taxonID	type	description	language	source
038B976CFF89FFECCA2DCAEE16A5CBF0.taxon	type_taxon	Type species Astaena tridentata Erichson, 1847 (by monotypy).	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF89FFECCA2DCAEE16A5CBF0.taxon	diagnosis	Diagnosis Brown or reddish-brown coloration, mostly unicolored. Antennae with eight or nine antennomeres; antennal club with three antennomeres. Abdomen with a lateral carina.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF89FFEDCA3DCF101466CB3B.taxon	description	Figs 1 A – D, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF89FFEDCA3DCF101466CB3B.taxon	materials_examined	Type material examined Holotype COLOMBIA • ♀; “ Paine viii Colum. / Apolinar-Maria / L. W. Saylor Collection / California Academy of Sciences Type nº 7904 / Holotype Astaena abaca L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF89FFEDCA3DCF101466CB3B.taxon	description	MEASUREMENTS. Length: 12 mm. Width: 5.8 mm. HEAD. Reddish-brown, surface with simple setae. Labroclypeus convex, shape of anterior margin convex, posteriorly with sparse punctation. Ratio ocular canthus (measured just the process on the eye) / eye (measured along longitudinal axis): 1 / 4. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum unicolored, reddishbrown, widest at middle, anterior marginal line present, lateral margin strongly convex and smooth (not serrated), antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface tegument opaque, microsculpture absent, punctation on intervals present, strial and interval punctation equal in size. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, setae on basal surface present, a laterally extended longitudinal shelf present. Metatibia, widest at apex, dorsal margin carinate, ventral margin smooth (not serrated), distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. Male Unknown.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF88FFEBCA28CCA5149DCB12.taxon	description	Figs 1 E – I, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF88FFEBCA28CCA5149DCB12.taxon	materials_examined	Type material examined Holotype PERU • ♂; “ ix- 20 - 35 M. Sani Beni Lima Peru, 35 / L. W. Saylor Collection / California Academy of Sciences Type nº 7907 / Holotype Astaena biciliata L. W. Saylor ”; CAS. Paratype PERU • 1 ♀; “ ix- 20 - 35 M. Sani Beni Lima Peru, 35 / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena biciliata L. W. Saylor / CASENT 8438128 ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF88FFEBCA28CCA5149DCB12.taxon	description	MEASUREMENTS. Length: 11 mm. Width: 5.6 mm. HEAD. Reddish-brown, surface with upright setae. Labroclypeus, shape of anterior margin convex, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 4. Eyes, ratio diameter / interocular width: 0.55. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 4; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, reddishbrown, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and smooth (not serrated), antero-lateral angle dorso-ventrally not flattened, posterolateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddishbrown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal, posterior edge straight. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 0.77. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width / length: 1 / 2.69, dorsal margin not carinated, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur 1 / 2. Dorsal surface of tarsi glabrous and without punctures. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin not serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 1 E – F.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8EFFE8CA11CC71149CCDFE.taxon	description	Figs 1 J – N, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8EFFE8CA11CC71149CCDFE.taxon	materials_examined	Type material examined Holotype COLOMBIA • ♂; “ Bogota vi- Colom. / Apolinar-Maria / L. W. Saylor Collection / California Academy of Sciences Type nº 7909 / Holotype Astaena bogotana L. W. Saylor ”; CAS. Paratype COLOMBIA • 1 ♂; “ Bogota vi- Colom. / L. W. Saylor Collection / Paratype Astaena bogotana L. W. Saylor / CASENT 8438130 ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8EFFE8CA11CC71149CCDFE.taxon	description	MEASUREMENTS. Length: 12 mm. Width: 5.8 mm. HEAD. Reddish-brown, surface with setae. Labroclypeus, shape of anterior margin convex. Ratio ocular canthus / eye: 1 / 4. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 4; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, reddishbrown, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and not serrated, antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally flattened, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra bicolored, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur 1 / 3. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 1 J – K.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8DFFE9CA2FCD1C107DCD54.taxon	description	Figs 2 A – D, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8DFFE9CA2FCD1C107DCD54.taxon	materials_examined	Type material examined Holotype ECUADOR • ♀; “ Ecuador Baron / Coll. Kraatz / L. W. Saylor Collection / California Academy of Sciences Type nº 7910 / Astaena producta Bates? / Moser det. / Holotype Astaena excisipes L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8DFFE9CA2FCD1C107DCD54.taxon	description	MEASUREMENTS. Length: 9.5 mm. Width: 4.7 mm. HEAD. Reddish-brown. Labroclypeus, shape of anterior margin weakly sinuated medially. Ocular canthus with terminal setae, ratio to eye: 1 / 4. Ratio third vs fourth antennomere: 1 / 4. Fourth antennomere, lateral projection absent. Ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, reddishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin straight and not serrated, two angles on postero-lateral margin present, antero-lateral angle dorsoventrally not flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddishbrown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, dorsal margin not carinated, ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid. Male Unknown.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8DFFE9CA2FCD1C107DCD54.taxon	discussion	Remarks The holotype specimen is a female; Saylor (1946) incorrectly stated that the holotype as a male.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8CFFE7CA1ECAB5149CCC3E.taxon	description	Figs 2 E – I, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8CFFE7CA1ECAB5149CCC3E.taxon	materials_examined	Type material examined Holotype ARGENTINA • ♂; “ Salta Prov. Arg. S. A. / L. W. Saylor Collection / California Academy of Sciences Type nº 7911 / Holotype Astaena explaniceps L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF8CFFE7CA1ECAB5149CCC3E.taxon	description	MEASUREMENTS. Length: 9.2 mm. Width: 3.8 mm. HEAD. Reddish-brown, surface with upright setae. Labroclypeus, shape of anterior margin convex, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 2. Ratio third vs fourth antennomere: 1 / 3. Fourth antennomere, lateral projection absent. Ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum unicolored, reddish-brown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf absent. Metatibia, widest at apex, dorsal margin carinated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, with punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 2 E – F.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF82FFE4CA11CB5D1466CF55.taxon	description	Figs 2 J – M, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF82FFE4CA11CB5D1466CF55.taxon	materials_examined	Type material examined Holotype COLOMBIA • ♀; “ Fusago x Colom / Apolinar-Maria / L. W. Saylor Collection / California Academy of Sciences Type nº 7912 / Holotype Astaena fusagona L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF82FFE4CA11CB5D1466CF55.taxon	description	MEASUREMENTS. Length: 13 mm. Width: 6 mm. HEAD. Blackish-brown, surface with upright setae. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 4; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with upright setae. Pronotum unicolored, blackish-brown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally flattened, posterolateral angle dorso-ventrally not flattened, punctation on disc dense, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures medially present. Elytra unicolored, reddishbrown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, dorsal margin carinated, with group of spines positioned at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. Male Unknown.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF81FFE4CA17C8AA1494C9EC.taxon	description	Figs 3 A – E, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF81FFE4CA17C8AA1494C9EC.taxon	materials_examined	Type material examined Holotype BOLIVIA • ♂; “ Incachaca, Bolivia, Alt. 2500 m, J. Steinbach, Acc. 6873 / L. W. Saylor Collection / California Academy of Sciences Type nº 7913 / Holotype Astaena incachaca L. W. Saylor ”; CAS. Paratype BOLIVIA • 1 ♀, Incachaca, Bolivia, Alt. 2500 m, J. Steinbach, Acc. 6873 / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena incachaca L. W. Saylor / CASENT 8438145 ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF81FFE4CA17C8AA1494C9EC.taxon	description	MEASUREMENTS. Length: 11 mm. Width: 5.6 mm. HEAD. Reddish-brown, surface with upright setae. Labroclypeus, shape of anterior margin straight, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Eyes, ratio diameter / interocular width: 0.56. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum unicolored, reddishbrown, widest at middle, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 0.95. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf absent. Metatibia, widest at apex, ratio of width / length: 1 / 4.27, dorsal margin not carinated, ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, with punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 3 A – B.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF80FFE5CA2DC82D1497C9EC.taxon	description	Figs 3 F – J, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF80FFE5CA2DC82D1497C9EC.taxon	materials_examined	Type material examined Holotype ARGENTINA • ♂; “ Macueta Salta Prov. Arg. S. A. x-xi- 32 / L. W. Saylor Collection / California Academy of Sciences Type nº 7914 / Holotype Astaena maqueta L. W. Saylor ”; CAS. Paratypes ARGENTINA • 1 ♀; “ Macueta Salta Prov. Arg. SA. x-xi- 33 / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena maqueta L. W. Saylor ”; CAS • 1 ♂; “ Macueta Salta Prov. Arg. SA. x-xi- 33 / L. W. Saylor Collection / Paratype Astaena maqueta L. W. Saylor / CASENT 8438172 ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF80FFE5CA2DC82D1497C9EC.taxon	description	MEASUREMENTS. Length: 12 mm. Width: 5.8 mm. HEAD. Reddish-brown, surface with upright setae. Labroclypeus, shape of anterior margin convex, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 2. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Pronotum and elytra surface with upright setae. Pronotum unicolored, reddishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and not serrated, antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc dense, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, dorsal margin not carinated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, with punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and glabrous. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 3 F – G.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF86FFE3CA29C82C148DC9EC.taxon	description	Figs 3 K – O, 7 A	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF86FFE3CA29C82C148DC9EC.taxon	materials_examined	Type material examined Holotype COLOMBIA • ♂; “ Rio Chil. Colum / L. W. Saylor Collection / California Academy of Sciences Type nº 7915 / Holotype Astaena nigrona L. W. Saylor ”; CAS. Paratype COLOMBIA • 1 ♀; “ Bogota VI- Colum / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena nigrona L. W. Saylor ”; CAS.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF86FFE3CA29C82C148DC9EC.taxon	description	Holotype redescription MEASUREMENTS. Length: 12 mm. Width: 5.8 mm. HEAD. Reddish-brown. Labroclypeus, shape of anterior margin weakly sinuated medially. Ocular canthus without terminal setae, ratio to eye: 1 / 4. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum bicolored, reddish-brown with two red spots on laterals, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, blackish-brown, surface opaque, microsculpture absent, punctation on intervals present, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 3 M – N.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE0CA25C82C1494C8BE.taxon	description	Figs 4 A – E, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE0CA25C82C1494C8BE.taxon	materials_examined	Type material examined Holotype PERU • ♂; “ Peru / Vitoc, 1400 to 1800 m ix- 22 - 40 / Coll of Bob Potts / L. W. Saylor Collection / California Academy of Sciences Type nº 7917 / Holotype Astaena pygidia L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE0CA25C82C1494C8BE.taxon	description	MEASUREMENTS. Length: 8 mm. Width: 4 mm. HEAD. Reddish-brown. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, brownishorange, widest at middle, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, brownish-orange, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin setose. Protibia with three teeth. Protarsi, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 4 A – B.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE1CA18CFDF1466CB9F.taxon	description	Figs 4 F – I, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE1CA18CFDF1466CB9F.taxon	materials_examined	Type material examined Holotype COLOMBIA • ♀; “ Colombia, S. A. / L. W. Saylor Collection / California Academy of Sciences Type nº 7918 / Holotype Astaena rugithorax L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF85FFE1CA18CFDF1466CB9F.taxon	description	MEASUREMENTS. Length: 14 mm. Width: 7 mm. HEAD. Blackish-brown, surface with setae. Labroclypeus, shape of anterior margin straight, posteriorly with dense punctation. Ocular canthus without terminal setae, ratio to eye: 1 / 3. Ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, blackishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strong convex and serrated, antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorsoventrally not flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, blackish-brown, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf absent. Metatibia, widest at apex, distal spines groups disposed at half size of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. Male Unknown.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF84FFFFCA35CCFC1466CA7A.taxon	description	Figs 4 J – M, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF84FFFFCA35CCFC1466CA7A.taxon	materials_examined	Type material examined Holotype ARGENTINA • ♀; “ Salta, Arg. viii / L. W. Saylor Collection / California Academy of Sciences Type nº 7919 / Holotype Astaena salta L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF84FFFFCA35CCFC1466CA7A.taxon	description	MEASUREMENTS. Length: 5 mm. Width: 3 mm. HEAD. Reddish-brown, surface with setae. Labroclypeus, shape of anterior margin convex, posteriorly with dense punctation. Ocular canthus without terminal setae, ratio to eye: 1 / 3. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum unicolored, reddishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and not serrated, antero-lateral angle dorso-ventrally not flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, ventral margin serrated and glabrous. Protibia with two teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow. Male Unknown.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9AFFFCCA2BCD981494CA46.taxon	description	Figs 5 A – E, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9AFFFCCA2BCD981494CA46.taxon	materials_examined	Type material examined Holotype BOLIVIA • ♂; “ Yungas, Bolivia / L. W. Saylor Collection / California Academy of Sciences Type nº 7920 / Holotype Astaena yungasa L. W. Saylor ”; CAS. Paratype BOLIVIA • 1 ♂; “ Yungas, Bolivia / L. W. Saylor Collection / Paratype Astaena yungasa L. W. Saylor / CASENT 8438307 ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9AFFFCCA2BCD981494CA46.taxon	description	MEASUREMENTS. Length: 14 mm. Width: 6.75 mm. HEAD. Blackish-brown, surface with upright setae. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Eyes, ratio diameter / interocular width: 0.68. Antennae with eight antennomeres. Antennal club with three antennomeres. Ratio antennal club / remaining antennomeres: 1 / 1. Ratio third vs fourth antennomere: 1 / 4. Fourth antennomere, lateral projection absent. Ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum unicolored, blackishbrown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally not flattened, posterolateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 0.86. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf absent. Metatibia, widest at apex, ratio of width / length: 1 / 4.83, dorsal margin not carinated, ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur 1 / 2. Dorsal surface of tarsi glabrous, with punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 5 A – B.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF99FFFACA3CCDA51497CD29.taxon	description	Figs 5 F – J, 7 B	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF99FFFACA3CCDA51497CD29.taxon	materials_examined	Type material examined Holotype GUIANA • ♂; “ Tumatumari VI Br Guiana / L. W. Saylor Collection / California Academy of Sciences Type nº 7921 / Holotype Astaena zyrota L. W. Saylor ”; CAS. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF99FFFACA3CCDA51497CD29.taxon	description	MEASUREMENTS. Length: 7.5 mm. Width: 4.5 mm. HEAD. Reddish-brown, surface without setae. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with sparse punctation. Ocular canthus without terminal setae, ratio to eye: 1 / 4. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Pronotum unicolored, reddish-brown, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and not serrated, antero-lateral angle dorso-ventrally not flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, reddish-brown, surface opaque, microsculpture absent, punctation on intervals present, punctation on intervals dense, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf absent. Metatibia, widest at apex, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere smaller than second and third combined, ventral margin glabrous. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 5 F – G.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFFACAEBCA4F179CCB53.taxon	type_taxon	Type species Astaena bicoloripes Saylor, 1946 (by monotypy).	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFFACAEBCA4F179CCB53.taxon	diagnosis	Diagnosis Brown, reddish-brown, blackish coloration, often bicolored. Antennae with eight antennomeres; antennal club of males with three antennomeres. Abdomen without a robust lateral carina. Mentum deeply sinuated medially at anterior margin.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFF8CA4BCCB01480CF71.taxon	description	Figs 5 K – O, 7 C	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFF8CA4BCCB01480CF71.taxon	materials_examined	Type material examined Holotype (Astaena bicoloripes) PERU • ♀; “ M. Sani Beni Lima Peru. 35 / L. W. Saylor Collection / California Academy of Sciences Type nº 7908 / Holotype Astaena bicoloripes L. W. Saylor / Symmela bicoloripes Saylor det. G. Frey ”; CAS. Paratype (Astaena bicoloripes) PERU • 1 ♂; “ M. Sani Beni Lima Peru. 35 / L. W. Saylor Collection / Paratype Astaena bicoloripes L. W. Saylor / Astaena bicoloripes Saylor / CASENT 8438129 ”; CAS. Holotype (Astaena postnodata) PERU • ♂; “ Mt. Alegre, Rio Pachitea O. Peru G. Tessmann / Type Astaena postnodata G. Frey 1973 ”; ZMHB. Paratypes (Astaena postnodata) PERU • 4 ♀♀; “ Mt. Alegre, Rio Pachitea O. Peru G. Tessmann / Paratype Astaena postnodata G. Frey 1973 ”; ZMHB. Additional material examined PERU • 1 ♂; “ Dept. Cusco, PERU Santa Isabel Cosnipata Valley Dec 5 1951 Felix Woytkowski / H. & A. Howden collection Ottawa Canada ”; CMNC • 6 ♂♂, 5 ♀♀; “ Mt. Alegro, Rio Pachitea o. Peru G. Tessmann ”; ZFMK, ZMHB, NHMB. Holotype redescription	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFF8CA4BCCB01480CF71.taxon	description	MEASUREMENTS. Length: 6.5 mm. Width: 4 mm. HEAD. Black, surface with upright setae. Labroclypeus convex, shape of anterior margin weakly sinuated medially, posteriorly with dense punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Eyes, ratio diameter / interocular width: 0.44. Antennae with nine antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1; ratio third vs fourth antennomere: 1 / 2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1.2 / 1. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, brownishorange, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally not flattened, posterolateral angle dorso-ventrally not flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation present. Scutellum, punctures medially present. Elytra bicolored, surface opaque, microsculpture absent, punctation on intervals present and sparse, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina present. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 0.90. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width / length: 1 / 3, dorsal margin not carinated. ventral margin serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate. AEDEAGUS. Fig. 5 K – L.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9FFFF8CA4BCCB01480CF71.taxon	discussion	Remarks Also, in this case, the holotype was a female specimen; Saylor mentioned in the original description that the holotype was a male.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9DFFF6CA50C8911116CA05.taxon	description	Figs 6 A – I, 7 C	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9DFFF6CA50C8911116CA05.taxon	materials_examined	Type material examined Holotype (Astaena abcora) COLOMBIA • ♂; “ Buenaventura alt. 3500 - 4000 ft. Colombia, ii- 3 / L. W. Saylor Collection / California Academy of Sciences Type nº 7905 / Holotype Astaena abcora L. W. Saylor ”; CAS. Paratype (Astaena abcora) COLOMBIA • 1 ♀; “ Buenaventura alt. 3500 - 4000 ft. Colombia, ii- 3 / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena abcora L. W. Saylor / CASENT 8438114 ”; CAS. Holotype (Astaena apolinarmaria) COLOMBIA • ♀; “ Pensilvania Colomb / L. W. Saylor Collection / California Academy of Sciences Type nº 7906 / Apolinar-Maria / Holotype Astaena apolinar-maria L. W. Saylor ”; CAS. Additional material examined COLOMBIA • 4 ♂♂, 1 ♀; “ COLOM: Antioquia 1800 m, nr. Yarumai 20. iv. 73, J. Helava ”; CMNC • 19 ♂♂, 11 ♀♀; “ Manizales A. M. Patino / Slg. R. Oberthür Eing. Nr. 4, 1956 ”; ZFMK, MNHN • 3 ♂♂, 1 ♀; “ S. America / Coll. C. Felsche Kauf 20, 1918 ”; SMTD • 1 ♀; “ Columbia / Gehr. W. Müller Vermächt., 1909 ”; SMTD • 1 ♀; “ Cauca / Coll. C. Felsche Kauf 20, 1918 ”; SMTD • 1 ♀; “ Columbia Cañon de Gallo / Astaena apollinarmaria Sayl. Det. G. Frey, 1972 / cum typo comparatum ”; NHMB.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9DFFF6CA50C8911116CA05.taxon	description	Holotype redescription MEASUREMENTS. Length: 7 mm. Width: 3.75 mm. HEAD. Black, surface with simple setae. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 3. Eyes, ratio diameter / interocular width: 0.40. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum bicolored, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and serrated, antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, brownish-orange, surface opaque, microsculpture absent, punctation on intervals present, sparse, diameter of strial and interval punctation equal, posterior edge straight. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 0.77. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width / length: 1 / 3.34, dorsal margin carinated, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and glabrous. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 6 A – B.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF9DFFF6CA50C8911116CA05.taxon	discussion	Remarks There is no difference between female type specimens of S. abcora and S. apolinarmaria except coloration, which is very variable. Therefore, we consider them synonymous.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF93FFF7CA69CD1B149CCA11.taxon	description	Figs 6 J – N, 7 C	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF93FFF7CA69CD1B149CCA11.taxon	materials_examined	Type material examined Holotype PERU • ♀; “ Peru / Vitoc, 1400 to 1800 m x- 6 - 40 / Coll. of Bob Potts / L. W. Saylor Collection / California Academy of Sciences Type nº 7916 / Holotype Astaena pottsi L. W. Saylor ”; CAS. Paratypes PERU • 1 ♀; “ Peru / Vitoc, 1400 to 1800 m ix- 22 - 40 / Coll. of Bob Potts / L. W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype Astaena pottsi L. W. Saylor / CASENT 8438229 ”; CAS • 1 ♂; “ Vitoc, 1400 to 1800 m ix- 13 - 40 / Coll. of Bob Potts / L. W. Saylor Collection / Paratype Astaena pottsi L. W. Saylor / Locality label should read Vitor, PERU / CASENT 8438221 ”; CAS • 1 ♀; “ Peru / Vitoc, 1400 to 1800 m ix- 3 - 40 / Coll. of Bob Potts / Paratype Astaena pottsi L. W. Saylor / Locality label should read Vitor, PERU / CASENT 8438222 ”; CAS.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
038B976CFF93FFF7CA69CD1B149CCA11.taxon	description	Holotype redescription MEASUREMENTS. Length: 11 mm. Width: 5 mm. HEAD. Reddish-brown, surface with setae. Labroclypeus, shape of anterior margin straight, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1 / 4. Eyes, ratio diameter / interocular width: 0.7. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club / remaining antennomeres: 1 / 1.5; ratio third vs fourth antennomere: 1 / 2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1 / 5. PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum unicolored, brownishorange, widest at middle, lateral margin weakly convex and not serrated, antero-lateral angle dorsoventrally not flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin sparse, median longitudinal excavation present. Scutellum, punctures absent medially. Elytra unicolored, brownish-orange, surface opaque, microsculpture absent, punctation on intervals present and dense, diameter of strial and interval punctation equal. ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite. LEGS. Ratio of length of metepisternum / metacoxa: 1 / 1.12. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width / length: 1 / 3.88, dorsal margin not carinated, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1 / 3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin setose. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute. AEDEAGUS. Fig. 6 J – K.	en	Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z., Ahrens, Dirk (2021): Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor. European Journal of Taxonomy 750 (750): 94-123, DOI: 10.5852/ejt.2021.750.1365, URL: http://dx.doi.org/10.5852/ejt.2021.750.1365
