apically, with curved suture. Tibiae straight. Abdomen with strigulate microsculpture and very finely punctate. Basal punctures of ventrite I rather coarse. Male segments I to III of protarsi distinctly widened, segment I narrower than apex of protibia. Aedeagus (Figs 11 – 12) 0.42 mm long.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E5DFF3D3815FC4355E3.taxon	diagnosis	Differential diagnosis: The aedeagal characters suggest relationships of this new species with the Philippine S. episternalis LÖbl, 2011, though it would fall under the couplet 18 in the key to the Philippine species (LÖbl, 2011 b). It differs drastically from S. episternalis by the much narrower metanepisterna, darker body, not iridescent elytra, and much wider parameres. The two species given under the couplet 18, S. pseudotenella LÖbl, 2011 and S. data LÖbl, 2011, may be easily distinguished by the shape of the internal sac. Prominent articular processes are present in several congeners, such as S. cognata LÖbl, 1984, S. difficilis LÖbl, 1979, S. jirkai LÖbl, 2018, S. ornata (Pic), S. papuana Csiki, 1909, S. stipes LÖbl, 1971, S. tenella LÖbl, 1990 and S. uncata LÖbl, 1990. These species differ conspicuously from S. lata sp. nov. either by the shape of the apical process of the median lobe or by the narrow basal section of the parameres (seen in lateral view).	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD602E52FC803B00FA3D50F9.taxon	discussion	Currently, two species of Scaphoxium are known from the Philippines, S. alesi LÖbl, 2011 from Luzon and S. taylori LÖbl, 1981 described from Sarawak and subsequently reported from the Philippine islands Luzon, Mindanao and Palawan. The new collections yielded an additional species.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD6F2E53FC803C3AFDEA5639.taxon	description	Figs 13 – 15)	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD6F2E53FC803C3AFDEA5639.taxon	materials_examined	Holotype ♂ MHNG: Mindanao, Araibo, Pantukan, Compostela Valley, 900 m, Candalaga / Mts., 7 ° 16 ’ 35.3 ’’ N, 126 ° 10 ’ 12.8 ’’ E, 4.5.2019 Shavrin A. V. leg.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD6F2E53FC803C3AFDEA5639.taxon	etymology	Derivatio nominis: Patronymic. The species is dedicated to its collector Alexei V. Shavrin (Daugavpils, Latvia), a well-known expert of rove beetles.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD6F2E53FC803C3AFDEA5639.taxon	description	Description: Length 1.54 mm, width 0.77 mm, dorsoventral diameter 0.86 mm. Head and most of body evenly very dark reddish-brown, apical abdominal segments, femora and tibiae lighter, tarsi lighter than tibiae, antennae brown. Length / width rations of antennomeres as: III 20 / 6: IV 19 / 6: V 26 / 6: VI 22 / 7: VII 39 / 11: VIII 25 / 11: IX 38 / 13: X 33 / 13: XI 48 / 15. Pronotal punctation very fine, hardly visible at 50 times magnification. Scutellum concealed. Elytral punctation distinct, fine and dense, sutural striae starting about 0.20 mm posterior of level of pronotal lobe. Mesoventrite not striate, impressed and with low mesal ridge narrowed apically. Median part of metaventrite slightly convex, with irregular transverse row of extremely fine punctures. Lateral parts of metaventrite appearing impunctate near metacoxae, each with dense patch of conspicuously coarse punctures and oblique ridge. Mesocoxal areas about 0.05 mm, almost as halves of shortest intervals to metacoxae. Metanepisterna slightly convex, about 0.06 mm wide, with curved, deep, apically shortened sutures. Ventrite I with few distinct basal punctures, not microsculptured. Following ventrites and exposed tergites with punctulate microsculpture and extremely fine punctation. Male segments I to III of protarsi slightly widened. Aedeagus (Figs 13 – 15) 0.68 mm long.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
5E4E87DAFD6F2E53FC803C3AFDEA5639.taxon	diagnosis	Differential diagnosis: This new species is likely related with S. alesi LÖbl, 2011. It may be distinguished by the dark body-colour, the shorter sutural striae of the elytra, the sides of the metaventrite bearing dense and coarse punctures, the shape of the angular subapical parameral lobes and the presence of strongly sclerotized denticles in the internal sac of the aedeagus.	en	Löbl, Ivan (2021): Contribution to the knowledge of the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of Mindanao, Philippines. In: Biodiversity, Biogeography and Nature Conservation in WALLACEA and NEW GUINEA. Riga: The Entomological Society of Latvia: 265-272, ISBN: 978-9984-9768-9-1, DOI: http://doi.org/10.5281/zenodo.5646291
