taxonID	type	description	language	source
03EBFE378313FFB715F4FC8AFE819789.taxon	description	(Figs. 1 − 9; Map 1) urn: lsid: zoobank. org: act: 986 EFB 79 - 5 AEE- 4 CD 4 - AEDE- 58 D 6 EDDE 58 E 5	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378313FFB715F4FC8AFE819789.taxon	materials_examined	Type material. Holotype: Turkey: ♂, “ TR. Çorum province, Osmancık, Danişment 4 km S, 1480 m, 41 ° 04 ’ 26 ’’ N, 34 ° 57 ’ 03 ’’ E, 01. V. 2021, leg. Örgel, Kacar & Çelik. / Holotypus ♂, Lathrobium (Lathrobium) kacari sp. n. det. S. Anlaş 2021 ” (AZMM). Paratypes: 8 ♂, 13 ♀, same data as holotype (AZMM). 1 ♂, 2 ♀, Osmancık, Danişment 3 km S, 1490 m, 41 ° 04 ’ 36 ’’ N, 34 ° 55 ’ 48 ’’ E, 01. V. 2021, leg. Örgel, Kacar & Çelik (AZMM). 4 ♂, 4 ♀, Osmancık, Danişment 3 km E, 1461 m, 41 ° 04 ’ 37 ’’ N, 34 ° 56 ’ 04 ’’ E, 01. V. 2021, leg. Örgel, Kacar & Çelik (AZMM).	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378313FFB715F4FC8AFE819789.taxon	description	Description. Habitus as in Figure 1. Body 5.4 − 5.9 mm long. Colouration: forebody uniformly reddish brown, abdominal segments III – VI blackish or dark brown, abdominal segments VII – X reddish brown; antennae reddish brown, and legs yellowish brown. Head slightly oblong, approximately 1.05 − 1.10 times as long as wide (Figs. 1 − 2); integument with shallow fine microreticulation; punctation non-umbilicate, relatively coarse and not very dense, interstices on average slightly wider than diameter of punctures in lateral areas and wider in median dorsal area; pubescence brownish and very sparse; eyes reduced, slightly protruding from lateral contours of head (Fig. 3). Antennae long and slender (Fig. 1), approximately 1.8 − 1.9 mm long, all antennomeres distinctly oblong. Pronotum with subparallel lateral margins in dorsal view, distinctly oblong, approximately 1.30 − 1.35 times as long as wide (Figs. 1 − 2), approximately 0.95 times as wide as head and narrowed posteriorly; punctation similar to that of head, but slightly denser and larger; pubescence brownish and very sparse; microsculpture absent. Elytra at suture distinctly shorter and broader than pronotum, approximately 1.15 times as wide as pronotum (Figs. 1 − 2), punctation similar to that of pronotum, but well-defined and slightly smaller; pubescence brownish and very sparse; microsculpture absent. Hind wings completely reduced. Abdomen wider than elytra (Fig. 1), approximately 1.15 - 1.20 times as wide as elytra, widest at segment VII; punctation fine, small and shallow, moderately dense and well-defined; all tergites with distinct microsculpture; posterior margin of tergite VII without palisade fringe. ♂: sternite VII distinctly modified, with semi-circular median impression and with broadly concave posterior margin, median impression bearing short, stout and black setae (Fig. 3), posterior margin of sternite VIII with triangular median emargination, along median line without pubescence (Fig. 4). Aedeagus large (Figs. 7 − 9), approximately 1.05 − 1.10 mm long, ventral process stout and longer, triangular-shaped apically in ventral view; apical sclerotized structure of internal sac in lateral view shaped like a spoon apically and basally bent (Fig. 7 − 8). ♀: posterior margin of tergite VIII distinctly convex (Fig. 5); sternite VIII increasingly narrowed, with convex projection; without micro-pubescence in median posterior area (Fig. 6); tergite IX separated in the middle, lateral processes long, with acute spine-like extension apically; posterior margin of tergite X densely and finely micropubescent. Comparative notes. The new species can be separated from all other congeners in Turkey by the distinctive morphology of the aedeagus in both ventral and lateral view. It is additionally distinguished as follows: From L. brignolii Bordoni, L. distortum Assing and L. deforme Assing by the only weakly reduced eyes: in L. brignolii, eyes completely reduced, without ommatidia or pigmentation; in L. distortum Assing and L. deforme, eyes very small, barely visible in dorsal view, without distinct ommatidia, but with pigmentation. The above species are also completely different in sexual characters. From L. bodemeyeri Bernhauer by the much less modified setae, which do not extend further anteriad on male sternite VII, by the lack of micro-pubescence on the median posterior area of female sternite VIII, and by the different shape of the apical sclerotized structure of the internal sac in the aedeagus. In L. bodemeyeri, the apical sclerotized structure of internal sac is shaped like a hook apically in lateral view. From L. newtoni Anlaş by the darker colouration of the forebody, by the much less modified setae, which do not extend further anteriad on male sternite VII, by the lack of micro-pubescence on the median posterior area of female sternite VIII, and by the different shape of the apical sclerotized structure of the internal sac in the aedeagus. In L. newtoni, the apical sclerotized structure of internal sac is basally curved and apically as a straight spine in lateral view. From L. uncinatum Assing by the darker colouration of the forebody, by the more distinct punctation of the whole body, by the much less modified setae, which do not extend further anteriad on male sternite VII, by the lack of micro-pubescence on the median posterior area of female sternite VIII, and by the different shape of the apical sclerotized structure of the internal sac in the aedeagus. In L. uncinatum, the apical sclerotized structure of internal sac is shaped like a barb apically in lateral view. For descriptions and illustrations of the above species, see Bordoni (1980), Assing (2001, 2007) and Anlaş (2018).	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378313FFB715F4FC8AFE819789.taxon	etymology	Etymology. The species is dedicated to Enes Zafer Kacar, Manisa, who collected some of the type specimens of this new species.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378313FFB715F4FC8AFE819789.taxon	distribution	Distribution and bionomics. The species was found at three localities from Çorum (Osmancık, Danişment). The type specimens were collected under stones in various grassland areas and sifted from leaf litter at altitudes of 1460 − 1490 m.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378311FFB015F4FC46FACB936F.taxon	description	(Figs. 10 − 18; Map 1) urn: lsid: zoobank. org: act: 1 F 5801 B 7 - 4423 - 4 DEA- 8 F 21 - AEC 5154 DE 128	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378311FFB015F4FC46FACB936F.taxon	materials_examined	Type material. Holotype: Turkey: ♂, “ TR. Amasya province, Hamamözü, Tekçam 1 km SE, 1562 m, 40 ° 42 ’ 50 ’’ N, 35 ° 06 ’ 12 ’’ E, 27. IV. 2021, leg. Anlaş, Kacar & Çelik. / Holotypus ♂, Lathrobium (Lathrobium) celiki sp. n. det. S. Anlaş 2021 ” (AZMM). Paratypes: 1 ♂, 2 ♀, same data as holotype (AZMM).	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378311FFB015F4FC46FACB936F.taxon	description	Description. Habitus as in Figure 10. Body 5.7 − 5.9 mm long. Colouration: forebody uniformly reddish brown, head and pronotum slightly darker than elytra, abdominal segments III – VI blackish, abdominal segments VII – X reddish brown; appendages somewhat lighter than body. In general appearance similar to L. kacari, but it can be distinguished as follows: colouration of body darker than L. kacari; head approximately 1.05 times as long as wide (Figs. 10 − 11), punctation slightly denser and much more distinct than that of L. kacari, antennae 1.88 − 1.94 mm long; pronotum longer than wide, approximately 1.20 − 1.25 times as long as wide, punctation slightly denser and much more distinct than that of L. kacari (Figs. 10 − 11); elytra broader than pronotum, 1.17 − 1.24 times as wide as pronotum (Figs. 10 − 11); abdomen wider than elytra (Fig. 10), approximately 1.15 times as wide as elytra. ♂: in general appearance similar to sternite VII and sternite VIII of L. kacari but sternite VII with emargination slightly smaller and region with modified setae extending further anteriad (Fig. 12); posterior margin of sternite VIII with triangular median emargination slightly smaller (Fig. 13). Aedeagus larger than that of L. kacari (Figs. 16 − 18), approximately 1.15 − 1.20 mm long, ventral process longer, basally broader and increasingly narrowed apically in ventral view; apical sclerotized structure of internal sac in lateral view protruding in two parts apically and basally curved, in ventral view oblique oval shaped (Figs. 7 − 8). ♀: in general appearance very similar to tergite VIII, sternite VIII, tergite IX and tergite X of L. kacari; with only very few differences in tergite VIII and sternite VIII (Figs. 14 − 15); sternite VIII increasingly narrowed, with convex projection, and fine and dense micro-pubescence in median posterior area (Fig. 15). Comparative notes. This new species is distinguished from other congeners in Turkey particularly by the distinctive shape of the sclerotized structure of internal sac, and also the different shape of the apex of the ventral process of the aedeagus in both ventral and lateral view (see key to the microphthalmous Lathrobium species of Turkey).	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378311FFB015F4FC46FACB936F.taxon	etymology	Etymology. The species is dedicated to Mutlu Çelik, Manisa, who collected some of the type specimens of this new species.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378311FFB015F4FC46FACB936F.taxon	distribution	Distribution and bionomics. The species was collected at only one locality in Amasya province in central northern Anatolia. The type specimens were collected under stones in grassland at an altitude of about 1560 m.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378317FFBD15F4FF03FE9B95A5.taxon	description	(Figs. 19 − 34; Map 1)	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378317FFBD15F4FF03FE9B95A5.taxon	materials_examined	Additional material examined: Turkey: Bartın: 1 ♂, 2 ♀, 07. IV. 2021, Ulus, Uluyayla, 41 ° 32 ’ 24 ’’ N, 32 ° 48 ’ 16 ’’ E, 1000 m, leg. Örgel, Kacar & Çelik (AZMM). Bolu: 2 ♂, 1 ♀, 18. IV. 2013, Mudurnu, Sülüklü Göl road, leg. Kunt (AZMM). 3 ♂, 2 ♀, 03. VII. 2020, Mudurnu, Ekinören 4 km NE, 40 ° 33 ’ 27 ’’ N, 31 ° 04 ’ 50 ’’ E, 830 m, leg. Örgel & Kacar (AZMM). 2 ♂, 1 ♀, 05. IV. 2021, Mengen, Köprübaşı 5 km N, 40 ° 57 ’ 16 ’’ N, 31 ° 51 ’ 57 ’’ E, 775 m, leg. Örgel, Kacar & Çelik (AZMM). 1 ♂, Yedigöller, 14. V. 1987, leg. Schönmann & Schillhammer, L. bodemeyeri V. Assing det. 2001 (NHMW). Çorum: 1 ♂, 22. VI. 2020, Kargı, Tepelice 3 km W, 41 ° 12 ’ 41 ’’ N, 34 ° 27 ’ 30 ’’ E, 1870 m, leg. Örgel & Kacar (AZMM). Düzce: 1 ♂, 1 ♀, 01. VII. 2021, Kaynaşlı, Biçkıyanı 5 km N, 40 ° 40 ’ 39 ’’ N, 31 ° 22 ’ 21 ’’ E, 1200 m, leg. Kacar & Çelik (AZMM). Eskişehir: 1 ♀, Bodemeyer, Asia-minor, Bos Dagh (= Sündiken Dağları) L. bodemeyeri V. Assing det. 2001 (NHMW). 1 ♂, 12. III. 2018, Sündiken Dağları, 39 ° 59 ’ 26 ” N, 31 ° 00 ’ 47 ” E, 1253 m, leg. Örgel & Yaman (AZMM). 2 ♀, 21. VI. 2016, Sarıcakaya, 40 ° 02 ’ 38 ” N, 30 ° 53 ’ 43 ” E, 336 m, MSS trap (AZMM). Karabük: 1 ♂, 1 ♀, 06. IV. 2021, Eskipazar, Sallar 1 km SW, 40 ° 57 ’ 41 ’’ N, 32 ° 45 ’ 44 ’’ E, 1277 m, leg. Örgel, Kacar & Çelik (AZMM). Kastamonu: 6 ♂, 3 ♀, 06. V. 2021, Azdavay, Topuk 3 km W, 41 ° 41 ’ 01 ’’ N, 33 ° 21 ’ 00 ’’ E, 1010 m, leg. Örgel, Kacar & Çelik (AZMM). 1 ♂, 2 ♀, 10. VII. 2020, Pınarbaşı, Uzla 4 km NW, 41 ° 43 ’ 12 ’’ N, 32 ° 56 ’ 36 ’’ E, 815 m, leg. Örgel & Kacar (AZMM). Sakarya: 1 ♂, Goek Dagh, leg. v. Bodemeyer, L. bodemeyeri V. Assing det. 2001 (NHMW). 1 ♀, klein-Asien, Goek Dagh, leg. Bodemeyer, coll. J. Fodor (cotype) (NMPC). 1 ♀, klein-Asien, Goek Dagh, leg. Bodemeyer, coll. Rambousek (det. C. Koch as L. elangatulum Kraatz) (NMPC). 3 ♂, 4 ♀, 01. VII. 2021, Akyazı, Boztepe 5 km N, 40 ° 30 ‘ 38 ‘‘ N, 30 ° 41 ‘ 44 ‘‘ E, 1058 m, leg. Kacar & Çelik (AZMM). 3 ♂, 6 ♀, 01. VII. 2021, Akyazı, Boztepe 5 km NE, 40 ° 31 ’ 07 ’’ N, 30 ° 42 ’ 32 ’’ E, 1010 m, leg. Kacar & Çelik (AZMM). Sinop: 1 ♀, 19. V. 2014, Tıngıroğlu village, 41 ° 47 ’ 32 ” N, 35 ° 00 ’ 12 ” E, leg. Koç (AZMM). 3 ♂, 3 ♀, 23. VI. 2020, Ayancık, Kurtköy 5 km W, Çangal Dağları, 41 ° 47 ’ 29 ’’ N, 34 ° 44 ’ 23 ’’ E, 1100 m, leg. Örgel & Kacar (AZMM).	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378317FFBD15F4FF03FE9B95A5.taxon	description	Zonguldak: 1 ♂, 1 ♀, 08. IV. 2021, Ayvatlar 2 km SW, 41 ° 17 ’ 54 ’’ N, 31 ° 49 ’ 19 ’’ E, 364 m, leg. Örgel, Kacar & Çelik. Doubtful record: Hatay: 1 ♂, 16. IV. 1966, Arsuz (Hatay), Kompost, Türkei-Exped. 1966, Naturhist. Mus. Wien, wahrscheinlich Fundortsverwechslung, Lathrobium bodemeyeri Bernhauer det. V. Assing 2002.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
03EBFE378317FFBD15F4FF03FE9B95A5.taxon	discussion	Remarks: Bernhauer (1903) described this species based on an unspecified number of syntypes from Goek Dagh (Gökdağ, Samanlı Dağları, Sapanca, Sakarya) in Turkey. A lectotype of this species was designated by Assing (2001) who also provided illustrations. This species is widely distributed in Northwestern Anatolia. The original description of L. paphlagonicum is based on a male holotype, and a male and female paratype from Cangal Dagh (Çangal Mountains) in Sinop province, Turkey (Assing, 2001). Later, Assing (2004, 2009, 2011) recorded this species from several localities in Sinop and Kastamonu provinces. I studied the type specimens of L. bodemeyeri and L. paphlagonicum in the collections of the NHMW during a visit in 2021 (Figs 19 − 34). Accoding to Assing (2001), L. paphlagonicum is distinguished from L. bodemeyeri by the larger body (L. paphlagonicum, 5.4 − 6.3 mm long; L. bodemeyeri, on avarage 6.0 mm long), by the different shape of pronotum in dorsal view (L. paphlagonicum, lateral margins weakly concave a short distance before middle; L. bodemeyeri, lateral margins mostly straight or weakly convex), by the much finer punctation on the abdomen, by the abdomen more distinctly darker than forebody and by the different shape of the aedeagus (L. paphlagonicum, in lateral view with ventral process weakly and smoothly bent, dorso-apical sclerite in lateral view wider; L. bodemeyeri, in lateral view with ventral process more distinctly and more abruptly bent, dorso-apical sclerite in lateral view narrow). An examination of material from various localities of the distribution revealed that both external (Figs 35 − 46) and aedeagal characters (Figs 47 − 64) of L. bodemeyeri are subject to remarkable intraspecific variation and overlap with those of L. paphlagonicum. Consequently, there is little doubt that L. paphlagonicum is conspecific with L. bodemeyeri. Thus, L. paphlagonicum is here proposed as a synonym of L. bodemeyeri. L. bodemeyeri is known from Bartın, Bolu, Çorum, Düzce, Eskişehir, Karabük, Kastamonu, Sakarya, Zonguldak provinces in Turkey (Bernhauer, 1903; Bordoni, 1980, Assing & Schülke, 2000; Assing, 2001, 2004, 2011, 2013, 2014; Anlaş, 2018 and present paper). It seems that this species is confined to northwestern and central western Anatolia. The species has also been recorded from Greece (Makhedonia, W Kilkis, Metallikon, 20. III. 1978, leg. Heinz) by Assing (2001) and from Hatay province in the southest Anatolia (see in the additional material examined). The record in Greece is situated 650 km to the nearest Turkish locality (Sakarya province). Thus, this species was recorded far outside of its known distribution in Turkey. Despite the extensive field studies I have carried out, L. bodemeyeri has never been recorded from further west of Sakarya province in Turkey, so this record seems doubtful. In the same way, since the record from Hatay province is far outside the known distribution, it requires confirmation. The most likely explanation for the records is that the specimens were mislabeled. It has been observed for material collected by Heinz before (for a precedent see the section on Leptobium syriacum (Saulcy) in Assing (2017 )). The collector was in Turkey in the same year. Thus, L. bodemeyeri should be removed from the list of Greek Lathrobium.	en	Anlaş, Sinan (2021): On the microphthalmous Lathrobium Gravenhorst species of Turkey II. Two new species, a new synonymy, and an updated key to Turkish species (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5052 (1): 117-127, DOI: https://doi.org/10.11646/zootaxa.5052.1.7
