taxonID	type	description	language	source
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	description	(Figs 1 A-D; 2 A-H; 3 A, B) urn: lsid: zoobank. org: act: ED 5952 DC- 3544 - 434 B-ADD 6 - CEC 5 BE 37 DF 67	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (CW 19.53 mm, CL 13.19 mm, CH 9.38 mm, FW 7.19 mm); Maharashtra: Satara district: Ramban Trail, Koyna Wildlife Sanctuary; 17 ° 25 ’ 58 ” N, 73 ° 42 ’ 32 ” E; alt. 940 m; 1. VII. 2017; Tejas Thackeray leg.; ZSI-WRC C. 1999. Paratypes. India • 3 ♂ (CW 14.18 - 17.84 mm, CL 9.17 - 12.22 mm, CH 6.01 - 7.86 mm, FW 5.42 - 6.83 mm); 3 ♀ (CW 16.06 - 22.83 mm, CL 10.66 - 15.04 mm, CH 7.46 - 10.64 mm, FW 6.06 - 8.40 mm); same data as for holotype; ZSI-WRC C. 2000. OTHER MATERIAL. — India • ♂ (CW 13.77 mm, CL 9.45 mm, CH 5.96 mm, FW 5.08 mm), 2 ♀ (CW 18.91 - 21.89 mm, CL 12.64 - 14.51 mm, CH 8.66 - 9.78 mm, FW 6.82 - 7.46 mm); Maharashtra: Kolhapur district: Kokan Darshan Point, Masnoli village, c. 5 km south of Amba; 16 ° 55 ’ 22 ” N, 73 ° 47 ’ 45 ” E; alt. 840 m; 15. VII. 2020; Tejas Thackeray leg.; ZSI-WRC C. 2001. TYPE LOCALITY. — India: Maharashtra: Satara district: RambanTrail, Koyna Wildlife Sanctuary; 17 ° 25 ’ 58.8 ” N, 73 ° 42 ’ 32.4 ” E; alt. 940 m.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	diagnosis	DIAGNOSIS. — Carapace in adult proportionately narrow (CW / CL = 1.5), moderately to strongly arched (CH / CL = 0.6 - 0.7); lateral margins strongly convex; anterolateral margins relatively short, cristate; epibranchial tooth visible as weak notch; branchial regions slightly rugose; frontal margin some distance from anterior margin of epistome, exposing antennular fossae (Fig. 1 A-C). First, second maxillipeds with long flagellum on exopod; third maxilliped lacking flagellum on exopod (Fig. 2 A). Major chela with pointed fingertips; palm in adult male relatively stout; ventral margin of fixed finger and distal half of palm in adult male strongly concave (Figs 1 A, D; 2 B). Ambulatory legs relatively long (P 3 length / CL = c. 2.3) (Fig. 1 A). Male pleonal somite 6 subquadrate, slightly broader than long (Figs 1 D; 2 C). Male telson elongated (Figs 1 D; 2 C). G 1 relatively slender, almost straight; terminal segment relatively stout, distally gently curved inwards, relatively long, c. 0.5 times length of subterminal segment; subterminal segment relatively slender (Fig. 2 D, E). G 2 very short, with very short distal segment (Fig. 2 F). Female pleon in adult broadly subtriangular (Fig. 2 G). Vulvae in adult positioned apart from each other (VD / SW = c. 0.3), each subovate in shape, relatively large, occupying c. 0.4 times length of S 6, positioned close to S 5 / S 6 (Fig. 2 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	etymology	ETYMOLOGY. — The species name honours British naturalist, Gerald Malcolm Durrell for his in situ and ex situ conservation efforts. The specific epithet is thus conceived as a noun in the genitive singular. COLOUR IN LIFE. — The crabs show colour variations. They are generally fluorescent red dorsally (Fig. 3 A) and relatively paler ventrally. Some crabs are reddish brown dorsally (Fig. 3 B).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	biology_ecology	ECOLOGICAL NOTES. — Ghatiana durrelli n. sp. is a nocturnal crab and is found in tree trunk holes and in holes or crevices of laterite rocks (locally known as “ Jambha ”). The species occurs at elevated mountains (840 - 940 m altitude). Some individuals were observed feeding on dead millipedes.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Ghatiana durrelli n. sp. is currently known only from two localities of Maharashtra state (Kolhapur and Satara districts), India. Both the localities are situated in the Western Ghats.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321D06FDF3FB3B4B85.taxon	discussion	REMARKS Ghatiana durrelli n. sp. most closely resembles to G. aurantiaca (type species) mainly due to the proportionately narrower adult carapace [CW / CL = 1.5 in G. durrelli n. sp. (Fig. 1 A, B); CW / CL = 1.4 in G. aurantiaca (see Pati & Sharma 2014: fig. 2 A; Pati & Thackeray 2018: fig. 4 A)], the exposed fossae due to the gap between the frontal margin and the anterior margin of the epistome (Fig. 1 C; see Pati & Sharma 2014: fig. 2 B; Pati & Thackeray 2018: fig. 4 B), and an almost straight G 1 (Fig. 2 D, E; see Pati & Sharma 2014: fig. 3 E, F; Pati & Thackeray 2018: fig. 4 D, E). Ghatiana durrelli n. sp., however, is differentiated from G. aurantiaca by the relatively broader male pleonal somite 6 (Figs 1 D; 2 C) (vs male pleonal somite 6 as long as broad; see Pati & Sharma 2014: figs 2 C, 3 B; Pati & Thackeray 2018: fig. 4 C), the relatively stouter G 1 terminal segment (Fig. 2 D, E) (vs G 1 terminal segment relatively slenderer; see Pati & Sharma 2014: fig. 3 E-G; Pati & Thackeray 2018: fig. 4 D, E), the relatively slenderer G 1 subterminal segment (Fig. 2 D, E) (vs G 1 subterminal segment relatively stouter; see Pati & Sharma 2014: fig. 3 E, F; Pati & Thackeray 2018: fig. 4 D, E), and the relatively closely positioned adult vulva in relation to S 5 / S 6 (Fig. 2 H) (vs adult vulva positioned a clear distance from S 5 / S 6; see Pati & Thackeray 2018: fig. 4 H). Among congeners, the stout G 1 terminal segment is unique in G. durrelli n. sp. and G. pulchra Pati & Thackeray, 2018 (Fig. 2 D; see Pati & Thackeray 2018: fig. 8 I, J). In fact, the G 1 structure of G. durrelli n. sp. is almost identical to that of G. pulchra except for the gently inwardly curved terminal segment (Fig. 2 D) (vs straight G 1 terminal segment in G. pulchra; see Pati & Thackeray 2018: fig. 8 I, J). Ghatiana durrelli n. sp. can nevertheless be separated from G. pulchra by the proportionately narrower adult carapace, CW / CL = 1.5 (Fig. 1 A, B) (vs proportionately broader adult carapace, CW / CL = 1.6; see Pati & Thackeray 2018: fig. 8 A), the strongly concave ventral margin of the fixed finger and the distal half of the palm of the major chela in adult males (Fig. 2 B) (vs gently concave ventral margin of the fixed finger and the distal half of the palm of the major chela in adult males; see Pati & Thackeray 2018: fig. 8 D), and the relatively longer ambulatory legs, P 3 length / CL = c. 2.3 (Fig. 1 A) (vs the relatively shorter ambulatory legs, P 3 length / CL = c. 2.0; see Pati & Thackeray 2018: fig. 8 A).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDEE3321CB5FEF1FE794E4A.taxon	type_taxon	TYPE SPECIES. — Ghatiana aurantiaca Pati & Sharma, 2014, by original designation; gender feminine.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	description	(Figs 3 C; 4 A-D; 5 A-H) urn: lsid: zoobank. org: act: 802 C 4 E 90 - F 54 C- 4060 - A 72 D- 23 C 1 C 42474 C 7	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (CW 13.38 mm, CL 10.95 mm, CH 7.28 mm, FW 6.13 mm); Karnataka: Uttara Kannada district: Shirley Waterfall; 14 ° 52 ’ 55 ” N, 74 ° 39 ’ 46 ” E; alt. 168 m; 6. X. 2016; Tejas Thackeray and Swapnil Pawar leg.; ZSI-WRC C. 2002. Paratypes. India • 2 ♂ (CW 12.39 - 12.41 mm, CL 9.77 - 9.83 mm, CH 6.60 - 6.76 mm, FW 5.50 - 5.55 mm), ♀ (CW 14.11 mm, CL 11.19 mm, CH 7.32 mm, FW 6.18 mm); same data as for holotype; ZSI-WRC C. 2003 • 2 ♀ (CW 12.87 - 13.16 mm, CL 10.44 - 10.56 mm, CH 7.42 - 7.56 mm, FW 5.81 - 6.03 mm); Karnataka: Uttara Kannada district: Mavinagundi, near Jog Falls; 14 ° 15 ’ 18 ” N, 74 ° 48 ’ 21 ” E; alt. 601 m; 7. X. 2016; Tejas Thackeray and Swapnil Pawar leg.; ZSI-WRC C. 2004 • ♂ (CW 12.99 mm, CL 10.37 mm, CH 6.93 mm, FW 5.70 mm), ♀ (CW 14.81 mm, CL 12.14 mm, CH 8.29 mm, FW 6.75 mm); Goa: North Goa district: Chorla; 15 ° 38 ’ 5 ” N, 74 ° 7 ’ 12 ” E; alt. 423 m; 18. IX. 2016; Tejas Thackeray and Swapnil Pawar leg.; ZSI-WRC C. 2005. OTHER MATERIAL. — India • 4 ♂ (CW 8.20 - 12.50 mm, CL 6.53 - 9.62 mm, CH 4.06 - 6.45 mm, FW 3.81 - 5.50 mm), 6 ♀ (CW 10.63 - 14.10 mm, CL 8.20 - 11.16 mm, CH 5.60 - 7.39 mm, FW 4.69 - 6.15 mm); Karnataka: Uttara Kannada district: Mavinagundi, near Jog Falls; 14 ° 15 ’ 18 ” N, 74 ° 48 ’ 21 ” E; alt. 601 m; 7. X. 2016; Tejas Thackeray and Swapnil Pawar leg.; ZSI-WRC C. 2006. TYPE LOCALITY. — India: Karnataka: Uttara Kannada district: Shirley Waterfall; 14 ° 52 ’ 55 ” N, 74 ° 39 ’ 46 ” E; alt. 168 m.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	diagnosis	DIAGNOSIS. — Carapace in adult proportionately narrow (CW / CL = 1.2 - 1.3), moderately to strongly arched (CH / CL = 0.6 - 0.7); lateral margins gently convex; anterolateral margins relatively short, cristate; epibranchial tooth visible as distinct notch; branchial regions rugose; frontal margin very close to anterior margin of epistome, hiding antennular fossae (Fig. 4 A-C). First, second maxillipeds with long flagellum on exopod; third maxilliped lacking flagellum on exopod (Fig. 5 A). Major chela with pointed fingertips; palm in adult male relatively stout; ventral margin of fixed finger and distal half of palm in adult male strongly concave (Figs 4 A, D; 5 B). Ambulatory legs relatively short (P 3 length / CL = c. 2.0) (Fig. 1 A). Male pleonal somite 6 subquadrate, slightly longer than broad (Figs 4 D; 5 C). Male telson elongated (Figs 4 D; 5 C). G 1 relatively stout, medially gently curved outwards; terminal segment relatively slender, distally gently curved inwards, relatively long, c. 0.5 times length of subterminal segment; subterminal segment relatively stout (Fig. 5 D, E). G 2 very short, with very short distal segment (Fig. 5 F). Female pleon in adult broadly subtriangular (Fig. 5 G). Vulvae in adult relatively closely positioned (VD / SW = c. 0.2), each subovate in shape, relatively large, occupying c. 0.4 times length of S 6, positioned some distance from S 5 / S 6 (Fig. 5 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	etymology	ETYMOLOGY. — The species is named after the Swiss zoologist, Dr Jean Roux for his contributions on the study of crabs of the Western Ghats. The species name is conceived as a noun in the genitive singular.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	description	COLOUR IN LIFE. — Carapace, chelipeds, and ambulatory legs all are deep purple dorsally (Fig. 3 C). The carapace and pereiopods are relatively paler on the ventral surface.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	biology_ecology	ECOLOGICAL NOTES. — Ghatiana rouxi n. sp. was found under small boulders adjacent to waterfalls and along stream banks. They can be also seen underneath small boulders of forest trails clearly away from water bodies. On the basis of the collected material, the species occurs at both low and high elevations (168 - 601 m altitude).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Ghatiana rouxi n. sp. is currently known only from the Western Ghats of the Goa and Karnataka states, India.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFDDE3311D12FF51FAA54B85.taxon	discussion	REMARKS Ghatiana rouxi n. sp. is a relatively smaller species (the largest known adult female with a fully developed pleon measures 14.81 mm in CW), which has a proportionately narrower adult carapace (CW / CL = 1.2 - 1.3) with the lateral margins gently convex (Fig. 4 A, B). The gently convex lateral margins of the carapace is characteristic to G. rouxi n. sp. (Fig. 4 A, B); the remaining species of the genus have a carapace with the lateral margins strongly convex (Fig. 1 A, B; Pati & Sharma 2014: fig. 4 A; Pati & Thackeray 2018: figs 3 A; 4 A; 5 A; 6 A; 7 A; 8 A; 9 A; 10 A). Ghatiana rouxi n. sp. has an elongated male pleonal somite 6 (Figs 4 D; 5 C), which is also characteristic to G. botti Pati & Thackeray, 2018, G. rathbunae Pati & Thackeray, 2018, and G. splendida Pati, Thackeray & Khaire, 2016 (see Pati et al. 2016: fig. 5 C, G; Pati & Thackeray 2018: figs 6 C, G; 9 C, G; 10 C). Ghatiana rouxi n. sp. is immediately distinguished from G. botti, G. rathbunae, and G. splendida by the basally broad subterminal segment of the G 1 (Fig. 5 D, E) (vs G 1 subterminal segment basally narrow; see Pati et al. 2016: fig. 6 A, B; Pati & Thackeray 2018: figs 6 I, K; 9 I, K; 10 D, E) in addition to the gently convex lateral margins of its carapace (Fig. 4 A, B) (vs carapace with strongly convex lateral margins; see Pati et al. 2016: fig. 5 A; Pati & Thackeray 2018: figs 6 A; 9 A; 10 A). The stout, basally broad G 1 subterminal segment of G. rouxi n. sp. is a character shared only with G. aurantiaca Pati & Sharma, 2014 (Fig. 5 D, E; see Pati & Thackeray 2018: fig. 4 D, E). In addition, their G 1 s are relatively slenderer, distally gently curved inwards, with the terminal segment relatively longer, c. 0.5 times the length of the subterminal segment (Fig. 5 D, E; see Pati & Thackeray 2018: fig. 4 D, E). The G 1 is nevertheless gently curved outwards medially in G. rouxi n. sp. (Fig. 5 D, E) as compared to the straight G 1 of G. aurantiaca (see Pati & Thackeray 2018: fig. 4 D, E). Moreover, the structure of the adult vulvae and most of the features of the carapace are similar in both. The antennular fossae, however, are being hidden by the closely located frontal margin and the anterior margin of the epistome in G. rouxi n. sp. (Fig. 4 C) (vs exposed fossae due to the gap between the frontal margin and the anterior margin of the epistome in G. aurantiaca; see Pati & Sharma 2014: fig. 2 B; Pati & Thackeray 2018: fig. 4 B). Again, their male pleonal somite 6 is different, being subquadrate, slightly longer than broad in G. rouxi n. sp. (Figs 4 D; 5 C) and quadrate, as long as broad in G. aurantiaca (see Pati & Sharma 2014: figs 2 C; 3 B; Pati & Thackeray 2018: fig. 4 C).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E33D1C98F974FD944B85.taxon	type_taxon	TYPE SPECIES. — Sahyadriana sahyadriensis Pati & Thackeray, 2018, by original designation; gender feminine.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	description	(Figs 3 D; 6 A-D; 7 A-H) urn: lsid: zoobank. org: act: A 64 D 966 D-A 859 - 48 C 1 - 9949 - 092 B 1 D 7288 EB	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (CW 10.29 mm, CL 7.86 mm, CH 4.47 mm, FW 4.16 mm); Maharashtra: Satara district: Dhobi Waterfall, Mahabaleshwar; 17 ° 56 ’ 13 ” N, 73 ° 38 ’ 45 ” E; alt. 1075 m; 30. VI. 2018; Tejas Thackeray leg.; ZSI-WRC C. 2015. Paratypes. India • ♂ (CW 10.30 mm, CL 7.89 mm, CH 4.55 mm, FW 4.14 mm); ♀ (CW 11.93 mm, CL 9.05 mm, CH 5.61 mm, FW 4.58 mm); same data as for holotype; ZSI-WRC C. 2016 • 2 ♂ (CW 9.20 - 10.24 mm, CL 6.92 - 7.78 mm, CH 4.00 - 4.82 mm, FW 3.62 - 4.03 mm); 3 ♀ (CW 10.97 - 12.33 mm, CL 8.27 - 9.14 mm, CH 5.13 - 5.56 mm, FW 4.24 - 4.84 mm); same data as for holotype; ZSI-WRC C. 2017. TYPE LOCALITY. — India: Maharashtra: Satara district: Dhobi Waterfall, Mahabaleshwar; 17 ° 56 ’ 13 ” N, 73 ° 38 ’ 45 ” E; alt. 1075 m.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	diagnosis	DIAGNOSIS. — Carapace in adult slightly broader than long (CW / CL = 1.3), moderately deep (CH / CL = 0.6); epibranchial tooth visible as relative distinct notch; branchial regions inflated; epistome posterior margin with strongly concave lateral lobes (Fig. 6 A-C). Third maxilliped lacking flagellum on exopod (Fig. 7 A). Chelipeds with pointed fingertips (Figs 6 A, D; 7 B). Ambulatory legs with densely setose dactylus and propodus (Fig. 6 A, D). Male sternopleonal cavity long, extending beyond imaginary line joining bases of third maxillipeds (Figs 6 D; 7 C). Male pleon relatively broad, T-shaped; pleonal somite 5 with concave lateral margins; pleonal somite 6 subquadrate, slightly broader than long, subequal in length to telson, lateral margins convex (Figs 6 D; 7 C). Male telson short (Figs 6 D; 7 C). G 1 stout, straight; terminal segment relatively less stout, proximal two-thirds stouter than distal third, inverted funnel-shaped, relatively long, c. 0.5 times length of subterminal segment; subterminal segment relatively slender (Fig. 7 D, E). G 2 very short, with very short distal segment (Fig. 7 F). Female pleon in adult broadly subtriangular (Fig. 7 G). Vulvae in adult positioned apart from each other (VD / SW = c. 0.3), each subovate in shape, relatively small, occupying c. 0.4 times length of S 6, positioned some distance from S 5 / S 6 (Fig. 7 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	etymology	ETYMOLOGY. — The species name is an adjective in the Latin nominative singular meaning unexpected, referring to the unexpected discovery of the species from the type locality of a congener, i. e., S. triangulus (Pati & Sharma, 2014).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	description	COLOUR IN LIFE. — Crabs have chocolate brown coloured carapace and ambulatory legs, whereas their chelipeds are orangish brown (Fig. 3 D). Their ventral surface is relatively paler.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	biology_ecology	ECOLOGICAL NOTES. — Sahyadriana inopinata n. sp. was found under cobblestones adjacent to a waterfall. The species dwells at higher altitude (1075 m above a. s. l.). The sympatric species, S. triangulus is so far known only from preserved specimens, with no information on its ecological requirements (Pati & Thackeray 2018).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Sahyadriana inopinata n. sp. is known only from the type locality (Dhobi Waterfall, Mahabaleshwar) in Satara district of Maharashtra, India. The type locality is situated in the Western Ghats.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD1E3381F8DF977FA2D4AC3.taxon	discussion	REMARKS Among congeners, S. inopinata n. sp. is most similar to S. billyarjani Pati & Thackeray, 2018 and the type species, S. sahyadriensis, due to strongly concave lateral lobes of the epistome posterior margin (Fig. 6 C; see Pati & Thackeray 2018: figs 20 B; 24 B), the pointed fingertips of the chelipeds (Figs 6 A, D; 7 B; see Pati & Thackeray 2018: figs 20 A, C, D; 24 A, C, D), the relatively stouter, inverted funnel-shaped terminal segment of the G 1 with the distal portion abruptly narrow (Fig. 7 D, E; see Pati & Thackeray 2018: figs 20 I-K, M; 24 I-K), and the relatively slenderer subterminal segment of the G 1 (Fig. 7 D, E; see Pati & Thackeray 2018: figs 20 I, K, M; 24 I, K). Sahyadriana inopinata n. sp. is nevertheless distinguished from S. billyarjani and S. sahyadriensis by the relatively distinct notch between the epibranchial tooth and the external orbital angle (Fig. 7 A, B) (vs indistinct notch between the epibranchial tooth and the external orbital angle; see Pati & Thackeray 2018: figs 20 A; 24 A), the relatively slenderer and relatively longer terminal segment of the G 1, i. e., c. 0.5 times the length of the subterminal segment (Fig. 7 D, E) (vs relatively stouter and relatively shorter terminal segment of the G 1, i. e., c. 0.3 - 0.4 times the length of the subterminal segment; see Pati & Thackeray 2018: figs 20 I-K, M; 24 I-K), and the subovate-shaped vulva, which is relatively smaller, occupying c. 0.4 times the length of the S 6 and is positioned a clear distance from S 5 / S 6 (Fig. 7 H) (vs suborbicular-shaped vulva, which is relatively larger, occupying c. 0.5 times the length of the S 6 and is positioned close to S 5 / S 6; Pati & Thackeray 2018: figs 20 O; 24 N). The strongly concave lateral lobes of the epistome posterior margin and the inverted-funnel shaped terminal segment of the G 1 of S. inopinata n. sp. are also similar to those of S. pilosipes (Alcock, 1909) (Figs 6 C; 7 D, E; see Pati & Thackeray 2018: figs 22 B, E, F; 23 B, D, E). Both the species have a relatively broader male pleon with the concave lateral margins of the pleonal somite 5 (Figs 6 D; 7 C; see Pati & Thackeray 2018: figs 22 C; 23 C). Moreover, the G 1 is stout and straight, with the terminal segment relatively longer (c. 0.5 times the length of the subterminal segment) and the subterminal segment relatively slenderer in S. inopinata n. sp. and S. pilosipes (Fig. 7 D, E; see Pati & Thackeray 2018: figs 22 E, F; 23 D, E). Sahyadriana inopinata n. sp., however, is easily distinguished from S. pilosipes by the pointed fingertips of the chelipeds (Figs 6 A, D; 7 B) (vs broadly rounded or spoon-shaped fingertips of the chelipeds; see Pati & Thackeray 2018: figs 22 C; 23 C), the relatively stouter terminal segment of the G 1, i. e., the proximal two-thirds stouter than the distal third (Fig. 7 D, E) (vs relatively slenderer G 1 terminal segment, i. e., the proximal half stouter than the distal half; see Pati & Thackeray 2018: figs 22 E, F; 23 D, E), and the subovateshaped and relatively smaller vulva in adult that occupies c. 0.4 times the length of the S 6 (Fig. 7 H) (vs suborbicular-shaped and relatively larger vulvae in adult, occupying c. 0.5 times the length of the S 6; see Pati & Thackeray 2018: fig. 23 H). Sahyadriana inopinata n. sp. need not be confused with the sympatric species, S. triangulus, because of the relatively distinct epibranchial tooth (Fig. 6 A, B) (vs indistinct epibranchial tooth; see Pati & Thackeray 2018: fig. 28 A), the straight terminal segment of the G 1 (Fig. 7 D, E) (vs distally distinctly curved terminal segment of the G 1; see Pati & Thackeray 2018: fig. 28 D, E), and the basally narrow subterminal segment of the G 1 (Fig. 7 D, E) (vs basally broad subterminal segment of the G 1; see Pati & Thackeray 2018: fig. 28 D, E).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	description	(Figs 3 E; 8 A-D; 9 A-H) urn: lsid: zoobank. org: act: 548 CA 624 - C 80 A- 48 BF- 86 D 0 - 19 F 9 B 835609 E	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (CW 13.96 mm, CL 10.88 mm, CH 6.38 mm, FW 4.98 mm); Maharashtra: Nashik district: Bramhagiri, near Trimbak; 19 ° 54 ’ 46 ” N, 73 ° 31 ’ 4 ” E; alt. 1048 m; 17. VIII. 2017; Tejas Thackeray leg.; ZSI-WRC C. 2007. Paratypes. India • ♂ (CW 13.97 mm, CL 11.02 mm, CH 6.24 mm, FW 5.06 mm), ♀ (CW 15.92 mm, CL 12.16 mm, CH 6.35 mm, FW 5.74 mm); same data as for holotype; ZSI-WRC C. 2008 • 5 ♂ (CW 12.44 - 14.18 mm, CL 9.55 - 11.01 mm, CH 5.58 - 7.16 mm, FW 4.46 - 4.94 mm), 5 ♀ (CW 12.11 - 15.53 mm, CL 9.39 - 11.73 mm, CH 6.14 - 7.23 mm, FW 4.44 - 5.44 mm); same data as for holotype; ZSI-WRC C. 2009. TYPE LOCALITY. — India: Maharashtra: Nashik district: Bramhagiri, near Trimbak; 19 ° 54 ’ 46 ” N, 73 ° 31 ’ 4 ” E; alt. 1048 m.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	diagnosis	DIAGNOSIS. — Carapace in adult slightly broader than long (CW / CL = 1.3), moderately to strongly deep (CH / CL = 0.5 - 0.7); epibranchial tooth visible as relative distinct notch; branchial regions inflated; epistome posterior margin with gently concave lateral lobes (Fig. 8 A-C). Third maxilliped lacking flagellum on exopod (Fig. 9 A). Chelipeds with pointed fingertips (Figs 8 A, D; 9 B). Ambulatory legs with densely setose dactylus and propodus (Fig. 8 A, D). Male sternopleonal cavity long, extending beyond imaginary line joining bases of third maxillipeds (Figs 8 D; 9 C). Male pleon relatively broad, T-shaped; pleonal somite 5 with concave lateral margins; pleonal somite 6 subquadrate, broader than long, subequal in length to telson, lateral margins strongly convex (Figs 8 D; 9 C). Male telson short (Figs 8 D; 9 C). G 1 distinctly stout, almost straight; terminal segment relatively slender, subcylindrical, distally gently curved inwards, relatively short, c. 0.4 times length of subterminal segment; subterminal segment distinctly stout, triangular, distally relatively broad, with convex inner margin (Fig. 9 D, E). G 2 very short, with very short distal segment (Fig. 9 F). Female pleon in adult broadly subtriangular (Fig. 9 G). Vulvae in adult positioned apart from each other (VD / SW = c. 0.4), each suborbicular in shape, relatively large, occupying c. 0.5 times length of S 6, positioned close to S 5 / S 6 (Fig. 9 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	etymology	ETYMOLOGY. — The species name, derived from the Marathi language for orange coloured, refers to live colouration of the crab. The name is used as a Latin noun in apposition.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	description	COLOUR IN LIFE. — Carapace, chelipeds, and ambulatory legs all are completely orange in colour (Fig. 3 E). The orange colour on the ventral side of the carapace, however, is relatively faint.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	biology_ecology	ECOLOGICAL NOTES. — Sahyadriana keshari n. sp. dwells on the walls of high mountainous cliffs (1048 m above a. s. l.), which host ephemeral waterfalls. Crabs gather in large numbers during the daytime on these exposed walls between the downpour.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Sahyadriana keshari n. sp. is currently known only from Bramhagiri, an isolated high mountain of the Western Ghats in Nashik district of Maharashtra state, India. The new species is very likely to occur on the adjacent mountainous peak “ Anjneri ” of the same district because the second author (TT) once saw a dead crab of the species there.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFD4E3271F9CF977FAD749C3.taxon	discussion	REMARKS The carapace of S. keshari n. sp. most resembles to that of S. waghi (Pati in Pati, Thackeray & Khaire, 2016) mainly due to the relatively distinct notch between the epibranchial tooth and the external orbital angle (Fig. 8 A, B; see Pati et al. 2016: fig. 11 A; Pati & Thackeray 2018: fig. 29 A), the gently concave lateral lobes of the epistome posterior margin (Fig. 8 C; see Pati et al. 2016: fig. 11 B; Pati & Thackeray 2018: fig. 29 B), and the pointed fingertips of the chelipeds (Figs 8 A, D; 9 B; see Pati et al. 2016: fig. 11 A, C, D; Pati & Thackeray 2018: fig. 29 A, C). In the G 1 structure, S. keshari n. sp. is most similar to S. waghi and S. triangulus because all have a distinctly stout G 1, with the relatively slenderer, subcylindrical terminal segment (Fig. 9 D, E; see Pati & Sharma 2014: fig. 7 E-G; Pati et al. 2016: fig. 12 A-C; Pati & Thackeray 2018: figs 28 D, E, H; 29 D-G, I-K). Both S. keshari n. sp. and S. waghi can be differentiated from S. triangulus by the relatively distinct notch between the epibranchial tooth and the external orbital angle (Fig. 8 A, B; see Pati et al. 2016: fig. 11 A; Pati & Thackeray 2018: fig. 29 A) (vs indistinct notch between the epibranchial tooth and the external orbital angle in S. triangulus; see Pati & Sharma 2014: fig. 6 A; Pati & Thackeray 2018: fig. 28 A), the gently concave lateral lobes of the epistome posterior margin (Fig. 8 C; see Pati et al. 2016: fig. 11 B; Pati & Thackeray 2018: fig. 29 B) (vs strongly concave lateral lobes of the epistome posterior margin in S. triangulus; see Pati & Sharma 2014: fig. 6 B; Pati & Thackeray 2018: fig. 28 B), the relatively shorter G 1 terminal segment, c. 0.4 times the length of the subterminal segment, with the gently inwardly curved distal portion (Fig. 9 D; see Pati & Thackeray 2018: figs 29 I-K) (vs relatively longer G 1 terminal segment, c. 0.5 times the length of the subterminal segment, with the distinctly outwardly curved distal portion in S. triangulus; see Pati & Sharma 2014: fig. 7 E-G; Pati & Thackeray 2018: fig. 28 D, E, H), and the convex inner margin of the G 1 subterminal segment (Fig. 9 D, E; see Pati et al. 2016: fig. 12 A, B; Pati & Thackeray 2018: fig. 29 D, F, I-K) (vs almost straight inner margin of the G 1 subterminal segment in S. triangulus; see Pati & Sharma 2014: fig. 7 E, F; Pati & Thackeray 2018: fig. 28 D, E, H). Sahyadriana keshari n. sp. can be further separated from S. waghi by the relatively stouter G 1 terminal segment (Fig. 9 D) (vs relatively slenderer G 1 terminal segment; see Pati et al. 2016: fig. 12 A; Pati & Thackeray 2018: fig. 29 D, I-K), the relatively broader distal portion of the G 1 subterminal segment (Fig. 9 D) (vs relatively narrower distal portion of the G 1 subterminal segment; see Pati et al. 2016: fig. 12 A; Pati & Thackeray 2018: fig. 29 D, I-K), and the relatively larger adult vulva that occupies c. 0.5 times the length of the S 6 and is positioned close to S 5 / S 6 (Fig. 9 H) (vs relatively smaller adult vulva that occupies c. 0.4 times the length of the S 6 and is positioned a clear distance from S 5 / S 6; see Pati & Thackeray 2018: fig. 29 M).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	description	(Figs 3 F-H; 10 A-D; 11 A-H) urn: lsid: zoobank. org: act: 70189 F 64 - 7 C 11 - 495 C- 96 AB- 42 E 320 DFDF 78	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (CW 9.36 mm, CL 7.37 mm, CH 4.14 mm, FW 3.63 mm); Maharashtra: Pune district: Tamhini Ghat; 18 ° 28 ’ 37 ” N, 73 ° 25 ’ 1 ” E; alt. 621 m; 25. VII. 2017; V. D. Hegde et al. leg.; ZSI-WRC C. 2010. Paratypes. India • ♂ (CW 8.86 mm, CL 6.97 mm, CH 4.08 mm, FW 3.44 mm), ♀ (CW 9.02 mm, CL 6.96 mm, CH 3.74 mm, FW 3.53 mm); same data as for holotype; ZSI-WRC C. 2011 • ♂ (CW 9.21 mm, CL 7.30 mm, CH 4.15 mm, FW 3.68 mm), ♀ (CW 11.59 mm, CL 8.80 mm, CH 5.68 mm, FW 4.38 mm); Maharashtra: Pune district: Tamhini Ghat; 18 ° 28 ’ 22 ” N, 73 ° 25 ’ 8 ” E; alt. 621 m; 11. VIII. 2017; Tejas Thackeray leg.; ZSI-WRC C. 2012 • ♂ (CW 8.77 mm, CL 6.74 mm, CH 3.74 mm, FW 3.49 mm), ♀ (CW 10.02 mm, CL 7.60 mm, CH 4.02 mm, FW 3.94 mm); Maharashtra: Pune district: Tamhini Ghat; 18 ° 28 ’ 4 ” N, 73 ° 24 ’ 46 ” E; alt. 584 m; same collection date and collector as for holotype; ZSI-WRC C. 2013 • ♂ (CW 9.51 mm, CL 7.23 mm, CH 4.00 mm, FW 3.77 mm), ♀ (CW 11.06 mm, CL 8.11 mm, CH 4.83 mm, FW 4.17 mm); Maharashtra: Pune district: Tamhini Ghat; 18 ° 28 ’ 1 ” N, 73 ° 24 ’ 46 ” E; alt. 572 m; same collection date and collector as for ZSI-WRC C. 2012 paratypes; ZSI-WRC C. 2014. TYPE LOCALITY. — India: Maharashtra: Pune district: Tamhini Ghat; 18 ° 28 ’ 37 ” N, 73 ° 25 ’ 1 ” E; alt. 621 m.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	diagnosis	DIAGNOSIS. — Carapace in adult slightly broader than long (CW / CL = 1.3 - 1.4), moderately deep (CH / CL = 0.5 - 0.6); epibranchial tooth visible as relative distinct notch; branchial regions inflated; epistome posterior margin with strongly concave lateral lobes (Fig. 10 A-C). Third maxilliped lacking flagellum on exopod (Fig. 11 A). Chelipeds with pointed fingertips (Figs 10 A, D; 11 B). Ambulatory legs with densely setose dactylus and propodus (Fig. 10 A, D). Male sternopleonal cavity long, extending beyond imaginary line joining bases of third maxillipeds (Figs 10 D; 11 C). Male pleon relatively broad, T-shaped; pleonal somite 5 with concave lateral margins; pleonal somite 6 subquadrate, broader than long, subequal in length to telson, lateral margins gently convex (Figs 10 D; 11 C). Male telson short (Figs 10 D; 11 C). G 1 moderately stout, almost straight; terminal segment relatively slender, subcylindrical, distally distinctly narrow and gently curved inwards, relatively long, c. 0.5 times length of subterminal segment; subterminal segment relatively slender, with sinuous inner margin (Fig. 11 D, E). G 2 very short, with very short distal segment (Fig. 11 F). Female pleon in adult broadly subtriangular (Fig. 11 G). Vulvae in adult positioned apart from each other (VD / SW = c. 0.3), each suborbicular in shape, relatively large, occupying c. 0.5 times length of S 6, positioned some distance from S 5 / S 6 (Fig. 11 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	etymology	ETYMOLOGY. — The species is named after the type locality, Tamhini Ghat, a hotspot for wildlife in Maharashtra that hosts rich biodiversity and many endemic species, including that of the freshwater crabs. The species name is used as a Latin noun in apposition.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	description	COLOUR IN LIFE. — The species shows colour variation. The carapace and ambulatory legs are generally saddle brown, with orange coloured chelipeds on the dorsal surface (Fig. 3 F); the carapace and pereiopods are relatively paler ventrally. Some crabs have orange coloured carapace and chelipeds but brownish ambulatory legs dorsally (Fig. 3 G). Some individuals are completely brown (Fig. 3 H), with a paler ventral surface of the carapace.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	biology_ecology	ECOLOGICAL NOTES. — Sahyadriana tamhini n. sp. was found under cobblestones near the base of rainfed cascades in an elevated mountain (above 570 m altitude). Some individuals were also seen dwelling in the crevices of mountain cliffs. These crabs are very active during the monsoon season (June to September).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Sahyadriana tamhini n. sp. is known only from the type locality (Tamhini Ghat) in the Western Ghats of Pune district, Maharashtra, India.	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
4846381AFFCBE3221F9BFA74FB9A4983.taxon	discussion	REMARKS Among congeners, S. tamhini n. sp., is most closely related to S. alcocki (Pati in Pati, Thackeray & Khaire, 2016) mainly due to the strongly concave lateral lobes of the epistome posterior margin (Fig. 10 C; see Pati et al. 2016: fig. 7 B; Pati & Thackeray 2018: fig. 19 B), the pointed fingertips of the chelipeds (Figs 10 A, D; 11 B; see Pati et al. 2016: fig. 7 A- D; Pati & Thackeray 2018: fig. 19 A, C), the moderately stout and almost straight G 1, and the relatively slenderer, subcylindrical terminal segment of the G 1, with the gently inwardly curved distal portion (Fig. 11 D, E; see Pati et al. 2016: fig. 8 A-C; Pati & Thackeray 2018: fig. 19 D, E). These two species, however, can be differentiated by the terminal segment of the G 1, which is distinctly narrow distally and relatively longer, c. 0.5 times the length of the subterminal segment in S. tamhini n. sp. (Fig. 11 D, E), and gradually narrow distally and relatively shorter, c. 0.4 times the length of the subterminal segment in S. alcocki (see Pati et al. 2016: fig. 8 A-C; Pati & Thackeray 2018: fig. 19 D, E). Again, the shape of the subterminal segment of the G 1 looks different; the inner margin of the G 1 subterminal segment is sinuous in S. tamhini n. sp. (Fig. 11 D, E), while it is almost straight in S. alcocki (see Pati et al. 2016: fig. 8 A, B; Pati & Thackeray 2018: fig. 19 D, E). Although both the species have a suborbicular-shaped adult vulva that is situated clearly away from S 5 / S 6 (Fig. 11 H; see Pati & Thackeray 2018: fig. 19 H), the vulvae in adult are relatively closed positioned (VD / SW = c. 0.3) and relatively larger, occupying c. 0.5 times the length of the S 6 in S. tamhini n. sp. (Fig. 11 H), whereas they are clearly positioned apart from each other (VD / SW = c. 0.4) and relatively smaller, occupying c. 0.4 times the length of the S 6 in S. alcocki (see Pati & Thackeray 2018: fig. 19 H).	en	Pati, Sameer K., Thackeray, Tejas (2021): Five new species of freshwater crabs of the genera Ghatiana Pati & Sharma, 2014 and Sahyadriana Pati & Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema 43 (26): 627-647, DOI: 10.5252/zoosystema2021v43a26
