identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2A6687ABFFF18355EDDDF0A2FE10ACA0.text	2A6687ABFFF18355EDDDF0A2FE10ACA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium arvinum (Schluger 1960)	<div><p>Leptotrombidium arvinum (Schluger, 1960)</p> <p>(Figs. 1, 2)</p> <p>Trombicula (Leptotrombidium) arvina Schluger, 1960 (in Schluger et al. 1960b): 1792, fig. 2.</p> <p>Leptotrombidium (Leptotrombidium) arvina: Traub &amp; Lakshana 1966: 271, figs. 1–5; Lakshana 1973: 6; Chau et al. 2007: 40, fig. 15.</p> <p>Leptotrombidium (Leptotrombidium) miculum arvinum: Vercammen-Grandjean &amp; Langston 1976: 422, pl. 100; Xiang &amp; Wen 1984: 260; Wen 1984: 308.</p> <p>Leptotrombidium (Leptotrombidium) arvinum: Kudryashova 2004: 13.</p> <p>Leptotrombidium arvinum: Chaisiri et al. 2016: 323; Stekolnikov 2013: 69; 2021: 112.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNN; fSc: PL&gt; AM&gt; AL; Ip = 601–625; fD = 2H-8-6-6-2-2; DS = 26–28; V = 23–24; NDV = 49–52. Standard measurements of type series given in Table 1.</p> <p>Description (larva) (based on lectotype and 2 paralectotypes, fD recorded in one specimen). IDIOSOMA (Figs. 1A, 2B–G). Eyes 2 + 2; 26–28 barbed dorsal idiosomal setae, including one pair of humeral setae, arranged 2H-8-6-6-2-2; 4 sternal setae; 23–24 ventral setae; NDV = 49–52.</p> <p>GNATHOSOMA (Fig. 2H, I). Cheliceral blade with tricuspid cap; gnathobase with sparse puncta and 1 pair of branched gnathocoxal (tritorostral) setae; proximal part of cheliceral base and palpal femur with sparse puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude; dorsal palpal tibial seta branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 1, 2A). Rectangular, moderately covered with rather large puncta, greatly projected posterior to PL, posterior margin almost straight in middle part; AM posterior to level of AL, sensillary (trichobothrial) bases posterior to level of PL (P-PL – PSB = 6–8 µm); AL and PL densely covered with rather long thin barbs, PL slightly expanded proximally, covered with shorter and wider barbs, similarly to dorsal idiosomal setae; PL&gt; AM&gt; AL; flagelliform sensilla (trichobothria), with about 11 long branches in distal half.</p> <p>LEGS (Fig. 2J–L). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 21B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Nghe An Province, ex Tupaia sp. (Schluger et al. 1960b). Later on, it was recorded in the following countries: in Thailand, from mammals Tupaia glis (Diard and Duvaucel), Callosciurus caniceps (Gray), C. erythraeus (Pallas), C. finlaysonii (Horsfield), Crocidura horsfieldii (Tomes), Hylomys suillus S. Müller, Maxomys rajah (Thomas) (= Rattus rajah), Menetes berdmorei (Blyth), Niviventer niviventer (Hodgson) (= Rattus niveiventer), Rattus rattus (L.), Berylmys berdmorei (Blyth) (= Rattus berdmorei), Paradoxurus hermaphroditus (Pallas), and birds Chalcophaps indica (L.), Pellorneum ruficeps Swainson (Traub &amp; Lakshana 1966), and Leopoldamys sabanus (= Rattus sabanus) (Lakshana 1973); in Laos, from T. glis and Dremomys rufigenis (Blanford); in Malaysia, from Tupaia sp. and Callosciurus caniceps (Vercammen- Grandjean &amp; Langston 1976), and in China, from the specimens collected with a black plate (Wen 1984).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3144, specimen 4, designated here) ex Tupaia sp. No 611, VIETNAM: Nghe An Province, Phu Quy, 15 September 1956, coll. I.M. Grochovskaja. One paralectotype larva (ZMMU Tdt-3144, specimen 2) with same data as lectotype; one paralectotype larva (ZMMU Tdt-3143, specimen 3) ex Tupaia sp. No 610, other data same as for lectotype.</p> <p>Remarks. According to the cluster analysis performed by Stekolnikov (2013), L. arvinum belongs to the arvinum species group, which includes also L. saigoni Stekolnikov, 2013. The latter was described as Leptotrombidium (Leptotrombidium) miculum arvinum var. saigoni Vercammen-Grandjean and Langston, 1976, from a single specimen, and raised to species by Stekolnikov (2013). This species differs from L. arvinum by the presence of 10 setae in 1 st posthumeral row, vs. 8, NDV = 58 vs. 49–52, and by slightly longer legs (Ip = 732 vs. 601–625).</p> <p>Vercammen-Grandjean &amp; Langston (1976) included L. arvinum in L. miculum, as a subspecies. However, according to Stekolnikov (2013), L. miculum belongs to the arenicola species group. The latter species differs from L. arvinum in having the sensillary bases situated at the level of PL vs. posterior to PL, shorter scutal setae (AM 32–38, AL 27–32, and PL 40–46, vs. 44–50, 35–39, 59–64, respectively), shorter idiosomal setae (H 41–43, D min 31–32, D max 37–40, and V max 31–33 vs. 60–61, 34–40, 52–55, and 33–45, respectively), and slightly shorter legs (Ip 554–602 vs. 601–625).</p> </div>	http://treatment.plazi.org/id/2A6687ABFFF18355EDDDF0A2FE10ACA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFF6835FEDDDF2E1FBEBAA36.text	2A6687ABFFF6835FEDDDF2E1FBEBAA36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium globosum (Schluger 1960)	<div><p>Leptotrombidium globosum (Schluger, 1960)</p> <p>(Figs. 3–5)</p> <p>Trombicula (Leptotrombidium) globosa Schluger, 1960 (in Schluger et al. 1960b): 1796, fig. 5.</p> <p>Leptotrombidium (Leptotrombidium) globosum: Lakshana 1973: 7; Vercammen-Grandjean &amp; Langston 1976: 337, pl. 55; Kudryashova 2004: 19; Chau et al. 2007: 47, fig. 19.</p> <p>Leptotrombidium globosum: Stekolnikov 2013: 119; 2021: 117.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNN; fSc: PL&gt; AM&gt; AL; Ip = 746–776; fD = 2H-(10–12)-(15–22)+(53–61); DS = 85–90; V = 67–73; NDV = 152–162. Standard measurements of type series given in Table 2.</p> <p>Description (larva) (based on lectotype and 3 paralectotypes, fD recorded in three specimens). IDIOSOMA (Figs. 3A, 4B–G). Eyes 2 + 2; 85–90 barbed dorsal idiosomal setae, including one pair of humeral setae, 10–12 setae in 1 st posthumeral row (C except humeral setae), 15–22 setae in 2 nd posthumeral row (D) plus 53–61 setae with unclear distribution by rows (sometimes 3 rd row, E, with 26–28 setae, recognized); 4 sternal setae; 67–73 ventral setae; NDV = 152–162.</p> <p>GNATHOSOMA (Fig. 5A, B). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base and palpal femur covered with puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude; dorsal palpal tibial seta branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 3, 4A). Nearly rectangular, moderately covered with rather large puncta, posterior margin broadly rounded; AM posterior to level of AL, sensillary (trichobothrial) bases at level of PL or slightly posterior (P-PL – PSB = 1–3 µm); all scutal setae barbed similarly to dorsal idiosomal setae; PL ≥ AM&gt; AL; flagelliform sensilla (trichobothria), with about 7–8 long branches in distal half.</p> <p>LEGS (Fig. 5C–E). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Leopoldamys sabanus (Thomas) (= Rattus sabanus) (Schluger et al. 1960b). Vercammen-Grandjean &amp; Langston (1976) added to the list of hosts Leopoldamys edwardsi (Thomas) [= Rattus grochovskiae D. Tien (nomen nudum)] and Tupaia belangeri chinensis (Anderson) (= T. glis modesta), based on examination of syntypes. Later it was also recorded from Thailand, ex Mus sp., Niviventer niviventer (= Rattus niviventer), and T. glis (Lakshana 1973).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3276, specimen 4, designated here) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 36, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 2 February 1956, coll. I.M. Grochovskaja. Three paralectotypes larvae (ZMMU Tdt- 3260, specimen 1; ZMMU Tdt-3261, specimen 1; ZMMU Tdt-3262, specimen 5) with same data as lectotype.</p> <p>Remarks. Leptotrombidium globosum belongs to the magnum species group, which includes, in total, 15 species (Stekolnikov 2013). It differs from L. magnum (Schluger, 1960) (and two species synonymized with it in the present paper, L. dooleyi Nadchatram, 1970 and L. submagnum Wang, Li and Shi, 1988) in having the posterior scutal margin evenly rounded from the level of PL vs. greatly projected posterior to PL and straight in the middle part, the lesser number of setae in 1 st posthumeral row (10–12 vs. 14–17), greater NDV (152–162 vs. 118–130); smaller scutum (PW 74–77, SB 30–33, and SD 46–49 vs. 79–89, 33–39, and 51–58, respectively); PL&gt; AM vs. AM ≥ PL, and shorter legs (Ip = 746–776 vs. 851–946).</p> <p>Leptotrombidium globosum differs from L. keukenschrijveri (Walch, 1924) in having the evenly rounded posterior margin of scutum (vs. slightly bilobate and concave in center); by PL&gt; AM vs. AM&gt; PL; by greater NDV (152–162 vs. 114), by 10–12 setae in 1 st posthumeral row vs. 16 (8 + 8), and by larger scutum (AW 64–68, PW 74–77, and SD 46–49 vs. 59–61, 69–72, and 38–41, respectively).</p> <p>Remaining species of the magnum group were described in 1980s–1990s in China (L. arctonycis Xiang and Wen, 1984, L. baoshui Wen and Xiang, 1984, L. biji Wen and Xiang, 1984, L. caudatum Wen, Zhou, Chen, Wang and Zhang, 1984, L. huangchuanense Yang, 1994, L. kunshui Wen and Xiang, 1984, L. shuiqui Liao and Yuan, 1998, L. shuminense Zhang, Deng and Wang, 1996, L. sixinum Wen, Zhou, Chen, Wang and Zhang, 1984, and L. xianglinense Wen, 1984). Two of them, L. arctonycis and L. xianglinense, are known only from the holotypes. Differences of all these species from L. globosum, L. magnum, and L. keukenschrijveri require further investigation.</p> <p>Among these species from China, L. sixinum is the most probable candidate for the synonymization with L. globosum. The former species differs from L. globosum in PW 83–85 vs. 74–77, AM ≥ PL vs. PL&gt; AM, lesser NDV (132 vs. 152–162), and in slightly longer legs (Ip 835 vs. 746–776). Statistical and taxonomic significance of these differences can not be estimated without a direct comparison of the type series. Two other species, L. biji and L. caudatum, are also similar to L. globosum by the shape of scutum, but differ in its greater size (in L. biji, AW 70–80, PW 85–103, AP 33–38; in L. caudatum, AW 73–79, PW 85–88, ASB 35–42; in L. globosum, AW 64–68, PW 74–77, ASB 32–33, AP 26–30) and in AM ≥ PL vs. PL&gt; AM. Leptotrombidium biji also differs from L. globosum in much longer legs (Ip 1015–1049 vs. 746–776); L. caudatum has longer setae (AM 64–75, AL 48–56, PL 65–72, H 73–75 vs. 52–56, 44–47, 56–61, and 60–65, respectively) and lesser NDV (116 vs. 152–162).</p> </div>	http://treatment.plazi.org/id/2A6687ABFFF6835FEDDDF2E1FBEBAA36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFFA835CEDDDF1A7FF7AAC8D.text	2A6687ABFFFA835CEDDDF1A7FF7AAC8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium gracipalpe (Schluger 1960)	<div><p>Leptotrombidium gracipalpe (Schluger, 1960)</p> <p>(Figs. 6, 7)</p> <p>Trombicula (Leptotrombidium) gracipalpis Schluger, 1960 (in Schluger et al. 1960b): 1793, fig. 3.</p> <p>Leptotrombidium (Leptotrombidium) gracipalpe: Vercammen-Grandjean &amp; Langston 1976: 520, pl. 137; Kudryashova 2004: 19; Chau et al. 2007: 48, fig. 20.</p> <p>Leptotrombidium gracipalpe: Stekolnikov 2013: 80; 2021: 117.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/B/BNN; fSc: PL&gt; AM&gt; AL; Ip = 574–643; fD = 2H-8-6-6-4(6)-4; DS = 30–32; V = 24–38; NDV = 56–68. Standard measurements of type series given in Table 3.</p> <p>Description (larva) (based on lectotype and 12 paralectotypes, fD recorded in four specimens). IDIOSOMA (Figs. 6A, 7B–G). Eyes 2 + 2; 30–32 barbed dorsal idiosomal setae, including one pair of humeral setae, with distribution by rows 2H-8-6-6-4(6)-4; 4 sternal setae; 24–38 ventral setae; NDV = 56–68.</p> <p>GNATHOSOMA (Fig. 7H, I). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base covered with puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; seta on palpal femur nude, seta on palpal genu branched, dorsal palpal tibial seta branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 6, 7A). Rectangular, moderately covered with rather large puncta, greatly projected posterior to PL, posterior margin almost straight in middle part; AM posterior to level of AL, sensillary (trichobothrial) bases posterior to level of PL (P-PL – PSB = 1–5 µm); all scutal setae barbed similarly to dorsal idiosomal setae; PL&gt; AM&gt; AL; flagelliform sensilla (trichobothria) densely covered with small cilia in proximal half and with about 11 long branches in distal half.</p> <p>LEGS (Fig. 7J–L). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) at level or slightly posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Leopoldamys sabanus, Rattus tanezumi Temminck (= R. rattus tikos), and Tupaia belangeri chinensis (= T. glis modesta) (Schluger et al. 1960b).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3270, specimen 4, designated here) ex Tupaia belangeri chinensis (labeled as T. glis and T. ferruginea) No 29, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 1 February 1956, coll. I.M. Grochovskaja. Twelve paralectotypes larvae (ZMMU Tdt-3270, specimen 2; ZMMU Tdt-3271, specimens 3, 5; ZMMU Tdt-3272, specimen 2; ZMMU Tdt-3273, specimens 1, 3, 4, 5; ZMMU Tdt-3274, specimens 1, 4; ZMMU Tdt-3275, specimen 5; ZMMU Tdt-3266, specimen 6) with same data as lectotype.</p> <p>Remarks. Leptotrombidium gracipalpe belongs to the orientale group, which includes, in total, 17 species (Stekolnikov 2013). This species is unique in its palpal formula fPp = N/B/BNN, i.e. the palpal genual seta is branched that is opposed to all other members of the group and to the overwhelming majority of other Leptotrombidium species.</p></div> 	http://treatment.plazi.org/id/2A6687ABFFFA835CEDDDF1A7FF7AAC8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFFF8346EDDDF1C9FDF4AA12.text	2A6687ABFFFF8346EDDDF1C9FDF4AA12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium horridum (Schluger 1960)	<div><p>Leptotrombidium horridum (Schluger, 1960)</p> <p>(Figs. 8–10)</p> <p>Trombicula (Leptotrombidium) horrida Schluger, 1960 (in Schluger et al. 1960b): 1790, fig. 1.</p> <p>Leptotrombidium (Leptotrombidium) horridum: Vercammen-Grandjean &amp; Langston 1976: 560, pl. 152.</p> <p>Leptotrombidium horridum: Stekolnikov 2013: 88; 2021: 118.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNN; fSc: PL&gt; AM&gt; AL; Ip = 851–990; fD = 2H-(9–11)-8-(7–11)-6+(8–12); DS = 42–48; V = 38–46; NDV = 83–93. Standard measurements of type series given in Table 4.</p> <p>Description (larva) (based on lectotype and 24 paralectotypes, fD recorded in four specimens). IDIOSOMA (Figs. 8A, 9B–G). Eyes 2 + 2; 42–48 barbed dorsal idiosomal setae, including one pair of humeral setae, 9–11 setae in 1 st posthumeral row (C except humeral setae), 8 setae in 2 nd row (D), 7–11 setae in 3 rd row (E), 6 setae in 4 th row (F) plus 8–12 caudal setae; 4 sternal setae; 38–46 ventral setae; NDV = 83–93.</p> <p>GNATHOSOMA (Fig. 10A, B). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base and palpal femur covered with puncta, palpal genu with few puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude, dorsal palpal tibial seta branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 8, 9A). Rectangular, moderately covered with rather large puncta, greatly projected posterior to PL, posterior margin slightly concave in middle part; AM posterior to level of AL, sensillary (trichobothrial) bases at level of PL or slightly posterior (P-PL – PSB = 0–6 µm); all scutal setae barbed similarly to dorsal idiosomal setae; PL ≥ AM&gt; AL; flagelliform sensilla (trichobothria) with about 7 branches in distal half.</p> <p>LEGS (Fig. 10C–E).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 14B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Leopoldamys sabanus (= Rattus sabanus) and Tupaia belangeri chinensis (= T. glis modesta) (Schluger et al. 1960b). Vercammen- Grandjean &amp; Langston (1976) added also Leopoldamys edwardsi (Thomas) [= Rattus grochovskiae (nomen nudum)] and Tupaia glis (= T. ferruginea) to the list of hosts, on the base of examination of syntypes.</p> <p>Type material examined. Lectotype larva (ZMMU L. horridum _04, specimen 5, designated here) ex Leopoldamys edwardsi [labeled as R. grochovskii (nomen nudum)] No 36, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 2 February 1956, coll. I.M. Grochovskaja. Eight paralectotypes larvae (ZMMU L. horridum _01, specimens 1–3; ZMMU L. horridum _06, specimen 2; ZMMU L. horridum _08, specimens 4, 5; ZMMU L. horridum _09, specimens 2, 5; ZMMU Tdt-3248, specimen 2) ex L. edwardsi [labeled as R. grochovskii (nomen nudum)] No 40, 3 February 1956, other data same as for lectotype; three paralectotypes larvae (ZMMU Tdt-3245, specimen 5; ZMMU Tdt-3246, specimens 2, 3) ex Tupaia belangeri [labeled as T. glis and T. hongaiensis (nomen nudum)] No 42, 3 February 1956, other data same; one paralectotype larva (ZMMU Tdt-3249, specimen 2) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 44, 6 February 1956, other data same; one paralectotype larva (ZMMU Tdt-3250, specimen 4) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 45, 6 February 1956, other data same; five paralectotypes larvae (ZMMU Tdt-3260, specimen 2; ZMMU Tdt-3261, specimens 3, 4, 6; ZMMU Tdt-3262, specimen 1) with same data as lectotype; five paralectotypes larvae (ZMMU Tdt-3264, specimen 4; ZMMU Tdt-3265, specimens 3, 5; ZMMU Tdt-3268, specimen 5; ZMMU Tdt-3272, specimen 3) ex Tupaia belangeri (labeled as T. glis and T. ferruginea) No 29, 1 February 1956, other data same.</p> <p>Remarks. Leptotrombidium horridum belongs to the alopeciatum group, which includes five species (Stekolnikov 2013). It differs from L. spicatum Traub, 1960, L. baluense (Traub and Audy, 1954), and L. tupaianum Vercammen-Grandjean and Langston, 1976 in having the PL and dorsal idiosomal setae covered with usual uniform barbs vs. bearing 2–3 rows of short barbs and two lateral rows of large spikes. Leptotrombidium horridum differs from L. alopeciatum by the presence of nude ventral palpal tibial setae vs. branched (fPp = N/N/BNN vs. N/N/ BNB); by nearly doubled number of idiosomal setae (DS 42–48, V 38–46, and NDV 83–93 vs. 22, 22, and 44, respectively); by PL&gt; AM vs. AM&gt; PL; by sensillary bases situated at the level or posterior to SB vs. anterior; and by longer legs (Ip = 851–990 vs. 718–745).</p> </div>	http://treatment.plazi.org/id/2A6687ABFFFF8346EDDDF1C9FDF4AA12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFE38342EDDDF183FDB2ABA2.text	2A6687ABFFE38342EDDDF183FDB2ABA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium magnum (Schluger 1960) Nagayo, Miyagawa, Mitamura and Imamura 1916	<div><p>Leptotrombidium magnum (Schluger, 1960)</p> <p>(Figs. 11–13)</p> <p>Trombicula (Leptotrombidium) magna Schluger, 1960 (in Schluger et al. 1960b): 1796, fig. 4.</p> <p>Leptotrombidium (Leptotrombidium) magnum: Lakshana 1973: 8; Vercammen-Grandjean &amp; Langston 1976: 332, pl. 52; Kudryashova 2004: 25.</p> <p>Leptotrombidium magnum: Stekolnikov 2013: 120; 2021: 122.</p> <p>Leptotrombidium dooleyi Nadchatram, 1970: 150, figs. 17–24, syn. nov.; Vercammen-Grandjean &amp; Langston 1976: 338, pl. 56; Stekolnikov 2013: 120.</p> <p>Leptotrombidium (Leptotrombidium) submagnum Wang, Li and Shi, 1988: 140, figs. 9–12, syn. nov.; Li et al. 1997: 178, fig. 2-1-101; Stekolnikov 2013: 119.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNN; fSc: AM ≥ PL&gt; AL; Ip = 851–946; fD = 2H-(14–15)-(12–17)-(17–18)-(10–11)-(6–8)+(4–9); DS = 71–74; V = 55–56; NDV = 126–130. Standard measurements of type series given in Table 5.</p> <p>Description (larva) (based on lectotype and 17 paralectotypes, fD recorded in two specimens). IDIOSOMA (Figs. 11A, 12B–G). Eyes 2 + 2; 71–74 barbed dorsal idiosomal setae, including one pair of humeral setae, 14–15 setae in 1 st posthumeral row (C except humeral setae), 12–17 setae in 2 nd row (D), 17–18 setae in 3 rd row (E), 10–11 setae in 4 th row (F), 6–8 setae in 5 th row plus 4–9 caudal setae; 4 sternal setae; 55–56 ventral setae; NDV = 126–130.</p> <p>GNATHOSOMA (Fig. 13A–C). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base and palpal femur covered with puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude, dorsal palpal tibial seta branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 11, 12A). Rectangular, moderately covered with rather large puncta, greatly projected posterior to PL, posterior margin straight in middle part; AM posterior to level of AL, sensillary (trichobothrial) bases at level of PL or slightly posterior (P-PL – PSB = –5–4 µm); all scutal setae barbed similarly to dorsal idiosomal setae; AM ≥ PL&gt; AL; flagelliform sensilla (trichobothria) with about 5 branches in distal part.</p> <p>LEGS (Fig. 13D–G). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) at level of tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Rattus tanezumi Temminck (= R. rattus tikos), Leopoldamys sabanus (= Rattus sabanus) and Tupaia belangeri chinensis (= T. glis modesta) (Schluger et al. 1960b). Later it was recorded in Thailand, from birds Napothera macrodactyla (Strickland) and Stachyris poliocephala (Temminck) (Lakshana 1973). Vercammen-Grandjean &amp; Langston (1976) added also Tupaia glis (= T. ferruginea) to the list of hosts, based on examination of syntypes.</p> <p>Leptotrombidium dooleyi was described from Nepal, Daman, ex field mouse (Apodemus sp.), and from Nepal, Bokaikunde, 23 km NE Trisuli, ex Rattus nitidus (Hodgson). Leptotrombidium submagnum was described from China, Hubei Province, Yichang County, Jiangjiamiao, ex Rattus losea (Swinhoe).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3268, specimen 3, designated here) ex Tupaia belangeri (labeled as T. glis and T. ferruginea) No 29, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 1 February 1956, coll. I.M. Grochovskaja. Five paralectotypes larvae (ZMMU Tdt-3262, specimen 3; ZMMU Tdt-3276, specimens 1, 2, 3, 5) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 36, 2 February 1956, other data same; one paralectotype larva (ZMMU Tdt-3263, specimen 4) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 35, other data same; 11 paralectotypes larvae (ZMMU Tdt-3265, specimens 1, 4; ZMMU Tdt-3266, specimens 1, 2, 4; ZMMU Tdt-3267, specimen 5; ZMMU Tdt-3268, specimens 4, 6; ZMMU Tdt-3272, specimens 1, 4, 5) with same data as lectotype.</p> <p>Remarks. This species is a member of the magnum group (Stekolnikov 2013). Its differences from L. globosum are provided above. Leptotrombidium magnum differs from L. keukenschrijveri by a larger scutum (AW 69–79, PW 79–89, and SD 51–58 vs. 59–61, 69–72, and 38–41, respectively) and by longer legs (Ip 851–946 vs. 702–738).</p> <p>There are no evident differences between L. magnum and L. dooleyi, except for a longer AM (66–72 vs. 51–62) and, consequently, AM&gt; PL vs. AM ≥ PL in the latter species that was used for discrimination of these species in the key published by Stekolnikov (2013) (couplet 270). Therefore, we synonymize here L. dooleyi with L. magnum.</p> <p>Leptotrombidium submagnum is also definitely identical to L. magnum. The characters of the former species given by its authors to discriminate it from L. magnum and L. dooleyi were as follows: PL&gt; AM&gt; AL, PW/AP = 2.5, and 16–17 setae in 1 st posthumeral row (Wang et al. 1988). According to our data, PW/AP = 2.47–2.89 (mean 2.69) in the type series of L. magnum. The number of setae in 1 st posthumeral row was 14 and 15 in the two paralectotypes with fD recorded. Taking into account a high level of variation in the arrangement of idiosomal setae in all chigger species with large DS, the difference between 14–15 and 16–17 cannot be estimated as taxonomically significant. The difference in the length of PL (50–59 in L. magnum and 62–66 in L. submagnum) that resulted in AM ≥ PL vs. PL&gt; AM is also not weighty. In addition, we can note that NDV is 118–125 in L. submagnum and 126–130 in L. magnum. This slight difference does not constitute a basis for species discrimination. Therefore, we synonymize here L. submagnum with L. magnum.</p> </div>	http://treatment.plazi.org/id/2A6687ABFFE38342EDDDF183FDB2ABA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFE7834EEDDDF12AFAEEA94A.text	2A6687ABFFE7834EEDDDF12AFAEEA94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotrombidium monstrosum (Schluger 1960)	<div><p>Leptotrombidium monstrosum (Schluger, 1960)</p> <p>(Figs. 14–16)</p> <p>Trombicula (Leptotrombidium) monstrosa Schluger, 1960 (in Schluger et al. 1960b): 1798, fig. 6.</p> <p>Leptotrombidium (Leptotrombidium) monstrosa: Traub &amp; Lakshana 1966: 282.</p> <p>Leptotrombidium (Leptotrombidium) monstrosum: Lakshana 1973: 8; Vercammen-Grandjean &amp; Langston 1976: 569, pl. 155; Kudryashova 2004: 27; Chau et al. 2007: 54, fig. 23.</p> <p>Leptotrombidium monstrosum: Stekolnikov 2013: 47; 2021: 123.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNB; fSc: AM ≥ PL&gt; AL; Ip = 708–784; fD = 2H-8-6-6-4-2-2; DS = 30; V = 30–33; NDV = 60–63. Standard measurements of type series given in Table 6.</p> <p>Description (larva) (based on lectotype and 21 paralectotypes, fD recorded in three specimens). IDIOSOMA (Figs. 14A, 15B–G). Eyes 2 + 2; 30 barbed dorsal idiosomal setae, including one pair of humeral setae, arranged 2H-8-6-6-4-2-2; 4 sternal setae; 30–33 ventral setae; NDV = 60–63.</p> <p>GNATHOSOMA (Fig. 16A–E). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base covered with puncta, palpal femur with few puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude, dorsal palpal tibial seta branched, lateral palpal tibial seta nude; ventral palpal tibial seta branched; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 14, 15A). Rectangular, moderately covered with rather large puncta, projected posterior to PL, posterior margin slightly concave in middle part; AM posterior to level of AL, sensillary (trichobothrial) bases at level of PL or slightly anterior (P-PL – PSB = –4–1 µm); all scutal setae barbed similarly to dorsal idiosomal setae; AM ≥ PL&gt; AL; flagelliform sensilla (trichobothria) with about 8 branches in distal part.</p> <p>LEGS (Fig. 16F–H).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Leopoldamys sabanus (= Rattus sabanus) and Tupaia belangeri chinensis [= T. glis modesta; = T. hongaiensis (nomen nudum)] (Schluger et al. 1960b). Later on, it was recorded in Thailand (Lakshana 1973) from Menetes berdmorei. Vercammen-Grandjean &amp; Langston (1976) added also Leopoldamys edwardsi (Thomas) [= R. grochovskiae (nomen nudum)] to the list of hosts based on examination of syntypes. Chau et al. (2007) also reported this species from Atherurus macrourus (L.), Callosciurus erythraeus (Pallas), Dremomys rufigenis, Rattus andamanensis (Blyth) (= R. koratensis), Tamiops rodolphii (Milne-Edwards), Hylomys suillus, Paradoxurus hermaphroditus, and birds Gallus gallus domesticus (L.) and Francolinus pintadeanus (Scopoli), probably, from Vietnam.</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3250, specimen 1, designated here) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 45, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 6 February 1956, coll. I.M. Grochovskaja. Twelve paralectotypes larvae (ZMMU Tdt-3244, specimens 1–5; ZMMU Tdt-3245, specimens 1, 2, 4, 6; ZMMU Tdt-3246, specimens 1, 4, 5) ex Tupaia belangeri chinensis [labeled as T. glis and T. hongaiensis (nomen nudum)] No 42, 3 February 1956, other data same as for lectotype; one paralectotype larva (ZMMU Tdt-3248, specimen 4) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 40, 3 February 1956, other data same as for lectotype; one paralectotype larva (ZMMU Tdt-3249, specimen 1) ex L. edwardsi [labeled as Rattus sp. and R. grochovskii (nomen nudum)] No 44, other data same as for lectotype; five paralectotypes larvae (ZMMU Tdt-3250, specimens 2, 3, 5, 6, 7) with same data as lectotype; two paralectotypes larvae (ZMMU Tdt-3268, specimens 1, 2) ex Tupaia belangeri (labeled as T. glis and T. ferruginea) No 29, 1 February 1956, other data same.</p> <p>Remarks. This species is a member of the scanloni group (Stekolnikov 2013), which includes three species.</p> <p>Leptotrombidium monstrosum differs from L. scanloni Traub and Lakshana, 1966 in having the ventral palpal tibial seta branched (fPp = N/N/BNB vs. N/N/BNN); sensillary bases situated at the level of PL or slightly anterior to them vs. posterior; and larger scutum (AW 67–73, PW 78–89, and AP 26–30 vs. 62–66, 73–78, and 23–25, respectively). Leptotrombidium monstrosum differs from L. hanseni Traub and Lakshana, 1966 in having a slightly lesser number of idiosomal setae (DS 30, V 30–33, and NDV 60–63 vs. 34, 34, and 68, respectively); fD = 2H-8-6-6-4-2-2 vs. 2H-8-6-6-6-4-2; and shorter legs (Ip 708–784 vs. 847). Taxonomical significance of these rather slight differences is unclear, since no data on the variation of NDV and Ip are available for L. hanseni. Probably, the latter species should be synonymized with L. monstrosum, but this suggestion needs a verification based on a larger material.</p> <p>Traub &amp; Lakshana (1966) stated that L. hanseni differs from L. monstrosum, in addition to the difference by fD, also by a smaller scutum (PW 84, SB 36, and AP 25 vs. 100, 45, and 33, respectively), but our measurements of L. monstrosum do not support that observation (Table 6). The source of the values provided by Traub &amp; Lakshana as measurements of L. monstrosum is unclear. These data do not coincide with those in the original description (Schluger et al. 1960b), and these authors did not mention the examination of any material on L. monstrosum.</p> </div>	http://treatment.plazi.org/id/2A6687ABFFE7834EEDDDF12AFAEEA94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFEB834AEDDDF2E3FDEDA862.text	2A6687ABFFEB834AEDDDF2E3FDEDA862.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lorillatum attapinum (Schluger, Grochovskaja, Ngu, Hoe and Tung 1963)	<div><p>Lorillatum attapinum (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963)</p> <p>(Figs. 17–19)</p> <p>Trombicula attapina Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963: 692, figs. 8–15.</p> <p>Lorillatum attapinum: Vercammen-Grandjean &amp; Langston 1976: 947; Kudryashova 2004: 14; Stekolnikov 2021: 131. Lorillatum attapina: Chau et al. 2007: 71, figs. 33, 34.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0201; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/N/NNb; fSc: PL&gt; AM&gt; AL; Ip = 924–1056; fD = 2H-[8-(1–3)]-10-10+18; DS = 51–56; V = 34–40; NDV = 87–91. Standard measurements of type series given in Table 7.</p> <p>Description (larva) (based on lectotype and 4 paralectotypes, fD recorded in one specimen). IDIOSOMA (Figs. 17A, 18B–G). Eyes 2 + 2; 51–56 barbed dorsal idiosomal setae, including one pair of humeral setae, 1 st posthumeral row (C except humeral setae) double, with 8 anterior and 3 posterior setae, 2 nd row (D) and 3 rd row (E) with 10 setae each plus 18 setae with unclear distribution by rows; 4 sternal setae; 34–40 ventral setae; NDV = 87–91.</p> <p>GNATHOSOMA (Fig. 19A, B). Cheliceral blade with tricuspid cap; gnathobase densely covered with small puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base and palpal femur sparsely covered with puncta, palpal genu with few puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; seta on palpal femur with few branches, seta on palpal genu nude, dorsal and lateral palpal tibial setae nude, ventral palpal tibial seta with one branch; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 17, 18A). Rectangular, densely covered with small puncta, with prominent anterolateral shoulders anterior to AL, anterior scutal margin concave, posterior margin bilobate; AM at level of AL, sensillary (trichobothrial) bases slightly anterior to level of PL (PSB – P-PL = 1–3 µm); all scutal setae barbed similarly to dorsal idiosomal setae; PL very long, pointed; PL&gt; AM&gt; AL; flagelliform sensilla (trichobothria) with one branch or nude.</p> <p>LEGS (Fig. 19C–J).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 23B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω) II shorter and slender than tarsala I, microtarsala (ε) posterior to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 2B and one very long flagelliform seta with few cilia in basal part; genu 3B, genuala; tibia 4B and two long flagelliform setae, nude or with few cilia, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Ha Giang Province, ex Crocidura sp. [= Crocidura leucrura (nomen nudum? lapsus calami?)] (Schluger et al. 1963).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3141, specimen 4, designated here) ex Crocidura sp. No 218, VIETNAM: Ha Giang Province, Ha Giang, 25 May 1956, coll. I.M. Grochovskaja. Three paralectotypes larvae (ZMMU Tdt-3141, specimen 5; ZMMU Tdt-3140, specimens 1, 5) with same data as lectotype; one paralectotype larva (ZMMU Tdt-3142, specimen 1) ex Crocidura sp. No 247, 1 June 1956, other data same as for lectotype.</p> <p>Remarks. The genus Lorillatum includes 11 species distributed in Southeast and East Asia (Vercammen- Grandjean &amp; Langston 1976; Li et al. 1997; Stekolnikov 2021). Lorillatum attapinum differs from all other species by the absence of palpal subterminala (ζ). Vercammen-Grandjean &amp; Langston (1976), who did not see any material on this species, indicated the presence of the palpal subterminala in the diagnosis of L. attapinum obviously by a mistake. On the contrary, these authors did not notice the subterminala in L. mastigophorum (Nadchatram, 1963), as evident from their description and figures, although this seta is present according to the original description (Nadchatram 1963). Lorillatum attapinum differs from L. mastigophorum, in addition to the absence of palpal subterminala, by the presence of one mastifemorala and two mastitibialae, vs. one mastifemorala, one mastigenuala, one mastitibiala, and one mastitarsala. However, stability of these characters is not evident, since the level of ciliation of mastisetae can vary at least in L. attapinum. Thus, Schluger et al. (1963) reported on the presence of one mastifemorala, one mastigenuala, and one mastitibiala in this species, but did not mention mastitarsala. Measurements and numbers of idiosomal setae in these species are very similar.</p> <p>Lorillatum attapinum also shares its fPp = B/N/NNB(N) with L. kianjoei (Nadchatram and Traub, 1964) and L. panitae (Nadchatram and Traub, 1964), but differs from L. kianjoei in having the greater number of idiosomal setae (DS = 51–56 and fD = 2H-[8-2]-10-… vs. 42–45 and 2H-8-8-…) and the presence of at most one mastigenuala vs. two, and from L. panitae in having shorter legs (Ip = 924–1056 vs. 1248), lesser number of idiosomal setae (two humeral setae vs. four, fD = 2H-[8-2]-10-… vs. 4H-[10-2]-13-…, NDV = 87–91 vs. 112), and rounded anterolateral shoulders of scutum vs. sharply angulate.</p> </div>	http://treatment.plazi.org/id/2A6687ABFFEB834AEDDDF2E3FDEDA862	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
2A6687ABFFEF8376EDDDF39BFE6EAAA5.text	2A6687ABFFEF8376EDDDF39BFE6EAAA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trombiculindus griselda (Schluger, Grochovskaja, Ngu, Hoe and Tung 1963)	<div><p>Trombiculindus griselda (Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963)</p> <p>(Figs. 20–22)</p> <p>Trombicula (Trombiculindus) griselda Schluger, Grochovskaja, Ngu, Hoe and Tung, 1963: 699, figs. 36–42.</p> <p>Leptotrombidium (Trombiculindus) griseldae: Vercammen-Grandjean &amp; Langston 1976: 641, pl. 182.</p> <p>Leptotrombidium (Trombiculindus) griselda: Chau et al. 2007: 63, fig. 29.</p> <p>Trombiculindus griseldae: Kudryashova 2004: 20.</p> <p>Trombiculindus griselda: Stekolnikov 2021: 145.</p> <p>Diagnosis. SIF = 7B-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = N/N/BNN; fSc: AM ≥ PL&gt; AL; Ip = 713–803; fD = 2H-8-6-6-2-4-2(4-2-2); DS = 24–30; V = 21–27; NDV = 45–56. Standard measurements of type series given in Table 8.</p> <p>Description (larva) (based on lectotype and 13 paralectotypes, fD recorded in one specimen). IDIOSOMA (Figs. 20A, 21B–H). Eyes 2 + 2; 24–30 dorsal idiosomal setae, including one pair of humeral setae, arranged 2H- 8-6-6-2-4-2 (posterior 3 rows can be also recorded as 4-2-2); 4 sternal setae; 21–27 ventral setae; NDV = 45–56. Humeral and dorsal idiosomal setae broadly foliate, ovate, with pointed end, irregularly covered with thick scalelike setules; caudal and postanal setae less expanded, lanceolate, sparsely covered with small scalelike setules; preanal setae setiform, covered with long thin barbs.</p> <p>GNATHOSOMA (Fig. 22A, B). Cheliceral blade with tricuspid cap; gnathobase covered with puncta and bears 1 pair of branched gnathocoxal (tritorostral) setae; cheliceral base and palpal femur covered with puncta; galeal (deutorostral) seta branched; palpal claw with 3 prongs; setae on palpal femur and genu nude, dorsal palpal tibial setae branched, lateral and ventral palpal tibial setae nude; palpal tarsus with 7 branched setae and tarsala (ω).</p> <p>SCUTUM (Figs. 20, 21A). Nearly hexagonal, sparsely covered with small puncta, greatly projected posterior to PL, bases of AL and PL situated close to each other, posterior margin slightly concave in middle part; AM far posterior to level of AL, sensillary (trichobothrial) bases far posterior to level of PL (P-PL – PSB = 10–14 µm); AM and AL slightly expanded, covered with large barbs, PL broadly foliate, ovate, with pointed end, irregularly covered with thick scalelike setules; AM ≥ PL&gt; AL; flagelliform sensilla (trichobothria) nude or having few branches in distal part.</p> <p>LEGS (Fig. 22C–E).All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 2 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) proximal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) not observed, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 15B.</p> <p>Distribution and hosts. This species was described from North Vietnam, Quang Ninh Province, ex Rattus rattus (= R. rattus alexandrinus) and Leopoldamys edwardsi (Thomas) [= R. grochovskiae (nomen nudum)] (Schluger et al. 1963).</p> <p>Type material examined. Lectotype larva (ZMMU Tdt-3151, specimen 1, designated here) ex L. edwardsi [labeled as R. grochovskii (nomen nudum)] No 44, VIETNAM: Quang Ninh Province, Ha Lam District, Ha Long (= Hon Gai), 6 February 1956, coll. I.M. Grochovskaja. Seven paralectotypes larvae (ZMMU Tdt-3147, specimen 1; ZMMU Tdt-3148, specimens 2, 3, 4; ZMMU Tdt-3263, specimens 1, 2, 3) ex L. edwardsi [labeled as R. grochovskii (nomen nudum)] No 35, 2 February 1956, other data same as for lectotype; five paralectotypes larvae (ZMMU Tdt- 3149, specimens 1, 2, 3, 4, 6) ex L. edwardsi [labeled as R. grochovskii (nomen nudum)] No 37, other data same as for lectotype; one paralectotype larva (ZMMU Tdt-3150, specimen 1) ex L. edwardsi [labeled as R. grochovskii (nomen nudum)] No 39, 3 February 1956, other data same as for lectotype.</p> <p>Remarks. Trombiculindus includes 59 species distributed in Southeast, East, South, and Central Asia (Vercammen-Grandjean &amp; Langston 1976, Li et al. 1997, Stekolnikov 2021). Trombiculindus griselda shares the most part of characters with T. cuneatum Traub and Evans, 1951. The setation of palps and legs, number and arrangement of idiosomal setae, shape and measurements of scutum are almost identical in these species. Both species have slightly expanded AM and AL, but broadly foliate PL, H, and other dorsal idiosomal setae. Trombiculindus griselda differs from T. cuneatum in having foliate setae shorter and, therefore, relatively wider (PL 47–54, H 48–58, D max 53–64 vs. 60–70, 58–68, and 61–70, respectively) and irregularly covered with thick scalelike setules (vs. foliate setae with 8–9 longitudinal rows of setules); shorter AM (51–56 vs. 63–64) and AM&gt; PL vs. PL&gt; AM; and sensilla nude or with a few branches in distal half (vs. sensilla covered with small cilia in proximal part and up to 29–32 branches in distal two thirds).</p> </div>	http://treatment.plazi.org/id/2A6687ABFFEF8376EDDDF39BFE6EAAA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Stekolnikov, Alexandr A.;Antonovskaia, Anastasia A.	Stekolnikov, Alexandr A., Antonovskaia, Anastasia A. (2021): Re-descriptions of eight chigger mite species (Acariformes: Trombiculidae) of the Leptotrombidium generic complex from Vietnam. Zootaxa 5057 (3): 329-363, DOI: https://doi.org/10.11646/zootaxa.5057.3.2
