taxonID	type	description	language	source
15EEDF9997655977963EC1389D208105.taxon	etymology	Etymology. The species is dedicated to the memory of the collector of the type series Gaden S. Robinson (1949 - 2009), former researcher and curator of Lepidoptera at the then British Museum (Natural History). The specific epithet is a noun in the genitive case.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
15EEDF9997655977963EC1389D208105.taxon	diagnosis	Diagnosis. The new species looks superficially similar to species of the ophiusine genus Artena Walker, 1858, especially members of Artena rubida - group, namely A. reggiantii Zilli & Lourens, 2018, A. velutina (L. B. Prout, 1919) and A. rubida (Walker, [1863]), but is easily recognisable by the non-sexually dimorphic, slender rod-like third joints of labial palpi, rough instead of plain scaling of head and thorax, uniformly dark-coloured forewing without pale antemarginal field, larger, more centrally- and transversely positioned pale band of the hindwing and conspicuous scent tufts on the upperside of male metatibiae. Also somewhat resembling in habitus members of the poaphiline genus Achaea Huebner, 1823 and the yet incertae sedis genus Leistera Swinhoe, 1909, the latter showing another unusual combination of characters (type species, Catephia pulchristrigata Bethune-Baker, 1906, examined), e. g. no androteca as in Catephiini (Berio 1992) but unlike these with spines on male meso- (weakly) and metatibiae, uncus with dorsal peak and valva structured as in several members of the Ophiusini + Poaphilini clade, including also single tubular dorsal corema from its base. Nonetheless, no Achaea species with forewing completely deprived of pattern are known to date, and the arabesqued ornamentation of forewing of species of Leistera will prevent any confusion.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
15EEDF9997655977963EC1389D208105.taxon	description	Description. Male (Fig. 1). Habitus. Wingspan 74.5 - 78 mm (mean = 76 mm, n = 4). Ground colour of head, body trunk and forewing uniformly dark blackish brown, with some scattered sprinkles of silvery scales. Forewing broad subtriangular with no evident pattern, apex smoothly rounded, termen overall slightly convex, feebly undulated at veins, anal margin straight, fringe concolorous with wing disc; hindwing broad and rounded, termen as in forewing but slightly produced at vein 1 A + 2 A and with feeble white terminal line, colour dark chocolate brown, except for broad transverse white discal band, this widest at middle and with margins blurred, and with very feeble light-blue hue (visible if swinging the wing), ending before anal margin. Metatibial scent pencil blunt-ended and reaching in length almost the whole basal tarsomere, its basal third chocolate brown-coloured, mesially and distally brightly honey-coloured. Underside almost unicolorous dark blackish brown. Male genitalia (Fig. 5). Tegumen high, very tapered superiorly, with small thorn-like projections before apex; vinculum semicircular; valva oblong-obovate, with hairy distal membranous section, sacculus small, lozenge-shaped, with small apical outwardly-produced horn-like process, ventral sclerotised part centrally positioned, in shape of wide-based sub-rectangular transverse plate ending distally into two processes, inferior one (interpreted here as a clasper proper) thumb-like, perpendicular to longitudinal axis of valva, and superior one (interpreted as of costal derivation) similarly shaped but longer and much stouter, subparallel to costal margin, with reclinate tip, inflatable saccate corema on dorsal side with thick tuft of long hair-like scales; uncus long and reclinate, bulbous at base, its dorsal process subbasal and horn-like with forward projected tip, distally sinuous with middorsal hump and downcurved pointed tip; tuba analis wholly membranous. Juxta narrow, sub-rectangular, slightly sinuous at sides, with slightly divergent projections at corners, the inferior pair articulated with valvae. Phallus bent at basal third, vesica small, with paired opposite long thin diverticula and central, slightly bilobed bulge protruding from central corpus. Female (Fig. 2). Habitus. Wingspan 71.5 - 77.5 mm (x = 75.6 mm, N = 5). As in male, except for legs (described under generic diagnosis), and with white band of hindwing wider at middle of wing disc. Sternum A 7 (lodix) posteriorly bilobate (Fig. 6). Female genitalia (Fig. 6). Tergum A 8 wide and short, its ventro-lateral belts facing the sternum about half as high as the dorsal part, long and with rounded ends, separated only by narrow sternal section, apophyses anteriores thin; basal sterigmatal plate broad and rugulose, trapezoid, its ventral longitudinal outgrowth narrow, sub-rectangular, posteriorly bifid with conspicuous, pointed asymmetrical processes, and with anterior excision extending posteriorly as a narrow furrow for about one third of its whole length, ostium bursae opening at bottom of short infundibular depression in correspondence of posterior end of midventral furrow, sternum A 8 with narrow elongated sclerotisation proceeding from post-sterigmatal area. Ductus bursae very short, strongly sclerotised, joining with corpus bursae at junction between broad, dome-shaped cervix, which thus extends posteriorly to it, and large ovoid fundus bursae (note that the several irregular dark blotches visible in fig. 6 appear in completely other positions in another preparation, thus they seem tissue remains not pertaining to the bursa, though they appear tightly linked to it and not separable in the well cleaned preparations); appendix bursae conical. Intersegmental membrane A 8 - A 9 short. Papillae anales elongated sub-rectangular, apophyses posteriores approximately 1.5 times as long as anteriores.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
15EEDF9997655977963EC1389D208105.taxon	distribution	Distribution. Currently known only from the island of Espiritu Santo in the Vanuatu archipelago.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
7BF5F85461F2507F95175E720F7A12AD.taxon	etymology	Etymology. The generic epithet derives from the combination of " Tabwe- " (from Mt Tabwemasana in Espiritu Santo Island) and " - cala " (from the Greek " κᾰλή ", meaning " nice "); it is feminine in gender.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
7BF5F85461F2507F95175E720F7A12AD.taxon	diagnosis	Diagnosis. The absence of sexual dimorphism in the labial palpi, the presence of two anterior spines on the male profemur, the absence of spines on male tibiae, the presence of an androteca on male mesotibia, the strongly modified male metatibia bearing a dorsal hair pencil, the tuba analis without scaphium and with a deep, hood-like scaphial pocket, and the sterigma projected posteriorly into a free end, represent altogether a combination of characters not seen, at the author's best knowledge, in other genera of the Erebidae.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
7BF5F85461F2507F95175E720F7A12AD.taxon	description	Description. Male (Fig. 1). Head. Large, feebly sunk into thorax, frons slightly bulged, unscaled at middle (scales evidently brushed off by labial palps after emergence as in numerous other Erebinae), vertex roughly scaled, eye large, globular, antenna filiform, scape stout, barrel-like, covered by scales projected into small dorsal hood, pedicel small globular, flagellum consisting of short cylindrical, ventrally very shortly ciliate antennomeres, pilifer conspicuous, with long bristles, haustellum well developed, with tiny flat apical sensilla, labial palpus slender, upcurved, first joint roughly scaled, second joint compactly scaled, third joint long and thin, rod-like, two thirds as long as second, with no apical club. Thorax. Stout, both patagium and tegula broad and long, roughly scaled, meso-metanotum with hair-like scales, without crests; pectus conspicuously hairy. Wings broad; forewing vein R 1 from middle of cell, areole present, splitting between Rs 2 and Rs 3 close to termen in apical area of wing, but topology of Rs branches from cell very variable (see dedicated paragraph below), M visible in outer half of cell, other veins as in other Erebinae, with CuA 2 from before middle of cell; hindwing veins as in other Erebinae. Legs slender (Fig. 3); foreleg: profemur elongated, slightly wider at base and gradually tapered distally, with paired preapical spines, protibia not spined, barely shorter than femur, and feebly incrassate at middle, with epiphysis half length of tibia, protarsus conspicuously spined ventrally on tarsomeres I-IV, spines arranged into 3 rows, terminal spines of each tarsomere longest, spines replaced by bristles on tarsomere V, pretarsus with semi-circular arolium, onychia sharply bent, with midventral tooth, and pulvilli short; midleg: longest of the three legs, mesofemur long and slender, of almost uniform width, mesotibia greatly developed, longer and wider than mesofemur, its edges wrapping internally so as to originate a groove (androteca, cf. Berio 1955, 1959, 1965) enclosing a long hair-pencil kept in place by flat scales, mesotibial spurs one terminal pair, mesotarsus slender, spines and bristles on I-V as in protarsus, pretarsus as in protarsus; hindleg: metafemur slender and of almost uniform width, 7 / 10 as long as mesofemur, metatibia not spined and greatly modified, very short, about 2 / 3 length of metafemur, and swollen distally and dorsally so as to show a teardrop outline with dorso-apical bulge (respecting tibio-tarsal axis) for insertion of conspicuous pencil of long, hair-like androconial scales, this leaning middorsally along basal tarsomere, metatibial spurs two pairs, long and flimsy, first pair from middle of metatibia, second pair from well before apex, mesotarsus with basal tarsomere greatly developed, thick and longer than tarsomeres II-V altogether, widest at base and gradually tapered apically, clothed externally with thin long setae and showing midventrally in its distal two thirds comb of long bristles, tarsomeres II-V and pretarsus as in fore- and midleg. Abdomen. Sternum A 2 broad, subrectangular, with short, broad-based anterior apodemes, other segments of pregenital abdomen with subrectangular urotergites and urosternites, much wider than high, tergum A 8 trapezoid, sternum A 8 shallowly cleft on its posterior margin. Male genitalia (Fig. 5). Tegumen and vinculum robust, the latter shorter than tegumen and without inwardly produced saccus, valva oblong, with strongly sclerotised proximo-ventral portion, deriving from fusion between costal process and clasper, narrowly joined to sacculus, this produced into short saccular process, and dorso-distal membranous portion, which is broadly but shallowly corematous dorsally at base, where a conspicuous scent tuft is inserted. Uncus with dorsal process far from base. Tuba analis of complex configuration, dorsally membranous, showing distinct hood-like scaphial pocket (best seen in side view) above anus to accommodate tip of uncus. Juxta long, symmetrical, of the ' X' - type, articulated at very base of valvae. Phallus arched and compressed, with recurved foot-shaped coecum, showing small sclerotised platelet at base of vesica in distal excision of ventral edge of its shaft, vesica multi-lobed. Female (Fig. 2). Head. As in male, no sexual dimorphism detectable, including labial palpus, except for base of flagellum, slightly thinner, and periscapular scales, shorter and not forming hood. Thorax. Pro-metathorax and wings as in male, legs without the sexual secondary modifications seen in male (namely, profemural spines, mesotibial androteca, shortened and swollen metatibia bearing dorsal scent pencil and weak spurs, strongly developed basal segment of metatarsus with setae and comb), mesotibia irregularly spined externally, metatibia not spined, basal segment of metatarsus as long as tarsomeres II + III, with three rows of spines ventrally (Fig. 4); other features as in male. Abdomen. As in male but tergum A 7 broad and high, approximately square-shaped, and sternum A 7 greatly reduced to narrow and long subrectangular lodix plate (Fig. 6) overlapping ostium bursae. Female genitalia (Fig. 6). Complete sterigma present and of complex configuration, corresponding to broadly dilated midventral section of intersegmental membrane A 7 - A 8 produced midventrally into stout, strongly sclerotised longitudinal outgrowth that terminates posteriorly into free end overlapping midventral sclerotisation which extends between from post-sterigmatal area to sternum A 8; ostium bursae opening at approximately one third from anterior margin of sterigma. Bursa copulatrix with broad cervix and appendix anteriorly oriented originating from left of this. Ovipositor short, papillae anales soft, elongated, densely setose. Gonapophyses rod-like.	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
7BF5F85461F2507F95175E720F7A12AD.taxon	distribution	Distribution. The genus is currently known only from a restricted area of the Pacific (Vanuatu: Espiritu Santo).	en	Zilli, Alberto (2021): Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic position in the ' Ophiusini - Poaphilini ' clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193-211, DOI: http://dx.doi.org/10.3897/nl.44.70359, URL: http://dx.doi.org/10.3897/nl.44.70359
