taxonID	type	description	language	source
7247F8607839543890D1AA25132F08D1.taxon	description	Figs 3, 4, 5, 6, 7	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
7247F8607839543890D1AA25132F08D1.taxon	materials_examined	Other material examined. AUSTRALIA - Western Australia • 1 ♀; Carnarvon, 99 Gascoyne Road; 24 ° 53 ′ S 113 ° 39 ′ E; 23 July 2002; residents leg.; by hand; WAM T 46798 • 1 ♂; Cloudbreak Mining Lease, Fortescue Metals Group (site 25); 22 ° 20.1 ′ S 119 ° 24.23 ′ E; 6 Sept. 2006; S. Thompson leg.; WAM T 84005 • 1 ♀; Jimblebar minesite, 35 km E of Newman; 23 ° 22.5 ′ S 120 ° 12.58 ′ E; 6 Feb. 2009; P. Bolton and C. Weston leg.; active search; WAM T 95397 • 1 ♀; Murray Hills, Mulga Downs Station, Ecologia project 1142; 22 ° 07.67 ′ S 118 ° 30.92 ′ E; 19 Apr. 2009; N. Dight and L. Quinn leg.; dry pitfall trap; WAM T 97637 • 1 ♀; Davidson Creek, ca. 75 km E of Newman, vert site 6; 23 ° 25.73 ′ S 120 ° 26.8 ′ E; 9 Apr. 2010; J. Clark leg.; dry pitfall; WAM T 102165 • 1 ♀; South Parmelia, 52 km NW of Newman; 23 ° 5.13 ′ S 119 ° 19.08 ′ E; 16 Apr. 2011; R. Teale and M. Greenham leg.; dug from burrow; WAM T 113591 • 1 ♂; Southern Flank, 72 km NW of Newman; 23 ° 0.17 ′ S 119 ° 8.37 ′ E; 14 Apr. 2011; R. Teale and M. Greenham leg.; dug from burrow; WAM T 113596 • 1 ♂; same locality; 23 ° 0.18 ′ S 119 ° 8.35 ′ E, 14 Apr. 2011; R. Teale and M. Greenham leg.; dug from burrow; WAM T 113598 • 1 ♀; 113.8 km NW of Newman; 22 ° 39.39 ′ S 118 ° 55.09 ′ E; 26 May 2011; M. Greenham and R. Teale leg.; dug from burrow; WAM T 113626 • 1 ♀; 119.1 km NW of Newman; 22 ° 38.02 ′ S 118 ° 52.19 ′ E; 30 May 2011; M. Greenham and R. Teale leg.; dug from burrow; WAM T 113660 • 1 ♀; 117.6 km NW of Newman; 22 ° 37.66 ′ S 118 ° 53.76 ′ E; 31 May 2011; M. Greenham and R. Teale leg.; dug from burrow; WAM T 113665 • 1 ♀; Mudlark, 107 km W of Newman; 23 ° 5.63 ′ S 118 ° 43.17 ′ E; 30 June 2011; C. Cole and N. Watson leg.; dug from burrow; WAM T 116746 • 1 ♀; Mudlark, 111 km WNW of Newman; 23 ° 5.2 ′ S 118 ° 41.18 ′ E; 30 June 2011; M. Greenham and J. Cairnes leg.; dug from burrow; WAM T 116751 • 1 ♀; same data as for preceding; 23 ° 5.22 ′ S 118 ° 41.17 ′ E; WAM T 116755 • 1 ♀; Mudlark, 113 km W of Newman; 23 ° 2.28 ′ S 118 ° 40.97 ′ E; 1 July 2011; M. Greenham and J. Cairnes leg.; dug from burrow; WAM T 116776 • 1 ♀; Mudlark, 102 km W of Newman; 23 ° 5.4 ′ S 118 ° 48.67 ′ E; 3 July 2011; C. Cole and N. Watson leg.; dug from burrow; WAM T 116839 • 1 ♀; Mudlark, 94 km W. of Newman; 23 ° 4.78 ′ S 118 ° 51.48 ′ E; 6 July 2011; M. Greenham and J. Cairnes leg.; dug from burrow; WAM T 116866 • 1 ♀; same locality; 23 ° 4.77 ′ S 118 ° 51.47 ′ E; 26 July 2011; C. Cole and N. Watson leg.; dug from burrow; WAM T 116868 • 1 ♀; 84.2 km NW of Newman; 22 ° 40.5 ′ S 119 ° 20.95 ′ E; 26 July 2011; D. Kamien, M. Greenham and Z. Hamilton leg.; dug from burrow; WAM T 116873 • 1 ♀; same data as for preceding; 22 ° 40.48 ′ S 119 ° 20.9 ′ E; WAM T 116874 • 1 ♀; same data as for preceding; 89.3 km NW of Newman; 22 ° 38.45 ′ S 119 ° 19.23 ′ E; WAM T 116875 • 1 ♀; same data as for preceding; 89.3 km NW. of Newman; 22 ° 38.45 ′ S 119 ° 19.22 ′ E; WAM T 116881 • 1 ♀; Mulga Downs Station, Cowra, site 994 - 13; 22 ° 13.63 ′ S 119 ° 0.82 ′ E; 16 Apr. 2012; WAM T 118328 • 1 ♀; 84.8 km NW of Newman; 22 ° 40.1 ′ S 119 ° 22.48 ′ E; 27 July 2011; D. Kamien, M. Greenham and Z. Hamilton leg.; dug from burrow; WAM T 119975 • 1 ♀; same data as for preceding; 99.1 km NW of Newman; 22 ° 34.3 ′ S 119 ° 17.15 ′ E; WAM T 119979 • 1 ♀; same data as for preceding; 22 ° 34.3 ′ S 119 ° 17.17 ′ E; WAM T 119980 • 1 ♀; 105.3 km NW of Newman; 22 ° 30.72 ′ S 119 ° 15.55 ′ E; 28 July 2011; D. Kamien, M. Greenham and Z. Hamilton leg.; dug from burrow; WAM T 119984 • 1 ♀; same data as for preceding; 22 ° 30.72 ′ S 119 ° 15.53 ′ E; WAM T 119993 • 1 ♀; 81.2 km NW of Newman; 22 ° 42.02 ′ S 119 ° 22.68 ′ E; 29 July 2011; D. Kamien, M. Greenham and Z. Hamilton leg.; dug from burrow; WAM T 119995 • 1 ♀; 85.2 km NW. of Newman; 22 ° 39.2 ′ S 119 ° 24.82 ′ E; 31 July 2011; D. Kamien, M. Greenham and Z. Hamilton leg.; dug from burrow; WAM T 120018 • 1 ♀; same data as for preceding; 22 ° 39.2 ′ S 119 ° 24.83 ′ E; WAM T 120019 • 1 ♂; same data as for preceding; 18.9 km NE of Tom Price; 22 ° 39.2 ′ S 119 ° 24.82 ′ E; WAM T 120081 • 1 ♀; Koodaideri Corridor West, 93.7 km NE of Tom Price; 22 ° 19.7 ′ S 118 ° 36.61 ′ E; 20 Feb. 2012; C. Cole leg.; burrow search; WAM T 122209 • 1 ♀; same data as for preceding; 89.4 km NE of Tom Price; 22 ° 15.92 ′ S 118 ° 31.3 ′ E; WAM T 122217 • 1 ♀; same data as for preceding; 71.7 km NE of Tom Price; 22 ° 8.12 ′ S 118 ° 8.17 ′ E; WAM T 122224 • 1 ♀; same data as for preceding; 70.1 km NE of Tom Price; 22 ° 9.31 ′ S 118 ° 8.07 ′ E; WAM T 122226 • 1 ♀; same data as for preceding; 22 ° 9.37 ′ S 118 ° 8.07 ′ E; leaf litter rake; WAM T 122235 • 1 ♀; same data as for preceding; 82.1 km NE of Tom Price; 22 ° 13.95 ′ S 118 ° 24.88 ′ E; burrow search; WAM T 122252 • 1 ♀; same data as for preceding; 77.3 km NE of Tom Price; 22 ° 1.68 ′ S 118 ° 0.22 ′ E; WAM T 122254 • 1 ♀; 111.6 km NW of Newman; 22 ° 53.52 ′ S 118 ° 45.89 ′ E; 29 Mar. 2012; C. Cole and N. Watson leg.; dug from burrow; WAM T 122822 • 1 ♀; 115.4 km NW of Newman; 22 ° 54.52 ′ S 118 ° 43.05 ′ E; 31 Mar. 2012; C. Cole and N. Watson leg.; dug from burrow; WAM T 122843 • 1 ♀; 118.6 km NW of Newman; 22 ° 52.85 ′ S 118 ° 41.22 ′ E; 1 Apr. 2012; N. Watson and P. Brooshooft leg.; dug from burrow; WAM T 122865 • 1 ♀; 124 km NW of Newman; 22 ° 51.93 ′ S 118 ° 38.47 ′ E; 1 Apr. 2012; N. Watson leg.; dug from burrow; WAM T 122872 • 1 ♂; 63.5 km ESE of Paraburdoo, site 1000 - tur 01; 23 ° 17.31 ′ S 118 ° 17.1 ′ E; 27 Apr. 2012; E. S. Volschenk leg.; wet pitfall; WAM T 125176 • 1 ♀; Koodaideri Western Corridor, 217.5 km NW of Newman; 22 ° 7.71 ′ S 118 ° 5.57 ′ E; 28 Mar. 2012; G. Humphreys and M. Greenham leg.; dug from burrow; WAM T 125307 • 1 ♀; same data as for preceding; 194.8 km NW of Newman; 22 ° 7.71 ′ S 118 ° 7.71 ′ E; WAM T 125308 • 1 ♀; Koodaideri Western Corridor, 214 km NW of Newman; 22 ° 8.14 ′ S 118 ° 6.46 ′ E; 29 Mar. 2012; G. Humphreys and J. King leg.; dug from burrow; WAM T 125316 • 1 ♀; 118.2 km NW of Newman; 22 ° 36.32 ′ S 118 ° 55.15 ′ E; 19 Nov. 2011; M. Greenham and Z. Hamilton leg.; WAM T 126257 • 1 ♀; 118.3 km NW of Newman; 22 ° 36.67 ′ S 118 ° 54.43 ′ E; 18 Nov. 2011; M. Greenham and Z. Hamilton leg.; WAM T 126260 • 1 ♀; same data as for preceding; WAM T 126264 • 1 ♀; 114 km NW of Newman; 22 ° 36 ′ 54 ″ S 118 ° 57 ′ 18 ″ E; 21 Nov. 2011; M. Greenham and Z. Hamilton leg.; WAM T 126272 • 1 ♀; same data as for preceding; 22 ° 36.87 ′ S 118 ° 57.3 ′ E; WAM T 126276 • 1 ♀; Karijini National Park, ca. 20 km SW of Hancock Gorge; 22 ° 29.03 ′ S 118 ° 8.85 ′ E; 15 Mar. 2015; C. Stevenson, M. S. Harvey and M. Hillyer leg.; WAM T 135548 • 1 ♀; Karijini National Park, ca. 25 km SSW. of Dales Gorge; 22 ° 39.48 ′ S 118 ° 26.05 ′ E; 17 Mar. 2015; M. S. Harvey et al. leg.; WAM T 135563 • 1 ♀; Karijini National Park, ca. 6 km NW of Mt Bruce; 22 ° 34.12 ′ S 118 ° 5.98 ′ E; 15 Mar. 2015; J. Huey et al. leg.; WAM T 135591.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
7247F8607839543890D1AA25132F08D1.taxon	diagnosis	Diagnosis. Males of Missulena davidi sp. nov. share the red colouration of chelicerae and pars cephalica with M. langlandsi Harms and Harvey, 2013, M. occataria Walckenaer, 1805, M. insignis O. Pickard-Cambridge, 1877, M. iugum sp. nov. and M. manningensis sp. nov. that are morphologically most similar. They differ from M. langlandsi by having strong, conical spines of the rastellum (thin and not conical in the former) and a longer carapace (> 3.00 mm; M. langlandsi up to 2.8 mm). They differ from M. occataria and M. insignis by having spines on patellae III and IV only and not on all four legs (on patellae I and II 1 spine, respectively). Missulena davidi sp. nov. males have more cuspules on maxillae and labium than those of M. manningensis sp. nov. (M. manningensis sp. nov.: 5 at labium, 30 at maxillae; M. davidi sp. nov.: 15 - 10 at labium, 35 - 100 at maxillae). Missulena davidi sp. nov. males differ from M. iugum sp. nov. by the ridge present in the cheliceral groove. Females of Missulena davidi sp. nov. have uniformly red chelicerae that they share with M. insignis; however, the fourth leg of M. davidi sp. nov. is the longest of all legs, whilst in M. insignis the longest leg is the first. Additionally, there are no cuspules recorded on the labium or the maxillae in M. insignis females.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
7247F8607839543890D1AA25132F08D1.taxon	description	Description. MALE (based on holotype; WAM T 119725). Total length 9.8. Colour: pars cephalica and chelicerae reddish-orange (Fig. 3 C); a slim, black ring surrounding the PME (Fig. 4 E); pars thoracica brown with a light, metallic blue sheen (Fig. 3 C); abdomen greyish with a strong, metallic blue sheen on the dorsal side, ventrally more brownish with a faint hint of purple (Fig. 3 D, E); sternum orange, slightly fading into olive with 8 sigilla in similar colour (Fig. 4 D); labium and maxillae orange with a dark olive spot on the base of labium (Fig. 3 F); legs olive-yellowish fading into light brown ventrally, dorsally brown (Fig. 3 A, B); spinnerets beige-coloured (Fig. 3 E). Carapace: 3.86 long and 4.67 wide; clypeus 0.31; pars cephalica covers 2.45 of its length, is highly elevated and slightly granulated with very few setae (Fig. 3 G); pars thoracica also granulated with bands of faint, radial fissures and with two notches close to the abdomen (Fig. 3 C). Eyes: OQ 4 times wider than long; outer width of each eye pair AME 0.66, ALE 2.44, PME 1.48 and PLE 2.21; diameter of AME 0.19, ALE 0.22, PME 0.15, PLE 0.19; anterior eyes very slightly recurved; posterior eyes strongly recurved (Fig. 4 E). Chelicerae: 2.04 long and 1.47 wide on the base; edges rounded and recurved with the widest point being 1.55 close to the chelicerae base (Fig. 3 C); small, faint files along the outer margin of each chelicera; evenly spread setae along the inner margin and the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 7 (left 8) strong, conical spines (Fig. 4 F); over 25 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth and a general cheliceral teeth area in between those 2 clear rows (Fig. 4 A, J); prolateral row with approx. 9 teeth; retrolateral row with 4 teeth; intermediate area with 10 small teeth. Maxillae: 2.08 long and 1.44 wide; at least 80 - 100 weakly developed cuspules along entire anterior margin (Fig. 3 F). Labium: 0.96 long and 0.86 wide on the base; conical; at least 40 weakly developed cuspules anteriorly (Fig. 3 F); labiosternal junction visible (Fig. 4 D). Sternum: 2.6 long and 2.38 wide; ovoid (Fig. 4 D); setae of various length somewhat densely but disordered along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair smallest and hardly visible, second pair also very small and circular, third pair significantly larger than second (roughly 5 times bigger) in the shape of an elongated oval, and posterior pair biggest (roughly 1.5 times the size of the third pair) in the shape of a drop, all sigilla slightly depressed. Abdomen: 3.88 long and 3.4 wide; shape of a rounded trapezoid (Fig. 3 D); 4 spinnerets, PLS 1.12 long, 0.48 wide; PMS 0.43 long, 0.23 wide. Pedipalp: length of trochanter 1.49, femur 3.68, patella 1.68, tibia 4.01, tarsus 0.72; all segments with setae, tibia ventrally covered with comparably long setae (Fig. 4 I); tibia rather thin and slightly recurved, 1.00 wide on the widest point from dorsal and prolateral view (Fig. 4 G-I); bulb roughly pyriform (Fig. 4 G-I), two strongly sclerotized sections connected by a velar median structure (" haematodocha "); embolus short with an intumescence in proximal region; tip of embolus triangular with a small lamella and a tooth best visible from prolateral view (paratype, Fig. 3 H, I). Legs: brown setae of various sizes on all sides of the legs and bent strongly towards the exterior with the exceptions of some long, dorsal setae on tibia I and IV; ventral preening comb on tarsi and metatarsi III and IV. Leg spination: leg I: tibia rv 0, v 5, pl 0, d 0; metatarsus rv 3, v 8, pl 2, d 0; tarsus rv 3, v 3, pl 4, d 0; leg II: tibia rv 0, v 8, pl 0, d 0; metatarsus rv 0, v 11, pl 0, d 0; tarsus rv 3, v 3, pl 2, d 0; leg III: tibia rv 4, v 8, pl 0, d 11; metatarsus rv 4, v 11, pl 0, d 11; tarsus rv 5, v 9, pl 4, d 5; leg IV: tibia rv 0, v 5, pl 0, d 0; metatarsus rv 0, v 16, pl 0, d 2; tarsus rv 4, v 10, pl 3, d 3; patella I with one spine prolateral close to the tibia and patella II with one spine ventrally also close to the tibia; patella III with ca. 23 spines prolateral to dorsal (Fig. 4 B), 1 spine retrolateral; patella IV with one spine dorsal close to the tibia and approx. 12 small spines (Fig. 4 C) and ca. 18 prolateral, also very small. Leg measurements: Leg I: femur 3.72, patella 1.29, tibia 2.72, metatarsus 2.37, tarsus 1.37, total 11.47. Leg II: femur 3.31, patella 1.36, tibia 2.43, metatarsus 2.26, tarsus 1.38, total 10.74. Leg III: femur 2.72, patella 1.26, tibia 1.93, metatarsus 1.92, tarsus 1.35, total 9.18. Leg IV: femur 3.45, patella 1.47, tibia 2.76, metatarsus 2.41, tarsus 1.51, total 11.6. Formula: 4> 1> 2> 3. FEMALE (based on allotype; WAM T 107393). Total length 23.9. Colour: Carapace brown (Fig. 5 A); chelicerae reddish-orangish with a darker spot (dark reddish-brown) on each chelicerae base (Fig. 5 A); eye region light reddish colour similar to chelicerae (Fig. 5 A); abdomen greyish-brown with a faint, dorsal, metallic blue sheen (Fig. 5 B); sternum light brown fading into a reddish-brown towards labium (Fig. 5 C), sigilla darker brown (Fig. 5 C); labium und maxillae reddish-brown (Fig. 5 E); legs brown (Fig. 5 F, G); spinnerets lighter brown (Fig. 5 B). Carapace: 7.31 long and 9.88 wide; clypeus 0.74; pars cephalica covers 4.72 of its length, is highly elevated and smooth (Fig. 5 D) with some setae going along the margin of the chelicerae as well as vertically in a line from the AME to fovea plus some random setae (Fig. 5 A); pars thoracica smooth surface with bands of faint, radial fissures (Fig. 5 A). Eyes: OQ 4.9 times wider than long; width of each eye pair AME 0.91, APE 6.01, PME 3.59 and PLE 5.54; diameter of AME 0.33, ALE 0.36, PME 0.25, PLE 0.3; anterior eyes in one straight line; posterior eyes clearly recurved (Fig. 5 H). Chelicerae: 5.65 long and 4.15 wide on the base; edges rounded and recurved with the widest point being 4.73 close to the chelicerae base (Fig. 5 A); long setae along the inner margin increasing in amount towards rastellum; short and fewer setae along the outer margin and no setae in centre (Fig. 5 A); rastellum present with 10 - 14 conical spines on each chelicera and long, densely disordered setae (Fig. 5 I); approx. 25 setae cover anterior base of fang; inner margin of cheliceral furrow with two main rows of cheliceral teeth and a small cheliceral teeth area in between (Fig. 6 F); prolateral row with 11 teeth, the first 3 teeth next to the fang's base are partially grown together; retrolateral row with 10 teeth; intermediate area with approx. 32 small teeth. Maxillae: 4.48 long and 3.67 wide; at least 150 - 170 strongly developed cuspules along entire anterior margin (Fig. 5 E). Labium: 4 long and 2.27 wide on the base; conical; at least 60 cuspules (Fig. 5 E); anterior pair of sigilla reach labiosternal junction; labiosternal junction clearly developed (Fig. 5 C). Sternum: 6.32 long and 5.7 wide; oval (Fig. 5 C); setae of various length somewhat densely but disordered along the margin and a similar amount of setae of various sizes spread evenly over the sternum; 4 pairs of sigilla, anterior pair small and hardly visible, second pair (anterior-posterior) smallest and divided in circles, third pair larger than second and roughly in the shape of an elongated oval, and posterior pair biggest (roughly 4 times the size of the third pair); all sigilla depressed, two anterior pairs just slightly, two posterior pairs strongly. Abdomen: 10.87 long and 9.12 wide; surface covered with horizontal wrinkles and dense setae (Fig. 5 B); PLS 2.84 long and 1.5 wide; PMS 1.43 long and 0.6 wide. Pedipalp: Length of trochanter 1.5, femur 4.62, patella 1.79, tibia 3.23, tarsus 2.96; approx. 17 spines spread prolateral, retrolateral and ventral on tarsus. Genitalia: one pair of simple and rounded spermathecae, sperm ducts relatively short (Fig. 6 A-E). Legs: densely covered in brown setae of various sizes on all sides of the legs and bent towards the exterior with the exceptions of some long, dorsal setae on tibia, metatarsus and tarsus. Leg spination: leg I: tibia rv 0, v 0, pl 0, d 0; metatarsus rv 1, v 2, pl 0, d 0; tarsus rv 6, v 13, pl 6, d 0; leg II: tibia rv 0, v 0, pl 0, d 0; metatarsus rv 1, v 3, pl 0, d 0; tarsus rv 6, v 11, pl 3, d 0; leg III: tibia rv 1, v 0, pl 2, d 8; metatarsus rv 0, v 1, pl 0, d 18; tarsus rv 7, v 11, pl 7, d 6; leg IV: tibia rv 0, v 0, pl 1, d 5; metatarsus rv 0, v 3, pl 4, d 2; tarsus rv 1, v 15, pl 9, d 3; patellae I and II aspinose; patella III with ca. 28 spines prolateral to dorsal (Fig. 5 F); patella IV with ca. 18 spines prolateral to dorsal (Fig. 5 G). Leg measurement: leg I: femur 5.2, patella 2.16, tibia 3.4, metatarsus 2.5, tarsus 1.93, total 15.19. leg II: femur 5.32, patella 2.54, tibia 3.02, metatarsus 2.86, tarsus 2.04, total 15.78. leg III: femur 5.03, patella 2.64, tibia 2.39, metatarsus 3.08, tarsus 2.15, total 15.29. leg IV: femur 5.6, patella 2.61, tibia 3.66, metatarsus 3.31, tarsus 2.24, total 17.42. Formula: 4> 2> 3> 1.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
7247F8607839543890D1AA25132F08D1.taxon	etymology	Etymology. The specific epithet is a patronym in honour of the senior author's husband, David A. Greenberg.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
7247F8607839543890D1AA25132F08D1.taxon	distribution	Distribution. Pilbara region of Western Australia, excluding the northern Pilbara subregion, extending into the Little Sandy Desert region. The known linear range of this species is 295 km (Fig. 7).	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
32CE03EE0BAB5C5091ACC132BD79FB5F.taxon	description	Figs 8, 9, 10	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
32CE03EE0BAB5C5091ACC132BD79FB5F.taxon	diagnosis	Diagnosis. Males of Missulena iugum sp. nov. share the red chelicerae and pars cephalica with M. davidi sp. nov., M. manningensis sp. nov., M. langlandsi, M. occataria and M. insignis that are morphologically most similar. They differ from M. langlandsi by a longer carapace (> 3.00 mm; M. langlandsi up to 2.8 mm) and a rastellum with strong, conical spines (simple in M. langlandsi). Rastellum and cuspules on labium and maxillae stronger than in M. insignis (rastellum: 8 - 9 spines; M. insignis 2 - 5; cuspules: M. insignis none). Pars cephalica lower than in M. occatoria (up to 1.69; M. occataria approx. 3.0) and carapace shorter (3.87 long, 4.98 wide; M. occataria approx. 5.0 long, 7.0 wide). Differs from M. davidi sp. nov. and M. manningensis sp. nov. by the presence of a ridge in the cheliceral groove (Fig. 9 G, I).	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
32CE03EE0BAB5C5091ACC132BD79FB5F.taxon	description	Description. MALE (based on holotype; WAM T 123110). Total length 9.89. Colour: pars cephalica and chelicerae orange (Fig. 8 C); a slim, black ring surrounding the PME (Fig. 8 G); pars thoracica brown with a light, metallic blue sheen (Fig. 8 C); abdomen greyish with a very strong, metallic blue and green sheen on the dorsal side (Fig. 8 D), ventrally more brownish with a faint hint of purple (Fig. 8 E); sternum orange, slightly fading into olive with 8 sigilla in different shades of orange (Fig. 9 C); labium and maxillae orange with a slightly darker reddish-orange spot on the base of labium (Fig. 8 F); legs olive fading into light brown ventrally, dorsally brown (Fig. 8 A-B); spinnerets beige-coloured (Fig. 8 E). Carapace: 3.87 long, 4.98 wide and 1.67 high; clypeus 0.34; pars cephalica covers 2.34 of its length, is highly elevated and faintly granulated with very few setae (Fig. 9 H); pars thoracica also granulated with bands of faint, radial fissures and with two notches close to the abdomen (Fig. 8 C). Eyes: OQ 3.7 times wider than long; outer width of each eye pair AME 0.53, ALE 2.54, PME 1.66 and PLE 2.2; diameter of AME 0.2, ALE 0.21, PME 0.11, PLE 0.19; anterior eyes slightly recurved; posterior eyes strongly recurved (Fig. 8 G). Chelicerae: 2.28 long and 1.55 wide on the base; edges rounded and curved with the widest point being 1.57 very close to the chelicerae base (Fig. 8 C); few setae along the inner margin and slightly more evenly spread setae along the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 8 strong, conical spines (Fig. 8 H); approx. 20 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth with each a ridge along the row and a general cheliceral teeth area in between those 2 clear rows (Fig. 9 G, I); prolateral row with approx. 10 teeth; retrolateral row with 3 teeth (paratype has 4 teeth); intermediate area with 8 small teeth. Maxillae: 2.24 long and 1.52 wide; at least 85 - 100 weakly developed cuspules along entire anterior margin (Fig. 8 F). Labium: 0.94 long and 0.91 wide on the base; conical; at least 35 weakly developed cuspules anteriorly (Fig. 8 F); labiosternal junction visible (Fig. 9 C). Sternum: 2.84 long and 2.77 wide; ovoid (Fig. 9 C); setae of various length somewhat densely but disordered along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair small and hardly visible, second pair smallest (roughly half the size of the first pair) and circular, third pair larger than first and second pair (roughly 2 times bigger than first pair) in the shape of a drop, and posterior pair biggest (roughly 2.5 times the size of the third pair) in the shape of an elongated drop, all sigilla slightly depressed. Abdomen: 3.7 long and 3.41 wide; shape of a rounded trapezoid (Fig. 8 D); 4 spinnerets, PLS 0.88 long, 0.55 wide; PMS 0.45 long, 0.19 wide (Fig. 8 E). Pedipalp: length of trochanter 1.37, femur 4.17, patella 1.88, tibia 4.22, tarsus 0.75; all segments with setae, tibia ventrally densely covered with comparably long setae (Fig. 9 D-F); tibia rather thin and slightly recurved from lateral view, 1.00 wide on the widest point from prolateral and 1.08 from dorsal view (Fig. 9 D-F); bulb roughly pyriform, two strongly sclerotized sections connected by a velar median structure (" haematodocha "); embolus rather short with an intumescence in proximal region; tip of embolus triangular with a small lamella and a tooth best visible from prolateral view (paratype, Fig. 9 J, K). Legs: brown setae of various sizes on all sides of the legs and bent towards the exterior; some comparably long setae dorsally on most segments double the length of the other setae; ventral preening comb on tarsi and metatarsi III and IV. Leg spination: leg I: tibia rl 1, v 13, pl 0, d 0; metatarsus rl 2, v 10, pl 1, d 0; tarsus rl 2, v 7, pl 5, d 0; leg II: tibia rl 1, v 11, pl 1, d 0; metatarsus rl 2, v 11, pl 0, d 0; tarsus rl 5, v 9, pl 1, d 0; leg III: tibia rl 6, v 5, pl 5, d 6; metatarsus rl 5, v 7, pl 5, d 9; tarsus rl 10, v 9, pl 2, d 6; leg IV: tibia rl 1, v 10, pl 1, d 1; metatarsus rl 1, v 9, pl 3, d 2; tarsus rl 14, v 13, pl 5, d 5; patella I with 9 spines spread out prolaterally and three spine ventrally in a vertical row; patella II two spines ventrally in a vertical row; patella III with 27 spines spread out dorsally and prolaterally, 4 spines retrolaterally (Fig. 9 A) and 3 spines ventrally in a vertical row; patella IV with approx. 7 spines on the dorsal side (Fig. 9 B), approx. 6 spines on the prolateral side, all very small, and 3 spines ventrally in a vertical row. Leg measurement: Leg I: femur 4.2, patella 1.63, tibia 3.06, metatarsus 2.76, tarsus 1.58, total 13.23. Leg II: femur 3.47, patella 1.53, tibia 2.66, metatarsus 2.24, tarsus 1.48, total 11.38. Leg III: femur 3.07, patella 1.35, tibia 1.88, metatarsus 2.37, tarsus 1.52, total 10.19. Leg IV: femur 3.91, patella 1.59, tibia 2.99, metatarsus 2.78, tarsus 1.64, total 12.91. Formula: 1> 4> 2> 3.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
32CE03EE0BAB5C5091ACC132BD79FB5F.taxon	etymology	Etymology. The specific epithet is a Latin noun (Missulena iugum = ridge) in apposition, referring to the strongly developed ridges along the cheliceral groove of the males.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
32CE03EE0BAB5C5091ACC132BD79FB5F.taxon	distribution	Distribution. Known only from the Mt Ida region approximately 16 km east of Ularring in the Goldfields region of Western Australia (Fig. 10). The habitat of the holotype comprises Acacia shrubland.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
6294FFA5B7095301A471739A872F3A1B.taxon	description	Figs 10, 11, 12	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
6294FFA5B7095301A471739A872F3A1B.taxon	diagnosis	Diagnosis. Males share with M. davidi sp. nov., M. iugum sp. nov., M. langlandsi, M. occatoria and M. insignis, the closest morphological matches, the red colouration of the chelicerae and pars cephalica. They differ from M. langlandsi by a longer carapace (> 3.00 mm; M. langlandsi up to 2.8 mm) and the presence of strong, conical spines of the rastellum (simple in M. langlandsi). They differ from M. occataria and M. insignis by the lack of spines ventrally on patellae III and IV (at the most 1 thickened seta). Pars cephalica lower than in M. occataria (up to 1.96; M. occataria approx. 3.0) and carapace shorter (3.6 long, 4.61 wide; M. occataria approx. 5.0 long, 7.0 wide). More cuspules on the labium and maxillae than in M. insignis but less than in M. davidi sp. nov. (M. insignis: none; M. manningensis sp. nov.: 5 at labium, 30 at maxillae; M. davidi sp. nov.: 15 - 10 at labium, 35 - 100 at maxillae). Lacks a ridged cheliceral groove which is present in M. iugum sp. nov.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
6294FFA5B7095301A471739A872F3A1B.taxon	description	Description. MALE (based on holotype; WAMT 92071). Total length 8.95. Colour: pars cephalica and chelicerae orange (Fig. 11 C); a slim, black ring surrounding the PME (Fig. 11 H); pars thoracica brown with a light, purplish sheen (Fig. 11 C); abdomen greyish with a light, metallic blue sheen on the dorsal side (Fig. 11 D), ventrally more brownish with a faint hint of purple (Fig. 11 E); sternum orange, slightly fading into olive with 8 sigilla in different shades of orange (Fig. 11 G); labium and maxillae orange with a dark olive spot on the base of labium (Fig. 11 F); legs olive fading into light brown ventrally, dorsally brown (Fig. 11 A, B); spinnerets beige (Fig. 11 E). Carapace: 3.6 long, 4.61 wide and 1.96 high; clypeus 0.31; pars cephalica covers 2.25 of its length, is highly elevated and slightly granulated with very few setae (Fig. 12 D); pars thoracica also granulated with bands of faint, radial fissures and with two deeply expressed notches close to the abdomen (Fig. 11 C). Eyes: OQ 3.4 times wider than long; outer width of each eye pair AME 0.48, ALE 2.26, PME 1.44 and PLE 2.14; diameter of AME 0.19, ALE 0.19, PME 0.13, PLE 0.17; anterior eyes in a straight line; posterior eyes strongly recurved (Fig. 11 H). Chelicerae: 2.17 long and 1.41 wide on the base; edges rounded and recurved with the widest point being 1.51 close to the chelicerae base (Fig. 11 C); few setae along the inner margin and slightly more evenly spread setae along the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 5 (left 8) strong, conical spines (Fig. 11 I); over 20 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth and a general cheliceral teeth area in between those 2 clear rows (Fig. 12 G); prolateral row with approx. 11 teeth; retrolateral row with 4 teeth; intermediate area with 5 small teeth. Maxillae: 2.09 long and 1.5 wide; at least 30 extremely weakly developed cuspules along anterior margin (Fig. 11 F). Labium: 0.79 long and 0.86 wide on the base; conical; at least 5 extremely weak developed cuspules anteriorly (Fig. 11 F); labiosternal junction visible (Fig. 11 G). Sternum: 2.62 long and 2.29 wide; ovoid (Fig. 11 G); setae of various length somewhat densely but irregular along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair very small and hardly visible, second pair also hardly visible, smallest of all pairs and divided into two circles, third pair significantly larger than second (roughly 4 times bigger) in the shape of an elongated oval, and posterior pair biggest (roughly 2 times the size of the third pair) and drop-shaped, all sigilla slightly depressed. Abdomen: 3.12 long and 2.74 wide; shape of a rounded trapezoid (but collapsed through preservation; Fig. 11 D); 4 spinnerets, PLS 0.43 long (part of it broken off), 0.35 wide; PMS 0.37 long, 0.19 wide (Fig. 11 E). Pedipalp: length of trochanter 1.64, femur 4.46, patella 1.82, tibia 3.75, tarsus 0.64; all segments with setae, tibia ventrally covered with comparably long and dense setae (Fig. 12 A-C); tibia rather thin and slightly recurved, 1.00 wide on the widest point from dorsal / ventral and prolateral / retrolateral view (Fig. 12 A-C); bulb roughly pyriform, two strongly sclerotized sections connected by a velar median structure (" haematodocha "); embolus rather short and bend with an intumescence in proximal region; tip of embolus triangular with a small lamella, best visible retrolateral (Fig. 12 I), and a tooth, best visible prolateral (Fig. 12 H). Legs: brown setae of various sizes on all sides of the legs and bent towards the exterior with the exception of the femur setae on ventral position which are mostly vertically; ventral preening comb on tarsi and metatarsi III and IV. Leg spination: leg I: tibia rl 0, v 17, pl 0, d 0; metatarsus rl 0, v 9, pl 0, d 0; tarsus rl 0, v 154 - 8 - 3, pl 0, d 0; leg II: tibia rl 1, v 12, pl 0, d 0; metatarsus rl 0, v 8, pl 0, d 0; tarsus rl 0, v 10, pl 2, d 0; leg III: tibia rl 3, v 13, pl 0, d 7; metatarsus rl 3, v 6, pl 3, d 9; tarsus rl 3, v 11, pl 4, d 5; leg IV: tibia rl 0, v 12, pl 0, d 0; metatarsus rl 0, v 12, pl 5, d 1; tarsus rl 8, v 13, pl 7, d 5; patella I with 8 spines spread out prolaterally and one spine ventrally close to tibia; patella II with one spine prolaterally and one ventrally both close to tibia; patella III with 26 spines spread out dorsally and prolaterally and one spine retrolaterally (Fig. 12 E); patella IV with approx. 14 spines on the dorsal side (Fig. 12 F) and approx. 15 spines on the prolateral side, all very small. Leg measurement: Leg I: femur 4.0, patella 1.37, tibia 2.97, metatarsus 2.69, tarsus 1.56, total 12.59. Leg II: femur 3.4, patella 1.12, tibia 2.5, metatarsus 2.38, tarsus 1.47, total 10.87. Leg III: femur 2.81, patella 1.05, tibia 1.94, metatarsus 2.26, tarsus 1.54, total 9.6. Leg IV: femur 3.28, patella 1.13, tibia 2.83, metatarsus 2.41, tarsus 1.59, total 11.24. Formula: 1> 4> 2> 3.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
6294FFA5B7095301A471739A872F3A1B.taxon	etymology	Etymology. The specific epithet refers to the type locality, Mt Manning, in the Goldfields region of Western Australia.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
6294FFA5B7095301A471739A872F3A1B.taxon	distribution	Distribution. Known only from the Mt Manning area approximately 47 km northwest of Boorabbin in the Goldfields region of Western Australia (Fig. 10). The habitat of the holotype comprises open tall eucalypt woodland with mixed shrubs.	en	Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W., Harms, Danilo (2021): Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome. Arthropod Systematics & amp; Phylogeny 79: 509-533, DOI: http://dx.doi.org/10.3897/asp.79.e62332, URL: http://dx.doi.org/10.3897/asp.79.e62332
