taxonID	type	description	language	source
8260172F77D25A7FA4358B520450C65F.taxon	description	Figs 13, 14	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
8260172F77D25A7FA4358B520450C65F.taxon	materials_examined	Material examined. Holotype, ♂, Vietnam, | Cha Val, Nam Giang, Quang | Nam, 15 ° 35 ' 21.9 " N, | 107 ° 29 ' 01.9 " E, RTS, 546 m, VD, | 28. V. 2009, Tran Thieu Du leg. (coll. S. Pugaev) | [RTS means " secondary forest " and VD means " come to light trap "] (MWM, GU 35.635). Paratypes, ♂, Vietnam, Cha Val vill., Nam Giang, Quang Nam, 15 ° 35 ' 21.9 " N, 107 ° 29 ' 01.9 " E, RTS, 546 m, VD, 28. V. 2009, Tran Thieu Du leg (IEBR); ♂, Vietnam, S. Vietnam, Quang Nam Prov., Nam Giang Distr., Cha Val vill., secondary forest, 400 m [15 ° 35 ' 21.9 " N, 107 ° 29 ' 01.9 " E], 24. VI. 2009, Tran Thieu Du leg (MWM, GU 17.744); ♂, Central Vietnam, Quang Nam Prov., Phuoc Son Distr., Phuos My Comm., Deo Lo Xo, 17. VII 2009, Du Thieu Tran leg. (coll. S. Pugaev); ♂, N. Vietnam, Cao Bang Prov., Phi Oak Mts., Nguyen Binh Distr., Thanh Cang comm., Phia Den vill., 22 ° 34 ' N, 105 ° 52 ' E, 1030 m, pupa 10. XI. 2009, ex pupa 22. XII. 2009, leg. S. Pugaev (MWM, GU 35.634). Eggs: C. Vietnam, Prov. Da Nang, comm. Hoa Vang, Ba Na Mts, Da Nang, 15 ° 59.5 ' N, 107 ° 59 ' E, 1.500 m, 08. XII 2009, leg. Tran Thieu.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
8260172F77D25A7FA4358B520450C65F.taxon	description	Description. Male (Figs 13, 14). Fore wing slightly rounded at apex. Brown to blackish basal fascia broad, distinct and prominent; the outer margin of the basal fascia is light coloured and blurred. Discal dot distinct, round, dark brown. Antemedial fascia crenulate. Postmedial fascia is yellow, not well expressed, straight, and with contrastingly dark brown basal shadow. Apical patch in the fore wings is yellow, sometimes enclosing a brown, large, semi-ovoid spot. Submarginal fascia dentate, weakly expressed. At the apex, the outer margin of the submarginal fascia is densely irrorated with dark brown scales. On the hind wing, the postmedial fascia is yellow, not clearly visible, with narrow, light brown inner shadow; submarginal fascia almost absent and only noticeable as dots on veins. Male genitalia (Figs 26, 27). Uncus lobes pointed, boomerang-shaped; their bases separated. Valvae wide at the base and narrowed distally, apically terminating with hook-shaped spurs. Saccus not developed. Aedeagus short and thick; its coecum is not longer than the width. Vesica slightly curved, with a ring lacking scobination in the medial zone where cornuti change their direction. The cornuti are of equal length.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
8260172F77D25A7FA4358B520450C65F.taxon	diagnosis	Diagnosis. The paler yellow ground colour distinguishes the species clearly from all other congeners except the externally similar A. kantonensis. Both species are found flying allopatrically and can only be reliably separated genitalically. The apex of the uncus lobes is tapered whereas it is spatulate in A. kantonensis, the distance between two first spurs on the valvae apex is much greater than in A. kantonensis has. The apical lobes of the uncus of A. kantonensis are more slender with apex expanded and round while in A. chloralis they are stouter, with medial widening and pointed apex; the valve of A. kantonensis has 3 spines of the same size while in A. chloralis the ventral spine is much larger than the dorsal and medial spines.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
8260172F77D25A7FA4358B520450C65F.taxon	etymology	Etymology. In Greek mythology, Chloris (or Flora in Roman mythology) was one of the wives of the god of the West Wind Zephyrus. " Chloris " also means a greenish yellow colour typical of the ground colour of the moth.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
8260172F77D25A7FA4358B520450C65F.taxon	distribution	Distribution. Vietnam: Cao Bang, Da Nang, Quang Nam (Fig. 59).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
68503DEC735755D39D29636EC224072D.taxon	description	Description. Moderate sized moths (fore wing length 24 - 29 mm in males, 30 - 35 mm in females), characterized by bright, contrasting pattern. The basal fascia of the fore wing is dark brown, dentate and sometimes indistinct. The discal spot is round, dark brown and is always present. The antemedial fascia is angled or curved, towards the costa approximately at a right angle. The apical patch of the fore wing is elongated, semicircular, and extends along the costa. It is pale in colour, usually with a dark centre. The postmedial fascia is yellow, usually of the same colour as the apical forewing patch, straight, and running towards the apex of the wing; it is fused with the apical patch in A. floralis, A. zephyrus, A. witti, A. kantonensis, and A. chloralis. The postmedial fascia is bordered basally by a well expressed, thin, dark shadow giving the impression that it is a double line; in some species the postmedial fascia is not clearly visible and weakly separable in colour from the distal part of the wing, but its basal shadow is always well defined. The submarginal fascia is zigzag-shaped, brown, sometimes indistinct. The hind wings are darker basally while the external field is bright yellow, usually without prominent pattern. The postmedial fasciae of the hind wings are of the same colour as those of the fore wings.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
68503DEC735755D39D29636EC224072D.taxon	diagnosis	Diagnosis of the group. Valvae are without saccular lobes, except for A. witti. The aedeagus is truncated. The antemedial fascia with angle of curvature about 90 °.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
68503DEC735755D39D29636EC224072D.taxon	distribution	Distribution. The group occupies the southern area of the generic range and is known from northern India, Nepal, southern China, north-eastern Myanmar, Cambodia, and Vietnam (Figs 39, 40).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
38D5D4B9F02D544C91158FF813429B48.taxon	description	Figs 1 - 4	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
38D5D4B9F02D544C91158FF813429B48.taxon	materials_examined	Material examined. Lectotype, ♂, Darjiling, 79. 57 (BMNH). Paralectotype, ♀, Darjiling, 79. 57 (BMNH); f #, Darjeeling (ZMHU); ♀, India, Himalaya, Darj [eeling], 15. X [19] 18 (MWM); ♂, India W. B., Darjeeling, 2.000 m, 17 - 22. VII. 1989, leg. W. Thomas (SMFL); ♂, Sikkim, Harmand, 1890 (MNHN); 12 ♂♂, Nepal, Annapurna Himal, 1.700 m, 1 km N of Tal, 84 ° 23 ' E, 28 ° 28 ' N, 08. VI. 1996, leg. Hreblay & Szaboky (MWM); ♀, Nepal Annapurna Himal, 2.450 m, 83 ° 43 ' E, 28 ° 22 ' N, 23 - 24. VI 1996, leg. Gy. M. Lszl & G. Ronkay (MWM); 2 ♂♂, East-Nepal, Surke Danda, 3 km NE Sukeyer, Lali Kharka 18. V. 1997, 2.600 m, lg. Hreblay & Szecsenyi (MWM); 3 ♂♂, East-Nepal, Deorali Danda, Anpan, 1.900 m, 18. VI. 1998, leg. M. Hreblay & B. Benedek (MWM); 2 ♂♂, E. Nepal, Janakpur, Dolakha Deolari, 2.800 m, 25. V. - 7. VI. 1994, M. S. Limbu leg. (NSMT); 14 ♂♂, 4 ♀♀, C. Nepal, Kali-Gandaki-Tal, Kalopani-Dhumpu, 2.500 m, 30. V. - 15. VI. 1973, leg. Dierl-Lehmann (ZSM).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
38D5D4B9F02D544C91158FF813429B48.taxon	description	Description. Male (Figs 1, 2). The forewing length is 25 - 29 mm. The costa is straight; the fore wing apex is weakly falcate. The wing pattern is contrasting, with yellow and bright brown (brick-red) colours predominating. Discal dot is always distinct, dark coloured and rounded. The antemedial fascia is brown to brick-red or light-brown, serrate. The outer field bordering the antemedial fascia can have duplicating serrate fasciae (from one to three). The postmedial fascia is straight, bright yellow in colour and has the distal margin extending to the apex. The fore wing apical patch is elongated, yellow or pale yellow. The postmedial fascia of the fore wing is fused with the distal part of the apical patch. It is bordered basally by a thin dark shadow which does not reach the wing apex, but is angled towards the costa. The submarginal fascia is zigzag-shaped, with expansion along the veins, terminating as a dark field at the wing apex. The postmedial fascia situated across the middle of the hind wing separates the outer yellow field of the wing from the inner brown one. The latter itself becoming more yellowish basally; submarginal fascia brown, zigzag-shaped, with corresponding separate dots on veins. Male genitalia (Figs 15, 16). Lobes of uncus are narrow, sabre-shaped with rounded apex, their bases fused at one third of their length. The valvae are roundly triangular; without saccular lobes but with three small spurs which are directed inwards (sometimes hardly visible depending on the preparation). The coecum is short, not longer than its width; the opening of vesica is usually more sclerotized. The vesica is dome-shaped, its base is completely covered with short cornuti which are poorly extended to the aedeagus tube. Female (Figs 3, 4). Length of the fore wing is 31 - 33 mm; similar in pattern to the male but is generally larger and paler. Additionally, the female is not so contrastingly coloured. The wings are wider with a more blunt apex and with vague basal fascia and light brown discal dot. The abdomen is light red laterally and terminally. Female genitalia (Figs 21 - 23). Ovipositor lobes are bean-shaped. Anterior apophyses are slender with well-marked, sclerotized base extending to the base of posterior apophyses which are a bit shorter. Postvaginal plate with acute margins and well-marked medio-caudal dent. Ostium is sheltered by the antevaginal plate which consists of two lobes laterally. Antrum is wide, calycine, slightly sclerotized, standing proud of antevaginal plate ventrally. Ductus bursae is not sclerotized, wide up to the curve and narrowing beyond that. A single thorn-shaped signum is situated at the equator of the corpus bursae, long, slightly curved, with broader base.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
38D5D4B9F02D544C91158FF813429B48.taxon	diagnosis	Diagnosis. The males of the species differ from some of the other members of the group by having a well expressed " double " postmedial fascia on the hind wing. This " double " postmedial fascia can be also found in A. zephyrus and A. witti, but in both the latter species this fascia (not its basal shadow) is much more weakly expressed. The species A. floralis can be distinguished from A. zephyrus sp. nov. and A. witti sp. nov. by the smaller distance between the postmedial fascia and outer wing margin. Lobes of uncus are more slender and longer than in other species. Valvae with small apical spurs visible only under magnification. Aedeagus is straight in contrast to faintly curved in the other three species, cornuti on vesica are sparse, not dense as in other species. Saccus is small and not substantially distinguished.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
38D5D4B9F02D544C91158FF813429B48.taxon	distribution	Distribution. Northern India and Nepal (Fig. 58).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
4BBF49DCED515060ABDF0B74D318B7BD.taxon	description	Figs 9 - 12	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
4BBF49DCED515060ABDF0B74D318B7BD.taxon	materials_examined	Material examined. Holotype, ♂, China, Kwangtung, [Lo fao shan] (ZMHU). Paratypes, 2 ♂♂, 3 ♀♀, [China] [mostly unreadable because of Mell's abbreviations or given by him in Chinese, some are pointed from Lo Fao], ex Raupe, ex coll. R. Mell (ZMHU); 3 ♂♂, China, Hong Kong, 100 m, Sekkong, Jan. 1978, leg. Allen (MWM); ♂, China, Hongkong, New Territories, leg. Uk Tau, 27. IV. 1998 (SMFL); 2 ♂♂, China, Hongkong, New Territories, leg. Uk Tau, 21. XI. 1998 (SMFL); ♂, N. Vietnam, Prov. Ninh Binh, Nho Quan Distr., Bong - Cuc Phuong vill., 20 ° 21 ' N, 105 ° 36 ' E, 6 - 9. X. 2008, 360 m, leg. Zolotuhin (MWM); ♂, Tonkin [Northern Vietnam], An Chau, coll. L. & J. de Joannis (MNHN); ♂, Central Vietnam, Quang Nam Prov., Phuoc Son Distr., Phuos My Comm., Deo Lo Xo, 17. VII 2009, Du Thieu Tran leg. (coll. S. Pugaev); 2 ♂♂, Cambodia, Kampot Prov., Bokor N. P., Hill Station, 1.025 m, 10 ° 37,37 ' N, 104 ° 01,33 ' E, 19 - 21. I. 2006, leg. G. Csorba & G. Ronkay (MWM).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
4BBF49DCED515060ABDF0B74D318B7BD.taxon	description	Description. Male (Figs 9, 10; Figs 52, 53). Fore wing length 23 - 29 mm. Costal margin slightly curved, wing apex rounded or weakly acute. There is a dark spot in the yellow basal area of the fore wing. Basal fascia is dentate, inner margin distinct, vague distally. Discal dot is well expressed, round, brown or black. Antemedial fascia is crenulate, thick, with two such duplicating lines on the outer margin. Postmedial fascia is yellow, not well expressed, but with conspicuous inner dark-brown or black shadow which terminates at the wing apex, anterially to the apical patch. The apical patch is yellow, with an enclosed large ovoid dark spot. Submarginal fascia brown, dentate, hardly visible, but widened into well expressed dots on the veins. In the hind wing, the shadow of the postmedial fascia is terracotta, straight, with a slight apical curve, while the yellow postmedial fascia is not well expressed. Medial field is uniformly coloured citron-yellow. Moths emerging in the dry summer season are smaller than those of the wet winter season. Male genitalia (Figs 24, 25). Bases of uncus lobes situated tightly against each other, their lobes are spatulate. In the valva, the costa is straight, the sacculus margin curved, the apex scoop-shaped and bearing 2 - 3 spurs. Saccus is not expressed. Aedeagus short, coecum almost the same length of the tube of the aedeagus. Vesica cone-shaped and angled at which point the cornuti change direction forming a bare membranous, circular zone lacking cornuti. Cornuti are of equal length. Female (Figs 11, 12). Fore wing length 30 - 35 mm. Apart from the much wider wings, the female is markedly different to the male in the colour of the outer margins which are not yellow but light brown. The postmedial fascia on the fore wing is yellow and falls short of the apex by about 3 mm. Female genitalia (Fig. 30). Papillae anales are bean-shaped. Anterior apophyses are more slender and longer than the posterior. The postvaginal plate is wide with two rounded, lateral lobes in its upper half. The antevaginal plate has a deep, rounded, medial incision, which divides the plate into two lobes, each lobe is elongated, curved, strongly widened distally to form a fishtail shape and with rounded apical corners. Antrum is wide and short, slightly sclerotized. Ductus is wide at antrum and distinctly narrowed at the teardrop-shaped corpus bursae which has a thorn-shaped signum with a round base at its equator.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
4BBF49DCED515060ABDF0B74D318B7BD.taxon	diagnosis	Diagnosis. The brown shadow of the postmedial fascia reaches the wing margin at the apex in contrast to A. floralis, A. zephyrus and A. witti which terminates approximately 5 mm from the wing apex. It is very similar to A. chloralis in colour and pattern, but in the latter species the submarginal fasciae are vague or absent in the hind wings; in the male genitalia the lobes of the uncus are spatulate.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
4BBF49DCED515060ABDF0B74D318B7BD.taxon	distribution	Distribution. South-eastern China: Guangdong; Vietnam; Cambodia (Fig. 59).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	description	Figs 7, 8	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	materials_examined	Material examined. Holotype, ♂, China / Sichuan, | Qionglai Shan, | 31 ° 13 ' N, 102 ° 23 ' E, | May 2006, 1.400 m, | leg. Siniaev & his team (MWM). Allotype, ♀, China / Sichuan, Qionglai Shan, 31 ° 13 ' N, 102 ° 23 ' E, May 2006, 1400 m, leg. Siniaev & his team (MWM). Paratypes, 6 ♂♂, China, Sichuan, Qionglai Shan, 31 ° 13 ' N, 102 ° 23 ' E, May 2006, 1.400 m, leg. Siniaev & his team (MWM); 4 ♂♂, China, Sichuan, Wolong Reserve, Siguliang Shan, 31 ° 09 ' N, 103 ° 06 ' E, May 2006, 1.500 - 1.800 m, leg. Sinaev & his team (MWM); ♂, China, Sichuan, Dayu, Xiling, 1.900 m, 22. IV. 2001, Nativ. coll. (NSMT).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	description	Description. Male (Fig. 7). The antennae have long rami and their length is a bit more than 1 mm. The forewing length is 26 - 27 mm. The forewing costa is straight, with pointed apex. The basal fascia is broad, brown, irregularly shaped. The discal dot is round, brown. The antemedial fascia is crenulate and curved towards the costa. The postmedial fascia is straight and ends in the apex zone, but not reaching the apex; it is a " double " line and similar in this respect to A. floralis and A. zephyrus. The apical fore wing patch is yellow or pale yellow. The submarginal fascia is dentate, brown. In the hind wings the postmedial fascia is straight. The submarginal fascia is curved and consists of dots situated on the veins. Male genitalia (Figs 19, 20). The lobes of the uncus are similar to those of A. zephyrus. The valvae are broad basally; their costa is concave. The cucculus is dome-shaped, with two apical spurs. The sacculus is moderately developed, triangular, pointed apically. The saccus is weakly developed, almost not protruding. The aedeagus is short, broad, with rounded coecum. The vesica is in the shape of an inverted dome, with the opening on the top; it is covered with small cornuti, which form a single, round field. Female (Fig. 8). Fore wing length 30 mm. The females generally resemble the males of the species, but are paler, with wider wings. The female differs from A. floralis in the location of the postmedial fascia of the hind wing which is aligned with postmedial fascia on the fore wing (it does not transect the wing medially as in A. floralis, but is found closer to the base). Female genitalia (Fig. 29). The papillae anales are bean-shaped. The posterior and anterior apophyses are of almost equal size, similar both in length, and width. The postvaginal plate has a medio-caudal dentition. The antevaginal plate consists of two high, rounded lobes with a slit between them. The antrum is bulb-shaped, equal to the antevaginal plate in height, slightly sclerotized. The ductus bursae is straight, membranous. The corpus bursae is circular-shaped, with a claw-shaped signum on the equator, somewhat shorter than in A. floralis.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	diagnosis	Diagnosis. A sister species to A. zephyrus and A. floralis; all of them very similar externally. The species A. witti can be separated from A. floralis by the position of the postmedial fascia of the hind wing which is located more basally in A. witti; the submarginal fascia of the hind wing is excavate in A. witti, but is dentate in A. floralis. In A. witti the rami of the male antennae are twice as long as those of A. zephyrus (Figs 56, 57). The new species is easily distinguished by the male genitalia; although the shape of the uncus lobes resemble those of A. zephyrus and the shape of the aedeagus and vesica are similar to those of A. floralis, A. witti ' s valvae are of diagnostic shape, quite dissimilar to those of either A. floralis or A. zephyrus.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	distribution	Distribution. China: Sichuan (Fig. 58).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
EF2CD2A427725172A2124D6B1141A445.taxon	etymology	Etymology. The species is named in honor of the eminent entomologist late Dr. Thomas J. Witt (Munich, Germany) He has made many outstanding contributions to the study of Lepidoptera worldwide and is well known for his support of young scientists; he is the founder of the largest Lepidoptera collection in Europe and it is thanks to this collection that the new species was revealed.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	description	Figs 5, 6	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	materials_examined	Material examined. Holotype, ♂, " China / Yunnan, | Dianceng Shan, | 25 ° 41 ' N, 100 ° 05 ' E, | 2.200 m, 15 - 30. VII. 2004, | leg. Sinaev & his team " (MWM). Allotype, ♀, China, Prov. Nord-Yuennan, Li-kiang, 2.000 m, 3. VII. 1935, H. Hoene (ZFMK). Paratypes, 6 ♂♂, China, prov. Yunnan, Dali Bali aut. Pref., Yunlong, 13 km N of Caojian, Feng Shuing Mts, 2.400 m, 20. V. - 23. VI. 1999, leg. R. Brechlin (MWM); ♂, China, Yunnan, 18 km SW of Baoshan, 2.100 m, 16 - 19. V. 2003, leg. Murzin (MWM); ♂, China, Yunnan, Wudingshan, N-Xiangyan, 3.200 m, VI. 2000, leg. native collector (MWM); 2 ♂♂, China, W-Yunnan, Wudingshan, N-Xiangyan, 3.200 m, E. VI. 2000, leg. local collector (SMFL); 2 ♂♂, China, Yunnan, Dian Cang Shan, 25 ° 41 ' N, 100 ° 05 ' E, 2.200 m, 15 - 30. VII. 2004, leg. Siniaev & his team (MWM); m #, China, South-Yunnan, Yueson Ximeng, 1.200 m, Mitte V. 2000, leg. native collector (MWM); 2 ♂♂, China, E-Yunnan, Huangcaotai, Malong, 2.450 m, EVI. 2000, leg. local collector (SMFL); ♂, China, E-Yunnan, West Qiubei, Yangxiong Mts. 2.500 m, VI. 2000, leg. native collector (MWM); 3 ♂♂, China, Prov. Nord-Yuennan, Li-kiang, 9., 14., 16. VII. 1934, H. Hoene (ZFMK); ♂, China, N-Yunnan, Sanfengshan, Yaoan, 2.200 m, VII. 2000, leg. local collector (SMFL); 3 ♂♂, China, SE-Sichuan, Gaomushan, Guling (near west Guizhou border), about 1.900 m, leg. Ying et al. (SMFL); ♂, China, prov. Jiangxi / Fujian, Wuyi Mt., Zhod, 1.490 m, VIII. 2000, leg. native collector (MWM); ♂, Thibet, H. Testont (MNHN); m #, East-Tibet, Tonge-Pelung, 2.000 m, 24. VI. [19] 96 (SMFL); m #, Thibet Oriental, Chasseurs thibetains du P. Dejean, 1903 (MNHN); ♂, Tibet, Vrianatong, E. Le-Moult, coll. Sheljuzhko (ZMSU); ♂, Myanmar, VIII. 2006, Chudu Raji, 2.800 m, Kachin (SMFL); ♂, N. Vietnam, Mt. Fan-si-pan, 1.600 - 1.800 m, near Chapa, 22 ° 20 ' N, 103 ° 40 ' E, secondary forest, May 1996, leg. local collectors (MWM).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	description	Description. Male (Fig. 5). The fore wing length is 25 - 29 mm. The costa is straight; the fore wing apex is weakly falcate. The antemedial fascia is dimpled and curved. The postmedial fascia is straight and ends in the apex zone, but does not reach the apex. The submarginal fascia is sinuate and joins with the postmedial fascia at the wing apex. The postmedial fascia is a " double " line. The fore wing yellow apical patch encloses a distinct black spot. The discal dot is ovoid, dark brown and distinct. The basal spot is wide, dark brown with ragged margins, sometimes indistinct or completely absent. In the hind wing, the postmedial fascia is straight until approaching the costa where it angles outwards, it has a sharp inner boundary and is distinctly separated from the basal dark shadow, but the outer boundary of the fascia is not well defined and blurred. The submarginal fascia is present as a line of dashes on the veins terminating near the costa as an indistinct patch. Male genitalia (Figs 17, 18). The bases of uncus lobes are slender and distinctly separated; their apices are pointed and directed inwards. The base of the valvae is wide but narrows towards the apex and terminates in two small spurs. The costa is straight with a little, medial depression. The saccus is not large, its length not greater than its width. The aedeagus is short, thick; its coecum is rounded and twice as long as it is wide. Distally the vesica is curved at a right angle to the utricular basal part carrying cornuti. All cornuti are of equal length. Female (Fig. 6). The fore wing length is about 33 mm. The females are paler and the wings broader than the males, but the wing pattern is similar. In the hind wings the indistinct patch situated near the costa on the submarginal fascia is developed as in males. Female genitalia (Fig. 28). The anterior apophyses are 3 times shorter than the posterior apophyses, with small apical rounded thickening. The postvaginal plate has a broad medio-caudal dent. The antevaginal plate consists of a pair of small plates, which are situated close together. The antrum is rectangular, sclerotized, slightly longer than the antevaginal plate. The ductus bursae has constant width along the full length. The corpus bursae is ball-shaped, with a medial thorn-shaped signum.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	diagnosis	Diagnosis. This taxon is a sister species of A. floralis and is its vicariant, being found allopatrically in southern China, northern Vietnam and north-eastern Myanmar. The costal margin of the fore wing is straight as in A. floralis, it distinguishes them from A. kantonensis and A. chloralis. The yellow postmedial fascia of the hind wing is not distinctly visible (but its basal shadow is well expressed) and does not transect the middle of the wing as in A. floralis but is closer to a wing base. The apical fore wing patch is yellow, and encloses a distinct black spot. The submarginal fascia terminates apically as two dots on the vein. It also differs from related species in genitalic characters, especially by the wide valvae base, the narrowed apex which terminates with one, two or three small spurs, and the small, medial depression on a straight costa; the lobes of the uncus are narrowly separated from each other in contrast to A. floralis and A. kantonensis. From A. chloralis it also differs in the shape of the uncus lobes. Vesica with cornuti different from A. floralis but similar to A. kantonensis and A. chloralis; the field with cornuti is divided into two parts by a ring lacking any cornuti. Position of the postmedial fascia in the hind wing distinguishes the females of the species from those of A. floralis: in A. floralis it divides the wing into two, rather equal halves and therefore its end point on the hind wing costa does not correspond to the fore wing's postmedial fascia at the tornus. In contrast, in A. zephyrus, both fore and hind wing fasciae come together at almost the same point. The signum is a bit shorter than in A. floralis. The species A. zephyrus differs from A. witti by the presence of a dark small spot near the forewing apex situated close to the costa and by the blurred dark submarginal marking near the costa of the hind wing (A. witti has no such markings); the length of the rami in the A. zephyrus male antennae are diagnostically half the length of those of A. witti. Transversal hind wing fascia diagnostically angled in M 1 zone in both sexes in A. zephyrus.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	distribution	Distribution. China: Yunnan, Jiangxi, Sichuan, Xizang; northern Vietnam; north-eastern Myanmar (Fig. 58).	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
80E76B4D68375D7485A881D68FF1B41D.taxon	etymology	Etymology. In Greek mythology, Zephyrus or Zephyr (Greek: Ζέφυρος), is the god of the West Wind. He was reported to have had several wives in different stories, one of which was the nymph Chloris, upon whom he bestowed the domain of the flowers. The Romans later raised her status to goddess and gave her the name of Flora - the Latin name for flowers.	en	Zolotuhin, Vadim V., Pugaev, Sergey N., Du, Tran Thieu (2020): A review of Apha floralis species group (Lepidoptera: Eupterotidae). Acta Biologica Sibirica 6: 611-635, DOI: http://dx.doi.org/10.3897/abs.6.e59529, URL: http://dx.doi.org/10.3897/abs.6.e59529
