taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577239/files/figure.png	https://doi.org/10.5281/zenodo.5577239	FIGURE 1. (A) Opheliidae species reported in the Arabian Gulf including references; square showing area detailed in (B); (B) location of samples collected in the coast of Kuwait with presence of Polyophthalmus zhadanae sp. nov.; circles showing ranges of abundance; report of Polyophthalmus pictus (Dujardin, 1839) by Mohammad (1971) marked with red circle.	FIGURE 1. (A) Opheliidae species reported in the Arabian Gulf including references; square showing area detailed in (B); (B) location of samples collected in the coast of Kuwait with presence of Polyophthalmus zhadanae sp. nov.; circles showing ranges of abundance; report of Polyophthalmus pictus (Dujardin, 1839) by Mohammad (1971) marked with red circle.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577241/files/figure.png	https://doi.org/10.5281/zenodo.5577241	FIGURE 2. Polyophthalmus zhadanae sp. nov. Line drawings. (A) complete specimen, lateral view (sample P11072; holotype MNCN 16.01/19108); (B–C) prostomium, dorsal and ventral view respectively (sample P11298; paratype MNCN 16.01/19104); (D–E) posterior end, left and right views respectively (sample P11072; holotype MNCN 16.01/19108). Abbreviations: bo— buccal opening; lae—lateral eyes; lg—lateral groove; nuo—nuchal organ; pmp—paired marginal papillae; pr—prostomium; vg—ventral groove.	FIGURE 2. Polyophthalmus zhadanae sp. nov. Line drawings. (A) complete specimen, lateral view (sample P11072; holotype MNCN 16.01/19108); (B–C) prostomium, dorsal and ventral view respectively (sample P11298; paratype MNCN 16.01/19104); (D–E) posterior end, left and right views respectively (sample P11072; holotype MNCN 16.01/19108). Abbreviations: bo— buccal opening; lae—lateral eyes; lg—lateral groove; nuo—nuchal organ; pmp—paired marginal papillae; pr—prostomium; vg—ventral groove.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577243/files/figure.png	https://doi.org/10.5281/zenodo.5577243	FIGURE3. Polyophthalmuszhadanae sp.nov. Stereomicroscopeimages oflive specimensfrom Kuwait.(A)completespecimen, dorsal view (sample P13418; A. Zhadan leg.); (B) same specimen, anterior end, dorsal view (body chaetigers numbered); (C) complete specimen, dorsal view (no sample number; Al Nuwaiseeb; 05.11.2016); (D–E) same specimen, anterior end, dorsal and ventral view respectively (lateral eyes numbered in (E)). Abbreviations: abp—anterior chaetigers pigmentation pattern; lae—lateral eyes; mpp—midbody and posterior chaetigers pigmentation pattern; nuo—nuchal organ; po—proboscis opening; pr—prostomium; prb—proboscis.	FIGURE3. Polyophthalmuszhadanae sp.nov. Stereomicroscopeimages oflive specimensfrom Kuwait.(A)completespecimen, dorsal view (sample P13418; A. Zhadan leg.); (B) same specimen, anterior end, dorsal view (body chaetigers numbered); (C) complete specimen, dorsal view (no sample number; Al Nuwaiseeb; 05.11.2016); (D–E) same specimen, anterior end, dorsal and ventral view respectively (lateral eyes numbered in (E)). Abbreviations: abp—anterior chaetigers pigmentation pattern; lae—lateral eyes; mpp—midbody and posterior chaetigers pigmentation pattern; nuo—nuchal organ; po—proboscis opening; pr—prostomium; prb—proboscis.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577245/files/figure.png	https://doi.org/10.5281/zenodo.5577245	FIGURE 4. Polyophthalmus zhadanae sp. nov. Stereomicroscope images of fixed specimens from Pole Reef, Saudi Arabia (paratype SMF30263). (A) anterior end, dorsal view (body chaetigers numbered); (B) midbody region, dorsal view; (C) posterior end, dorsal view. Abbreviations: abp—anterior chaetigers pigmentation pattern; mpp—midbody and posterior chaetigers pigmentation pattern.	FIGURE 4. Polyophthalmus zhadanae sp. nov. Stereomicroscope images of fixed specimens from Pole Reef, Saudi Arabia (paratype SMF30263). (A) anterior end, dorsal view (body chaetigers numbered); (B) midbody region, dorsal view; (C) posterior end, dorsal view. Abbreviations: abp—anterior chaetigers pigmentation pattern; mpp—midbody and posterior chaetigers pigmentation pattern.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577247/files/figure.png	https://doi.org/10.5281/zenodo.5577247	FIGURE 5. Polyophthalmus zhadanae sp. nov. SEM micrographs of anterior end of several specimens. (A) sample P13063 (paratype MNCN16.01/19114); (B) prostomium, detail of specimen figured in (A); (C) sample P12629 (paratype MNCN 16.01/19100); (D) sample P12977 (paratype MNCN 16.01/19117); (E) sample P12977 (paratype MNCN 16.01/19117); (F) prostomium, detail of specimen figured in (E). Abbreviations: bo—buccal opening; CH—chaetiger; nuo—nuchal organ; po— proboscis opening; ppa—prostomial papilla; pr—prostomium; prb—proboscis.	FIGURE 5. Polyophthalmus zhadanae sp. nov. SEM micrographs of anterior end of several specimens. (A) sample P13063 (paratype MNCN16.01/19114); (B) prostomium, detail of specimen figured in (A); (C) sample P12629 (paratype MNCN 16.01/19100); (D) sample P12977 (paratype MNCN 16.01/19117); (E) sample P12977 (paratype MNCN 16.01/19117); (F) prostomium, detail of specimen figured in (E). Abbreviations: bo—buccal opening; CH—chaetiger; nuo—nuchal organ; po— proboscis opening; ppa—prostomial papilla; pr—prostomium; prb—proboscis.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577249/files/figure.png	https://doi.org/10.5281/zenodo.5577249	FIGURE 6. Polyophthalmus zhadanae sp. nov. SEM micrographs showingchaetiger parapodial features from severalspecimens. (A) CH1 (sample P13063; paratype MNCN 16.01/19114); (B) CH2 (sample P12977; paratype MNCN 16.01/19117); (C) CH3 (sample P12977; paratype MNCN 16.01/19117); (D) CH10 (sample P12629; paratype MNCN 16.01/19100); (E) parapodial ventral cirrus, detail of chaetiger figured in (D); (F) CH20 (sample P12629; paratype MNCN 16.01/19100); (G) CH28 (sample P12629; paratype MNCN 16.01/19100). Abbreviations: CH—chaetiger; ip—interramal papilla; lcc—long capillary chaetae; pdc—parapodia dorsal cirrus; pvc—parapodial ventral cirrus; scc—short capillary chaetae.	FIGURE 6. Polyophthalmus zhadanae sp. nov. SEM micrographs showingchaetiger parapodial features from severalspecimens. (A) CH1 (sample P13063; paratype MNCN 16.01/19114); (B) CH2 (sample P12977; paratype MNCN 16.01/19117); (C) CH3 (sample P12977; paratype MNCN 16.01/19117); (D) CH10 (sample P12629; paratype MNCN 16.01/19100); (E) parapodial ventral cirrus, detail of chaetiger figured in (D); (F) CH20 (sample P12629; paratype MNCN 16.01/19100); (G) CH28 (sample P12629; paratype MNCN 16.01/19100). Abbreviations: CH—chaetiger; ip—interramal papilla; lcc—long capillary chaetae; pdc—parapodia dorsal cirrus; pvc—parapodial ventral cirrus; scc—short capillary chaetae.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577251/files/figure.png	https://doi.org/10.5281/zenodo.5577251	FIGURE 7. Polyophthalmus zhadanae sp. nov. SEM micrographs of posterior body end showing pygidium in different state of contraction. (A) pygidial tube not contracted (sample P12977; paratype MNCN 16.01/19117); (B) pygidial tube semicontracted (sample P13063; paratype MNCN 16.01/19114); (C) pygidial tube fully contracted (sample P12629; paratype MNCN 16.01/19100); (D) pygidial tube fully contracted (sample P12629; paratype MNCN 16.01/19100). (C) and (D) showing several pygidial papillae enlarged. Abbreviations: CH—chaetiger; pmp—paired marginal papillae.	FIGURE 7. Polyophthalmus zhadanae sp. nov. SEM micrographs of posterior body end showing pygidium in different state of contraction. (A) pygidial tube not contracted (sample P12977; paratype MNCN 16.01/19117); (B) pygidial tube semicontracted (sample P13063; paratype MNCN 16.01/19114); (C) pygidial tube fully contracted (sample P12629; paratype MNCN 16.01/19100); (D) pygidial tube fully contracted (sample P12629; paratype MNCN 16.01/19100). (C) and (D) showing several pygidial papillae enlarged. Abbreviations: CH—chaetiger; pmp—paired marginal papillae.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577253/files/figure.png	https://doi.org/10.5281/zenodo.5577253	FIGURE 8. Polyophthalmus zhadanae sp. nov. Micro-CT images of internal anatomy. (A) anterior end, 3D sagittal section (sample P13063; paratype MNCN 16.01/19115); (B) anterior end, 2D sagittal section (sample P12629; paratype MNCN 16.01/19102); (C) anterior end, 2D frontal section (sample P12629; paratype MNCN 16.01/19102). Abbreviations: adbv— antero-dorsal blood vessel/s; bc—body cavity; bcc—buccal cavity; bo—buccal opening; br—brain; nuo—nuchal organ; obm— oblique muscles; oes—oesophagus; po—proboscis opening; ppa—prostomial papilla; pr—prostomium; prb—proboscis; sep— septum; sto—stomach; vlml—ventral longitudinal muscle layer.	FIGURE 8. Polyophthalmus zhadanae sp. nov. Micro-CT images of internal anatomy. (A) anterior end, 3D sagittal section (sample P13063; paratype MNCN 16.01/19115); (B) anterior end, 2D sagittal section (sample P12629; paratype MNCN 16.01/19102); (C) anterior end, 2D frontal section (sample P12629; paratype MNCN 16.01/19102). Abbreviations: adbv— antero-dorsal blood vessel/s; bc—body cavity; bcc—buccal cavity; bo—buccal opening; br—brain; nuo—nuchal organ; obm— oblique muscles; oes—oesophagus; po—proboscis opening; ppa—prostomial papilla; pr—prostomium; prb—proboscis; sep— septum; sto—stomach; vlml—ventral longitudinal muscle layer.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577255/files/figure.png	https://doi.org/10.5281/zenodo.5577255	FIGURE 9. Polyophthalmus zhadanae sp. nov. Micro-CT images of anterior end of a specimen from sample P12977 (paratype MNCN 16.01/19119). (A) 2D sagittal section; (B) 3D sagittal section. Broken lines in (B) showing transverse sections as found in Fig. 10. Abbreviations: adbv—antero-dorsal blood vessel/s; bc—body cavity; br—brain; obm—oblique muscles; oes—oesophagus; ppa—prostomial papilla; ppabv—prostomial papilla blood vessel; po—proboscis opening; pr—prostomium; prb—proboscis; vnc—ventral nerve cord.	FIGURE 9. Polyophthalmus zhadanae sp. nov. Micro-CT images of anterior end of a specimen from sample P12977 (paratype MNCN 16.01/19119). (A) 2D sagittal section; (B) 3D sagittal section. Broken lines in (B) showing transverse sections as found in Fig. 10. Abbreviations: adbv—antero-dorsal blood vessel/s; bc—body cavity; br—brain; obm—oblique muscles; oes—oesophagus; ppa—prostomial papilla; ppabv—prostomial papilla blood vessel; po—proboscis opening; pr—prostomium; prb—proboscis; vnc—ventral nerve cord.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577257/files/figure.png	https://doi.org/10.5281/zenodo.5577257	FIGURE 10. Polyophthalmus zhadanae sp. nov. Micro-CT images of a specimen from sample P12977 (paratype MNCN 16.01/19119). (A) anterior end, 3D frontal view; (B–E) prostomium, 3D transverse sections; (F) anterior end, 3D transverse section. Abbreviations: adbv—antero-dorsal blood vessel/s; br—brain; dlml—dorsal longitudinal muscle layer; norm—nuchal organ retractor muscle; nuo—nuchal organ; obm—oblique muscles; oes—oesophagus; ppa—prostomial papilla; ppabr— prostomial papilla blood ring; ppabv—prostomial papilla blood vessel; prb—proboscis; sto—stomach; stobpl—stomach blood plexus; vlbv—ventral longitudinal blood vessel; vlml—ventral longitudinal muscle layer; vnc—ventral nerve cord.	FIGURE 10. Polyophthalmus zhadanae sp. nov. Micro-CT images of a specimen from sample P12977 (paratype MNCN 16.01/19119). (A) anterior end, 3D frontal view; (B–E) prostomium, 3D transverse sections; (F) anterior end, 3D transverse section. Abbreviations: adbv—antero-dorsal blood vessel/s; br—brain; dlml—dorsal longitudinal muscle layer; norm—nuchal organ retractor muscle; nuo—nuchal organ; obm—oblique muscles; oes—oesophagus; ppa—prostomial papilla; ppabr— prostomial papilla blood ring; ppabv—prostomial papilla blood vessel; prb—proboscis; sto—stomach; stobpl—stomach blood plexus; vlbv—ventral longitudinal blood vessel; vlml—ventral longitudinal muscle layer; vnc—ventral nerve cord.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577259/files/figure.png	https://doi.org/10.5281/zenodo.5577259	FIGURE 11. Polyophthalmus zhadanae sp. nov. Micro-CT images of a specimen from Abu Ali (paratype SMF30263). (A) 3D sagittal section; (B–E) 3D frontal sections of (A) at several levels.Abbreviations: adbv—antero-dorsal blood vessel/s; bc—body cavity; br—brain; brn—brain nerves; dbv—dorsal blood vessel; dlml—dorsal longitudinal muscle layer; norm—nuchal organ retractor muscle; nuo—nuchal organ; obm—oblique muscles; oes—oesophagus; ppabv—prostomial papilla blood vessel; pr— prostomium; prb—proboscis; prm—proboscis retractor muscle/s; prpr—proboscis posterior region; vlml—ventral longitudinal muscle layer.	FIGURE 11. Polyophthalmus zhadanae sp. nov. Micro-CT images of a specimen from Abu Ali (paratype SMF30263). (A) 3D sagittal section; (B–E) 3D frontal sections of (A) at several levels.Abbreviations: adbv—antero-dorsal blood vessel/s; bc—body cavity; br—brain; brn—brain nerves; dbv—dorsal blood vessel; dlml—dorsal longitudinal muscle layer; norm—nuchal organ retractor muscle; nuo—nuchal organ; obm—oblique muscles; oes—oesophagus; ppabv—prostomial papilla blood vessel; pr— prostomium; prb—proboscis; prm—proboscis retractor muscle/s; prpr—proboscis posterior region; vlml—ventral longitudinal muscle layer.	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577263/files/figure.png	https://doi.org/10.5281/zenodo.5577263	FIGURE 13. World map showing type localities of all Polyophthalmus species described. Black boxes: species considered in the identification key; Grey boxes: species considered as synonyms of P. pictus (Dujardin, 1839).	FIGURE 13. World map showing type localities of all Polyophthalmus species described. Black boxes: species considered in the identification key; Grey boxes: species considered as synonyms of P. pictus (Dujardin, 1839).	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
E81287982C58F763FF518B20FB2BFC0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5577261/files/figure.png	https://doi.org/10.5281/zenodo.5577261	FIGURE 12. Stereomicrographs of fixed specimens identified as Polyophthalmus pictus (Dujardin, 1839) from the coast of Spain. (A) midbody chaetigers, lateral view (three specimens, MNCN 16.01/9398; Almería, Mediterranean coast); (B–C) midbody region and posterior end, lateral view (same specimen, MNCN 16.01/19100; Galicia, Atlantic coast); (D–E) posterior end, lateral view (two specimens, MNCN 16.01/13213; Cantabria, Atlantic coast).	FIGURE 12. Stereomicrographs of fixed specimens identified as Polyophthalmus pictus (Dujardin, 1839) from the coast of Spain. (A) midbody chaetigers, lateral view (three specimens, MNCN 16.01/9398; Almería, Mediterranean coast); (B–C) midbody region and posterior end, lateral view (same specimen, MNCN 16.01/19100; Galicia, Atlantic coast); (D–E) posterior end, lateral view (two specimens, MNCN 16.01/13213; Cantabria, Atlantic coast).	2021-10-18	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan		Zenodo	biologists	Parapar, Julio;Al-Kandari, Manal;Candás, María;Moreira, Juan			
