taxonID	type	description	language	source
B16C7280B117562A924B36F7203A17ED.taxon	description	Figs 2 A, 3 A, 4 A, 13 A, B, 14 C, 15 C, 16 C, 17 C, 18 C, 19 C, 20 A, B, 25, 27 C	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B16C7280B117562A924B36F7203A17ED.taxon	description	Description. Male. n = 71. Length 1.6 - 2.5 mm. Colouration (Fig. 4 A). Head dull, blackish brown; pronotum and postpronotum dark brown; postpronotal lobe brown with dark brown markings, light brown around anterior spiracle; prescutum, mesoscutum and pleura shiny, blackish brown; scutellum and mediotergite shiny, blackish brown; katepisternum dark brown with blackish brown markings, remaining pteropleuron mainly blackish brown with dispersed markings of brown to light brown; base of halter black, knob creamy yellow; legs greyish brown, apex of tarsi darker; abdomen blackish brown; terminalia concolourous with abdomen. Head. Eyes above antennae broadly joined, with small triangular frons visible above antennae; frons with three to five strong setae. Flagellomeres 1 - 3 subquadrate, with flagellomere 1 expanded, 3 x as wide as next segment, equal to lengths of 2 and 3 combined; flagellomeres 4 - 10 cylindrical, becoming progressively thinner and elongate. Vertex with black setae of uniform length, with longer, black orbital setae. Thorax. Mesoscutum with prominent antealar ridge, bearing thee pronounced setae. Scutum clothed dorsally in short, black setulae; notopleural, supra-alar and postsutural setae long, black. Pteropleuron bare. All legs with tarsi simple. Wing. Wing length: 1.8 - 2.5 mm. Dark, infuscate throughout, apex somewhat narrowed; C fringed in small setulae, with a few microtrichia scattered throughout; posterior wing margin with closely spaced fringe of microtrichia; Sc incomplete; R 1 and R 1 (+ R 2 + 3) with three weakenings or depigmented gaps, first slightly beyond R 2 + 3, second and third closely approximated, near C; microtrichia of R 1 (+ R 2 + 3) confined to base near humeral crossvein, remaining veins bare; R flexed into cell br; R 2 + 3 distinct, situated in basal third of R 1 (+ R 2 + 3); bend in R 4 + 5 gentle; R 4 + 5 and M 1 running parallel toward margin; M 1 straight; M 2 with gentle bend in apical third; M 4 with slight bend. Abdomen. Abdominal sternite 1 narrow, spectacle-shaped; sternite 2 reduced to slender median sclerite, a few setae restricted to laterad on posterior third and medially beneath sclerite; sternites 3 - 7 rectangular, lacking distinct sclerites, setae restricted to posterior two-thirds; sternite 8 strongly reduced, anterior margin well sclerotised, arched slightly into preceding segment, lacking setae. Terminalia (Figs 2 A, 3 A). Epandrium quadrate in ventral view, posterior margin rounded with large, medial indentation; long, extending well beyond gonostyli; without lobes or projections. Gonocoxites conical, one-third longer than wide, anterior margin rounded, expanded dorsally behind gonocoxal plate, nearly meeting medially, extended anteriorly toward sternite 8; posterior inner margin produced into rounded projection, outer margin without notch; inner margin densely setose. Gonostylus short, three-quarters length of cercus, strongly curved laterally throughout; widest at base, tapered toward bifurcated apex; outer margin bearing laterally directed setae. Parameres fused at gonocoxal apodeme, widest at point of fusion; extended beyond cerci; forming canal-like structure, flattened apically, tapered to off-centre sharp point (occasionally specimens with broken apex). Gonocoxal plate well sclerotised; tongue-like plate extended anteroventrally; hollow medially, pitcher plant-like; gonocoxal apodeme with secondary structure comprising three arms fused medially, running along interior of paramere canal, flaring into three or four filaments, projected ventrally at apex, with pair of lateral flanges near point of fusion. Cercus large, prominent; subquadrate; projected posteriorly; situated within epandrial indentation. Female. n = 15. Similar to male except as follows: Terminalia (Fig. 13 A, B). Hypogynial valve not projecting beyond tergite 9; posterior margin deeply emarginated in ventral view, nearly dividing sternite in half, forming two subtriangular lobes; lobes densely setose. Tergite 9 subtriangular in lateral view, 3 x as wide as tergite 8, lacking lateral projections; posterior margin heavily sclerotised at base of cerci. Sternite 9 (genital fork) slender, Y-shaped anteriorly and posteriorly; lateral arms extended slightly beyond hypogynial valve, divergent toward apex. Hypoproct sclerotised, narrow. Cercus rounded, projected posteroventrally; bearing numerous setae. Pupa. n = 8 (Figs 14 C, 15 C, 16 C). Length 3.0 - 4.0 mm. Colouration. Brown; with black spot above eye in developing adult. Head. Maxillary sheath short, posteromedially directed; gently tapered toward truncate apex; apices of palpi separated medially. Three short, slender setae above black spot over eye. Thorax. Width nearly subequal to abdomen at widest point. Foreleg sheath projecting straight, slightly longer than wing sheaths, reaching posterior margin of sternite 2; anterior half of midleg visible anterior to wing sheath, then hidden behind foreleg, slightly shorter than foreleg; hind leg concealed behind wing sheath, only apex visible between apex of foreleg and wing sheath, shorter than foreleg. Wing sheaths not reaching posterior margin of abdominal sternite 2; large tubercle at base bearing pair of short, slender setae. Respiratory organ short and squat, much shorter than maxillary sheath, broadest subapically; bulbous; spiracular openings encircling apex; stalk wide, emerging from small tubercle. Tubercle situated posterodorsally to respiratory organ, rounded, projected slightly laterally; apex nearly touching or touching respiratory organ. Tubercle situated posterolaterally to respiratory organ crenulate, projected slightly anteriorly. Ridge located anteroventrally to respiratory organ with single, thin midlateral seta; mesothorax with group of four short, slender dorsocentral setae near ridge; single seta on humeral lobe. Abdomen. Subcylindrical, strongly tapered at caudal segment. Spiracles weakly developed, not projected or distinctly visible. Tergites 1 - 8 rectangular, without ridges; bearing pair of slender lateral setae above lateral margins and pair of slender dorsolateral setae. Tergite 9 rounded, posterior with dorsolateral ridges bearing pair of lateral setae and hind margin emarginated; projection directed posteriorly in lateral view. Sternites 3 - 8 rectangular, without row of faint setulae along anterior margin; lateral margins crenulate, bearing a few thin, short setae. Sternites 3 and 4 bearing pair of small lateral adhesive structures, sternite 5 bearing pair of large lateral adhesive structures on lateral margin. Caudal sternite subquadrate, with pair of posteriorly projected medial lobes; posterior margin with pair of medial ridges, curved dorsally forming small, dorsally projected tubercle in lateral view; without distinct caudal hooks. Larva. n = 8 (Figs 17 C, 18 C, 19 C, 20 A, B). Length of final instar 6.2 - 6.7 mm. Colouration. Head capsule usually black or dark brown, sometimes black with light brown markings. Body mottled with various shades of grey and brown. Head capsule (Fig. 20 A, B). Two large, circular eye spots, elevated on tubercle; antenna on largest tubercle, with three finger-like processes; with five pairs of tubercles outside of ecdysial lines (not including antennal and ocular tubercle), two outermost tubercles bifurcate; single tubercle between ecdysial line, bi- or trifurcate; 15 pairs of unbranched setae; five pairs of sensory pits (13, 14, 18, 20, 21), sensory pit 13 above antennal tubercle. Thorax. Prothorax with pair of anterodorsolateral protuberances bare; anterolateral protuberances with one long and two short setae; spiracular protuberance bearing two protuberances, inner protuberance with pair of setae, outer with single seta; pair of midlateral setae below anterolateral protuberance; three closely approximated setae near base of prothoracic leg (Keilin's organ). Mesothorax and metathorax with pair of dorsolateral protuberances bearing pair of closely approximated setae, one thickened, one slender; mesothorax with additional seta beneath protuberance; lateral protuberance on both segments bearing four setae; one long seta slightly ventral to lateral protuberance; three mid-ventrolateral setae directed ventrally. Prothorax bearing proleg, posterior half with rectangular adhesive structure; meso- and metathoracic sternites with rectangular adhesive structures. Abdomen. Sternites 1 - 7 modified into circular, suction cup-like adhesive structures; sternite 8 with quadrate adhesive structure, extending over sternite 9; sternite 9 smooth, bearing anal proleg. Tergites 1 - 7 with single anterolateral protuberance on each side with single seta, and pair of posterodorsolateral protuberances, each bearing two closely approximated short, thin setae; lateral adhesive structure swelling bearing four setae, two lateral, two basal; additional single seta located anterior to lateral swelling. Segment 8 with dorsolateral protuberance on either side of posterior spiracular plate, each bearing pair of small setae; lateral protuberance with three setae; single short ventrolateral seta; ventral surface bearing pair of setae. Posterior spiracular plate with sclerite encircling procerci; procercus shorter than length of spiracular plate, bearing four setae, two thick, two slender; without cone-like protuberance on either side of procerci. Terminal segment with anterior dorsal protuberance bare; pair of posterior lateral protuberances with pair of setae; five lateral setae; two pairs of long setae on posterior margin, above pair of anal papillae; ventral sternite bearing single pair of setae.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B16C7280B117562A924B36F7203A17ED.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B16C7280B117562A924B36F7203A17ED.taxon	distribution	Distribution. Known from the south-central Andes of Chile (Fig. 24 A); this species includes the northern-most record for Niphta in South America.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B16C7280B117562A924B36F7203A17ED.taxon	etymology	Etymology. The species name is from the Latin acu (needle, pin) in allusion to the needle-like tip of the paramere.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	description	Figs 6 A, 8 A, 9 A, 10 A, 11 A, 12 A, 24 B	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	description	Description. Male. n = 3. Length 1.7 - 1.9 mm. Colouration (Figs 9 A, 10 A). Head dull, dark brown; pronotum and postpronotum dark brown; postpronotal lobe and lateral margins of prescutum light brown; scutum shiny with three distinct dark brown stripes, pleura light brown; postscutum brown; scutellum shiny, light brown; mediotergite shiny, anterior half light brown, posterior half brown; katepisternum mainly dark brown, light brown near coxa 1; paratergite brown; remaining pteropleuron mainly brown with dispersed light brown and black markings; base of halter pale brown, knob pale yellow; legs pale brown, apex of tarsi darker; abdomen brown; terminalia pale brown. Head. Eyes above antennae broadly joined, with small triangular frons visible above antennae; frons with two strong setae. Flagellomeres 1 - 3 subquadrate, 1 expanded, twice as wide as next segment, shorter in length than 2 and 3 combined; flagellomeres 4 - 10 cylindrical, becoming progressively thinner and elongate. Vertex with black setae of uniform length, with longer, black orbital setae. Thorax. Mesoscutum with prominent antealar ridge, bearing three setae, middle seta most pronounced. Scutum clothed dorsally in short, black setulae; notopleural, supra-alar and postsutural setae long, black. Pteropleuron bare. All legs with tarsi simple. Wing. Wing length: 2.1 - 2.4 mm. Infuscate throughout, apex somewhat narrowed; C fringed in small setulae, with widely spaced microtrichia; posterior wing margin with closely spaced fringe of microtrichia; Sc incomplete; R 1 and R 1 (+ R 2 + 3) with three weakenings or depigmented gaps, first slightly beyond R 2 + 3, second and third closely approximated, near C; microtrichia of R 1 (+ R 2 + 3) confined to base near humeral crossvein, remaining veins bare; R flexed into cell br; R 2 + 3 distinct, situated in basal third of R 1 (+ R 2 + 3); bend in R 4 + 5 strong; R 4 + 5 and M 1 running parallel toward margin; M 1 straight; M 2 with gentle bend in apical third; M 4 with slight bend. Abdomen. Abdominal sternite 1 narrow, spectacle-shaped; sternite 2 reduced to slender median sclerite, a few setae restricted to posterior third; sternites 3 - 7 rectangular, lightly sclerotised, setae restricted to posterior half; sternite 8 strongly reduced, anterior margin well sclerotised, arched slightly into preceding segment, a few setae restricted to laterad. Terminalia (Figs 6 A, 8 A). Epandrium quadrate in ventral view, posterior margin rounded, with medial cleft; long, extended beyond gonostyli; without lobes or projections. Gonocoxites oblong, longer than wide; anterior margin rounded, somewhat expanded dorsally behind gonocoxal plate, not closely approximated; with two spine-like projections; anterior projection wide, bifurcate; posterior projection long, slender, slightly sinuous, tapered to single apex, nearly twice as long as anterior projection; inner margin with numerous long, thin setae. Gonostylus cheliform, dorsoventrally flattened anteriorly, swollen posteriorly; anterior apex with a few setae; posterior apex bifurcate, setose. Parameres medially fused, attached basally to arms of gonocoxal plate; divided distally into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally toward gonocoxal plate, strongly curved anteriorly, sickle-shaped, surface textured with tiny bumps, except for smooth apex; ventral arm extends posteroventrally presumably to aid in copulation; when retracted, rests ventrally between dorsal arm of gonocoxal plate and dorsal to anterior gonocoxal projection. Gonocoxal plate broad, well sclerotised; anterior margin subquadrate, basal margin cleft; pair of dorsal arms connected to parameres; medial aedeagal guide projected ventrally between posterior margins of gonocoxites, well sclerotised, comprising two parts: anterior Y-shaped structure with five finger-like projections protruded from posterior margin and dorsal triangular plate. Cercus ovoid, only slightly visible in lateral view; projected anteroventrally; situated within epandrial indentation. Female. n = 1. Similar to male except as follows: Abdomen. Tergite 9 noticeably more sclerotised than preceding tergites; sternite 8 well sclerotised, with distinct blunt projection at base of hypogynial valve. Terminalia (Figs 11 A, 12 A). Hypogynial valve not projected beyond tergite 9; posterior margin deeply cleft in ventral view, forming two triangular lobes; lobes densely setose, with both stout, thickened setae and thinner, long setae with slight apical bend; hypogynial protuberance between valves. Tergite 9 subquadrate in lateral view, 1.5 x as wide as tergite 8, lacking lateral projections. Sternite 9 (genital fork) slender, Y-shaped at both ends; lateral arms forming complex of highly modified structures: medial heavily sclerotised circular opening, dorsal to posterior cleft of hypogynial valve, with pair of lateral sclerotised triangular expansions; triangular expansions expanded dorsally into pair of circular plates, those further expanded anteriorly into pair of heavily sclerotised plates, strongly recurved posteroventrally toward circular opening, remaining dorsal to genital fork; dorsal surface of recurved plates with tiny grooves and indentations. Hypoproct lightly sclerotised, narrow. Cercus quadrate, projected posteroventrally; bearing numerous setae. Spermathecae not observed; two spermathecal ducts visible in centre of lateral arm circle.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	distribution	Distribution. Known only from the type locality in the Chilean Coastal Range (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	etymology	Etymology. Niphta bifurcata is named in reference to the posterior apex of the gonostylus and the anterior projection of the gonocoxite, both of which are bifurcate.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F969A23FE28D5676B17F9C9C60CF5032.taxon	description	Figs 6 C, 8 C, 9 F, 10 F, 11 C, 12 C, 24 B, 27 B, D	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F969A23FE28D5676B17F9C9C60CF5032.taxon	description	Description. The descriptions of N. bispinosa differ from that of N. bifurcata in the following regards: Male. n = 1. Length 1.9 - 2.4 mm. Colouration (Figs 9 F, 10 F). Pronotum and postpronotum brown; postpronotal lobe and lateral margins of prescutum pale yellow; remaining scutum shiny, brown, pleura yellow; postscutum with medial brown spot above scutoscutellar suture, encircled by yellowish brown margin; scutellum shiny, yellow; mediotergite shiny, dark brown except anterior margin yellow; katepisternum mainly brown, lighter near anterior spiracle; remaining pteropleuron mainly pale yellow with dispersed brown markings; base of halter yellow, distal half of stalk and knob grey; legs pale yellow, apex of tarsi darker; terminalia light brown. Head. Frons with three strong setae. Flagellomere 1 expanded, 1.5 x as wide as next segment, equal in length to 2 and 3 combined. Wing. Wing length: 2.0 - 2.4 mm. Lightly infuscate throughout; bend in R 4 + 5 gentle. Terminalia (Figs 6 C, 8 C). Posterior margin of epandrium rounded, with narrow medial cleft. Gonocoxites oblong, wider than long; anterior margin rounded, somewhat expanded dorsally behind gonocoxal plate, not closely approximated; with two spine-like projections; anterior projection bifurcate, projected posteriorly, 3 x longer than posterior projection; posterior projection short, tooth-like; margin around gonostylus setose. Gonostylus cheliform, dorsoventrally flattened, margins concave creating bowl-like appearance; anterior apex with a few indistinct setae, posterior apex with strong setae. Parameres medially fused, attached basally to arms of gonocoxal plate; divided distally into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally toward gonocoxal plate, strongly curved anteriorly, blade-like, ventral margin serrate; ventral arm extended posteroventrally presumably to aid in copulation; when retracted, rests ventrally between dorsal arm of gonocoxal plate and inner margin of gonocoxite, apex reaching base of anterior gonocoxal projection. Gonocoxal plate broad, well sclerotised; anterior margin subquadrate, basal margin cleft; pair of dorsal arms connected to parameres; medial aedeagal guide projected ventrally between gonostyli, well sclerotised, comprising two parts: anterior Y-shaped structure and posterior triangular plate. Cercus prominent, ovoid. Female. n = 2. Similar to male except as follows: Terminalia (Figs 11 C, 12 C). Posterior margin of hypogynial valve deeply cleft in ventral view, forming two quadrate lobes. Tergite 9 twice as wide as tergite 8. Sternite 9 (genital fork) slender, Y-shaped at both ends; lateral arms forming complex of highly modified structures: medial heavily sclerotised circular opening, dorsal to posterior opening of hypogynial valve, with pair of lateral sclerotised triangular expansions. Spermathecae not observed.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F969A23FE28D5676B17F9C9C60CF5032.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F969A23FE28D5676B17F9C9C60CF5032.taxon	distribution	Distribution. Known only from the type locality in central Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F969A23FE28D5676B17F9C9C60CF5032.taxon	etymology	Etymology. Niphta bispinosa is named in reference to the two projections from the gonocoxite.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
D9103CED43025A22A61F64491363B418.taxon	description	Description. The descriptions of N. brunnea differ from that of N. bifurcata in the following regards: Male. n = 5. Length 1.6 - 1.9 mm. Colouration (Figs 9 B, 10 B). Pronotum and postpronotum brown; remaining scutum shiny, brown, pleura light brown; scutellum shiny, brown; mediotergite shiny, anterior half light brown, posterior half dark brown; katepisternum mainly brown, lighter near anterior spiracle; remaining pteropleuron mainly brown with dispersed markings of dark / light brown; base of halter creamy, distal half of stalk and knob light brown; abdomen brown; terminalia light brown. Head. Flagellomere 1 expanded, 1.5 x as wide as next segment, shorter in length than 2 and 3 combined. Wing. Wing length: 1.9 - 2.2 mm. Terminalia (Figs 6 D, 8 D). Epandrium quadrate in ventral view, posterior margin rounded, with narrow medial cleft; long, extended beyond gonostyli; without lobes or projections. Gonocoxites oblong, longer than wide; anterior margin rounded, somewhat expanded dorsally, not closely approximated; with three spine-like projections; large anterior projection nearly equal in length to posterior projection, gradually tapered toward apex; smaller anterior projection positioned somewhat anterior to large tooth, may be small and inconspicuous in some specimens; posterior projection strongly tapered toward apex, some specimens with second basal tooth on outer margin of projection; margin around anterior projection with long, thin setae. Gonostylus cheliform, dorsoventrally flattened, margins concave creating bowl-like appearance; anterior apex with a few indistinct setae, posterior margin with strong setae. Parameres medially fused, attached basally to arms of gonocoxal plate; divided distally into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally toward gonocoxal plate, knife-shaped, ventral margin serrate; when retracted, resting ventrally between dorsal arm of gonocoxal plate and inner margin of gonocoxite, apex reaching base of anterior gonocoxal projection. Gonocoxal plate broad, well sclerotised; anterior margin triangular, basal margin cleft; pair of dorsal arms connect to parameres; medial aedeagal guide projected ventrally between gonostyli, well sclerotised, consisting of two parts, anterior structure with five projections and posterior rounded plate bearing minute setulae. Cercus ovoid, only slightly visible in lateral view; projected anteroventrally; situated within epandrial indentation. Female. Unknown. Pupa. n = 6 (Figs 14 A, 15 A, 16 A, 22). Length 2.7 - 2.9 mm. Colouration. Brown; sometimes with black spot above eyes in developing adult. Head. Maxillary sheath short, posteromedially directed, gently tapered toward truncate apex, apices of palpi separated medially; devoid of tubercles and setae. Thorax. 1.5 x wider than abdomen at widest point. Foreleg sheath projected straight and slightly beyond wing sheaths, reaching hind margin of sternite 2; anterior half of midleg visible anterior to wing sheath, then hidden behind foreleg, not projected beyond wing sheath; hindleg concealed beneath wing sheath, only small triangular apex visible between apex of foreleg and wing sheath, not extended beyond wing sheath. Wing sheath extended to posterior margin of abdominal sternite 2. Respiratory organ slightly shorter than maxillary sheath, broadest subapically; ovate, slightly arched medially, tapered toward apex; spiracular openings encircling apex; stalk thin, emerging from small tubercle. Tubercle situated posterodorsally to respiratory organ, rounded, projected laterally; apex nearly touching or touching respiratory organ. Thorax devoid of setae. Abdomen. Subcylindrical, evenly tapered toward caudal segment. Spiracles well developed, distinct on segments 3 - 7, projected anterodorsolaterally; all bearing minute spine-like setulae. Segment 8 with short lateral projection, less than half length of preceding spiracles, projected dorsolaterally. Tergites 1 - 8 quadrate, devoid of setae, with pair of dorsolateral ridges and faint medial transverse groove, groove not meeting dorsolateral ridges. Tergite 9 rounded, posterior margin emarginated, laterally compressed compared to preceding segments; small, rounded projection pointing posteriorly in lateral view. Sternites 3 - 8 rectangular, with row of faint setulae along anterior margin, not connecting to lateral margin; lateral margins crenulate, lacking setae. Sternites 3 and 4 bearing pair of small lateral adhesive structures, sternite 5 bearing pair of large lateral adhesive structures on lateral margin. Caudal sternite triangular, with medial sclerotised groove and pair of medial rounded, posteromedially projected lobes; posterior margin with medial longitudinal ridge; without distinct caudal hooks. Larva. n = 6 (Figs 17 A, 18 A, 19 A, 21 A, B, 23). Length of final instar 4.8 - 5.1 mm. Colouration. Head capsule pale brown, anterolateral margin of ecdysial line black. Body creamy brown. Head capsule (Fig. 28 A, B). Two large, circular eye spots, elevated on tubercle; antenna with three finger-like processes, elevated on largest tubercle; with five pairs of smaller tubercles outside of ecdysial lines (not including antennal and ocular tubercle); single tubercle between ecdysial lines, about same size as ocular tubercle; 15 pairs of unbranched setae; six sensory pits (13, 14, 18, 19, 20, 21), sensory pit 13 above antennal tubercle. Thorax. Prothorax with single pair of protuberances bearing single spiracle; spiracular protuberance bearing one pair of dorsal setae anterior to spiracle and single dorsolateral seta; three mid-lateral setae, two long, one short and fine; two closely approximated setae near base of prothoracic leg (Keilin's organ). Mesothorax and metathorax with pair of small dorsolateral protuberances and pair of large lateral protuberances; mesothoracic dorsolateral protuberance bearing single thickened seta, metathoracic lateral protuberance bearing pair of closely approximated setae, anterior seta thickened and longer than posterior seta; lateral protuberance on both segments bearing three setae, two short, one long; single long seta slightly ventral to lateral protuberance; one pair of mid-ventrolateral setae. Prothorax bearing proleg, posterior half with rectangular adhesive structure; meso- and metathoracic sternites with rectangular adhesive structure. Abdomen. Sternites 1 - 7 modified into circular, suction cup-like adhesive structure; sternite 8 with quadrate adhesive structure; sternite 9 smooth, bearing anal proleg. Segments 1 - 7 lacking distinct protuberances, at most, pair of dorsolateral swellings bearing single or paired short, thin setae; single seta situated anterolaterally; lateral adhesive structure swelling bearing numerous setae, two anterolateral, two midlateral, four basalateral. Segment 8 with small dorsolateral protuberance on either side of posterior spiracular plate, each bearing pair of small setae; lateral protuberance with single seta; single short ventrolateral seta; ventral sternite bearing pair of setae. Posterior spiracular plate with sclerite encircling procerci; procercus shorter than length of spiracular plate, bearing four setae, two thick, two slender; without cone-like protuberance on either side of procerci. Terminal segment with pair of protuberances, no setae; pair of dorsolateral setae; five lateral setae; two pairs of long setae on posterior margin, above pair of anal papillae; ventral sternite lacking setae.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
D9103CED43025A22A61F64491363B418.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
D9103CED43025A22A61F64491363B418.taxon	distribution	Distribution. Known only from two localities in the Andes of southern Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
D9103CED43025A22A61F64491363B418.taxon	etymology	Etymology. Niphta brunnea is from the Latin brunneus (brown) in allusion to its brown colouration, the darkest of the N. nudipennis group.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
3BEF2BECF04251A2802D7689F4AF6B53.taxon	description	Figs 6 B, 8 B, 9 C, H, 10 C, 11 B, 12 B, 24 B	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
3BEF2BECF04251A2802D7689F4AF6B53.taxon	description	Description. The descriptions of N. courtneyi differ from that of N. bifurcata in the following regards: Male. n = 1. Length 2.1 - 2.3 mm. Colouration (Figs 9 C, 10 C). Head dull, brown; pronotum and postpronotum brown; postpronotal lobe and lateral margins of prescutum yellow; scutum shiny with three distinct dark brown stripes, pleura yellow; postscutum yellow, two lateral brown spots above scutoscutellar suture; scutellum shiny, yellow; mediotergite shiny, anterior half yellow, posterior half brown; katepisternum mainly pale brown, yellow near fore coxa; remaining pteropleuron mainly yellow with dispersed brown markings; halter creamy yellow; legs pale yellow, apex of tarsi darker; abdomen brown; terminalia yellow. Head. Frons with three strong setae. Flagellomere 1 expanded, 1.5 x as wide as next segment, shorter in length than 2 and 3 combined. Wing. Wing length: 1.9 - 2.2 mm. Lightly infuscate throughout. Terminalia (Figs 6 B, 8 B). Epandrium quadrate in ventral view, posterior margin rounded, with narrow medial cleft; long, extended beyond gonostyli; without lobes or projections. Gonocoxites oblong, longer than wide; anterior margin rounded, somewhat expanded dorsally, not closely approximated; with three spine-like projections; two anterior projections, with inner projection shorter than outer; posterior projection long, slender, tapered to single apex, 3.5 x longer than shortest projection; margin around gonostylus with long, dense thin setae. Gonostylus cheliform, dorsoventrally flattened anteriorly, swollen posteriorly; anterior apex with a few setae; posterior apex bifurcate, setose. Parameres medially fused, attached basally to arms of gonocoxal plate; divided distally into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally toward gonocoxal plate, strongly curved anteriorly, sickle-shaped, surface textured with tiny bumps, except for smooth apex; ventral arm, when retracted, rests ventrally between dorsal arm of gonocoxal plate and inner margin of gonocoxite. Gonocoxal plate broad, well sclerotised; anterior margin subquadrate, basal margin cleft; pair of dorsal arms connected to parameres; medial aedeagal guide projected ventrally between posterior margins of gonocoxites, well sclerotised, consisting of two parts: anterior Y-shaped structure and posterior triangular plate. Cercus ovoid, only slightly visible in lateral view; projected anteroventrally; situated within epandrial indentation. Female. n = 2. Similar to male except as follows: Terminalia (Figs 11 B, 12 B). Tergite 9 subquadrate in lateral view, 2 x as wide as tergite 8, lacking lateral projections.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
3BEF2BECF04251A2802D7689F4AF6B53.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
3BEF2BECF04251A2802D7689F4AF6B53.taxon	distribution	Distribution. Known only from the type locality, the East side of Lago Llanquihue in Southern Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
3BEF2BECF04251A2802D7689F4AF6B53.taxon	etymology	Etymology. Niphta courtneyi is named in honour of its collector, Gregory W. Courtney (Iowa State University). Courtney collected three new species of Thaumaleidae from Chile (A. fredericki Pivar, N. courtneyi, and N. mapuche), as well as immature material, prompting us to further investigate the Chilean fauna.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
5A4DA9ABABB951669931265A83DAA531.taxon	description	Figs 5 A, 7 A, 9 D, 10 D, 24 B, 26 A	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
5A4DA9ABABB951669931265A83DAA531.taxon	description	Description. The description of N. daniellae differs from that of N. bifurcata in the following regards: Male. n = 2. Length 1.7 - 2.0 mm. Colouration (Figs 9 D, 10 D). Postpronotal lobe and lateral margins of prescutum yellow; scutum shiny with three distinct brown stripes, pleura yellow; postscutum yellow, two lateral brown spots above scutoscutellar suture; scutellum shiny, yellowish; mediotergite shiny, anterior half yellow, posterior half brown; katepisternum dark brown, except yellow at base of fore coxa; anepisternum and paratergite brown; remaining pteropleuron yellow; halter entirely creamy yellow; legs yellowish brown, tarsi dark brown; abdominal tergites brown, posterior margin pale brown, sternites mainly yellow with scattered brown markings; terminalia yellowish brown. Head. Frons with two to three strong setae. Flagellomere 1 expanded, 1.5 x as wide as next segment, subequal in length to 2 and 3 combined. Wing. Wing length: 2.2 - 2.3 mm. Lightly infuscate throughout; bend in R 4 + 5 gentle; M 4 with slight apical bend. Abdomen. Abdominal sternite 2 reduced to slender median sclerite, lacking setae; sternites 3 - 7 rectangular, setae restricted to posterior two-thirds; sternite 8 strongly reduced, lacking setae. Terminalia (Figs 5 A, 7 A). Epandrium quadrate in ventral view, posterior margin rounded, with narrow medial cleft; long, extended beyond gonostyli; without lobes or projections. Gonocoxites oblong, longer than wide, inner margin setose, expanded anterodorsally above gonocoxal plate, closely approximated but not fused; two posteromedially directed spine-like projections, one anterior to gonostylus, one posterior; anterior projection pointed, bare; posterior projection blunt, setose. Gonostylus longer than wide, distal half strongly tapered to pointed apex; dorsoventrally compressed, margins curved slightly ventrally, scoopula-like, heavily sclerotised; a few setae scattered throughout, apex bare. Parameres medially fused, attached basally to arms of gonocoxal plate; surface textured with tiny bumps, except for smooth apex; divided medially into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally, expanded medially, apical third rapidly tapered to pointed apex, sharply curved anterodorsally; ventral arm, when retracted, rests ventrally between lateral margin of gonocoxal plate and gonocoxite. Gonocoxal plate broad, well sclerotised; anteroventral margin subtriangular, basal margin cleft forming two ventrally directed projections; pair of dorsal arms connect to parameres; with median expansion projected ventrally between posterior margins of gonocoxites. Cercus ovoid, visible in lateral view; projected anteroventrally; situated within epandrial indentation. Female. Unknown.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
5A4DA9ABABB951669931265A83DAA531.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
5A4DA9ABABB951669931265A83DAA531.taxon	distribution	Distribution. Known from two localities in the Andes of south-central Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
5A4DA9ABABB951669931265A83DAA531.taxon	etymology	Etymology. This species is named in honour of RJP's wife, Danielle Lombardi, for her support during Pivar's graduate research and entomological endeavours, and for playing an important role in organising the Chilean expedition. Raised in northern Chile (Arica), Danielle's Spanish skills were critical for translating all communications with government and national park contacts, as well as translating our requests for collecting permits.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
623605C366905BF79A46F3E509F742F8.taxon	description	Figs 2 B, 3 B, 4 B, 24 A, 27 A	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
623605C366905BF79A46F3E509F742F8.taxon	description	Description. The description of N. downesi differs from that of N. acus in the following regards: Male. n = 3. Length 1.9 - 2.4 mm. Colouration (Fig. 4 B). Head dull, black; pronotum and postpronotum black; postpronotal lobe, prescutum and mesoscutum shiny, blackish brown, lateral face of postpronotal lobe brown, creamy around anterior spiracle; scutellum blackish brown dorsally, light brown ventrally; mediotergite shiny, blackish brown; pteropleuron mainly brown with dispersed markings of black and dark brown; base of halter blackish, knob creamy yellow; abdomen blackish brown. Head. Frons with two strong setae. Flagellomeres 1 - 3 subquadrate, expanded, 1.5 x as wide as next segment, as long as 2 and 3 combined. Thorax. Antealar ridge bearing single pronounced, medial seta flanked by two smaller setae. Wing. Wing length: 2.2 - 2.4 mm. C and posterior wing margin with fringe of microtrichia. Abdomen. Sternites 3 - 7 with setae restricted to posterior two-thirds and laterad; sternite 8 strongly arched into preceding segment, lacking setae. Terminalia (Figs 2 B, 3 B). Gonocoxite subquadrate, posterior inner margin produced into pointed projection, outer margin notched. Gonostylus short, slightly more than half-length of cercus, strongly arched outwards; apex pointed. Parameres distally fused, widest medially, as wide as gonocoxite; extended past posterior margin of epandrium; lateral margins curved ventrally, canal-like medially in ventral view, then tapered rapidly to filamentous, pointed apex projected posterodorsally, extended beyond epandrium; medial structure cradled within canal, protruded ventrally; in lateral view ending at halfway point of longest filament. Gonocoxal apodeme with secondary structure comprising single filament, running along interior of paramere canal, projected ventrally at apex. Cercus trapezoidal; projected posteroventrally; situated within epandrial indentation. Female. Unknown.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
623605C366905BF79A46F3E509F742F8.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
623605C366905BF79A46F3E509F742F8.taxon	distribution	Distribution. Known from the foothills of the southern Andes in Chile (Fig. 24 A).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
623605C366905BF79A46F3E509F742F8.taxon	etymology	Etymology. Niphta downesi is named in honour of veterinary and medical entomologist J. A. Downes, who collected the first specimen of this species in 1985.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
C274435D50125285AC6262C1F5109C80.taxon	description	Figs 5 B, 7 B, 9 E, 10 E, 24 B, 27 A	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
C274435D50125285AC6262C1F5109C80.taxon	description	Description. The description of N. eurydactyla differs from that of N. bifurcata in the following regards: Male. n = 17. Length 1.5 - 1.8 mm. Colouration (Figs 9 E, 10 E). Variable colouration, even among specimens from the same population; base colouration of thorax either yellow or orange as follows: postpronotal lobe and lateral margins of prescutum orangey / yellowish brown; scutum shiny with three distinct brown stripes, pleura yellow to yellowish brown; postscutum orangey / yellowish brown, two lateral brown spots above scutoscutellar suture; scutellum shiny, orangey / yellowish brown; mediotergite shiny, anterior half orangey / yellowish brown, posterior half brown; katepisternum mainly pale brown with scattered orange / yellow and black markings, or mostly yellowish brown with brown lateral markings above mid coxae; paratergite brown; remaining pteropleuron mainly orangey-brown to yellowish brown with dispersed markings of brown and black; base of halter creamy grey, knob pale yellow; legs pale brown, apex of tarsi darker; abdomen brown; terminalia pale brown. Thorax. Antealar ridge bearing three to four setae, middle seta most pronounced. Wing. Wing length: 1.8 - 2.2 mm. Abdomen. Abdominal sternites with setae restricted to posterior two-thirds. Terminalia (Figs 5 B, 7 B). Epandrium quadrate in ventral view, posterior margin rounded, with narrow medial cleft; long, extended beyond gonostyli; without lobes or projections. Gonocoxites oblong, longer than wide, large C-shaped scallop where gonostyli inserted; anterior margin rounded, somewhat expanded dorsally, not closely approximated; anterior inner margin with stout spine-like projection; margin around gonostylus setose. Gonostylus subquadrate; with finger-like projection emerging from posterolateral corner, gently tapered toward apex, bearing a few setae; inner anterior margin with small, rounded projection bearing a few fine setae. Parameres medially fused, attached basally to arms of gonocoxal plate; divided distally into dorsal parameral apodeme and ventral arm; ventral arm projected anteroventrally toward gonocoxal plate, strongly curved anteriorly, sickle-shaped, surface textured with tiny bumps, except for smooth apex; ventral arm, when retracted, rests ventrally between lateral margin of gonocoxal plate and gonostylus. Gonocoxal plate broad, well sclerotised; anterior margin rounded; pair of dorsal arms connect to parameres; median aedeagal guide trident-shaped, weakly sclerotised. Cercus prominent; ovoid; projected anteroventrally; situated within epandrial indentation. Female. Unknown.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
C274435D50125285AC6262C1F5109C80.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
C274435D50125285AC6262C1F5109C80.taxon	distribution	Distribution. Known from both the Chilean Coastal Range and Andes of southern Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
C274435D50125285AC6262C1F5109C80.taxon	etymology	Etymology. The specific name is from the Greek eury (broad, wide) and daktylos (finger), in allusion to the broad, finger-like projection on the gonostylus.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B94E2F8451705AEBAE9F01E1821A88BC.taxon	description	Figs 2 C, 3 C, 4 C, 13 C, D, 24 A, 27 F	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B94E2F8451705AEBAE9F01E1821A88BC.taxon	materials_examined	Additional material examined. Chile: Region X (Los Lagos): Camino de Penetracion @ Hwy. 7 sign, 16. xii. 2013, 42 ° 07 ' 57.5 " S 72 ° 27 ' 45.3 " W, seep, sweeping veg., G. R. Curler (1 ♂, 1 ♀ *); Camino de Penetracion @ km post 125.600, 16. xii. 2013, 42 ° 03 ' 33.3 " S 72 ° 27 ' 07.4 " W, rock seep, G. R. Curler (1 ♂); Casa Pangue, Llanquihue, 12.1926, R & E Shannon, USNMENT 01115811 (1 ♂, USNM); Ensenada, nr. Banos de Petrohue, 12. i. 1985, J. A. Downes (2 ♂, CNC; 1 ♂, USNM (USNMENT 01115812 )); Rte. 215, 12. xii. 2016, 40 ° 40 ' 32.4 " S 72 ° 17 ' 35.6 " W, elev. 252 m, trickle falls, J. K. Moulton & R. J. Pivar (1 ♂); Rte. U- 99, 10. xii. 2016, 41 ° 08 ' 09.6 " S 72 ° 35 ' 43.3 " W, elev. 81 m, roadside falls, J. K. Moulton & R. J. Pivar (15 ♂; 4 ♀ *); Rte. U- 99, 10. xii. 2016, 41 ° 08 ' 28.2 " S 72 ° 35 ' 16.8 " W, elev. 101 m, roadside seeps / creek, J. K. Moulton & R. J. Pivar (11 ♂, 1 ♀ *); Rte. V- 69, 12. xii. 2016, 41 ° 26 ' 37.7 " S 72 ° 17 ' 42.2 " W, elev. 34 m, cascading stream, J. K. Moulton & R. J. Pivar (1 ♂); Rte. V- 69, Puente El Salto, 12. xii. 2016, 41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W, elev. 37 m, splash zone above falls, J. K. Moulton & R. J. Pivar (2 ♂); Region XIV (Los Rios): Antilhue, Rte. T- 35, 9. xii. 2016, 39 ° 49 ' 09.8 " S 72 ° 56 ' 22.6 " W, elev. 40 m, roadside creek, J. K. Moulton & R. J. Pivar (3 ♂); Valdivia, Los Ulmos Rd., 15. i. 1985, J. A. Downes (5 ♂, CNC).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B94E2F8451705AEBAE9F01E1821A88BC.taxon	distribution	Distribution. Known from both the Andes and Chilean Coastal Range in southern Chile (Fig. 24 A).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	description	Figs 2 D, 3 D, 4 D, 13 E, F, 20 C, D, 24 A	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	description	Description. The descriptions of N. mapuche differ from that of N. acus in the following regards: Male. n = 10. Length 1.4 - 2.5 mm. Colouration (Fig. 4 D). Head dull, black; pronotum and postpronotum brown; prescutum and mesoscutum shiny, blackish brown, pleura brown; katepisternum brown with blackish brown margins; base of halter light brown turning black medially, knob creamy yellow; legs ranging from pale brown to blackish brown; abdomen blackish brown, posterior margins creamy; terminalia variable in colour from blackish brown to grey. Head. Flagellomeres 1 - 3 subquadrate, 1 expanded, 2 x as wide as next segment, as long as 2 and 3 combined. Abdomen. Abdominal sternite 2 with few setae restricted to laterad on posterior third; sternites 3 - 7 with setae restricted to lateral margins and middle third; sternite 8 with three or fewer setae medially and on lateral margins. Terminalia (Figs 2 D, 3 D). Gonocoxite subquadrate; posterior inner margin slightly produced into pointed projection, outer margin notched. Gonostylus short, less than half-length of cercus, strongly curved lateroventrally throughout; apex pointed; outer margin bearing laterally directed setae. Parameres distally fused, widest at point of fusion; not reaching posterior margin of epandrium; dividing into two pointed, filamentous projections medially; shorter ventral filament not extended beyond gonostyli, apex directed posteriorly; longer dorsal filament not extended to posterior margin of epandrium, at most slightly beyond base of cercus, apex projected slightly posterodorsally. Gonocoxal plate broad, well sclerotised, tongue-like plate extending anteroventrally; gonocoxal apodeme with secondary structure comprising single short, indistinct filament, running along interior of paramere, ending before apical margin of gonostylus. Cercus trapezoidal. Female. n = 10. Similar to male except as follows: Terminalia (Fig. 13 E, F). Hypogynial valve not projected beyond tergite 9; posterior margin deeply emarginated in ventral view, forming two triangular lobes; lateral margin sclerotised at base. Tergite 9 triangular in lateral view. Sternite 9 (genital fork) slender, Y-shaped; lateral arms not extended beyond hypogynial valve, fused distally forming rounded posterior margin; lightly sclerotised ventral plate at base of hypogynial valve. Two sclerotised, tube-like structures dorsal to genital fork, spermathecal pumps perhaps; spermathecal receptacles and ducts not observed. Pupa. n = 4 (not figured due to condition of specimens). Length 3.5 - 4.0 mm. Head. Setae not observed. Thorax. Hindleg concealed behind wing sheath, only apex visible between apex of foreleg and wing sheath, slightly shorter than foreleg, but longer than wing sheath. Wing sheaths with large tubercle at base, setae not visible. Abdomen. Setae not visible on tergites 1 - 8. Larva. n = 17. Length of final instar 6.3 - 6.7 mm. Colouration. Head capsule variable, ranging from light brown to black. Body mottled brown and grey, possibly pale brown to creamy; cream coloured ventrally. Head capsule (Fig. 20 C, D). Tubercles below and beside eye bifurcate; two tubercles between ecdysial line, upper tubercle bi- or trifurcate. Abdomen. Tergites 1 - 7 with lateral adhesive structure swelling bearing five setae, two lateral, three basal.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	materials_examined	Additional material examined. Known only from the type series.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	distribution	Distribution. Known from the southern Andes of Chile (Fig. 24 A).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	etymology	Etymology. This species is named after the Mapuche (mapu, land, che, people) indigenous peoples, who since ~ 500 B. C., have inhabited the regions of southern Chile, where N. mapuche is known.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
A1B7EFD258385CEBB4420F467047ADE0.taxon	description	Figs 1, 5 C, D, 7 C, D, 9 G, 10 G, 11 D, 12 D, 14 B, 15 B, 16 B, 17 B, 18 B, 19 B, 21 C, D, 24 B, 26 D, E, 27 A, E	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
A1B7EFD258385CEBB4420F467047ADE0.taxon	materials_examined	Additional material examined. Chile: Region X (Los Lagos): Chiloe, Cascada Tocoihue, 10. xii. 2016, 42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W, elev. 32 m, smaller falls, J. K. Moulton & R. J. Pivar (2 ♂, 4 ♀ *); Ensenada, nr. Banos de Petrohue, 12. i. 1985, J. A. Downes (13 ♂, 2 ♀, CNC); Isla Chiloe, Ancud, 12.1926, R. & E. Shannon, USNMENT 01115824 (1 ♂, USNM) [Note: There is an additional USNM specimen identified as N. nudipennis from this same collection event; however, abdomen is missing and species identification cannot be confirmed (USNMENT 01115825)]; Rte. V- 69, 12. xii. 2016, 41 ° 19 ' 51.5 " S 72 ° 24 ' 40.0 " W, elev. 129 m, roadside seep, J. K. Moulton & R. J. Pivar (2 ♂); Rte. V- 69, 12. xii. 2016, 41 ° 31 ' 48.8 " S 72 ° 17 ' 31.2 " W, elev. 39 m, trickling creek, J. K. Moulton & R. J. Pivar (5 ♂); same data as previous except, larvae / pupae on foliage in splash zone (27 larvae *, 4 pupae, 3 pupal exuviae); Region XIV (Los Rios): Rte. T- 85, 13. xii. 2016, 40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W, elev. 145 m, foliage around waterfalls, J. K. Moulton & R. J. Pivar (1 ♂).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
A1B7EFD258385CEBB4420F467047ADE0.taxon	distribution	Distribution. Known from both the Chilean Coastal Range and the Andes of southern Chile (Fig. 24 B).	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B3593280B79159BE9BD27F67C51A9C21.taxon	diagnosis	Diagnosis. Niphta is characterised as follows: presence of a distinct antealar ridge; proepisternal setae absent; microtrichia of R 1 (+ R 2 + 3) confined to base near humeral crossvein; R 2 + 3 crossvein situated closer to apex of R 1 (+ R 2 + 3) than to origin of R 4 + 5; R 1 and R 1 (+ R 2 + 3) with three weakenings or depigmented gaps; R 4 + 5 often with arch not strongly produced; basal spur of CuA absent; gonocoxites broad, not much longer than wide; hypandrium absent; gonocoxal plate extended posterodorsally forming a medial process; parameres fused medially, emerging from gonocoxal plate complex.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
B3593280B79159BE9BD27F67C51A9C21.taxon	distribution	Distribution. Chile and Australia.	en	Pivar, Robert J., Sinclair, Bradley J., Moulton, John K. (2021): Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype. ZooKeys 1063: 49-104, DOI: http://dx.doi.org/10.3897/zookeys.1063.71180, URL: http://dx.doi.org/10.3897/zookeys.1063.71180
