taxonID	type	description	language	source
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	description	urn: lsid: zoobank. org: act: F 3840017 - E 3 FA- 4 A 8 A- 99 EC- 15 EB 8 C 79310 D Figs 2, 5 C	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	diagnosis	Diagnosis Homotropus hellqvisti sp. nov. (Fig. 2 A – B) is similar to Homotropus collinus (Stelfox, 1941) (Fig. 5 D, F), but is primarily distinguished based on the irregular microsculpture on the 3 – 7 th tergites (Fig. 2 B) and the shorter and more weakly excised clypeus (Fig. 5 C). The new species is also similar to Homotropus melanogaster (Holmgren, 1872) (Fig. 5 E) from which it differs by the only weakly laterally compressed metasoma (in H. melanogaster the metasoma is strongly compressed, knife-like from the 5 th tergite onwards), the irregular microsculpture on the 3 – 7 th tergites (Fig. 2 B), the absence of a subbasal dark spot on the hind tibia and the usually more numerous flagellomeres. Note that while the females of H. melanogaster usually have the mid and hind coxae infuscate, the species sometimes occurs in a colour variety with red hind coxae in Sweden (Klopfstein 2014 b: 68 – 69) (see Remarks). The number of flagellomeres ranges from 17 – 19 in H. melanogaster (n = 20) and 19 – 20 in H. hellqvisti sp. nov. Two of the three known females all have distinct, quite large shoulder marks, while only one of the studied females of H. melanogaster (n = 17) has small and diffuse shoulder marks.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	etymology	Etymology The specific epithet hellqvisti (masculine name in genitive case) is derived from the name of one of the collectors of the specimens who revealed the presence of this new species, Sven Hellqvist (Umeå, Sweden).	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	materials_examined	Type material Holotype SWEDEN • ♀; Västerbotten, Åselet, Hästholmen; 65.233 ° N, 20.414 ° E; 25 May – 19 Aug. 2019; R. Mugerwa Pettersson and S. Hellqvist leg.; Malaise trap in riverside, herb rich, mixed forest; NHRS- HEVA 000011635. Paratypes SWEDEN • 2 ♀♀; same collection data as for holotype; NHRS-HEVA 000011636, NHRS- HEVA 000011637.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	description	Description Female Fore wing length 4.2 – 4.3 mm. Body length 5.5 mm. Head transverse in anterior view. Inner orbits parallel. Face entirely coriaceous and matt. Clypeus with apical margin bilobed and thin, weakly excised centrally (Fig. 5 C). Antennae with 19 – 20 flagellomeres. First flagellomere about four times as long as wide. Apical flagellomeres with multiporous plate sensillae also present ventrally. Head behind eyes slightly narrowed. Temple in lateral view about 0.6 times as wide as compound eye. Mesoscutum matt with fine rugulose microsculpture and very weak punctures dissolving against background. Mesopleuron in lower half irregularly rugulose, in upper half polished, below subtegular ridge with irregular punctures. Metapleuron coriaceous. Propodeum evenly coriaceous without carinae. Coxae coriaceous, matt. Fore wing with areolet closed and vein 3 rs-m unpigmented (Fig. 2 A). Metasoma laterally weakly compressed from the 4 th tergite onwards (Fig. 2 A), with dense hairs. First tergite stout, slightly longer than wide, evenly coriaceous with a faint indication of longitudinal striae in apical corners. Second tergite evenly coriaceous. Third tergite onwards with fine irregular microsculpture (Fig. 2 B). Spiracle of second and third tergite above lateral fold. Ovipositor sheath apically rounded. COLOURATION. Body and head black. Mandibles, palpi, occasionally subtegular ridge, hind corner of pronotum and tegulae whitish. Clypeus yellowish. Pale central face patch absent or diffuse. Yellowish shoulder mark small or absent. Scutellum black. Pterostigma transparent brownish. Legs orange, fore coxae white, basally more or less black. Mid and hind coxae orange. Trochanter and trochantelli yellow except for hind trochanter, which is orange. Tibiae whitish. Fore and mid tibia with yellow apices. Hind tibia with apical 0.25 black and base very narrowly infuscate. Hind tarsi black. Mid and fore tarsi whitish, except for brownish apical segment. Metasoma black. Male Unknown.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	discussion	Remarks Klopfstein (2014 b) noted that some females of Homotropus melanogaster collected in Sweden had orange hind and mid coxae, thereby differing in colour from the nominate form with black coxae. Since both forms also often occurred together at the same localities, they were treated as varieties of the same species. The actual specimens with the red coxae studied by Klopfstein bear a handwritten label “ sp. X ”. I have studied 5 ♀♀ and 2 ³³ of this form of H. melanogaster (NHRS, SMTP) and it is not conspecific with the species here described and apart from the paler coxae there are no reliable distinguishing characters between the two forms. In the key to the species of Homotropus Förster, 1869 provided by Klopfstein (2014 b), Homotropus hellqvisti sp. nov. will run to either couplet 6 together with Homotropus melanogaster or to couplet 8 with H. collinus. For distinguishing characters, see Diagnosis.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBAFFF26D75CEF2FD8EF966.taxon	biology_ecology	Ecology The three known females were collected by a Malaise trap situated in a herb rich riverside boreal forest dominated by birch Betula spp. along the forest river Byskeälven The area (Fig. 12 A – B) rests on sandy ground and is periodically flooded. The trap also collected several rarely recorded diplazontines such as Syrphoctonus idari Diller, 1985, Homotropus melanogaster, Diplazon neoalpinus Zwakhals, 1979 and Syrphophilus asperatus Dasch, 1964.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	description	urn: lsid: zoobank. org: act: F 9 C 92543 - EED 5 - 40 A 7 - AF 51 - DCFC 79 DCE 49 D Figs 3 – 4	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	diagnosis	Diagnosis Homotropus klopfsteinae sp. nov. (Figs 3, 4 A – E) is easily distinguished from other diplazontines by the combination of the strongly sculptured first and second tergites, the strongly convex clypeus which is protruding in lateral view, the unique constitution of the propodeal carinae, with four strongly raised longitudinal carinae and a central arcuate formation, and the unique colour combination with the entire scutellum, part of inner orbits and the basal half of the hind tibia pale yellowish. The known female is also strikingly larger than most species of Diplazontinae known from the Western Palaearctic.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	etymology	Etymology The species epithet klopfsteinae (feminine name in genitive case) is in honour of Seraina Klopfstein (Bern, Switzerland), for her contribution to the systematics of Diplazontinae wasps.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	materials_examined	Type material Holotype SWEDEN • ♀; Norrbotten, Haparanda, Seskarö, Santasaari; 65.777 ° N, 23.846 ° E; 5 Aug. – 6 Sep. 2017; N. Ryrholm and C. Källander leg.; MV-light trap in herb rich coastal meadow on sandy ground; Barcode deposited under GenBank accession number MT 268209; NHRS-HEVA 000011629.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	description	Description Female Fore wing length 7.8 mm. Body length: 9.0 mm. Head transverse in anterior view (Fig. 4 A). Inner orbits parallel. Face entirely coriaceous with weak, scarce punctures (Fig. 4 A). Central area below antennal sockets distinctly elevated. Clypeus with apical margin weakly bilobed, strongly convex in lateral view. Antenna with 23 flagellomeres. First flagellomere about 4 times as long as wide. Mesopleuron weakly punctate in lower half, punctures ventrally dissolving against strongly coriaceous background. Speculum and area around mesopleural furrow shining, impunctate (Fig. 4 E). Epicnemial carina widely interrupted centrally behind fore coxae. Metapleuron in lower part strongly coriaceous, along upper margin more weakly sculptured. Mesoscutum laterally strongly polished with large distinct punctures, interstices between punctures almost equal to their diameter. Mesoscutum centrally with more or less distinct microsculpture between punctures. Scutellum with weak punctures. Propodeum weakly rugose with rather irregular, but distinct arcuate carinae emanating from the metasomal incision (Fig. 4 D), with two strongly raised, almost flange-like longitudinal carinae laterally on each side. Fore wing with areolet closed and vein 3 rs-m pigmented. Metasoma elongate, dorsoventrally depressed. Hind margins of tergites almost straight. First tergite about two times as long as wide in dorsal view, with strong coriaceous sculpture and partly longitudinally striate laterally and basally. Second tergite about 1.3 times as long as wide in dorsal view. Dorsally with strong striation over most of its surface (Fig. 4 B). Third tergite polished with weak microsculpture, in basal half coriaceous and matt. Tergites 4 – 7 polished with weak reticulate-coriaceous microsculpture (Fig. 4 C). Spiracle on second and third tergites above lateral fold. Ovipositor sheath apically rounded. COLOURATION. Body and head black. Clypeus in ventral 0.8, mandibles and palpi yellow. Face black with two elongate pale spots along inner orbits. Antennae black. Large shoulder mark, tegulae, small mark on subtegular ridge and scutellum yellow. Legs yellow. Fore coxae black basally, yellow in apical half. Mid and hind coxae entirely black. Hind femur black, basally and apically narrowly yellow. Hind tibia yellow with apical 0.4 – 0.5 black. Hind tarsus black. Metasoma black with posterior margins of tergites 2 – 4 paler. Ovipositor sheath pale brownish. Male Unknown.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	biology_ecology	Ecology The only known female was collected by a MV-light trap placed on a herb rich coastal meadow on sand (Fig. 13) between the 5 th of August and the 6 th of September.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBCFFF16D46CEEAFA84F9FD.taxon	discussion	Remarks According to the phylogenetic analysis (Fig. 1), Homotropus klopfsteinae sp. nov. belongs to the Syphoctonus genus group, where it is situated between some rather atypical species of Homotropus (H. crassicornis Thomson, 1890 and H. vitreus Dasch, 1964) and the genera Phthorima Förster, 1869 and Bioblapsis Förster, 1869. This, in combination with the unique combination of characters, suggests that the species possibly should be assigned to a new genus. However, considering the present, unresolved situation in the genus group, a generic revision, preferably including molecular methods, would be needed to establish more strongly supported generic delimitations. The new species is here placed in the genus Homotropus, mainly based on the phylogenetic analysis in combination with the general appearance of the species, i. e., the absence of a strongly elongated and laterally compressed metasoma with distinctly concave hind margins of the 3 – 5 th tergites, typical of females belonging to Bioblapsis or Phthorima. Given the quite distinct characteristics of the species it runs to couplet 13 in the key to genera in Klopfstein (2014 b), where it is distinguished from Fossatyloides gracilentus (Holmgren, 1858) by the colouration, larger size, the more convex clypeus and the more distinct carination of the propodeum.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	description	urn: lsid: zoobank. org: act: 314328 E 3 - 4 E 48 - 45 CD-A 609 - E 0 BBA 965 B 5 ED Figs 5 B, 6	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	diagnosis	Diagnosis Homotropus mugerwai sp. nov. (Fig. 6) is similar to and easily confused with Homotropus pallipes (Gravenhorst, 1829), but has the mesoscutum polished with quite distinct punctures (Fig. 5 B), while it is shagreened with indistinct punctures in H. pallipes (Fig. 5 A). The area covered by microsculpture on the mesopleuron is ending lower than in Homotropus pallipes. The two known females lack pale shoulder marks anteriorly on the lateral part of mesoscutum, a feature which is present in all but one female specimen of H. pallipes studied by the author (n = 89), even if the mark is smaller and less distinct in northern specimens. The pterostigma is brownish with the basal part paler, but entirely yellowish in H. pallipes. The mid-coxae in the two known females are almost entirely yellow, while they are orange at least basally in H. pallipes. The species is also similar to Homotropus signatus (Gravenhorst, 1829), but has more numerous flagellomeres, 20 in Homotropus mugerwai sp. nov. and 19 in H. signatus. The metasoma is also less strongly compressed apically than in H. signatus. The scutellum is entirely yellow in Homotropus mugerwai sp. nov. and the hind trochanter has a dark spot while the scutellum in H. signatus is black or only with a small yellow spot apically and the hind trochanter lacks any dark markings.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	materials_examined	Type material Holotype SWEDEN • ♀; Västerbotten, Åselet, Hästholmen; 65.233 ° N, 20.414 ° E; 25 May – 19 Aug. 2019; R. Mugerwa Pettersson and S. Hellqvist leg.; Malaise trap in riverside, herb rich, mixed forest; NHRS- HEVA 000011634. Paratype SWEDEN • 1 ♀; Västerbotten, Fällfors, Marranäset; 65.109 ° N, 20.790 ° E; 25 May – 19 Aug. 2019; R. Mugerwa Pettersson and S. Hellqvist leg.; Malaise trap in riverside, decidous forest. NHRS- HEVA 000011633.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	etymology	Etymology The specific epithet mugerwai (masculine name in genitive case) is derived from the name of one of the the collectors, Roger Mugerwa Pettersson (Robertsfors, Sweden).	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	description	Description Female Fore wing length 4 mm. Body length 4.5 mm. Head transverse in anterior view. Inner orbits almost parallel. Face entirely coriaceous and matt. Clypeus with apical margin bilobed and thin, weakly excised centrally. Antennae with 20 flagellomeres. First flagellomere about 5.5 – 6.0 times as long as wide. Apical flagellomeres with multiporous plate sensilla also present ventrally. Head behind eyes slightly narrowed. Temple in lateral view about 0.5 times as wide as compound eye. Mesopleuron in lower half with weak rugulose microsculpture, in upper half polished. Metapleuron strongly coriaceous. Mesoscutum shining with small punctures, centrally with very weak microsculpture (Fig. 5 B). Propodeum evenly coriaceous with only pleural carinae present. Fore wing with areolet closed and vein 3 rs-m weakly pigmented. Coxae coriaceous, matt. Pterostigma light brownish, basally pale (Fig. 6). Metasoma laterally weakly compressed from the 4 th tergite (Fig. 6). First tergite about 1.5 times as long as wide, coriaceous with weak longitudinal striae. Second tergite coriaceous with weak striation basally, striae reaching about 0.5 times length of tergite. Third tergite coriaceous, apically becoming more or less reticulate-coriaceous. Fourth tergite onwards reticulate-coriaceus. Spiracle of second and third tergite above lateral fold. Ovipositor sheath apically rounded. COLOURATION. Body and head black. Mandibles, lower part of clypeus, palpi, subtegular ridge, hind corner of pronotum, tegulae and weak central spot on face whitish. Scutellum entirely yellow. Legs orange. Fore and mid coxae, trochanter and trochantelli yellow, fore and mid coxae at most with very small dark spot basally. Hind coxae orange. Hind trochantellus with small black spot. Tibae whitish, with apical 0.3 and base narrowly black. Hind tibia with subbasal black spot dorsally. Metasoma black. Apical third of tergite 2 and the entire tergites 3 and 4 orange. Male Unknown.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	discussion	Remarks In the key to the species of Homotropus provided by Klopfstein (2014 b), Homotropus mugerwai sp. nov. will run to couplet 30 as H. pallipes, from which it is distinguished by the characters given in the diagnosis.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFBFFFFC6D4BCE7CFE77F954.taxon	biology_ecology	Ecology The two known females were collected by Malaise traps situated in herb rich riverside boreal forests dominated by Betula spp. (Fig. 12 A – B). Both areas are periodically flooded. The trap at Hästholmen also collected several other rarely recorded diplazonines such as Syrphoctonus idari, Homotropus melanogaster, Diplazon neoalpinus, Syrphophilus asperatus and the here described Homotropus hellqvisti sp. nov.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	description	urn: lsid: zoobank. org: act: 015 B 88 D 4 - B 3 AA- 4 A 41 - B 694 - 425 A 00662113 Figs 7 – 8	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	diagnosis	Diagnosis Separated from all other species of Homotropus primarily by the unique colour pattern with the legs, metasoma and antennae almost entirely red and the mesosoma with extensive red markings (Fig. 7). Also distinguished by the strong sculpture and carination of the propodeum (Fig. 8 B – C). Perhaps superficially most similar to Homotropus haemorrhoidalis Szépligeti, 1898, but that species has a slightly different colouration with for example the antennae and most of metasoma black and all coxae orange. Furthermore, the apical flagellomeres have short thick bristles in H. haemorrhoidalis and the propodeum has less prominent carinae.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	etymology	Etymology The specific epithet riedeli (masculine name in genitive case) is in honour of Matthias Riedel (Bad Fallingbostel, Germany), for his contributions to the systematics of ichneumonid wasps.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	materials_examined	Type material Holotype SWEDEN • ♀; Skåne, Ystad, Kåseberga; 55.385 ° N, 14.066 ° E; 17 Jul. – 14 Sep. 2013; N. Ryrholm and C. Källander leg.; MV-light trap in warm coastal sandslopes; NHRS-HEVA 000011630.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	description	Description Female Fore wing length 5.0 mm. Body length 6.9 mm. Head transverse in anterior view. Inner orbits almost parallel. Face entirely coriaceous and matt (Fig. 8 A). Clypeus with apical margin bilobed and thin. Clypeus in basal 0.66 weakly convex. Antennae with 20 flagellomeres. Head behind eyes strongly narrowed. Temple in lateral view about 0.3 times as wide as compound eye. Mesopleuron centrally anteriorly with shallow punctures. The interstices about equal to diameter of punctures. In lower part, punctures become indistinct against coriaceous background. In upper part, punctures partly fading to irregular, almost rugose sculpture (Fig. 8 C). Area around mesopleural furrow polished without punctures. Metapleuron coriaceous, shining. Mesoscutum shining with weak microsculpture between punctures, centrally becoming more strongly coriaceous. Scutellum weakly coriaceous with distinct punctures, interstices between punctures slightly larger than diameter of punctures. Propodeum strongly irregularly rugose, with area petiolaris and indicated area superomedia delimited by strong, irregular carinae (Fig. 8 B). Indicated area superomedia with surface covered with irregular transverse carinae. Coxae coriaceous, matt. Fore wing with areolet closed and vein 3 rs-m pigmented (Fig. 7). Metasoma laterally compressed from 4 th tergite. First tergite strongly longitudinally striate. Second tergite strongly coriaceous with weak striation basally. Third tergite polished with weak microsculpture, in basal half coriaceous and matt. Tergites 4 – 7 polished with weak microsculpture. Spiracle of second tergite above lateral fold and on third tergite on lateral fold. COLOURATION. Body and head black. Antennae reddish brown. Scapus brown, fading to brownish yellow ventrally. Clypeus, mandibles except mandibular teeth, palpi and large central face patch yellow. Face below antennal sockets dark reddish brown. Lower part of mesopleuron, metapleuron and a large spot on propodeum laterally orange. Scutellum orange. Large shoulder mark, subtegular ridge, tegulae, hind margin of mesopleuron and large spot at the hind corner of pronotum yellow. Legs orange. Fore and mid coxae yellow, orange basally. Trochanters and trochantelli on fore and mid leg yellow. Hind trochanter and trochantelli with large yellow spot. Hind legs apart from trochanter and trochantelli, entirely orange. Metasoma orange. First tergite centrally black with lateral and apical margins widely orange. Tergites 7 and 8 black. Male Unknown.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB2FFFB6D65CEC5FC8DF9F9.taxon	biology_ecology	Ecology The only known specimen was collected by a MV-light trap placed in an area characterized by warm steppe-like sandy heaths and herb-rich coastal slopes at the southern tip of Sweden. The female was collected between the 7 th of July and the 14 th of September.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	description	urn: lsid: zoobank. org: act: 5130 CD 6 D-CB 68 - 403 E-AF 7 F- 1 F 76 C 8 E 5 B 558 Figs 9 – 10	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	materials_examined	Type material Holotype SWEDEN • ³; Härjedalen, Hede, Synderkölen; 62.294 ° N, 13.406 ° E; 5 – 7 Aug. 2016; N. Johansson leg.; Sweepnet along alpine stream / bog; Barcode deposited under GenBank accession number MT 268212; NHRS-HEVA 000011631. Paratypes SWEDEN • 1 ♀; Lappland; 10 Aug. (year unknown); C. H. Boheman (1796 – 1868) leg.; NHRS- HEVA 000010749 • 1 ♀; Härjedalen, Hede, Synderkölen; 62.292 ° N, 13.405 ° E; 28 Jul. 2019; N. Johansson leg.; sweepnet along alpine stream / bog; NHRS-HEVA 000011632. NORWAY • 1 ♀; Oppland, Dovre; C. H. Boheman (1796 – 1868) leg.; NHRS-HEVA 000010750.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	diagnosis	Diagnosis Woldstedtius aureotibialis sp. nov. (Figs 9 A – C, 10 A – C) is distinguished from Woldstedtius patei (Dasch, 1964) (Fig. 11) by the orange legs, the less extensive pale colouration of the mesosoma and the less distinct carination of the propodeum (Fig. 10 C). Woldstedtius patei, known from two specimens collected in the southern part of the Rocky Mountains in North America, has the fore and mid coxae orange, basally infuscate with the apices largely yellow (Fig. 11 A – C). The hind femur is orange with the inner side infuscate and the hind tibiae are whitish, apically infuscate and with a small subbasal black spot. Furthermore, the yellow shoulder mark, the yellow apices of the scutellum and postscutellum are more extensive in W. patei (Fig. 11 A – C) while the orange spot on the metapleuron is considerably smaller.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	etymology	Etymology The specific epithet aureotibialis (adjective) is derived from the colour of the hind tibia, aureo = golden.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	description	Description Fore wing length 3.1 – 4.2 mm. Body length 4.5 – 5.5 mm. Head in anterior view strongly transverse, central area strongly elevated. Inner orbits almost parallel (Figs 9 B, 10 B). Antennae with 22 flagellomeres. First flagellomere about 5 times as long as wide. Second flagellomere about 3.5 – 4 times as long as wide. Clypeus in basal third with a transverse swelling, separated from face by a shallow groove. Face evenly and finely coriaceous, matt (Figs 9 B, 10 B). Clypeus with apical margin bilobed, in basal 0.3 distinctly convex. Temple in dorsal view strongly narrowed behind eye. Mesopleuron almost entirely coriaceous, only a small area around mesopleural furrow shining. Sternauli weakly impressed. Epicnemial carina complete, metapleuron strongly coriaceous, matt. Mesoscutum and scutellum evenly and entirely coriaceous. Propodeum strongly coriaceous with only lateral longitudinal carinae present basally, centrally in anterior part with small depression covered by irregular transverse carinae (Fig. 10 C). Hind coxae coriaceous. Fore wing areolet open, hind wing with three basal hamuli. Metasoma dorsoventrally depressed. Tergites coriaceous, weakly shining. Second tergite basally with longitudinal striae. Third tergite with spiracle slightly above lateral fold. Ovipositor apically truncate. Colouration of male Body and head black. Antennae ventrally orange. Scapus yellow ventrally. Face, clypeus, palpi and temple in ventral part entirely yellow. Lower part of propleuron and mesopleuron yellow. Suture between meta- and mesopleuron yellow (Fig. 9 C). Large shoulder mark, tegulae, subtegular ridge and hind corner of pronotum yellow. Scutellum with small yellow apical spot. Legs orange. Coxae yellow. Hind coxae at base dorsally black and with an irregular black stripe on outer side. Trochanters and trochantelli yellow. Hind trochantelli basally with small black spot. Hind femur on inner side with a large black stripe, which covers almost the entire inner surface. Hind tibia orange, infuscate apically. Hind tarsi infuscate. Metasoma black. Second tergite with a small central yellow spot posteriorly. Third and fourth tergites with large t-shaped yellow spot. Fifth tergite basally yellow (Fig. 9 A). Laterotergites and parameres yellow. Colouration of female The three known females are quite consistent regarding colouration (Fig. 10 A – C). Body and head black. Tegulae and very small spot on hind corner of pronotum whitish. Palpi yellow. Mandibles yellow with more or less extensive black mark. Metapleuron basally with large red mark and posterior basal corner on mesopleuron with smaller red mark, the latter may be absent.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	discussion	Remarks The taxonomic status of the paratype of Woldstedtius patei from Alaska (1 ♀; Independence Mine; 6 Aug. 1956; W. C. Frohne leg.; USUC) is somewhat uncertain. The orange legs and the slightly weaker carination of the propodeum would suggest that the specimen also belongs to the species here described, but the more abundant pale markings of the coxae and mesosoma are features shared by Woldstedtius patei. It is therefore not included in the type series. Klopfstein (2014 b) based the decision to assign the then two known Scandinavian females to W. patei on the study of the Alaskan paratype and was, according to the notes (p. 113), unaware of a second, non-type female from Colorado (1 ♀; Colorado, Mount Evans; 26 Jul. 1964; C. Dasch leg.; 9800 ’ trap; USUC), thereby overlooking the identity of the species. The morphological differences between W. patei and W. aureotibialis are subtle. The shape and structure of the propodeum is consistent in the three Scandinavian females, which all have the carination of propodeum less distinct (Fig. 10 C) than in the two Nearctic females, including the holotype, which both have a diamond-shaped area surrounded by weak carinae centrally (Fig. 11 B). Furthermore, the colouration of the legs, in which the two species differ considerably, is regarded as a stable character in diplazontines, commonly used in species diagnosis and delimitation (Dasch 1964; Klopfstein 2014 b). The phylogenetic analysis places Woldstedtius aureotibialis sp. nov. at a rather basal position within the genus, as sister to a clade including W. flavolineatus and two Neotropical species (Fig. 1).	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFB5FFE76D5BCE70FE7AFB99.taxon	biology_ecology	Ecology The male holotype and one of the female paratypes were sweep netted between the 5 th and 10 th of August by an open bog with scattered Salix bushes (primarily Salix lapponum L.) surrounding a small stream in an alpine area.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFA9FFE76D43CCC3FACBF969.taxon	materials_examined	Material examined SWEDEN • 1 ♀; Halland, Kungsbacka, Förlanda, Äskhult; 57.430 ° N, 12.282 ° E; 2 Jun. – 15 Jul. 2016; K. Larsson leg.; Malaise trap in rocky grazed, small scale farmland; NJ.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFA9FFE76D43CCC3FACBF969.taxon	discussion	Remarks Previously known from Switzerland, Austria, Russia, Ukraine, Germany, Italy, Macedonia and Turkey (Klopfstein 2014 b). The trap where the single female was collected was situated in a graze meadow in the culture and nature reserve Äskhult, where an ancient agricultural landscape has been restored.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFA9FFE56D25CE9EFEE9FD95.taxon	materials_examined	Material examined SWEDEN • 1 ♀; Åsele lappmark, Åsele, Björnlandets nationalpark, Häggsjöbäcken; 63.949 ° N, 17.999 ° E; 29 Jul. – 26 Aug. 2014; A. Garpebring leg.; Malaise trap in boreal forest; NJ • 1 ♀; Östergötland, Boxholm, Rinna, Björneberg; 58.195 ° N, 14.911 ° E; 15 – 27 May 2018; N. Johansson leg.; NJ.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
03A287DEFFA9FFE56D25CE9EFEE9FD95.taxon	discussion	Remarks Described by the Swedish Entomologist C. G. Thomson from material collected by J. Kriechbaumer from Bavaria, Germany (Thomson 1890). The distribution covers the Palaearctic, but the species seems to be rarely recorded (Klopfstein 2014 b). The Swedish records are both from mature, semi-open taiga forests.	en	Johansson, Niklas (2020): Additions to the Swedish fauna of Diplazontinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 724: 70-92, DOI: https://doi.org/10.5852/ejt.2020.724.1159
