taxonID	type	description	language	source
038E87BCA6495405546CFC58FC79FEF8.taxon	description	urn: lsid: zoobank. org: act: FF 3 B 0077 - C 40 A- 464 C- 9 D 52 - B 0 F 43 FA 82497	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6495405546CFC58FC79FEF8.taxon	etymology	Etymology The generic name is given in respectful remembrance of the collector of the species, Prof. Dr Francis Dov Por (Israel) and is a compound of the collector’s name and the Greek ψύλλος [psillos] (‘ flea’), a term often used in the naming of taxa of the Ceratonotus group (e. g., Dendropsyllus, Echinopsyllus, Pseudechinopsyllus). Gender: masculine.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6495405546CFC58FC79FEF8.taxon	type_taxon	Type and only species Poropsyllus menzelae gen. et sp. nov.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6495405546CFC58FC79FEF8.taxon	diagnosis	Diagnosis Member of Cletodidae T. Scott, 1905, Cletodinae T. Scott, 1905 and Ceratonotus group sensu Conroy-Dalton (2001). Body elongate, cylindrical, podoplean boundary between pro- and urosoma inconspicuous. Sexual dimorphism in A 1, P 3, P 5, and P 6; female with GDS due to fusion of last (P 6 - bearing) thoracic and first abdominal (genital) somites. Cphth about ¼ of total body length, with some sensilla and long tube pores laterally and dorsally; moreover, with small, sensilla-bearing frontolateral horns (FLH), ventrally directed cephalothoracic anterior lateroventral processes (CLVPa), and bifurcate cephalothoracic posterior lateroventral processes (CLVPp). Cephalic front produced into a massive diverging peak. P 2 – P 4 - bearing pedigerous somites with dorsal cuticular processes. Rostrum absent, original central position indicated by small paired membranous projections accompanied by a pair of sensilla and 1 tube pore. Body somites except last thoracic somite (= anterior half of female GDS) and telson laterally with 1 long tube pore. Thoracic body somites dorsally also with 1 long tube pore. Telson broader than long, trapezoid in dorsal view, anal operculum flanked by 2 sensilla, posterior margin with row of fine spinules. FR long and slender, diverging, with 7 setae. Female A 1 slender, 4 - segmented, male A 1 sexually dimorphic, number of segments unknown. A 2 without exopod, allobasis with 2 abexopodal setae; endopod 1 - segmented, with 2 spines and 1 slender additional seta; apically with 5 setae, 3 of which geniculate. Intercoxal sclerites of P 1 – P 4 slender, bow-like; P 1 with transversely elongated basis and 2 - segmented exopod, endopod absent, represented by 1 long bare seta; bases of P 2 – P 4 showing extreme transverse elongation, without endopods, except male P 3, which bears a 3 - segmented endopod with an apophysis on enp- 2. P 5 also considerably elongated, reaching anterior margin of telson; in the female with exopod fused to baseoendopod, in male exopod distinct.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64E540D5422FEA3FD2CFBC8.taxon	description	urn: lsid: zoobank. org: act: D 3681697 - E 1 C 8 - 40 A 4 - 8 CC 4 - 7 DEB 1705903 E Figs 1 – 7; Table 1	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64E540D5422FEA3FD2CFBC8.taxon	etymology	Etymology The epitheton menzelae is given in friendly dedication to Dr Lena Menzel (Bremerhaven, Germany), who obtained the material from Prof. Por and placed it at the author’s disposal for further study.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64E540D5422FEA3FD2CFBC8.taxon	materials_examined	Type material Holotype CYPRUS • ♀; Akrotiri Bay (locus typicus), E of Cape Gata (S Cyprus); geographic transect: 34 ° 33 ' 54 " N / 33 ° 02 ' 45 " E – 34 ° 34 ' 20 " N / 33 ° 04 ' 06 " E; depth 115 fathoms (= 210.5 m); 26 Oct. 1968; F. D. Por leg.; Ockelmann detritus sledge (Por et al. 1972); HUJ INVCRUCOP 808, on 1 slide. Paratype (allotype) CYPRUS • ♂; Coral Bay (SW Cyprus); geographic transect: 34 ° 48 ' 24 " N / 32 ° 22 ' 57 " E – 34 ° 48 ' 06 " N / 32 ° 21 ' 24 " E; depth 100 fathoms (= 183.0 m); 31 Oct. 1968 F. D. Por leg.; Ockelmann detritus sledge (Por et al. 1972); HUJ INVCRUCOP 809, on 1 slide.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64E540D5422FEA3FD2CFBC8.taxon	description	Both specimens show several instances of damage, mainly loss of setae, segments of appendages, and both antennular segments and cuticular processes (male allotype). Dissection, which would have caused further destruction of the individuals, was not conducted, but all parts except the maxillula were drawn and described. Description Female HABITUS (Fig. 3 A). Elongate, cylindrical (body length including FR 550 µm), podoplean boundary between pro- and urosoma inconspicuous. Cphth about ¼ of total body length, with some sensilla and long tube pores laterally and dorsally; moreover, with small frontolateral horns (FLH), ventrally directed cephalothoracic anterior lateroventral processes (CLVPa) (arrowheads in Fig. 3 A), and bifurcate cephalothoracic posterior lateroventral processes (CLVPp). Cephalic front produced into a massive diverging peak, from which the A 1 arises. Rostrum (Figs 3 A, 4 A) absent, original central position indicated by small paired membranous projections accompanied by a pair of sensilla and 1 tube pore. P 2 – P 4 - bearing pedigerous somites with small dorsal cuticular processes (DP 1 – DP 3); each process carrying a sensillum at its tip (most of them broken in Fig. 3 A). Body somites except last thoracic somite (= anterior half of female GDS) and telson laterally with 1 long tube pore. Thoracic body somites with additional dorsal long tube pore. First abdominal somite fused with last thoracic body somite to form a GDS. Thoracic part of GDS dorsally with pair of diagonal cuticular ridges; abdominal part of GDS and abdominal somites except telson with abdominal lateral setulose tufts (AST 1 – AST 3). Telson broader than long, trapezoidal in dorsal view, anal operculum flanked by 2 sensilla, posterior margin with row of fine spinules. FR (Fig. 3 A – B). Diverging, rami slender, about 11 times as long as broad and longer than abdomen and GDS together, bearing 7 bare setae (Fig. 3 B). Setae I and II inserting laterally, standing close together, I minute, II twice as long as I; III inserting subapically on outer margin; setae IV and V located apically, IV slightly smaller than III, and V as long as fifth pedigerous somite, abdomen, and FR combined (Fig. 3 A); VI as long as IV, inserting apically on inner margin; VII tri-articulate, arising dorsally from small knob. A 1 (Fig. 4 B). 4 - segmented, long and slender. First segment longest, second to fourth segments of almost the same size. First 3 segments with long spinules (several spinules broken in Fig. 4 B). Several setae broken in Fig. 4 B; first segment distally with 1 seta (broken); second segment with 7 setae (6 broken, 1 biplumose); third segment with at least 6 setae (all but 1 bare seta broken); pedestal for acrothek discernible close to apical margin (elements broken); fourth segment with 11 bare setae (1 apical seta broken), two of which fused with 1 small aes. Armature formula: I – 1; II – 7; III – 6 (7?) + aes, IV – 11 + aes. A 2. See description of allotype. MD (Fig. 4 C). Gnathobase small in comparison with palp, apically with 3 multicuspidate teeth; palp unilobed, subapically with some fine, long spinules, and with 2 lateral (basal) and 3 apical (endopodal) setae, all biplumose. MXL. Because dissection, which would have caused destruction of the individuals, was not conducted, no in situ description could be made, and the mxl was not drawn. MX (Fig. 4 D). Syncoxa and allobasis separate, syncoxa with 2 endites. Proximal endite with 1 strong unipinnate spine fused to the segment, and with 1 bare and 1 biplumose seta. Distal endite with 1 strong bare spine fused to the segment, and with 1 unipinnate seta. Allobasis with 1 claw-like element fused to the segment; additionally with 1 strong and 2 slender bare setae. Endopod distinct, small, 1 - segmented, with 2 bare setae. MXP (Fig. 4 E). Prehensile, syncoxa with few long spinules and 1 strong multipinnate seta. Basis with 3 rows of long spinules. Endopod produced into long biplumose claw, basally accompanied by 1 tiny seta. P 1 – P 4 (Figs 5 B, 6 A – C). With slender, bow-like intercoxal sclerites (Figs 5 B, 6 A); bases extraordinarily elongated transversely (Figs 3 A, 5 B, 6 A), without any spinulose ornamentation, except the presence of 1 tube pore at mid-length on the anterior margin (Figs 5 B, 6 A). P 1 (Fig. 5 B) basis with 1 inner bare and 1 outer biplumose seta; with tube pore on anterior margin; endopod represented by 1 bare seta; exopod 2 - segmented, exp- 2 about 3 times as long as exp- 1; exp- 1 with 1 outer bipinnate seta; exp- 2 with 1 outer bipinnate seta; apically with 4 geniculate setae (2 broken in Fig. 5 B), one of which unipinnate. P 2 – P 4 (Fig. 6 A – C) without endopods; bases with 1 short bare outer seta (Fig. 6 A) and with tube pore on anterior margin; exopods 3 - segmented, segments with few or without spinules as depicted; exp- 1 with outer bipinnate seta (Fig. 6 A); exp- 2 with 1 outer and 1 inner seta; exp- 3 with 2 outer bipinnate setae, the posterior one shifted to an apical (P 2, P 3; Fig. 6 A – B) or subapical position (P 4; Fig. 6 C); exp- 3 additionally with 2 apical setae. P 2 exp- 3 (Fig. 6 A) in addition subapically with long tube pore. P 2 and P 4 exp- 3 with 1, P 3 exp- 3 with 2 inner setae. Inner setae and innermost apical seta biplumose, but outermost apical seta bare. Setation of P 2 – P 4 as in Table 1. P 5 (Fig. 5 C). Baseoendopod and exopod fused. Baseoendopod with outer bipinnate seta arising from minute setophore and accompanied by 1 long tube pore. Endopodal lobe completely reduced, represented by 1 long and 1 small bare seta. Exopodal lobe with 1 bipinnate outer seta and 3 apical setae, two of which bipinnate, the remaining one bare; subapically with long tube pore. GF (Fig. 7 A). With fused gonopores covered by slender genital operculum derived from the medially fused P 6, whose minute lobes each end in a bare seta. Male The male resembles the female in most characters. Sexual dimorphism is expressed in a slightly smaller body size (480 µm incl. FR), in the A 1, the P 3 endopod, the P 5, and the absence of a P 6. A 1 (Fig. 7 B). Damaged, but sexual dimorphism still discernible. First segment as in female; second segment half as long as first, with 7 setae and long spinules on anterior margin; third segment about ⅓ the length of previous segment, with 1 bare seta. Following segments broken, not illustrated. A 2 (Fig. 5 A). Without exp. Allobasis with 2 abexopodal bare setae. Endopod laterally on distal half with 2 well-developed setae, one of which bipinnate, and with 1 small bare seta; additionally, with frill-like row of long fine spinules. Apically with 3 long geniculate setae (one broken); moreover with 1 bipinnate and 1 bare slender seta. Subapically with spinulose frill. P 3 ENDOPOD (Fig. 6 D). 3 - segmented, first segment small and unarmed, second segment about 6 times as long as first, ending in an acute apophysis; third segment slightly longer than first, apically with 1 small bare seta. P 5 (Fig. 7 C). With exopod separated from baseoendopod, carrying 2 outer bipinnate and 3 bare apical setae; subapically with tube pore. Outer baseoendopodal seta bare, accompanied by small tube pore; endopodal lobe represented by 1 bare seta.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA646540C5454FB53FC2BFCE8.taxon	description	urn: lsid: zoobank. org: act: 883121 EC- 5626 - 40 EA-BC 42 - CC 2 A 7 FE 7 B 166	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA646540C5454FB53FC2BFCE8.taxon	type_taxon	Type and only species Pa. aaroni sp. nov.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA646540C5454FB53FC2BFCE8.taxon	etymology	Etymology The generic name ‘ Paratouphapleura ’ is a combination of the Greek παρά [pará], meaning ‘ besides’, ‘ beyond’, and the genus name Touphapleura, pointing to the general morphological resemblance of Touphapleura and Paratouphapleura gen. nov. Gender: feminine.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA646540C5454FB53FC2BFCE8.taxon	diagnosis	Diagnosis Membor of Cletodidae T. Scott, 1904, Cletodinae T. Scott, 1904, and the Ceratonotus group sensu Conroy-Dalton (2001). Body more or less cylindrical, tapering slightly posteriorly, without clear demarcation between pro- and urosoma. In female fusion of last thoracic (P 6 - bearing) and first abdominal (= genital) somite to form GDS. Cephalothorax with weakly developed cephalothoracic anterior lateroventral processes (CLVPa); cephalothoracic posterior lateroventral processes (CLVPp) well-developed, jet-wing-like, with 1 sensillum each on the anterior margin and the tip; cephalothoracic anterodorsal processes (CADP) and cephalothoracic laterodorsal processes (CLDPp) not developed; anterior cephalothoracic setulose tufts (ACST) present, posterior cephalothoracic setulose tufts (PCST) absent. Rostrum small, broader than long, fused to cephalothorax; with paired sensilla, 1 tube pore, paired membranous projections, and rostral setulose tuft (RST). Pedigerous somites dorsally with long tube pore, those bearing P 2 – P 4 additionally with pairs of small and unarmed dorsal cuticular processes (DP 1 – DP 3), each carrying a sensillum at its tip, and laterally with setulose tufts (TST 1 – TST 3); P 6 - bearing somite laterally with setulose tufts (GST). Urosomal somites laterally with tube pores accompanied by abdominal lateral spinulose tufts (AST 1 – AST 3); first abdominal somite (= posterior part of female GDS) and following 2 abdominal somites with dorsal row of spinules on posterior margin, the former and the second somite additionally with pair of tube pores dorsally. Telson half as long as penultimate somite, with spinulose anal operculum flanked by 2 sensilla and spinulose tufts (AST 4) running lateroventrally at the bases of the furcal rami. The latter about three times as long as broad, with 7 setae (I – VII). Antennule 4 - segmented in the female, 7 - segmented and subchirocer in the male; first segment elongated and equipped with long spinules along anterior margin. Antenna with allobasis and 1 - segmented endopod, without exopod; allobasis with 2 abexopodal setae; endopod with 2 spines, apically with 5 setae, 4 of which geniculate. Swimming legs with bow-like intercoxal sclerites, small coxae, and transversely elongated bases; P 1 with 2 - segmented lobes, non-prehensile; basis with inner and outer seta; exp- 1 with 1 outer bipinnate spine, exp- 2 with all elements located apically: 1 bipinnate outer spine and 4 bare or unipinnate geniculated setae. P 2 – P 4 bases carrying 1 outer seta, with 3 - segmented exopods; P 2, P 4, and female P 3 with 2 - segmented endopods, male P 3 endopod 3 - segmented, enp- 2 with apophysis on inner apical edge. P 5 with endopodal lobe reduced to a minute pedestal with 1 seta. Female GDS with P 6 strongly reduced, not forming a genital operculum.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64754165439FCB3FA81FD22.taxon	description	urn: lsid: zoobank. org: act: 6 F 7 C 1 F 0 C- 7 A 6 F- 473 A-B 5 B 4 - 78859 FE 44 D 59 Figs 8 – 14; Table 2	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64754165439FCB3FA81FD22.taxon	etymology	Etymology The epitheton aaroni is given in fond dedication to my brother, Mr Aaron Martin George (Münster / Selters, Germany).	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64754165439FCB3FA81FD22.taxon	materials_examined	Type material 3 ♀♀, 3 ♂♂, collected during research cruise ANT-XIX / 4 (ANDEEP II) of RV POLARSTERN in 2002. Holotype ANTARCTICA • ♀; Weddell Sea, E of Antarctic Peninsula; station # 133; 65 ° 20.30 ' S / 54 ° 14.67 ' W; depth 1109.6 m (locus typicus); multiple corer; SMF 37270 / 1, on 1 slide. Paratypes ANTARCTICA • 2 ♀♀, 3 ♂♂; same collection data as for holotype; SMF 37271 / 1 (♂ paratype 1 / allotype, on 1 slide), SMF 37272 / 1 (♀ paratype 2, on 1 slide), SMF 37273 / 1 – 9 (♀ paratype 3, on 9 slides), SMF 37274 / 1 – 8 (♂ paratype 4, on 8 slides), SMF 37275 / 1 (♂ paratype 5, on 1 slide).	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA64754165439FCB3FA81FD22.taxon	description	Description Female HABITUS (Fig. 8 A). Elongate, cylindrical; body length including FR: 493 µm (holotype), 562 µm (paratype 2), 515 µm (paratype 3). Podoplean boundary between pro- and urosoma inconspicuous. Cphth about ⅓ of total body length, with some sensilla and long tube pores laterally and dorsally, and with anterior cephalothoracic setulose tufts (ACST). Cephalic front produced into a moderate peak, from which the A 1 arises. Without frontolateral horns (FLH), but cephalothoracic anterior lateroventral processes (CLVPa) weakly developed, ending in long tube pore; sensilla not discernible. Cphth additionally with 3 pairs of tube pores, two of which located dorsally, the third laterally. Cephalothoracic posterior lateroventral processes (CLVPp) (Fig. 9 B) long and jet-wing-like, with 1 sensillum halfway on anterior margin and 1 sensillum on its tip. Rostrum (Fig. 9 A) small, broader than long, fused to cephalothorax; with pair of sensilla, 1 tube pore, paired membranous projections, and rostral setulose tuft (RST). P 2 – P 4 - bearing thoracic somites (Fig. 8 A) dorsally with 1 tube pore and paired unarmoured small cuticular processes (DP 1 – DP 3), each carrying a sensillum at its tip; laterally with thoracic setulose tufts TST 1 – TST 3. Following body somites without cuticular processes. P 5 - bearing somite dorsally with tube pore and 2 sensilla. GDS dorsally with fine suture indicating former separation; anterior part of GDS dorsally with 1 tube pore, laterally with setulose tufts (GST) accompanied by 1 tube pore; posterior part of GDS as well as subsequent abdominal somite dorsally with 2 tube pores and 2 sensilla on posterior margin, which in addition presents a row of spinules; laterally with abdominal lateral setulose tufts AST 1 and AST 2, each accompanied by 1 tube pore. Penultimate abdominal somite dorsally with spinulose posterior margin and with AST 3 (only discernible on the right side in Fig. 8 A) but without sensilla and tube pores. Telson half as long as previous somite, slightly trapezoidal, with AST 4 and spinulose anal operculum flanked by 2 sensilla. FR (Figs 8 A, 9 C). Diverging, rami slender, about 3.5 times as long as broad and as long as telson and preceding somite together, with 1 small tube pore proximally on outer margin and bearing 7 bare setae (Fig. 9 C): setae I and II of almost the same length, inserting halfway on outer margin and set close together, I bipinnate, II bare; III bipinnate, inserting subapically on outer margin; setae IV and V located apically, IV bare, visibly smaller than III, V longest element, bipinnate; VI bare, even smaller than IV, inserting apically on inner margin; VII bare, tri-articulate, arising dorsally from small knob. A 1 (Fig. 10 A). Partly or completely broken in all three females. Most complete condition present in holotype: slender, 4 - segmented, first and third segments of almost the same size, about twice as long as second and fourth segment. First and second segments with several long spinules, following segments without spinulose ornamentation. First segment with biplumose seta subapically; second segment with 8 setae (5 broken); third segment with 8 setae (2 broken), 2 of which forming acrothek together with aes; fourth segment with 3 lateral setae; apically with at least 5 setae; acrothek that might consist of 2 additional setae and aes broken, not discernible. Armature formula: I – 1; II – 8; III – 6 (+ 2 + aes), IV – 8 (+ 2? + aes?). A 2 (Fig. 11 A). Without exp. Allobasis with row of spinules and 2 abexopodal bipinnate setae. Endopod laterally with few spinules, on distal half with 2 well-developed bipinnate setae, third small bare seta absent. Apically with 4 long geniculate setae and 1 smaller seta, all bare, and with spinulose frill. MD (Fig. 11 B). Gnathobase apically with 2 multicuspidate teeth; palp unilobed, with some fine, long spinules, and with 2 lateral (basal), 3 apical (endopodal), and 1 outer (exopodal) setae, all biplumose. MXL (Fig. 11 C). Praecoxal arthrite with set of spinules, apically with 6 spines and on each edge with 1 seta; additionally, with 2 surface setae; all elements bare. Coxal endite apically with 1 bare seta. Basis, endopod and exopod fused to single lobe, carrying 4 lateral setae, the proximal one bipinnate, the remaining three bare; apically with 3 bare setae and 2 unipinnate spines. MX (Fig. 11 D). Syncoxa and allobasis separated; syncoxa apically with cluster of spinules, and with 2 endites; proximal endite with 1 strong bipinnate spine, fused to segment, and with 2 bare setae; distal endite with bare spine fused to the segment, and with 2 bare setae. Allobasis with 1 claw-like element fused to the segment; additionally, with 2 bare setae. Endopod 1 - segmented, small, with 2 bare setae. MXP (Fig. 11 E). Prehensile, syncoxa without spinules but with 1 biplumose seta on apical edge; basis with 2 rows of long spinules; endopod produced into long bare claw, basally accompanied by 1 tiny seta. P 1 (Fig. 10 B). With transversely elongated basis and 2 - segmented exo- and endopod. Praecoxa small, triangular; coxa small, more or less rectangular. Basis with several long spinules on anterior margin, and with 1 inner and 1 outer element (broken in Fig. 10 B). Exopodal segments of nearly the same size, exp- 1 with row of spinules on outer distal margin, and with 1 outer bipinnate seta; exp- 2 without spinules, with 1 bipinnate outer seta, and (sub) apically with 4 geniculate elements, all with row of pinnules previous to geniculation. P 2 – P 4 (Fig. 12 A – G). With slender, bow-like intercoxal sclerites (P 3: Fig. 12 F; P 4: Fig. 12 G) and transversely elongated bases that carry long spinules and 1 tube pore on their anterior margins, as shown for P 2 (Fig. 12 A). Exp- 1 with few spinules on inner and / or outer margin, posteriorly with 1 bipinnate spine; exp- 2 also with few spinules as depicted, and with 1 inner biplumose seta and 1 bipinnate outer spine; exp- 3 with 2 outer bipinnate elements, short and spine-like in P 2 but long and seta-like in P 3 and P 4; additionally, with 2 apical biplumose setae; inner margin of exp- 3 with 1 biplumose seta in P 2 and P 4, but with 2 biplumose setae in P 3. Endopods small and 2 - segmented, enp- 1 shorter than enp- 2, without ornamentation; P 2 enp- 2 with few spinules and 2 apical setae, outermost pinnate on the outer but plumose on the inner margin, innermost seta biplumose and not reaching half the length of the outer one; enp- 2 of P 3 and P 4 each with 2 biplumose apical setae, and with 1 outer bipinnate seta reaching at most ⅓ (P 3) or half (P 4) the length of the apical elements. Setation of P 2 – P 4 as in Table 2. P 5 (Fig. 13 A). Baseoendopod and exopod distinct. Baseoendopod with outer bipinnate seta arising from long slender setophore (broken in Fig. 13 A) and accompanied by 1 long tube pore and long spinules. Endopodal lobe minute, with 1 long bipinnate seta on its distal part, and with 1 tube pore. Exopodal lobe slightly longer than baseoendopod, with 2 outer, 1 subapical, and 2 apical bipinnate setae; subapically with long tube pore. GF (Fig. 9 D). With single gonopore; P 6 not forming genital operculum; lobes strongly reduced, each ending in a small bare seta (1 seta broken in Fig. 9 D). Male The male (Fig. 8 B) resembles the female in most characters. Sexual dimorphism consists of a slightly smaller body size (allotype: 410 µm, paratype 4: 417 µm, paratype 5: 459 µm), smaller dorsal cuticular processes DP 1 – DP 3, the A 1, the P 3 endopod, the P 5, and the absence of a P 6. A 1 (Fig. 14 A). 7 - segmented, subchirocer. First segment with long spinules on surface and anterior margin, and with 1 seta apically (broken in Fig. 14 A); second segment as long as first, with 8 bare setae (two of which broken in Fig. 14 A); third segment small, almost triangular, with 4 bare setae; fifth segment minute, with 2 small bare setae; sixth segment swollen, setation not discernible because of position on the slide and the risk of breaking the A 1 if it is turned (its description is therefore postponed until additional material is found) sixth segment small and squarish, without ornamentation; seventh segment twice as long as sixth, tapering posteriorly, with 9 bare setae, 2 of which form an apical acrothek together with 1 aes. P 3 ENDOPOD (Fig. 14 B). 3 - segmented, first segment small and unarmed, second segment about three times as long as first, ending in acute apophysis; third segment slightly shorter and narrower than first, not reaching end of apophysis, with 2 long biplumose setae apically. P 5 (Fig. 13 B). Exopod separated from baseoendopod, with several long spinules and 2 outer and 2 apical bipinnate setae; additionally, with 1 subapical bare seta but without tube pore; baseoendopod as in female but endopodal lobe smaller, likewise the single endopodal seta, which is small and bare.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA65D5416544BFD0DFC87F85A.taxon	description	urn: lsid: zoobank. org: act: D 9585661 - 5 A 33 - 40 AF- 938 F-CA 881880 B 7 E 9	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA65D5416544BFD0DFC87F85A.taxon	type_taxon	Type and only species Pseudopolyascophorus monoceratus (George et al., 2013) gen. et comb. nov. Polyascophorus monoceratus George et al., 2013: 20 – 33, figs 2 – 8.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA65D5416544BFD0DFC87F85A.taxon	etymology	Etymology The genus name is compound by the Greek ψευδής [pseudes] (‘ misleading’, ‘ deceiving’) and the genus name Polyascophorus, indicating an ostensible morphological similarity of Polyascophorus and Pseudopolyascophorus gen. nov. Gender: masculine.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA65D5416544BFD0DFC87F85A.taxon	diagnosis	Diagnosis Member of Cletodidae T. Scott, 1904, Cletodinae T. Scott, 1904 and the Ceratonotus group sensu Conroy- Dalton (2001). Body more or less cylindrical, tapering slightly posteriorly, without clear demarcation between pro- and urosoma. Sexual dimorphism in A 1, P 3, P 4, P 5, and P 6; females with GDS due to fusion of last (P 6 - bearing) thoracic and first abdominal (genital) somite. Cphth comprising approximately ¼ of body length; rostrum fused to cephalothorax, enlarged and Y-shaped; with long tube pore but no rostral setulose tuft (RST). Cphth with paired knobbed frontolateral horns (FLH), cephalothoracic posterior lateroventral (CLVPp) and laterodorsal processes (CLDPp); CLVPp bifurcated. Thoracic somites bearing P 2 – P 5 with dorsal cuticular processes DP 1 – DP 4. First abdominal somite (= posterior part of female GDS) with single centred DP 5. Cphth, body somites, and furca with tube pores dorsally and / or laterally. Furcal rami barely 4 times as long as broad, rounded at their apical outer margin; with 7 setae (I – VII). Female A 1 4 - segmented, with first segment elongated and bearing long spinules on inner margin, and with aesthetasc on third and fourth segment. Male A 1 6 - segmented, subchirocer. A 2 with allobasis, without endopod; allobasis with 2 abexopodal setae. Endopod as long as allobasis, with 2 spinulose frills (sub) apically. P 1 with 2 - segmented exopod; exp- 1 with 1 bipinnate outer spine, exp- 2 with 4 apical, geniculate setae and 1 outer spine; endopod 2 - segmented in female, with 1 apical seta, absent in male. P 2 – P 4 with 3 - segmented exopods; exp- 1 with 1 outer spine, exp- 2 with 1 outer spine and 1 inner seta; exp- 3 with 2 outer spines and 2 apical setae, and with 1 (P 2, P 4) or 2 (P 3) inner setae. P 2 without endopod. P 3 endopod in female 2 - segmented, carrying 2 apical setae; in male 3 - segmented with a small bare enp- 1, elongated enp- 2 with a curved apophysis on inner apical margin, and 2 apical setae on small enp- 3. P 4 endopod 2 - segmented; enp- 1 small, unarmed, enp- 2 in female with 1 apical seta; in male with 2 apical setae and 1 outer spine. P 5 with baseoendopod and exopod fused, sexually dimorphic; endopodal lobe completely reduced and represented by 1 seta and 1 tube pore in female, and by 2 setae and 2 tube pores in male. Female genital field with gonopore covered by genital operculum formed by the reduced P 6, with 1 seta on each side.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6525419548CFEC7FC04F857.taxon	description	urn: lsid: zoobank. org: act: 7 A 9 A 1 E 20 - 76 EB- 4 AD 2 - 9795 - 8 C 09 DFD 9854 B	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6525419548CFEC7FC04F857.taxon	type_taxon	Type species Tauroceratus tauroides (George, 2006) gen. et comb. nov. (by designation). Ceratonotus tauroides George, 2006: 89 – 100, figs 1 – 9.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6525419548CFEC7FC04F857.taxon	etymology	Etymology The genus name Tauroceratus is a combination of the Greek ταύρος [távros], meaning ‘ bull’, and the Greek κέρας [kéras], meaning ’ horn’, referring to the presence of cephalothoracic frontal horns (FH) in the assigned species. Gender: masculine.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
038E87BCA6525419548CFEC7FC04F857.taxon	diagnosis	Diagnosis Member of Cletodidae T. Scott, 1904, Cletodinae T. Scott, 1904, and the Ceratonotus group sensu Conroy-Dalton (2001). Body cylindrical, tapering slightly posteriorly, without clear demarcation between pro- and urosoma. Sexual dimorphism in A 1, P 3, P 5, and P 6; females with GDS due to fusion of last (P 6 - bearing) thoracic and first abdominal (genital) somites. Cphth about ⅓ of total body length, with some sensilla laterally and dorsally; cephalic front produced into a massive peak. Cphth frontally with pair of frontal horns (FH) that carry the rostral setae, and dentate frontolateral horns (FLH); posteriorly with dendroid cephalothoracic laterodorsal processes (CLDPp). Cephalothoracic anterior lateroventral processes (CLVPa) and cephalothoracic posterior lateroventral processes (CLVPp) developed. Rostrum absent, original position indicated by tube pore. P 2 – P 5 - bearing pedigerous somites dorsally with dendroid cuticular processes (DP 1 – DP 4). Last pedigerous somite (= anterior half of female GDS) with sensilla on posterior margin that arise from tubercles dorsally (DT 1) and small conical sensillate tubercles (LT 1) laterally (not confirmed for T. tauroides gen. et comb. nov.). First abdominal somite (i. e., posterior half of female GDS) dorsally with dendroid DP 5, second abdominal somite dorsally with conical DP 6, and laterally with small conical lateral sensillate tubercles (LT 2) (not confirmed for T. steiningeri gen. et comb. nov.). Telson broader than long, trapezoidal in dorsal view, anal operculum flanked by 2 sensilla, posterior margin bare or with row of spinules. Furcal rami long and slender, diverging, with 7 setae. Female A 1 slender, 3 - segmented, 5 – 6 - segmented and (sub) chirocer in male. A 2 without exopod, allobasis with 1 abexopodal seta on endopodal half. Intercoxal sclerites of P 1 – P 4 slender, bow-like; P 1 with transversely elongated basis and 2 - segmented exo- and endopod; exp- 2 longer than exp- 1, with 1 outer and 1 subapical bipinnate spine and 3 geniculated apical setae. P 2 – P 4 with transversely elongated bases, 3 - segmented exopods and 2 - segmented endopods except male P 3 endopod, which is 3 - segmented. Exp- 1 with 1 outer bipinnate spine, without inner seta; exp- 2 with 1 bipinnate outer spine and 1 biplumose inner seta, exp- 3 with 2 bipinnate outer spines, 2 biplumose apical and 1 (P 2, P 4) or 2 (P 3) biplumose inner setae. Enp- 1 small and bare, enp- 2 elongate, with 1 biplumose apical seta in P 2 and 1 – 2 apical elements in P 3 (female) and P 4; male P 3 enp- 1 as in female, enp- 2 elongate and bearing cuspidate apophysis on inner apical margin; enp- 3 nearly as small as enp- 1, with 1 – 2 setae. P 5 with distinct exopod in both sexes, baseoendopod smaller than exopod, with outer seta arising from small projection; endopodal lobe completely absorbed into basis, with 2 setae in the female and 1 seta in the male. Exopod apically with 3 bipinnate setae.	en	George, Kai Horst (2021): Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species. European Journal of Taxonomy 774: 1-41, DOI: https://doi.org/10.5852/ejt.2021.774.1525, URL: http://dx.doi.org/10.5852/ejt.2021.774.1525
