taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600671	http://doi.org/10.3897/zookeys.1063.71180.figure2	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600672	http://doi.org/10.3897/zookeys.1063.71180.figure3	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600673	http://doi.org/10.3897/zookeys.1063.71180.figure4	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600682	http://doi.org/10.3897/zookeys.1063.71180.figure13	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600683	http://doi.org/10.3897/zookeys.1063.71180.figure14	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600684	http://doi.org/10.3897/zookeys.1063.71180.figure15	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600685	http://doi.org/10.3897/zookeys.1063.71180.figure16	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600686	http://doi.org/10.3897/zookeys.1063.71180.figure17	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600687	http://doi.org/10.3897/zookeys.1063.71180.figure18	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600688	http://doi.org/10.3897/zookeys.1063.71180.figure19	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600689	http://doi.org/10.3897/zookeys.1063.71180.figure20	Figure 20. Niphta halteralis group larval head capsules with setae and sensory pits numbered A anterior, N. acus sp. nov. B lateral, N. acus sp. nov. C anterior, N. mapuche sp. nov. D lateral, N. mapuche sp. nov. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	Figure 20. Niphta halteralis group larval head capsules with setae and sensory pits numbered A anterior, N. acus sp. nov. B lateral, N. acus sp. nov. C anterior, N. mapuche sp. nov. D lateral, N. mapuche sp. nov. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600694	http://doi.org/10.3897/zookeys.1063.71180.figure25	Figure 25. Habitat and larvae of Niphta acus sp. nov. (36 ° 55 ' 02.7 " S 71 ° 25 ' 49.6 " W) A Moulton shown next to the falls for scale B close up of falls with box indicating where immatures were captured (Note: they were not found in high flow zones) C lateral view of larva, with adhesive structures visibly in contact with substrate D dorsal view of larva illustrating camouflage E larva and pupal exuviae on rock face.	Figure 25. Habitat and larvae of Niphta acus sp. nov. (36 ° 55 ' 02.7 " S 71 ° 25 ' 49.6 " W) A Moulton shown next to the falls for scale B close up of falls with box indicating where immatures were captured (Note: they were not found in high flow zones) C lateral view of larva, with adhesive structures visibly in contact with substrate D dorsal view of larva illustrating camouflage E larva and pupal exuviae on rock face.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B16C7280B117562A924B36F7203A17ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600675	http://doi.org/10.3897/zookeys.1063.71180.figure6	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600677	http://doi.org/10.3897/zookeys.1063.71180.figure8	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600680	http://doi.org/10.3897/zookeys.1063.71180.figure11	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600681	http://doi.org/10.3897/zookeys.1063.71180.figure12	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
9EF2ABFCED8F513197FBBAD6DCC06080.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600675	http://doi.org/10.3897/zookeys.1063.71180.figure6	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600677	http://doi.org/10.3897/zookeys.1063.71180.figure8	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600680	http://doi.org/10.3897/zookeys.1063.71180.figure11	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600681	http://doi.org/10.3897/zookeys.1063.71180.figure12	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F969A23FE28D5676B17F9C9C60CF5032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600675	http://doi.org/10.3897/zookeys.1063.71180.figure6	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600677	http://doi.org/10.3897/zookeys.1063.71180.figure8	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600683	http://doi.org/10.3897/zookeys.1063.71180.figure14	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600684	http://doi.org/10.3897/zookeys.1063.71180.figure15	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600685	http://doi.org/10.3897/zookeys.1063.71180.figure16	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600691	http://doi.org/10.3897/zookeys.1063.71180.figure22	Figure 22. Scanning electron micrographs of ventral view of pupae of Niphta brunnea sp. nov. A habitus (scale bar: 1.0 mm) B head and thorax (scale bar: 1.0 mm) C posterior segments (scale bar = 0.5 mm) D adhesive structures (scale bar: 0.5 mm). Abbreviations: S, sternite.	Figure 22. Scanning electron micrographs of ventral view of pupae of Niphta brunnea sp. nov. A habitus (scale bar: 1.0 mm) B head and thorax (scale bar: 1.0 mm) C posterior segments (scale bar = 0.5 mm) D adhesive structures (scale bar: 0.5 mm). Abbreviations: S, sternite.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600686	http://doi.org/10.3897/zookeys.1063.71180.figure17	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600687	http://doi.org/10.3897/zookeys.1063.71180.figure18	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600688	http://doi.org/10.3897/zookeys.1063.71180.figure19	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600690	http://doi.org/10.3897/zookeys.1063.71180.figure21	Figure 21. Niphta nudipennis group larval head capsules with setae and sensory pits numbered A anterior, N. brunnea sp. nov. B lateral, N. brunnea sp. nov. C anterior, N. nudipennis D lateral, N. nudipennis. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	Figure 21. Niphta nudipennis group larval head capsules with setae and sensory pits numbered A anterior, N. brunnea sp. nov. B lateral, N. brunnea sp. nov. C anterior, N. nudipennis D lateral, N. nudipennis. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600692	http://doi.org/10.3897/zookeys.1063.71180.figure23	Figure 23. Scanning electron micrographs of ventral view of larvae of Niphta brunnea sp. nov. A habitus B head and thoracic segments 1 & 2 C thoracic segments 2 & 3 and abdominal segment 1 D abdominal segments 2 & 3 E abdominal segments 6, 7, 8 & 9. Abbreviations: A, abdominal segment; T, thoracic segment. Scale bars: 1.0 mm.	Figure 23. Scanning electron micrographs of ventral view of larvae of Niphta brunnea sp. nov. A habitus B head and thoracic segments 1 & 2 C thoracic segments 2 & 3 and abdominal segment 1 D abdominal segments 2 & 3 E abdominal segments 6, 7, 8 & 9. Abbreviations: A, abdominal segment; T, thoracic segment. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
D9103CED43025A22A61F64491363B418.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600695	http://doi.org/10.3897/zookeys.1063.71180.figure26	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600675	http://doi.org/10.3897/zookeys.1063.71180.figure6	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 6. Ventral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600677	http://doi.org/10.3897/zookeys.1063.71180.figure8	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 8. Lateral views of male Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. brunnea sp. nov. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600680	http://doi.org/10.3897/zookeys.1063.71180.figure11	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600681	http://doi.org/10.3897/zookeys.1063.71180.figure12	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
3BEF2BECF04251A2802D7689F4AF6B53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600674	http://doi.org/10.3897/zookeys.1063.71180.figure5	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600676	http://doi.org/10.3897/zookeys.1063.71180.figure7	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
5A4DA9ABABB951669931265A83DAA531.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600695	http://doi.org/10.3897/zookeys.1063.71180.figure26	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
623605C366905BF79A46F3E509F742F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600671	http://doi.org/10.3897/zookeys.1063.71180.figure2	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
623605C366905BF79A46F3E509F742F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600672	http://doi.org/10.3897/zookeys.1063.71180.figure3	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
623605C366905BF79A46F3E509F742F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600673	http://doi.org/10.3897/zookeys.1063.71180.figure4	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
623605C366905BF79A46F3E509F742F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
623605C366905BF79A46F3E509F742F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600674	http://doi.org/10.3897/zookeys.1063.71180.figure5	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600676	http://doi.org/10.3897/zookeys.1063.71180.figure7	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
C274435D50125285AC6262C1F5109C80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600671	http://doi.org/10.3897/zookeys.1063.71180.figure2	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600672	http://doi.org/10.3897/zookeys.1063.71180.figure3	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600673	http://doi.org/10.3897/zookeys.1063.71180.figure4	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600682	http://doi.org/10.3897/zookeys.1063.71180.figure13	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B94E2F8451705AEBAE9F01E1821A88BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600671	http://doi.org/10.3897/zookeys.1063.71180.figure2	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 2. Ventral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600672	http://doi.org/10.3897/zookeys.1063.71180.figure3	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 3. Lateral views of male Niphta halteralis group terminalia A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gcx pl fl, gonocoxal plate filament; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600673	http://doi.org/10.3897/zookeys.1063.71180.figure4	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	Figure 4. Adult male lateral habitus micrographs of the Niphta halteralis group A N. acus sp. nov. B N. downesi sp. nov. C N. halteralis D N. mapuche sp. nov. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600682	http://doi.org/10.3897/zookeys.1063.71180.figure13	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	Figure 13. Female terminalia of the Niphta halteralis group A lateral, N. acus sp. nov. B ventral, N. acus sp. nov. C lateral, N. halteralis D ventral, N. halteralis E lateral, N. mapuche sp. nov. F ventral, N. mapuche sp. nov. Abbreviations: cerc, cercus; gen fk, genital fork; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm, lateral arm; S, sternite; spthc p, spermathecal pump; T, tergite; v scl, ventral sclerite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600689	http://doi.org/10.3897/zookeys.1063.71180.figure20	Figure 20. Niphta halteralis group larval head capsules with setae and sensory pits numbered A anterior, N. acus sp. nov. B lateral, N. acus sp. nov. C anterior, N. mapuche sp. nov. D lateral, N. mapuche sp. nov. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	Figure 20. Niphta halteralis group larval head capsules with setae and sensory pits numbered A anterior, N. acus sp. nov. B lateral, N. acus sp. nov. C anterior, N. mapuche sp. nov. D lateral, N. mapuche sp. nov. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
F6C00F4B089451EB9C4F8FC2B3C87AA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600670	http://doi.org/10.3897/zookeys.1063.71180.figure1	Figure 1. Left wing of Niphta nudipennis with arrows indicating three depigmented gaps. Abbreviations: C, costa; CuA, anterior branch of cubital vein; M, medial veins; m-cu, medial-cubital crossvein; R, radial veins; r-m, radial-medial crossvein; Sc, subcosta. Scale bar: 1.0 mm.	Figure 1. Left wing of Niphta nudipennis with arrows indicating three depigmented gaps. Abbreviations: C, costa; CuA, anterior branch of cubital vein; M, medial veins; m-cu, medial-cubital crossvein; R, radial veins; r-m, radial-medial crossvein; Sc, subcosta. Scale bar: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600674	http://doi.org/10.3897/zookeys.1063.71180.figure5	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 5. Ventral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600676	http://doi.org/10.3897/zookeys.1063.71180.figure7	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	Figure 7. Lateral views of male Niphta nudipennis group terminalia A N. daniellae sp. nov. B N. eurydactyla sp. nov. C N. nudipennis with parameres retracted D N. nudipennis with parameres extended. Arrows indicate range of motion of parameres. Abbreviations: aed gd, aedeagal guide; cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; pm, paramere. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600679	http://doi.org/10.3897/zookeys.1063.71180.figure10	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	Figure 10. Adult dorsal habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600680	http://doi.org/10.3897/zookeys.1063.71180.figure11	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	Figure 11. Ventral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; spthc p, spermathecal pump. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600681	http://doi.org/10.3897/zookeys.1063.71180.figure12	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	Figure 12. Lateral views of female Niphta nudipennis group terminalia A N. bifurcata sp. nov. B N. courtneyi sp. nov. C N. bispinosa sp. nov. D N. nudipennis. Abbreviations: cerc, cercus; gen fk, genital fork; hyp pr, hypogynial protuberance; hyp vlv, hypogynial valve; hypct, hypoproct; lat arm crcl, lateral arm circle; lat arm cmplx, lateral arm complex; S, sternite; spthc p, spermathecal pump; T, tergite. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600683	http://doi.org/10.3897/zookeys.1063.71180.figure14	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 14. Dorsal views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600684	http://doi.org/10.3897/zookeys.1063.71180.figure15	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	Figure 15. Lateral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; c tbr, caudal tubercle; resp org, respiratory organ. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600685	http://doi.org/10.3897/zookeys.1063.71180.figure16	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	Figure 16. Ventral views of Niphta pupae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: ad strct, adhesive structure; spr, spiracle. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600686	http://doi.org/10.3897/zookeys.1063.71180.figure17	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	Figure 17. Dorsal views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A, abdominal segment; Hc, head capsule; Mst, mesothorax; Mtt, metathorax; Pt, prothorax; pro spr, prothoracic spiracle; prtb, protuberance; post spr pl, posterior spiracular plate. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600687	http://doi.org/10.3897/zookeys.1063.71180.figure18	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	Figure 18. Lateral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: anl prl, anal proleg; pro prl, prothoracic proleg. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600688	http://doi.org/10.3897/zookeys.1063.71180.figure19	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	Figure 19. Ventral views of Niphta larvae A N. brunnea sp. nov. B N. nudipennis C N. acus sp. nov. Abbreviations: A ad strct, abdominal adhesive structure; T ad strct, thoracic adhesive structure. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600690	http://doi.org/10.3897/zookeys.1063.71180.figure21	Figure 21. Niphta nudipennis group larval head capsules with setae and sensory pits numbered A anterior, N. brunnea sp. nov. B lateral, N. brunnea sp. nov. C anterior, N. nudipennis D lateral, N. nudipennis. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	Figure 21. Niphta nudipennis group larval head capsules with setae and sensory pits numbered A anterior, N. brunnea sp. nov. B lateral, N. brunnea sp. nov. C anterior, N. nudipennis D lateral, N. nudipennis. Abbreviations: sp, sensory pit. Scale bars: 0.1 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600693	http://doi.org/10.3897/zookeys.1063.71180.figure24	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	Figure 24. Known distribution of Chilean Niphta A N. halteralis group B N. nudipennis group.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600695	http://doi.org/10.3897/zookeys.1063.71180.figure26	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	Figure 26. Habitat and immatures of members of the Niphta nudipennis group A type locality of N. brunnea sp. nov. and N. daniellae sp. nov., box indicating where plant stem in images B, C was taken from (38 ° 14 ' 20.6 " S 71 ° 53 ' 46.6 " W) B larvae of N. brunnea sp. nov. on plant stem from splash zone C close up of N. brunnea sp. nov. larva, adhesive structure visibly in contact with substrate D habitat of N. nudipennis, box indicating foliage in splash zone where immatures were found E pupa of N. nudipennis affixed to leaf with final instar larval exuviae visible.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
A1B7EFD258385CEBB4420F467047ADE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600696	http://doi.org/10.3897/zookeys.1063.71180.figure27	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	Figure 27. Examples of thaumaleid habitats in Chile A waterfalls where Niphta downesi sp. nov., N. eurydactyla sp. nov. and N. nudipennis were collected by sweeping adjacent foliage (40 ° 19 ' 58.7 " S 72 ° 16 ' 54.8 " W) B collection site of N. bispinosa sp. nov. (34 ° 59 ' 48.8 " S 70 ° 48 ' 37.0 " W) (after Pivar et al. 2020) C type locality of N. acus sp. nov., specimens were mainly collected from plants on left (37 ° 46 ' 30.8 " S 71 ° 42 ' 03.9 " W) D rock face seep, type locality of N. bispinosa sp. nov. (34 ° 59 ' 46.7 " S 70 ° 49 ' 19.2 " W) E Chiloe, N. nudipennis was collected at a small waterfall next to the larger Cascada Tocoihue, Pivar is shown for scale (42 ° 18 ' 20.3 " S 73 ° 26 ' 08.9 " W); F, Puente El Salto, N. halteralis was collected above the falls, Pivar is shown above the falls for scale (41 ° 31 ' 29.2 " S 72 ° 17 ' 14.6 " W).	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
B3593280B79159BE9BD27F67C51A9C21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/600678	http://doi.org/10.3897/zookeys.1063.71180.figure9	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	Figure 9. Adult lateral habitus micrographs of the Niphta nudipennis group A N. bifurcata sp. nov. (♂) B N. brunnea sp. nov. (♂) C N. courtneyi sp. nov. (♀), abdomen dissected D N. daniellae sp. nov. (♂) E N. eurydactyla sp. nov. (♂) F N. bispinosa sp. nov. (♂), abdomen dissected G N. nudipennis (♂) H N. courtneyi sp. nov. (♂), inset with arrow indicating antealar ridge. Scale bars: 1.0 mm.	2021-10-19	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.		Zenodo	biologists	Pivar, Robert J.;Sinclair, Bradley J.;Moulton, John K.			
