taxonID	type	description	language	source
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	description	Type species: Alpiscaptulus medogensis Jiang & Chen, sp. nov. Diagnosis: The following combination of characters defines the new genus. Medium in size (HB = 100 mm; GLS = 29.08 mm). The rostrum is long and slender. The tail is more than one-quarter the total length or twice the length of the hind foot and is moderately covered with pale brown to white hairs (Fig. 4). The chin is covered with white hairs. Fore toes and hind toes are not webbed, and the first toe of hind foot is curved. The dental formula is 3.1.4.3 / 3.1.4.3 = 44. The zygomatic plates are less developed. The interorbital region is broad, without constriction and appears rectangular. The pterygoid region and the coronoid valley are shallow, with a weak and triangular coronoid process. The auditory bullae are incomplete. Description: Because the new genus is monotypic, the description is the same as in the species description, presented below. Etymology: The genus name Alpiscaptulus is derived from the Greek αλΠ, alp or high mountain, and σκαΠτύλος, a small digger. The genus name refers to a small digging animal living in the mountains. Comparisons: Among the Scalopini tribe, Alpiscaptulus is most similar to the monotypic genus Scapanulus, and they are also the only two genera distributed in Asia. The two species have similar body sizes and share some unique characters in contrast with other species, such as curved thumbs of hind feet and relatively long tail, which is more than one-quarter the total length. However, they are strikingly dissimilar in many other external characters. The tail of Alpiscaptulus is moderately covered with pale brown to white hairs, unlike the dense dark brown hairs in Scapanulus. Thus, the tail of Alpiscaptulus is more slender than Scapanulus in appearance. The rostrum of Alpiscaptulus is much longer and more slender than that of Scapanulus. The most distinctive cranial difference between Alpiscaptulus and Scapanulus is the dentition. The possession of 44 teeth in Alpiscaptulus is distinguishable from Scapanulus (36 teeth), and the teeth of Alpiscaptulus are weaker and smaller than in Scapanulus. It can further be distinguished from Scapanulus by the less developed zygomatic plates, broader interorbital region, much shallower pterygoid region and coronoid valley, the reduced, triangular coronoid process and the relatively smaller measurements of UTL, M 2 - M 2, UML, P 4 M 3, ZB and CH (Table 3). Alpiscaptulus can be distinguished from the North American Scalopus, Scapanus and Parascalops by its smaller size, more slender rostrum, curved thumbs of hind feet and by the relatively longer (more than onequarter the total length), hairy and pale brown tail. Furthermore, the reduced and triangular coronoid process is much weaker than that of Parascalops and Scapanus, while the incomplete auditory bullae of Alpiscaptulus is noticeably different from Scalopus and Scapanus. The 44 teeth of Alpiscaptulus are distinct from Scalopus (36 teeth) and the webbed fore toes and hind toes of Scalopus are unique among scalopines. ALPISCAPTULUS MEDOGENSIS JIANG & CHEN, SP. NOV. Suggested common name: Medog mole, • µ. (motuo yan). Zoobank registration: urn: lsid: zoobank. org: act: 292 B 5 D 9 C- 4 F 54 - 4 C 58 - B 0 F 8 - 2 E 7 BF 7 DE 0 D 1 D Holotype: KIZ: 037966, an adult female collected by Kang Luo at on 2 June 2019. Dried skin, cleaned skull and alcohol-preserved carcass. Type locality: Mt Namjagbarwa, Damu Town, Medog County, south-east Tibet, China (29 ° 44 ’ 32 ” N, 95 ° 40 ’ 59 ” E), elevation approximately 3650 m a. s. l. Paratype: KIZ: 037965, an adult male collected by Changzhe Pu on 24 May 2019 at an elevation of 2400 m a. s. l. from Gedang Town, Medog (29 ° 29 ’ 31 ” N, 95 ° 45 ’ 11 ” E). Dried skin, cleaned skull (braincase damaged) and alcohol-preserved carcass. Measurements: See Table 3. Diagnosis: As for genus (see above). Description: A medium-sized mole (HB = 100 mm, TL = 41 mm, HF = 18 mm). Dorsal pelage dark grey, ventral pelage imperceptibly lighter. Eyes minute, concealed in fur; external ears absent. The nose is long and develops into a conical snout; the nose and chin are covered in white hairs (Fig. 4). The tail is about twice the length of the hind foot and is moderately covered in pale brown to white hairs, with a tuft of longer hairs located on the tip. The dorsal surfaces of hands and feet are light brown, white at the margin. Fore toes and hind toes not webbed, and the palms are nearly equal in length and width, with the flattened and slender claws. The hind foot is long and narrow, and the first toe is curved like that of Scapanulus oweni, which is set outward at a slight angle against remaining toes. We observed the curved second toe of the right hind foot in specimen KIZ: 037965, but not in specimen KIZ: 037966. The skull is triangular and flat (Fig. 2). The rostrum is long and slender; the braincase is broad with welldeveloped mastoid processes. The zygomatic arch is moderately heavy, and slightly convergent anteriorly; zygomatic plates are small. The orbits are narrow and situated in the middle of the skull. The interorbital region is broad, strong, and nearly rectangular. The infraorbital foramen is small. The pterygoid region is shallow, and auditory bullae are incomplete. The coronoid process is weak and triangular; the condyloid process is short and angled upward at roughly 45 °, so the coronoid valley appears shallow. The angular process is well developed, curved upward at the posterior. The dental formula is I 3 / 3, C 1 / 1, P 4 / 4, M 3 / 3, totalling 44, and the incisors, canines and premolars are all well-spaced. The I 1 is enlarged and flat and twice the height of I 2, while the I 3 is the smallest of all. The canine is larger than I 2 and I 3, P 1 and P 2 are subequal and smaller than the canine, P 3 is almost as high as canine but broader, and P 4 is much larger than P 3. The transverse section of the upper molar is W-shaped, M 1> M 2> M 3; M 1 and M 2 broad with a trilobed basal shelf, while M 3 is much smaller and bilobed. I 1 and I 2 are enlarged and flattened, and I 3 is much smaller. The canine is slightly larger than I 3. All three incisors and the canine lean forward. All premolars are subequal; the P 2 of the right side of the type specimen has two equal tooth tips (Figs 2, 3), but only one tip on the other specimen. M 1 and M 2 are almost equal, M 3 is slightly smaller. Etymology: The species name medogensis derives from Medog, the type locality of the new species, and the Latin adjectival suffix - ensis, meaning “ belonging to ”. Comparisons: Because all genera of the Scalopini except Scapanus are monotypic, the comparisons are the same as the genus comparisons presented above. Distribution: Alpiscaptulus medogensis is only known from the type locality at Mt Namjagbarwa in Medog County, south-east Tibet, China. The known elevational range is 2400 m a. s. l. to 3700 m a. s. l .. Ecology and habitat: From 20 April to 3 June 2019, we sampled small mammals along the elevational gradients from 650 to 3800 m a. s. l. at Mt Namjagbarwa. The two specimens of A. medogensis were captured at relatively high elevations, 2400 m a. s. l. and 3700 m a. s. l., respectively. During the survey, we did not see mole runways and mounds of excavated earth in this area. At 2400 m a. s. l., the mole was captured in a small hole using a mole trap. The habitat at this area was brushy, dominated by oak forest, with abundant grass cover on the ground. The mole from 3700 m a. s. l. was captured by a plastic bucket pitfall (15 cm in diameter and 20 cm in height). The vegetation of this area was dominated by heath forest, with mixed bamboo forests.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	type_taxon	Type species: Alpiscaptulus medogensis Jiang & Chen, sp. nov.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	diagnosis	Diagnosis: The following combination of characters defines the new genus. Medium in size (HB = 100 mm; GLS = 29.08 mm). The rostrum is long and slender. The tail is more than one-quarter the total length or twice the length of the hind foot and is moderately covered with pale brown to white hairs (Fig. 4). The chin is covered with white hairs. Fore toes and hind toes are not webbed, and the first toe of hind foot is curved. The dental formula is 3.1.4.3 / 3.1.4.3 = 44. The zygomatic plates are less developed. The interorbital region is broad, without constriction and appears rectangular. The pterygoid region and the coronoid valley are shallow, with a weak and triangular coronoid process. The auditory bullae are incomplete.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	description	Description: Because the new genus is monotypic, the description is the same as in the species description, presented below.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	etymology	Etymology: The genus name Alpiscaptulus is derived from the Greek αλΠ, alp or high mountain, and σκαΠτύλος, a small digger. The genus name refers to a small digging animal living in the mountains. Comparisons: Among the Scalopini tribe, Alpiscaptulus is most similar to the monotypic genus Scapanulus, and they are also the only two genera distributed in Asia. The two species have similar body sizes and share some unique characters in contrast with other species, such as curved thumbs of hind feet and relatively long tail, which is more than one-quarter the total length. However, they are strikingly dissimilar in many other external characters. The tail of Alpiscaptulus is moderately covered with pale brown to white hairs, unlike the dense dark brown hairs in Scapanulus. Thus, the tail of Alpiscaptulus is more slender than Scapanulus in appearance. The rostrum of Alpiscaptulus is much longer and more slender than that of Scapanulus. The most distinctive cranial difference between Alpiscaptulus and Scapanulus is the dentition. The possession of 44 teeth in Alpiscaptulus is distinguishable from Scapanulus (36 teeth), and the teeth of Alpiscaptulus are weaker and smaller than in Scapanulus. It can further be distinguished from Scapanulus by the less developed zygomatic plates, broader interorbital region, much shallower pterygoid region and coronoid valley, the reduced, triangular coronoid process and the relatively smaller measurements of UTL, M 2 - M 2, UML, P 4 M 3, ZB and CH (Table 3).	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	discussion	Alpiscaptulus can be distinguished from the North American Scalopus, Scapanus and Parascalops by its smaller size, more slender rostrum, curved thumbs of hind feet and by the relatively longer (more than onequarter the total length), hairy and pale brown tail. Furthermore, the reduced and triangular coronoid process is much weaker than that of Parascalops and Scapanus, while the incomplete auditory bullae of Alpiscaptulus is noticeably different from Scalopus and Scapanus. The 44 teeth of Alpiscaptulus are distinct from Scalopus (36 teeth) and the webbed fore toes and hind toes of Scalopus are unique among scalopines.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	materials_examined	Zoobank registration: urn: lsid: zoobank. org: act: 292 B 5 D 9 C- 4 F 54 - 4 C 58 - B 0 F 8 - 2 E 7 BF 7 DE 0 D 1 D Holotype: KIZ: 037966, an adult female collected by Kang Luo at on 2 June 2019. Dried skin, cleaned skull and alcohol-preserved carcass. Type locality: Mt Namjagbarwa, Damu Town, Medog County, south-east Tibet, China (29 ° 44 ’ 32 ” N, 95 ° 40 ’ 59 ” E), elevation approximately 3650 m a. s. l. Paratype: KIZ: 037965, an adult male collected by Changzhe Pu on 24 May 2019 at an elevation of 2400 m a. s. l. from Gedang Town, Medog (29 ° 29 ’ 31 ” N, 95 ° 45 ’ 11 ” E). Dried skin, cleaned skull (braincase damaged) and alcohol-preserved carcass.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	description	Measurements: See Table 3.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	diagnosis	Diagnosis: As for genus (see above).	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	description	Description: A medium-sized mole (HB = 100 mm, TL = 41 mm, HF = 18 mm). Dorsal pelage dark grey, ventral pelage imperceptibly lighter. Eyes minute, concealed in fur; external ears absent. The nose is long and develops into a conical snout; the nose and chin are covered in white hairs (Fig. 4). The tail is about twice the length of the hind foot and is moderately covered in pale brown to white hairs, with a tuft of longer hairs located on the tip. The dorsal surfaces of hands and feet are light brown, white at the margin. Fore toes and hind toes not webbed, and the palms are nearly equal in length and width, with the flattened and slender claws. The hind foot is long and narrow, and the first toe is curved like that of Scapanulus oweni, which is set outward at a slight angle against remaining toes. We observed the curved second toe of the right hind foot in specimen KIZ: 037965, but not in specimen KIZ: 037966. The skull is triangular and flat (Fig. 2). The rostrum is long and slender; the braincase is broad with welldeveloped mastoid processes. The zygomatic arch is moderately heavy, and slightly convergent anteriorly; zygomatic plates are small. The orbits are narrow and situated in the middle of the skull. The interorbital region is broad, strong, and nearly rectangular. The infraorbital foramen is small. The pterygoid region is shallow, and auditory bullae are incomplete. The coronoid process is weak and triangular; the condyloid process is short and angled upward at roughly 45 °, so the coronoid valley appears shallow. The angular process is well developed, curved upward at the posterior. The dental formula is I 3 / 3, C 1 / 1, P 4 / 4, M 3 / 3, totalling 44, and the incisors, canines and premolars are all well-spaced. The I 1 is enlarged and flat and twice the height of I 2, while the I 3 is the smallest of all. The canine is larger than I 2 and I 3, P 1 and P 2 are subequal and smaller than the canine, P 3 is almost as high as canine but broader, and P 4 is much larger than P 3. The transverse section of the upper molar is W-shaped, M 1> M 2> M 3; M 1 and M 2 broad with a trilobed basal shelf, while M 3 is much smaller and bilobed. I 1 and I 2 are enlarged and flattened, and I 3 is much smaller. The canine is slightly larger than I 3. All three incisors and the canine lean forward. All premolars are subequal; the P 2 of the right side of the type specimen has two equal tooth tips (Figs 2, 3), but only one tip on the other specimen. M 1 and M 2 are almost equal, M 3 is slightly smaller.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	etymology	Etymology: The species name medogensis derives from Medog, the type locality of the new species, and the Latin adjectival suffix - ensis, meaning “ belonging to ”.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	discussion	Comparisons: Because all genera of the Scalopini except Scapanus are monotypic, the comparisons are the same as the genus comparisons presented above.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	distribution	Distribution: Alpiscaptulus medogensis is only known from the type locality at Mt Namjagbarwa in Medog County, south-east Tibet, China. The known elevational range is 2400 m a. s. l. to 3700 m a. s. l ..	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B46755553FFFB2FCC4FAE8FC68F99E.taxon	biology_ecology	Ecology and habitat: From 20 April to 3 June 2019, we sampled small mammals along the elevational gradients from 650 to 3800 m a. s. l. at Mt Namjagbarwa. The two specimens of A. medogensis were captured at relatively high elevations, 2400 m a. s. l. and 3700 m a. s. l., respectively. During the survey, we did not see mole runways and mounds of excavated earth in this area. At 2400 m a. s. l., the mole was captured in a small hole using a mole trap. The habitat at this area was brushy, dominated by oak forest, with abundant grass cover on the ground. The mole from 3700 m a. s. l. was captured by a plastic bucket pitfall (15 cm in diameter and 20 cm in height). The vegetation of this area was dominated by heath forest, with mixed bamboo forests.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	materials_examined	Zoobank registration: urn: lsid: zoobank. org: act: 292 B 5 D 9 C- 4 F 54 - 4 C 58 - B 0 F 8 - 2 E 7 BF 7 DE 0 D 1 D Holotype: KIZ: 037966, an adult female collected by Kang Luo at on 2 June 2019. Dried skin, cleaned skull and alcohol-preserved carcass. Type locality: Mt Namjagbarwa, Damu Town, Medog County, south-east Tibet, China (29 ° 44 ’ 32 ” N, 95 ° 40 ’ 59 ” E), elevation approximately 3650 m a. s. l. Paratype: KIZ: 037965, an adult male collected by Changzhe Pu on 24 May 2019 at an elevation of 2400 m a. s. l. from Gedang Town, Medog (29 ° 29 ’ 31 ” N, 95 ° 45 ’ 11 ” E). Dried skin, cleaned skull (braincase damaged) and alcohol-preserved carcass.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	description	Measurements: See Table 3.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	diagnosis	Diagnosis: As for genus (see above).	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	description	Description: A medium-sized mole (HB = 100 mm, TL = 41 mm, HF = 18 mm). Dorsal pelage dark grey, ventral pelage imperceptibly lighter. Eyes minute, concealed in fur; external ears absent. The nose is long and develops into a conical snout; the nose and chin are covered in white hairs (Fig. 4). The tail is about twice the length of the hind foot and is moderately covered in pale brown to white hairs, with a tuft of longer hairs located on the tip. The dorsal surfaces of hands and feet are light brown, white at the margin. Fore toes and hind toes not webbed, and the palms are nearly equal in length and width, with the flattened and slender claws. The hind foot is long and narrow, and the first toe is curved like that of Scapanulus oweni, which is set outward at a slight angle against remaining toes. We observed the curved second toe of the right hind foot in specimen KIZ: 037965, but not in specimen KIZ: 037966. The skull is triangular and flat (Fig. 2). The rostrum is long and slender; the braincase is broad with welldeveloped mastoid processes. The zygomatic arch is moderately heavy, and slightly convergent anteriorly; zygomatic plates are small. The orbits are narrow and situated in the middle of the skull. The interorbital region is broad, strong, and nearly rectangular. The infraorbital foramen is small. The pterygoid region is shallow, and auditory bullae are incomplete. The coronoid process is weak and triangular; the condyloid process is short and angled upward at roughly 45 °, so the coronoid valley appears shallow. The angular process is well developed, curved upward at the posterior. The dental formula is I 3 / 3, C 1 / 1, P 4 / 4, M 3 / 3, totalling 44, and the incisors, canines and premolars are all well-spaced. The I 1 is enlarged and flat and twice the height of I 2, while the I 3 is the smallest of all. The canine is larger than I 2 and I 3, P 1 and P 2 are subequal and smaller than the canine, P 3 is almost as high as canine but broader, and P 4 is much larger than P 3. The transverse section of the upper molar is W-shaped, M 1> M 2> M 3; M 1 and M 2 broad with a trilobed basal shelf, while M 3 is much smaller and bilobed. I 1 and I 2 are enlarged and flattened, and I 3 is much smaller. The canine is slightly larger than I 3. All three incisors and the canine lean forward. All premolars are subequal; the P 2 of the right side of the type specimen has two equal tooth tips (Figs 2, 3), but only one tip on the other specimen. M 1 and M 2 are almost equal, M 3 is slightly smaller.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	etymology	Etymology: The species name medogensis derives from Medog, the type locality of the new species, and the Latin adjectival suffix - ensis, meaning “ belonging to ”.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	discussion	Comparisons: Because all genera of the Scalopini except Scapanus are monotypic, the comparisons are the same as the genus comparisons presented above.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	distribution	Distribution: Alpiscaptulus medogensis is only known from the type locality at Mt Namjagbarwa in Medog County, south-east Tibet, China. The known elevational range is 2400 m a. s. l. to 3700 m a. s. l ..	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
03B467555530FFB0FCC5F9FBFA74FE49.taxon	biology_ecology	Ecology and habitat: From 20 April to 3 June 2019, we sampled small mammals along the elevational gradients from 650 to 3800 m a. s. l. at Mt Namjagbarwa. The two specimens of A. medogensis were captured at relatively high elevations, 2400 m a. s. l. and 3700 m a. s. l., respectively. During the survey, we did not see mole runways and mounds of excavated earth in this area. At 2400 m a. s. l., the mole was captured in a small hole using a mole trap. The habitat at this area was brushy, dominated by oak forest, with abundant grass cover on the ground. The mole from 3700 m a. s. l. was captured by a plastic bucket pitfall (15 cm in diameter and 20 cm in height). The vegetation of this area was dominated by heath forest, with mixed bamboo forests.	en	Chen, Zhong-Zheng, He, Shui-Wang, Hu, Wen-Hao, Song, Wen-Yu, Onditi, Kenneth O., Li, Xue-You, Jiang, Xue-Long (2021): Morphology and phylogeny of scalopine moles (Eulipotyphla: Talpidae: Scalopini) from the eastern Himalayas, with descriptions of a new genus and species. Zoological Journal of the Linnean Society 193: 432-444, DOI: https://doi.org/10.1093/zoolinnean/zlaa172
