taxonID	type	description	language	source
023E39002426FFF16019FA2B2D0AC960.taxon	description	(Fig. 2, Clade E)	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	materials_examined	Holotype. ZFMK 88935 (adult male, Fig. 5 A-B, Clade E) from “ Vietnam, Dong Nai Province, Cat Tien National Park (11.6344444 ° N 107.456667 ° E), 104 m elevation, ” collected by Peter Geissler on May 10 th 2009.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	diagnosis	Original Diagnosis. Amedium-sized Calotes with a maximum SVL of 97 mm. It can be distinguished from all taxa of the C. mystaceus complex by the combination of the following characters: 1) head and body robust; 2) body scales homogeneous, relatively small, feebly keeled and arranged in regular rows; 3) 44 – 50 midbody scale rows; 4) upper dorsolateral scales pointing backwards and upwards; 5) two short and well separated spines, surrounded by three to four scales on either side of the upper head above the tympanum; 6) nuchal and dorsal crest continuous, composed of erected compressed scales, directed posteriorly; 7) vertebral spines and scales in males 35 – 42 and in females 43 – 46; 8) oblique fold of skin in front of fore limb insertion distinct, covered with small granular dark scales; 9) extremities and tail relatively long and slender (see comparison with C. mystaceus for details); 10) bluish to turquoise head and anterior body part, this coloration not well exceeding front limb insertion; 11) yellowish light stripe at upper lip reaching from below anterior corner of eye to posterior end of head; 12) no dorsolateral brownish blotches, sometimes faint medial brownish blotches across the vertebral crest.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	description	Male coloration. The brilliant coloration of adult males is characterized by a brightly colored bluish to turquoise head, with bluish coloration continuing posteriorly to fore limb insertion. There is a relatively faded light (bright yellowish, when under acute distress, a character typical for C. bachae only) stripe at the upper lip crossing the tympanum from beneath the eye to end of head. Gular pouch colored in darker blue, interscale skin black. Atriangular to crescent-shaped patch of small black scales is present in front of shoulder. Very faint brownish blotches extending mid-dorsally over the vertebral crest, from above fore limb insertion on to tail; posterior to fore limb insertion brownish orange in color on trunk and tail; hind limb in a slightly darker brown; venter cream.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	distribution	Distribution. Calotes bachae is known from southern Vietnam, with two specimens reported from the Vietnam-China border in northern Vietnam, and from eastern Cambodia (Fig. 1).	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	biology_ecology	Ecology. Calotes bachae is a diurnal, semi-arboreal lizard, often observed climbing on tree trunks at a height of 5 – 10 meters above the ground. The species mainly inhabits dipterocarp lowland forests and cultural landscapes up to 700 m a. s. l. The species seems to be quite heliophilic, preferring more open habitats without closed canopies (Hartmann et al. 2013), but was also observed in dense tropical monsoon forests with closed canopy, in open gallery forest, and anthropogenic habitats like roadsides within the forest or open park landscapes within the headquarters of the park. In these anthropogenic habitats C. bachae occurs in syntopy with Calotes versicolor (Hartmann et al. 2013). According to Hartmann et al. (2013) C. bachae feeds onnumerous arthropods including Formicidae, Coleoptera, Orthoptera, and Myriapoda. Males acquire their breeding coloration at the end of February, while gravid females have been found mid-April and egg-laying of a clutch of five eggs was observed in April and May. Juveniles hatched after 56 days while incubated at 22 to 25 ° Cin captivity.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002426FFF16019FA2B2D0AC960.taxon	materials_examined	Calotes bachae (n = 17): BMNH 1927.5.20.30, BMNH 1927.5.20.31, BMNH 1927.5.20.32, MNHN 1927.44, MNHN 1927.46, ZFMK 88935, ZFMK 88936 (now IEBR A. 2012.23), ZFMK 88937, ZFMK 92028, ZFMK 92029, ZFMK 94395, ZFMK 94396, ZFMK 94397, ZMMU NAP 01509, ZMMU NAP 01512, ZMMU NAP 02910, ZMMU NAP 02911.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	description	(Fig. 2, Clade B) urn: lsid: zoobank. org: act: EC 4 DA 90 A- 417 D- 48 E 6 - 8 EA 2 - FA 073 D 168425	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	materials_examined	Holotype. CAS 215539 (adult male, Fig. 6 A-B, Clade B) from Myanmar, Sagaing Division, Mon Ywa District, AK Park, Thabake Sae Camp [22.316806 ° N, 94.475556 ° E], collected by H. Win, T. Thin, S. L. Oo and H. Tun on June 9 th 2000. Paratypes. CAS 210270 from Myanmar, Alaungdaw Kathapa National Park, Thabakesay (Log Cabin Camp) [22.318194 ° N, 94.475722 ° E]; ZFMK 97991 (formerly CAS 243200) from Myanmar, Chin State, Phalum District, Simggial village [23.762583 ° N, 93.546167 ° E, 1362 m.]; CAS 243028, CAS 243050 both from Myanmar, Magway Division, Gangaw District, Gangaw Township, Mauk village [22.335861 ° N, 94.144583 ° E, 205 m.].	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	diagnosis	Diagnosis. Alarge sized Calotes species with a known maximum SVL of 122 mm in males and 114 mm in females. Tail relatively short, up to 270 mm in males and 223 mm in females. The new species can be distinguished from other species of the complex by the combination of the following characters: 1) head and body very robust; 2) nuchal and dorsal crests continuous, composed of erect compressed scales, directed posteriorly, larger on the nuchal crest than on the dorsal crest, becoming smaller towards the tail; 3) 50 – 62 scale rows around midbody; 4) 35 – 45 vertebral spines and scales in males, 49 – 50 in females; 5) body scales small, homogeneous, feebly keeled and arranged in regular rows; 6) a short row of separated spines on both sides of the head, directing from the tympanum to the first scale of the nuchal crest; 7) extremities relatively short and robust; 8) oblique skin fold in front of the fore limbs, 9) head and body bluish, with a white band from the tip of the mouth along the upper lip, the tympanum and prominently continuing between the dorsolateral brownish orange body blotches on the body reaching the hind limbs, band as broad as the height of the tympanum on the head and above the front legs, becoming gradually narrower until the insertion of the hindlimbs; 10) three or more large distinct brownish orange blotches on both sides of the body between the limbs.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	description	Description of the holotype. Moderately large male of 110 mm SVL. Tail relatively short (201 mm), extremities robust. Head large, distinct from the neck and lateral sides flat. Posterior parts of jaw angle swollen. Snout-tip blunt. Nostril in a single scale, separated from the labial scale by a single scale. Rostral and mental scales small. Canthus rostralis sharp and straight from the nostril to the posterior part of the eye, including six scales between the nostril and the eye and 12 supraciliary scales. Eleven supralabial scales on both sides of the head, separated from the orbit by five rows of small scales. Nine infralabial scales. Seven scales between the orbit and the tympanum, tympanum distinct, with a row of spiny scales from above the tympanum to the first scale of the nuchal crest. Scales on chin and throat keeled. Nuchal crest with 14 scales, dorsal crest with 21 scales. Spines of the nuchal crest larger than those of the dorsal crest, relatively uniform in height at the nuchal crest but gradually decreasing from posterior of the neck to the hind limbs. Dorsal and lateral body scales keeled, pointing upwards and backwards. Caudal scales keeled, directed backwards. Fore and hind limbs relatively robust, forth finger and fourth toe longest. Male coloration. Males in breeding color with blue head and body (Fig. 6 C). Awhite band is present from the tip of the snout along the upper lips and the tympanum, predominantly continuing between the dorsolateral blotches on the lateral sides of the body to the hind limbs. The band is as broad as the height of the tympanum on the head and above the fore legs, becoming gradually narrower towards the insertion of the hind limbs. Three or more large distinct brownish-orange blotches on the lateral sides of the body between the limbs. Non-display coloration unknown. Variation. Body measurements and meristic characters for adult individuals are given in Table 4. Specimens ranged in size from the smallest female with a SVL of 92 mm (CAS 210270) and the largest female with a SVL of 114 mm (CAS 243200) to the largest male with a SVL of 120 mm (CAS 243028). In general, adult males are larger than females, and have greater SVLs, tail lengths and head lengths and widths and fewer vertebral scales, including crest scales. While proportionally the heads of adult males and females are equal, males have proportionally wider heads than females, probably because of the swollen jaw angle. Dorsal coloration differs between sexes and both sexes are able to change coloration. Coloration characters (lateral stripe, blotches) are lighter or sometimes absent in females. Blue coloration on head and body. The stripe is distinctly present between the mental and the shoulder, becoming more indistinct between the blotches on the lateral sides of the body, and extending to above the hindlimbs. Irregular whitish dots of several scales are present on the lateral sides of the body in females.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	etymology	Etymology. The specific epithet is a patronym formed in the genitive singular honoring Dr. Peter Geissler, Museum Natur und Mensch, Freiburg, Germany, in recognition of his work on the Southeast Asian herpetofauna in general, and his collection of Calotes bachae specimens in 2009 in particular, which initiated research on the Calotes mystaceus complex.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	distribution	Distribution. The new species is documented only from Myanmar and India (e. g., BMNH 1956.1.11.98, specimens mentioned by Lalremsanga et al. [2010]).	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	biology_ecology	Ecology. Calotes geissleri sp. n. is diurnal and semi-arboreal. Preferred habitats are unknown. Lalremsanga et al. (2010) collected one individual on a branch of Pinus kesiya, about 3 m above the ground, in a secondary forest. The specimen was kept and remained greyish brown in coloration in captivity, but changed the color of the head and anterior portion of the trunk to bright blue minutes after exposure to the sun. Like other Calotes species, C. geissleri sp. n. feeds on arthropods like Coleoptera, Formicidae and others. In India (see Lalremsanga et al. 2010) the species occurs in sympatry with Calotes jerdoni and C. versicolor.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242DFFF163B1FDCE2E60C9C7.taxon	materials_examined	Calotes geissleri sp. n. (n = 6): BMNH 1856.1.11.98, CAS 210270, CAS 215539, CAS 243028, CAS 243050, CAS 243200 (now ZFMK 97991).	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	description	(Fig. 2, Clade A) urn: lsid: zoobank. org: act: 7827 D 48 E-E 121 - 4904 - 9636 - 3 A 46 AE 42 B 369	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	materials_examined	Holotype. ZFMK 92606 (adult male, Fig. 7 A-C, Clade A) from Cambodia, Siem Reap Province, near Kbal Spean within the Phnom Kulen National Park [13.699167 ° N, 103.998611 ° E]. Paratypes. ZFMK 88341 (adult male), ZFMK 92607 (adult female) from the same locality as the holotype.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	diagnosis	Diagnosis. Alarge species of Calotes, with a maximum SVL of 143 mm in males and 122 mm in females. It can be distinguished from other species of the group by the combination of the following characters: 1) head, body and limbs robust, tail long but not as long as in C. mystaceus; 2) body scales mid-sized, homogeneous, keeled, arranged in regular rows; 3) upper dorsolateral scales pointing up- and backwards; 4) body scales arranged in 45 – 60 rows around midbody; 5) two short and separated spines, surrounded by a ring of scales between the tympanum and the vertebral crest on both sides of the head; 6) vertebral crest continuous from above the tympanum to the hind limbs, composed of erected scales, directed posteriorly, highest slightly in front of the insertion of the front limbs, becoming gradually shorter towards the hind limbs; 7) vertebral scales, including vertebral spines, 37 – 52 in males and 43 – 60 in females; 8) oblique skin fold in front of the fore limbs; 9) head, body, and limbs bluish in males; 19) males with a white stripe from between nostril and orbit along the upper lip and the tympanum to the front limb insertion; 11) three to five distinct dark brown dorsolateral blotches.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	description	Description of the holotype. Large male (SVL 118 mm). Body robust, tail relatively short, 236 mm long. Fore- and hind limbs relatively slender, fourth finger and toe longest. Head distinct from the neck, posterior jaw angles heavily swollen. Tip of the snout blunt, rostral small. Nostril large, in a single scale, separated from the rostral by one elongated scale and from the first two supralabial scales by two rectangular scales. Canthus rostralis distinct, formed by nine scales from the nostril to above the orbit and continuous with supraciliary scales. Lateral sides of the head flat. Supralabial scales 9 / 9, infralabial scales 10 / 10. Supralabial scales separated from the orbit by four rows of small scales. Eight scales between the orbit and the tympanum, two of them slightly ridged. Tympanum distinct, two spines, surrounded by a ring of scales, between the tympanum and the vertebral crest. Mental scale small, bordered by two postmental scales which are separated from each other. First pair of the postmental scales in contact with the infralabial scale. Scales on the dorsal part of the chin smooth to feebly keeled, becoming strongly keeled towards the throat. Vertebral crest continuous from above the tympanum to the tail, spines highest above the insertion of the front limbs and gradually decreasing towards the tail. Dorsal and lateral scales strongly keeled, pointing up- and backwards. Ventral scales parallelly keeled. Caudal scales smooth to feebly keeled, directed backwards. Subcaudal scales parallel and strongly keeled. Male coloration. Head, body and limbs bluish. Awhite stripe, as high as the tympanum, is present from between nostril and orbit along the upper lip and the tympanum to the insertion of the front limb. The stripe is followed by three to five large reddish-brown to dark-brown blotches, with the first above the insertion of the fore limbs and the latest at around midbody or above the insertion of the hindlimbs, sometimes continuing on the tail. Throat coloration darker than the chin and the head coloration. From Thailand, males are known to have bright blue hindlimbs and crest scales. Intermediate males with a blue head and a white stripe, but body coloration brownish-black with indistinct light brown blotches. Non-brilliant coloration in males from Thailand reddish-brown, with a white non-continuous stripe from the orbit to above the hindlimbs. Blotches brownish, darker than the body coloration. Variation. Bodymeasurements and meristic characters for adult individuals are given in Tables 4. Specimens ranged in size between the smallest adult with a SVL of 84 mm (MNHN 1884.548) and the largest female with a SVL of 126 mm (ZFMK 44893) to the largest male measuring 145 mm (MNHN 1884.546). In general, adult males are larger than females, because of longer SVLs, tail lengths, head lengths, and widths and slightly fewer vertebral scales including the crest scales. Heads of adult males and females are proportionally equal in length, but male heads are wider than those of females, probably because of the swollen jaw angle. Dorsal coloration differs between sexes and both sexes are able to change the coloration. Coloration characters being lighter in females; head and body blue. Lateral stripe from between the nostril and orbit along the upper lips and the tympanum to the first blotch. Sometimes the first two brownish blotches are framed by scales in the same coloration as the lateral stripe. Three to five blotches present, from above the insertion of the fore limbs to about mid-body or even hindlimb insertion. Non-display coloration of females with lateral parts of the body gray and dorsal parts brown, with three to four darker crossbands between the lateral stripes on both sides of the body. Lateral stripe white, from the mental scale along the upper lips and the tympanum to the hindlimbs. Head light blue.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	etymology	Etymology. The specific epithet is a patronym formed in the genitive singular honoring Dr. Stephan Goetz, Munich, Germany, in recognition of his longtime support of species conservation efforts in Cambodia.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	distribution	Distribution. Calotes goetzi sp. n. is distributed in a large area of Indochina and inhabits most of the previous range of C. mystaceus. The new species is documented by museum specimens and photo vouchers from Cambodia, China (Yunnan province), Laos, Myanmar, and Thailand.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	biology_ecology	Ecology. Calotes goetzi sp. n. is diurnal and semi-arboreal to arboreal and can easily climb on tree trunks at a height of 5 – 10 meters or above. The species is known from dipterocarp lowland forests, cultural landscape and secondary forest. At its type locality the species was observed in more open habitats with a closed canopy, but also within dense monsoon tropical forests and solitary trees in rather open areas. Here it occurs sympatrically with Calotes versicolor. Hawkeswood & Sommung (2018) report it from farmland with e. g., rice, coconut, banana, and durian plantations. Chan-Ard et al. (2015) report the species from a wide range of forest types in Thailand, including tree-lined agricultural lands and grasslands. Similar to other Calotes species, C. goetzi sp. n. feeds on a variety of arthropods. Chan-Ard et al. (2015) report from Thailand, that mature individuals are territorial, chasing away possible intruders. Eggs are buried in soft soil and later guarded by the males. Amber et al. (2017) recognized a possible ontogenetic shift in defense strategies of C. goetzi sp. n. (C. mystaceus at the time of their publication), with older individuals utilizing color change, while juveniles do not.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E3900242FFFF163B1F9EC2CC9C9ED.taxon	materials_examined	Calotes goetzi sp. n. (n = 71): BMNH 1868.4.3.61, BMNH 1868.4.3.63, BMNH 1891.11.26.29, BMNH 1914.4.25.1, BMNH 1929.12.1.11, BMNH 1933.3.10.1, BMNH 1933.3.10.2, MNHN 1884.546, MNHN 1884.547, MNHN 1884.548, MNHN 1893.335, MNHN 1893.336, NME R 0581 / 09, NME R 0584 / 09, NME R 0585 / 09, NME R 0686 / 11, NME R 0751 / 12, NME R 0783 / 13, NME R 0784 / 13, NME R 0785 / 13, NME R 0786 / 13, NME R 0787 / 13, NME R 0790 / 14, ZFMK 16640, ZFMK 16641, ZFMK 40561, ZFMK 43906, ZFMK 43930, ZFMK 43931, ZFMK 43932, ZFMK 44893, ZFMK 44894, ZFMK 45490, ZFMK 45491, ZFMK 45492, ZFMK 45493, ZFMK 45494, ZFMK 45495, ZFMK 45496, ZFMK 45497, ZFMK 45498, ZFMK 45499, ZFMK 45500, ZFMK 45501, ZFMK 45502, ZFMK 45553, ZFMK 45554, ZFMK 47080, ZFMK 47081, ZFMK 49164, ZFMK 49202, ZFMK 49219, ZFMK 49220, ZFMK 49221, ZFMK 49242, ZFMK 49243, ZFMK 49244, ZFMK 49245, ZFMK 49246, ZFMK 55610, ZFMK 55611, ZFMK 55612, ZFMK 84867, ZFMK 88341, ZFMK 92606, ZFMK 92607, ZMB 11603 A, ZMB 30186, ZMB 30188 A, ZMB 30197, ZMB 6034.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	description	(Fig. 2, Clade D)	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	materials_examined	Holotype. MNHN 2557 (juvenile male, Fig. 8 A-B), from “ Indes orientales (…) pays de Birmans [= Myanmar]. ”	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	diagnosis	Original Diagnosis. “ Deux petites épines places l’une après l’autres de chaque côte de la nuque. Un pli oblique en longueur devant l’épaule. Écailles des côtes du tronc grandes; celles du ventre moitié plus petites. Dessus de la base de la queue subanguleux, garni d’écailles seulement un peu plus grandes que celles qui les avoisinent. Fauve en dessus; sous l’oeil une bande jaune qui se prolonge jusque sur l’épaule. ” Revised Diagnosis. Asmall sized Calotes with a maximum known SVL of 101 mm in males and 99 mm in females. Distinguished from all other species of the group by the combination of the following characters: 1) Head and body slender, with long tail and extremities; 2) body scales relatively large in respect to the body size, homogeneous, strongly keeled and arranged in regular rows; 3) upper dorsolateral scales pointing back- and upwards; 4) 44 – 56 scale rows around midbody; 5) no spines above the tympanum; 6) Vertebral crest, composed of erected spiny scales, directed posteriorly, continuous from above the tympanum to about the insertion of the hindlimbs, but spines becoming abruptly shorter above the insertion of the front limbs; 7) Vertebral scales, including crest spines 38 – 49 in males and 38 – 48 in females; 8) oblique skin fold in front of the fore limbs; 10) Head, chest, front limbs, and anterior dorsal crest turquoise; 11) whitish lateral stripe from the snout along the upper lip and the tympanum to behind the insertion of the fore limbs, behind tympanum becoming brownish beige and fusing with beige dorsolateral blotches above front limb insertion; 12) four faint beige dorsolateral blotches.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	description	Male coloration. According to the original description ventral and dorsal parts of the body, tail and limbs brownish. Upper parts of the head olive, chest and throat brownish to yellow. Orange-yellowish stripe from the upper lip crossing the tympanum to the shoulders. More recently collected males (Figure 8 C) show the head and the anterior of body to the shoulders blue, with a yellowish stripe from the mental towards the upper lips and the tympanum to above and in front of the insertion of the front limbs, followed by indistinct orange blotches. Variation. Body measurements and meristic characters for adult individuals are given in Table 4. Specimens ranged in size between the smallest juvenile male with a SVL of 58 mm (CAS 239398) to the largest male with a SVL of 101 mm (BMNH 1891.11.26.18) and the largest female measuring 99 mm (BMNH 1868.4.3.62). In general, adult males and females have the same body proportions. Dorsal coloration differs between sexes with coloration characters generally being lighter in females. Both sexes are able to change their color. Turquoise coloration is restricted to the throat, other parts of the head and body brownish. White lateral stripe present, extending from the mental along the upper lips, becoming beige posterior to tympanum, ending in the first lateral blotch of the same coloration, followed by three blotches of the same color but lighter.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	distribution	Distribution. With the description of the new species of the group, the distribution of Calotes mystaceus is restricted to the Irrawaddy delta region of coastal southern Myanmar. However, further research is needed to fully clarify the species distributional boundaries.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	biology_ecology	Ecology. Calotes mystaceus is diurnal and arboreal. The species inhabits forests and more open landscapes with a closed canopy. Mating was observed in September 2014 in a public garden within Yangon (pers. comm. Andreas Hellmann). As in other species of the genus the diet mainly consists of arthropods.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002434FFF16019FA682B79C84C.taxon	materials_examined	Calotes mystaceus (n = 9): BMNH 1868.4.3.60, BMNH 1868.4.3.62, BMNH 1891.11.26.18, CAS 206548, CAS 213300, CAS 239398, CAS 240287, CAS 240296, MNHN 2557.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	description	(Fig. 2, Clade C)	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	materials_examined	Holotype. CAS 232388 (adult male, Fig. 9 A-B, Clade C) from Myanmar, Kachin State, Myitkyina Township, Gat Shang Yang village [25.373421 ° N, 97.37475 ° E], collected by T. Z. Min on April 8 th 2003. Paratypes. CAS 232247 from Myanmar, Sagaing Division, Homalin Township, North of Swekawngaw [25.371694 ° N, 95.369028 ° E, 205 m.]; CAS 232387 from Myanmar, Kachin State: Myitkyina, Gat Shang Yang village; CAS 239206 from Myanmar, Sagaing Division, Hkanti District, Hkanti Township, Linpha village [25.803389 ° N, 95.528778 ° E, 155 m.]; CAS 232819 from Myanmar, Kachin State, Mohnyin Township, Hepu village [25.094528 ° N, 96.401833 ° E, 254 m.]; ZFMK 97990 (formerly CAS 232389) from Myanmar, Kachin State: Myitkyina, Gat Shang Yang village.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	diagnosis	Diagnosis. Asmall Calotes species of the complex, males with a known maximum SVL of 77 mm, females with a SVL of 73 mm. Tail length short, up to 140 mm. It can be distinguished from the other species of the complex by the combination of the following characters: 1) head slender, not as distinctfrom the body as in other species of the complex; 2) body scales small and homogeneous, smooth, and arranged in regular rows around the body. 3) Upper dorsolateral scales pointing backwards; 4) body scales arranged in 50 – 54 rows around midbody; 5) 40 – 47 vertebral spines and scales in males, 44 – 48 in females; two short and well separated spines above the tympanum; 6) low nuchal and dorsal crest continuous to the midpoint between the limbs, composed of erect compressed scales, which are larger on the nuchal and smaller on the dorsal crest, gradually decreasing towards the end of the crest; 7) oblique skinfold in front of the fore limb; 8) extremities and tail long, but shorter than in C. mystaceus; 9) bluish head and chest; 10) whitish stripe from about the nostril, along the upper lip to about the insertion of the fore limb, posteriorly the whitish stripe is laterally intersected by a dark reticulate pattern across vertebrae; 11) brownish blotches missing.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	description	Description of the holotype. Small male (SVL 77 mm). Extremities relatively slender with the fourth finger and toe longest, tail incomplete. Head slightly distinct from the neck, area posterior of jaw angle slightly swollen. Tip of the snout blunt, rostral small, nostril in a single scale, separated from the rostral and the first supralabial scale by two scales in a row. Canthus rostralis of five scales, distinct and straight, canthus scales continuous with supraciliary scales. Lateral sides of the head flat with 9 / 9 supralabial scales separated from the orbit by three rows of small scales. Five feebly ridged scales from the orbit to above the tympanum. Tympanum distinct, one spiny scale posteriorly above the tympanum. Mental scale small, bordered by two postmental scales which are separated from each other, only the first pair is in contact with the infralabial scales; 10 / 10 infralabial scales. Lateral scales on the chin and throat smooth, becoming keeled towards the ventral part. Acontinuous vertebral crest of elevated spiny scales from above the tympanum to about mid-body, spine height gradually decreasing posterior of the neck, in total 40 vertebral scales from the nape to above the cloaca. Dorsal and lateral scales keeled, pointing back- and upwards, in 50 rows around midbody. Caudal scales parallel keeled, directed backwards. Ventral scales parallel keeled. Male coloration. Brilliant coloration unknown (Fig. 9 C). Head most probably blue, with a whitish stripe, less broad than in the other species of the C. mystaceus complex, from behind the nasal scale along the upper lips and the tympanum to the shoulders. Followed by one distinct and one or more indistinct lateral blotches of the same color.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	materials_examined	Variation. Body measurements and meristic characters for adult individuals are given in Table 4. Specimens ranged in size from the smallest female with a SVL of 47 mm (CAS 232247) and the largest female with a SVL of 73 mm (CAS 232387) to the largest male with a SVL of 77 mm (CAS 232388). Adult males are slightly larger than females, because of longer SVLs and tail lengths, and have fewer vertebral scales including the crest scales. However, proportionally males and females are equal. Dorsal coloration of females unknown, but most probably similar to other taxa of the species group with females having lighter coloration characters (lateral stripe, blotches) than males. Most probably, both sexes are able to change their coloration.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	etymology	Etymology. The specific epithet is a patronym honoring Jens Vindum, retired staff of the California Academy of Sciences, in respect of his outstanding contributions to the herpetology of Myanmar and SE Asia. The patronym of his family name was used, in recognition of his impressive beard which has a direct link to “ mystaceus ” [Greek for bearded], together with the Latin word “ barbatus ” for bearded.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	distribution	Distribution. This new species is so far only known from northern Myanmar.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	biology_ecology	Ecology. Calotes vindumbarbatus sp. n. is diurnal and arboreal to semi-arboreal. It inhabits forests and more open habitats with a dense canopy, but detailed habitat preference is unknown. As in other Calotes species the diet most likely consists predominantly of arthropods, but details are not known.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
023E39002430FFF1639BFA682C4EC8AE.taxon	materials_examined	Calotes vindumbarbatus sp. n. (n = 6): CAS 232247, CAS 232387, CAS 232388, CAS 232389 (now ZFMK 97990), CAS 232819, CAS 239206.	en	Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas, Böhme, Wolfgang (2021): Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn zoological Bulletin 70 (1): 141-171, DOI: https://doi.org/10.20363/BZB-2021.70.1.141
