taxonID	type	description	language	source
03B17D7ED17EFFECFF3987ADFC1CF9D9.taxon	materials_examined	M a t e r i a l e x a m i n e d: Holotype ♂, MALAYSIA – Sabah Kinabalu Nat. Park Poring H. Spr. 600 m 11. VII. 92 Heiss (MHNG). Paratypes: 4 ♂♂, 8 ♀♀, with the same data as the holotype; 1 ♂, SABAH, Poring Hot Springs, 31. III. 1983, B. Hauser (MHNG); 2 ♂♂, BRUNEI, Temburong, Kuala Belalong F. s. c., mixed Dipterocarp for. 16 - 20. IV. 1993, E. Heiss (MHNG).	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17EFFECFF3987ADFC1CF9D9.taxon	description	D e s c r i p t i o n: Length 1.35 - 1.43 mm, width 0.65 - 070 mm, dorsoventral diameter 0.65 - 0.76 mm. Head and most of body dark brown to blackish with reddish shine, prothorax and abdomen sometimes lighter than elytra, mesoventer and metaventer. Apical abdominal segments, antennae and tarsi light, yellowish. Femora and tibiae reddish brown, darker than tarsi and antennae. Length / width rations of antennomeres as: III 22 / 4: IV 20 / 4: V 27 / 5: VI 24 / 6: VII 40 / 7: VIII 36 / 6: IX 39 / 10: X 43 / 10: XI 52 / 10. Pronotal punctation very fine, hardly visible at 25 times magnification. Scutellum concealed. Elytral punctation distinct, punctures well delimited, fine and dense, clearly visible at 20 times magnification, sutural striae parallel, shortened, starting about 0.25 mm posterior of level of pronotal lobe. Adsutural areas flat. Hypomeron very finely punctate, with short stria delimiting inferior vertical surface. Mesoventrite with shallow median impression, lacking mesal ridge, bearing longitudinally striate microsculpture; latter distinct between procoxae and mesocoxae, absent laterally of coxal cavities. Mesocoxal process of mesoventrite notched. Median part of metaventrite flattened or hardly impressed, impunctate, with microsculpture consisting of short oblique striae. Lateral parts of metaventrite not microsculptured, each with patch of fairly coarse punctures absent from posterior surface. Submesocoxal lines angulate, with oblique inner and outer sections, punctate only along outer section. Submesocoxal areas 0.04 - 0.05 mm long, as halves or somewhat longer than halves of shortest intervals to metacoxae. Metanepisternum flat, parallel-sided, with straight, wide, shortened, and punctate suture. Abdomen apparently impunctate, with exposed segments bearing transversally striae microsculpture. Male characters: Segments I to III of protarsi hardly widened. Aedeagus (Figs 1, 2) 0.59 - 0.66 mm long. Parameres narrow, each with rounded subapical lobe. Internal sac bearing pair of admesal, teeth-like sclerites, pair of narrow, weakly sclerotized basal filamentous structures, and broad apical vesicle covered by scales.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17EFFECFF3987ADFC1CF9D9.taxon	discussion	C o m m e n t s: This new species is likely related to S. taylori. It may be distinguished by the denser and more distinct elytral punctation, the strigulate microsculpture present on the mesoventrite, the metaventrite and the abdomen, the sides of the metaventrite bearing dense and coarse punctures, the internal sac of the aedeagus bearing two strongly sclerotized admesal tooth-like denticles and lacking a central sclerite.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17EFFECFF3987ADFC1CF9D9.taxon	etymology	E t y m o l o g y: The species epithet is a Latin adjective meaning dull.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF3986B0FBBBFB3C.taxon	materials_examined	M a t e r i a l e x a m i n e d: Holotype: ♂, [E Malaysia] 10 miles point from Keningau Sabah, Is. Borneo 7. iii. 1993 T. Ueno leg. (MHNG).	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF3986B0FBBBFB3C.taxon	description	D e s c r i p t i o n: Length 1.80 mm, width 1.0 mm, dorsoventral diameter 1.08 mm. Head and most of body dark brown, apical abdominal segments much lighter, femora and tibiae slightly lighter than thorax, tarsi and antennae yellowish. Length / width rations of antennomeres as: III 15 / 7: IV 19 / 8: V 30 / 8: VI 28 / 9: VII 45 / 13: VIII 35 / 11: IX 43 / 13: X 42 / 15: XI 58 / 15. Pronotal punctation very fine, hardly visible at 25 times magnification. Scutellum concealed. Elytral punctation distinct, punctures not well delimited, fine and dense, clearly visible at 20 times magnification, sutural striae parallel, shortened, starting about 0.20 mm posterior of level of pronotal lobe. Adsutural areas flat. Hypomeron very finely punctate, with short stria delimiting inferior vertical surface. Mesoventrite not microsculptured, impunctate, with low mesal ridge delimited by shallow impressions. Mesocoxal process of mesoventrite notched. Metaventrite not microsculptured, very finely and sparsely punctate, slightly convex in middle. Submesocoxal lines convex, punctate only along outer section. Submesocoxal areas 0.05 mm long, about as halves of shortest intervals to metacoxae. Metanepisternum flat, parallel-sided, with straight, narrow, shortened, and punctate suture. Abdomen very finely and sparsely punctate, dense basal puncture rows in middle of lateral margins of ventrites I excepted. Ventrites I to IV lacking microsculpture, apical abdominal segments with punctulate microsculpture. Male characters: Segments I to III of protarsi hardly widened. Aedeagus (Figs 3 - 5) 0.86 mm long. Parameres narrow, each with subapical, mesally oriented, acute denticle. Proximal part of internal sac longitudinally striate. Middle of internal sac with two short, incurved sclerites surrounded by finely denticulate membranes and followed by two sclerites pointed apically and two narrow apical pieces.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF3986B0FBBBFB3C.taxon	discussion	C o m m e n t s: The aedeagal characters suggest relationship of this new species to S. grande LÖBL, 1986. Notable are the very similar parameres having each a narrow subapical denticles pointed mesally. The structures of the internal sacs are, however, significantly distinctive in these two species, in particular by the presence of a pair of sclerotized subapical teeth and two narrow, apical sclerites present in S. opertum, absent from S. grande. Only S. simulans (LÖBL, 1971) 1986 possesses narrow and acute subapical parameral denticles. It is readily distinguished by its significantly smaller body (1.3 mm long) and the internal sac lacking teeth-like sclerites.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF3986B0FBBBFB3C.taxon	etymology	E t y m o l o g y: The species epithet is a Latin adjective meaning secrete, mysterious.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF398348FE28F9F4.taxon	materials_examined	M a t e r i a l e x a m i n e d: 2 ♂♂, SABAH, Sabalangan River env., about 25 km SE Sapulut, 26. VI. 1988, J. Kodada & F. Čiampor; 1 ♀, SABAH, Poring Hot Springs, LangananRiver, 850 m, 14. V. 1987, D. Burckhardt & I. Löbl; 1 ♂, BRUNEI, Temburong, Kuala Belalong F. s. c., 60 - 300 m, 16 - 20. IV. 1993, E. Heiss (all MHNG).	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17DFFEFFF398348FE28F9F4.taxon	discussion	C o m m e n t s: The species was based on a single male from the Semengoh Forest Reserve in Sarawak, and subsequently reported from the Philippine islands Luzon, Palawan and Mindanao.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17CFFEEFF3986B0FD82FE41.taxon	materials_examined	M a t e r i a l e x a m i n e d: 1 ♀, SABAH, Mt. Kinabalu Nat. Park, Headquaters, 5. V. 1980, M. & A. Sakai (MHNG).	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
03B17D7ED17CFFEEFF3986B0FD82FE41.taxon	discussion	C o m m e n t s: This is likely an undescribed species, similar to the above described S. opertum, though its body is smaller and darker. It may be clearly distinguished by the characters given in the key below.	en	Löbl, Ivan (2021): On the Bornean species of Scaphoxium LÖBL, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beiträge 53 (1): 85-89, DOI: http://doi.org/10.5281/zenodo.5643309
