taxonID	type	description	language	source
03F48795DC5FFFB4A6C9A50FFBACFE80.taxon	description	Description BODY (Fig. 1). Medium-sized species, length 8 – 8.4 mm, humeral width 5.1 – 5.2 mm. HEAD (Fig. 2). Eyes large, inter-ocular distance seven to nine times width of one eye. Clypeal median emargination broadly u-shaped. Clypeal teeth separated approximately by 1.5 times the basal width of a tooth. Anterior margin of the clypeus, between clypeal teeth, concave and expanded posteriorly into triangular shape. PRONOTUM (Fig. 3). Margin between anterior and medial angle subconcave. Medial angle of pronotum rounded to projected. Punctures almost with the same size, basal punctures only slightly larger than discal punctures. Pronotal disc with shiny points well-defined (Fig. 3 F, arrows), separated between each other and separated or contiguous to punctures; or irregular (Fig. 3 G, arrows), contiguous between to each other and to punctures. ELYTRA (Fig. 4 – 5). Interstriae with shiny points mixed with the punctures. Interstriae VI and VII with basal carina almost identical in size. Carina of ninth interstria reaching middle of elytral length or slightly surpassing it. Striae I – VIII inconspicuous including apically, only in some parts striae slightly visible and very narrow or striae I – VII conspicuous and narrow, width of third stria 1 / 30 th or 1 / 40 th of the distance between stria II and III. If conspicuous, first stria widest and either striae III – VII subequal in width or striae III – VII ill-defined, successively narrower and more ill-defined, with VII almost inconspicuous, and stria VIII conspicuous apical and laterally or only laterally; in both discontinuous in some parts and reaching the apex of carina of the ninth interstria. Apex of elytra with tubercles on interstriae III – VII or III, V – VII (Fig. 5). METAVENTRITE (Fig. 6). With weak posterior excavation, occupying approximately the metaventral basal fourth. Disc with conspicuous punctures at 8 × magnification. Disc punctures at least twice smaller than punctures on anterior-lateral area of metaventral process and dispersed, separated at least by three diameters of a puncture. Punctures on anterior-lateral area of metaventral process separated by less than one diameter. Anterior-medial area of metaventral process with few and smaller punctures than anteriorlateral ones. LEGS. Ventral surface of protibia with a weak carina. Posterior edge of metafemur with two margins, one dorsal and other one ventral. MALE. Protibial spur broad and foliaceus. Mesofemur modified, with slight sinuosity on apical third. Apex of mesotibia wider than female and on ventral-internal margin with a small or large spatulate expansion. Metatrochanter modified or not, if modified with an expansion on distal third. Metafemur with steep tapering on basal third, with expansion on posterior-ventral margin, before that steep tapering. Internal margin of metatibia with small or large tubercles. Ventrite I expanded posteriorly, expansion reaching from the middle of ventrite IV to almost the distal margin of ventrite V; width of expansion on ventrite III variable, narrower to wider than distance between clypeal teeth. Genitalia (Figs 7 – 8). Paramera subtriangular, with dorsal and ventral edges straight in lateral view. Apex of paramera rounded in dorsal view. Paramera with short and thin apical-dorsal notch (Fig. 7, arrows). Medial area of endophallus with one endophallite (Fig. 8). Basal circular shape endophallite with ring very thin and handle strongly broadened medially. FEMALE (Fig. 9). Protibial spur thinner than that of male and spiniform. Apex of mesotibia narrower than that of male. Metatrochanter not modified. Meso- and metafemur not modified. Internal margin of metatibia without tubercles. Metasternal disc with posterior excavation smaller than male. Ventrite V as wide as ventrite VI medially. Ventrite VI narrowed medially.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5FFFB4A6C9A50FFBACFE80.taxon	discussion	Remarks This species group is only known from few (eight) specimens and their biology is poorly known. The specimens were collected above 680 m a. s. l. and the highest record is 1411 m a. s. l. (one specimen does not have altitude information). They were collected using pitfall traps baited with human dung (five specimens), carrion (one specimen) as well as via window trap (one specimen).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5AFFBAA5EEA6D7FC4AFDEB.taxon	description	Figs 1 A, 2 A, 3 A, 4 A, 5 A, 6 A, 7 A, 8 A, 9 A – B, 10	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5AFFBAA5EEA6D7FC4AFDEB.taxon	diagnosis	Diagnosis Elytra with apical tubercles on interstriae III – VII (Fig. 5 A), first stria almost twice as wide as than second stria, striae III – VII subequal in width and conspicuous (Figs 1 A, 4 A) and sub-medial area of endophallus with elongate scales (Fig. 8 A, arrow) distinguish this species (for a complete description see González-Alvarado & Vaz-de-Mello 2021).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5AFFBAA5EEA6D7FC4AFDEB.taxon	materials_examined	Material examined Holotype GUYANA • 1 ♂; District 8, Mount Wokomung; 5 º 06 ′ 34.8 ″ N, 59 º 49 ′ 15.3 ″ W; alt. 1234 m; 27 Oct. – 1 Nov. 2004; B. Hubley leg.; pitfall trap (human dung), primary forest; ROM 2004516; BDGC. [aedeagus and endophallus extracted]; CNC 379888. Paratype GUYANA • 1 ♀; same collection data as for preceding; BDGC.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5AFFBAA5EEA6D7FC4AFDEB.taxon	discussion	Remarks Within D. (Deltohyboma) only few species which possess the anterior margin of clypeus expanded posteriorly into a triangular shape have more than four elytral apical tubercles. Frequently, in species bearing over four apical tubercles, the anterior margin of the clypeus is not expanded into a triangular shape. D. gilli is one of those few exceptions.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC5AFFBAA5EEA6D7FC4AFDEB.taxon	distribution	Known distribution Guyana. District 8, Mount Wokomung. (Fig. 10, blue circle).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	description	urn: lsid: zoobank. org: act: 6 A 25 B 58 A- 9 A 61 - 4 C 0 D- 95 C 3 - BCC 48729 B 806 Figs 1 B, 2 B, 3 B, F, 4 B, 5 B, 6 B, 7 B, F, 8 B, 9 C – D, 10	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	diagnosis	Diagnosis Close to D. nonstriatum sp. nov. by having striae I – VII inconspicuous (Figs 1 B – C, 4 B – C) including apically (Fig. 5 B – C). However, it can be distinguished by presenting the smallest and most disperse interstrial punctures (Fig. 4 B) as well as most disperse punctures on head frons (Fig. 2 B) and on pronotal disc (Fig. 3 B, F) and finally, by the well-defined shiny points on the pronotal disc (Fig. 3 B, F).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	etymology	Etymology A patronym, noun in the genitive case, for Jocelyn Gill. See also the “ Acknowledgments ” section.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	materials_examined	Type material Holotype GUYANA • 1 ♂; District 8, Mount Wokomung; 5 ° 05 ′ 33.4 ″ N, 59 ° 50 ′ 34.4 ″ W; alt. 1411 m; 4 – 8 Nov. 2004; B. Hubley leg.; pitfall trap (human dung), primary forest; ROM 2004526; BDGC; [aedeagus and endophallus extracted]; CNC 379889. Paratypes GUYANA • 1 ♂; same data as for holotype; BDGC • 1 ♀; same data as for holotype; 5 º 07 ′ 53.2 ″ N, 59 º 48 ′ 31.4 ″ W; alt. 698 m; 21 – 26 Oct. 2004; ROM 2004509; BDGC.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	description	Description MEASUREMENTS AND COLOR. Holotype male, length 8.1 mm, humeral width 5.1 mm. Dark green with some red reflections dorsally (Fig. 1 B). Black ventrally, with shiny red reflections on anterior area of metaventral process, meso- and metafemora and ventrite VI (Fig. 6 B). HEAD (Fig. 2 B). Dorsal inter-ocular distance approximately seven times width of one eye. Punctures on frons separated by less than one diameter of each puncture. Punctures on head disc separated by less than one diameter of each puncture. PRONOTUM (Fig. 3 B, F). Medial angle slightly projected. Punctures on the disc separated by less than one diameter. Shiny points on disc well-defined and contiguous to punctures. ELYTRA (Figs 1 B, 4 B). Carina of the ninth interstria reaching middle of elytral length. Striae I – VIII inconspicuous including apically (Fig. 5 B), only in some parts striae slightly visible and very narrow. Punctures of second interstria on disc separated by one or less than one diameter, on third a slightly disperse. Punctures of third interstria on disc occupying about 1 / 14 th of the distance between striae II and III. Apical tubercles on interstriae III, V – VII (Fig. 5 B). ABDOMEN (Fig. 6 B). Width of expansion of the ventrite I, on ventrite III, subequal to distance between clypeal teeth; expansion reaching distal margin of ventrite IV. Margins of expansion between ventrites II – IV forming an acute angle. Apex of expansion truncate. Basal area of expansion with punctures separated by less than one diameter. LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur 1.8 × wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length. PYGIDIUM (Fig. 5 B). Most of the punctures separated by less than one diameter; punctures basally denser than punctures on disc. Discal punctures occupying approximately 1 / 30 th the width on middle of pygidium. G ENITALIA (Figs 7 B, 8 B). Aedeagus as described in the gilli species group. Medial endophallite slightly sinuate. Sub-medial area of endophallus with scales.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	discussion	Remarks Only three specimens are known for this species, all teneral. The holotype does not appear to be teneral externally, however, the aedeagus is poorly sclerotised (Fig. 7 B). The paratypes differ from the holotype by the sexual dimorphism and by having the elytra and the pygidium less sclerotised, with those structures light brown (Fig. 9 C – D). It appears that this species is sympatric with D. gilli since both species were collected a few kilometres apart (see Fig. 10, red square). Deltochilum gilli was collected almost 200 meters higher (altitude) than D. jocelynae sp. nov .. However, both species are easily separated via the elytra; D. gilli has conspicuous striae (Fig. 4 A), whereas these are inconspicuous in D. jocelynae sp. nov. (Fig. 4 B).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC57FFBEA63AA65BFC04FE94.taxon	distribution	Known distribution GUYANA. District 8, Mount Wokomung (Fig. 10, yellow square).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	description	urn: lsid: zoobank. org: act: 563 D 19 C 2 - 0450 - 4 A 9 E-A 4 DC-B 4 AD 395 F 25 F 0 Figs 1 C, 2 C, 3 C, G, 4 C, 5 C, 6 C, 7 C, G, 8 C, 10	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	diagnosis	Diagnosis Close to D. jocelynae sp. nov. by having the striae I – VII inconspicuous (Figs 1 B – C, 4 B – C) including apically (Fig. 5 B – C). However, it can be distinguished by presenting the largest and densest interstrial punctures (Fig. 4 C), as well as densest punctures on the head frons (Fig. 2 C) and on the pronotal disc (Fig. 3 C, G); finally, by the irregular shiny points on the pronotal disc (Fig. 3 C, G).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	etymology	Etymology From Latin non - ' not' + stria in reference to the inconspicuous elytral striae.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	materials_examined	Type material Holotype VENEZUELA • 1 ♂; Bolívar, km 40 Sta. [Santa] Elena, Icabaru Road; 7 ° 26 ′ 20 ″ N, 61 ° 33 ′ 30 ″ W; alt. 1000 m; 4 – 6 Aug. 1986; B. D. Gill leg.; [aedeagus and endophallus extracted]; CMNEN WSD 00041746.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	description	Description MEASUREMENTS AND COLOR. Holotype male, length 8.4 mm, humeral width 5.2 mm. Dark green with red reflections dorsally (Fig. 1 C). Black ventrally, with shiny red reflections on anterior area of metaventral process, meso- and metafemora and ventrite VI (Fig. 6 C). HEAD (Fig. 2 C). Dorsal inter-ocular distance approximately seven times width of one eye. Punctures on frons separated by less than one diameter of each puncture, almost contiguous. Punctures on head disc separated by less than one diameter of each puncture. PRONOTUM (Fig. 3 C, G). Medial angle projected. Punctures on the disc separated by less than one diameter, almost contiguous. Shiny points on disc irregular and contiguous to punctures. ELYTRA (Figs 1 C, 4 C). Carina of the ninth interstria surpassing middle of elytral length. Striae I – VIII inconspicuous including apically (Fig. 5 C), only in some parts striae slightly visible and very narrow. Punctures of second and third interstriae on disc separated by less than one diameter. Punctures of third interstria on disc occupying about 1 ⁄ 10 th the distance between striae II and III. Apical tubercles on interstriae III, V – VII (Fig. 5 C). ABDOMEN (Fig. 6 C). Width of expansion of ventrite I, on ventrite III, narrower than distance between clypeal teeth; expansion reaching distal margin of ventrite IV. Margins of expansion between ventrites II – IV forming an acute angle. Apex of expansion rounded. Basal area of expansion with punctures separated by less than one diameter, almost contiguous. LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur twice wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length. PYGIDIUM (Fig. 5 C). Most of the punctures separated by less than one diameter; punctures basally denser that punctures on disc; basal punctures contiguous. Discal punctures occupying approximately 1 / 28 th the width on middle of pygidium. GENITALIA (FigS 7 C, 8 C). Aedeagus as described in the gilli species group. Medial endophallite almost straight. Sub-medial area of endophallus with scales (Fig. 8 C, arrow).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	discussion	Remarks This species is only known from the holotype. The holotype has an identification label from François Génier (CMNEN), in which he recognised this specimen as probably being a new species since 1998. The transcription of the label as follows: [handwritten] DELTOCHILUM /? SP. NOV. / [printed] Dét. F. Génier, 199 [handwritten] 8.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC50FFBCA614A6C2FB8BFDF7.taxon	distribution	Known distribution VENEZUELA. Bolívar: km 40 Santa Elena, Icabarú Road (Fig. 10, red star).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	description	urn: lsid: zoobank. org: act: E 7290 E 3 C- 1482 - 481 C-AAAC-E 0 C 44 D 7432 DE Figs 1 D, 2 D, 3 D, 4 D, 5 D, 6 D, 7 D, 8 D, 10	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	diagnosis	Diagnosis Close to D. tenuistriatum sp. nov. by having striae I – VII conspicuous (Figs 1 D – E, 4 D – E), but III – VII successively thinner and more ill-defined and the pronotal disc with irregular shiny points contiguous with punctures (Fig. 3 D – E). However, it can be distinguished by having the first stria wider than second, largest interstrial punctures (Fig. 4 D) and by the pygidial punctures which are largest and densest (Fig. 5 D).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	etymology	Etymology From latin quasi - ' almost' + stria in reference to the ill-defined striae.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	materials_examined	Type material Holotype GUYANA • 1 ♂; Mazaruni-Potaro District, Takutu Mountains; 6 ° 15 ′ N, 59 ° 5 ′ W; 18 Dec. 1983; P. D. Perkins and W. E. Steiner leg.; window trap, mountain rainforest, near logging area; EARTHWATCH Research Expedition; [aedeagus and endophallus extracted]; CMNEN WSD 00041744.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	description	Description MEASUREMENTS AND COLOR. Holotype male, length 8.1 mm, humeral width 5.1 mm. Dark green with red reflections on pronotum, elytra dark brown with some red reflections (Fig. 1 D). Dark brown ventrally, with few shiny red reflections on ventrite VI and meso- and meta-legs light brown (Fig. 6 D). HEAD (Fig. 2 D). Dorsal inter-ocular distance approximately seven times width of one eye. Punctures on frons separated by less than one diameter of each puncture, almost contiguous. Punctures on head disc separated by one diameter of each puncture. PRONOTUM (Fig. 3 D). Medial angle projected. Punctures on the disc separated by less than one diameter, almost contiguous. Shiny points on disc irregular and contiguous to punctures. ELYTRA (Figs 1 D, 4 D). Carina of ninth interstria reaching middle of elytral length. Punctures on first stria subequal in size to second stria, but denser. Striae I – VII conspicuous. First stria slightly wider than second stria. Striae III – VII narrow and ill-defined, successively narrower and more ill-defined, with VII almost inconspicuous. Width third stria approximately 1 / 40 th of distance between striae II and III. Stria VIII conspicuous only laterally, discontinuous in some parts and reaching apex of carina of the ninth interstria. Punctures of second and third interstriae on disc separated by less than one diameter. Punctures of third interstria on disc occupying about 1 / 10 th of the distance between striae II and III. Apical tubercles on interstriae III, V – VII (Fig. 5 D). ABDOMEN (Fig. 6 D). Width of expansion of the ventrite I, on ventrite III, subequal to distance between clypeal teeth; expansion reaching middle of ventrite IV. Margins of expansion between ventrites II – IV forming an acute angle. Apex of expansion truncate. Basal area of expansion with punctures separated by less than one diameter, almost contiguous. LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur 1.8 × wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length. PYGIDIUM (Fig. 5 D). Most of the punctures separated by less than one diameter; punctures basally denser than punctures on disc; basal punctures almost contiguous. Discal punctures occupying approximately 1 / 29 th the width on middle of pygidium. GENITALIA (Figs 7 D, 8 D). Aedeagus as described in the gilli species group. Medial endophallite sinuate. Sub-medial area of endophallus with scales.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	discussion	Remarks This species is only known from the holotype.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC52FFA3A61FA53DFC32FBCE.taxon	distribution	Known distribution Guyana. Cuyuni-mazaruni, Takutu Mountains (Fig. 10, green triangle).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	description	urn: lsid: zoobank. org: act: 9 BBED 604 - 6552 - 43 D 2 - B 7 AF- 44 B 2 DF 95 DDF 7 Figs 1 E, 2 E, 3 E, 4 E, 5 E, 6 E, 7 E, 8 E, 10	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	diagnosis	Diagnosis Close to D. quasistriatum sp. nov. by having the striae I – VII conspicuous (Figs 1 D – E, 4 D – E), but III – VII successively thinner and more ill-defined and by the pronotal disc with irregular shiny points contiguous to punctures (Fig. 3 D – E). However, it can be distinguished by having the first stria subequal to second stria, smallest interstrial punctures (Fig. 4 E) and by pygidial punctures which are smallest and most disperse (Fig. 5 E).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	etymology	Etymology From Latin tenuis ' weak' + stria in reference to the ill-defined striae.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	materials_examined	Type material Holotype VENEZUELA • 1 ♂; Bolívar, 10 km E of S. [San] F. [Francisco] Yuruani; 5 ° 1 ′ 34 ″ N, 61 ° 2 ′ 34 ″ W; alt. 1300 m; 8 – 10 Jul. 1987; S. and J. Peck leg.; carrion traps, Gran Sabana, forest; [aedeagus and endophallus extracted]; CMNEN WSD 00041745.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	description	Description MEASUREMENTS AND COLOR. Holotype male, length 8 mm, humeral width 5.2 mm. Dark green with red reflections dorsally (Fig. 1 E). Black ventrally, with shiny red reflections on metaventrite, metaventral process, meso- and metafemora and ventrite VI (Fig. 6 E). HEAD (Fig. 2 E). Dorsal inter-ocular distance approximately eight times width of the eye. Punctures on frons separated by less than one diameter of each puncture. Punctures on head disc separated by less than one diameter of each puncture, almost contiguous. PRONOTUM (Fig. 3 E). Medial angle projected. Punctures on disc separated by less than one diameter, almost contiguous. Shiny points on disc irregular and contiguous to punctures. ELYTRA (Figs 1 E, 4 E). Carina of ninth interstria reaching middle of elytral length. Punctures on first stria subequal in size to second stria, but denser. Striae I – VII conspicuous. Width of first stria subequal to second stria. Striae III – VII narrow and ill-defined, successively narrower and more ill-defined, with VII almost inconspicuous. Width third stria approximately 1 / 40 th the distance between striae II and III. Stria VIII conspicuous only laterally, discontinuous in some parts and reaching apex of carina of the ninth interstria. Punctures of second interstria on disc separated by less than one diameter, on third by one diameter. Punctures of third interstria on disc occupying about 1 / 14 th the distance between striae II and III. Apical tubercles on interstriae III, V – VII (Fig. 5 E). ABDOMEN (Fig. 6 E). Width of expansion of ventrite I, on ventrite III, subequal to distance between clypeal teeth; expansion reaching distal margin of ventrite IV. Margins of expansion between ventrites II – III forming an acute angle. Expansion on ventrite IV narrower than on ventrite III, and margins almost parallel. Apex of expansion rounded. Basal area of expansion with punctures separated by less than one diameter, almost contiguous. LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur 1.7 × wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length. PYGIDIUM (Fig. 5 E). Most of the punctures separated by one diameter; punctures basally denser than punctures on disc. Discal punctures occupying approximately 1 / 38 th the width on middle of pygidium. GENITALIA (Figs 7 E, 8 E). Aedeagus as described in the gilli species group. Medial endophallite sinuate. Sub-medial area of endophallus with scales.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	discussion	Remarks This species is only known from the holotype.	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
03F48795DC4DFFA2A613A31BFB53F9ED.taxon	distribution	Known distribution Venezuela. Bolívar: 10 km E San Francisco Yuruani (Fig. 10, white eight-point star).	en	González-Alvarado, Arturo, Vaz-De, Fernando Z. (2021): Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum). European Journal of Taxonomy 775: 86-106, DOI: 10.5852/ejt.2021.775.1551
