taxonID	type	description	language	source
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	description	urn: lsid: zoobank. org: act: E 0 CD 76 FD- 36 A 6 - 48 D 6 - BDEE- 21 A 62 C 3 FDABD Figs 1 – 2, 3 D, 4 D, 5 C, 6	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	diagnosis	Diagnosis This new species is most similar to G. deharvengi Tian, 2014, which also occurs in Huanjiang County, by the similar character configuration of the head and elytra. It is easily separated from the latter species by the clypeal wings not projecting anteriorly. Moreover, its body shape is much smaller and more slender, the elytral stria punctures are larger and sparser, and intervals four and five are carinated near the base.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	etymology	Etymology This new species is dedicated to Mr. Guofu Wei (Center of World Natural Heritage, Huanjiang) for his support during our biospeleological investigations.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Huanjiang, Jiale, Dapo Dong; 25 ° 06 ′ 06.78 ″ N, 108 ° 31 ′ 12.48 ″ E; alt. 211 m; 5 Dec. 2016; A. Faille leg.; SCAU.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	description	Description Male MEASUREMENTS. Length: 4.5 mm; width: 1.1 mm. Habitus as in Fig. 2. BODY. Slightly depigmented, concolorous brown. HEAD. Stout (Fig. 3 D), from apex of mandible much longer than wide, HLm / HW = 1.41, or slightly longer than wide when measured from apex of labrum, HLl / HW = 1.06; distinctly narrower than pronotum, HW / PW = 0.67; clypeus with fused wings transverse, moderately convex medially, bisetose at base, anterior margin slightly bisinuate, bordered, clypeal wings not protruding anteriorly; supra-antennal plates well developed, gently rounded, reflexed, margined, strongly convex, smooth and glabrous; frons and vertex moderately convex, with longitudinal pore at middle; laterally with two setiferous pores, situated at posterior end of supra-antennal plates and at level of neck constriction, respectively; frontal furrows deep, wide, distinctly divergent posteriorly; frontal carinae distinct, slightly convergent to neck constriction; frontoclypeal sulcus not deep but distinct; eyes completely lacking; genae well developed, subparallelsided, sides nearly vertically truncated; neck constriction distinct, slightly stepped; labrum transverse, slightly wider than clypeus, seven-setose, ciliate laterally, straight at anterior margin; mandibles of moderate size. Palpomeres ensiform, glabrous, apical segments of both maxillary and labial palpomeres much longer than penultimate ones, respectively; labial palpomere bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pair situated beneath mental tooth, the other at base near lateral margin, and with two large and deep concavities near base; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, rather short, reaching posterior angles of pronotum; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; pedicellus slightly longer than antennomere three; antennomeres five to ten subelongate, terminal segment slightly longer than penultimate. Underside of head rough. THORAX. Pronotum peltate (Fig. 3 D), much wider than head, slightly longer than wide, PL / PW = 1.06; disc smooth, moderately convex; anterior margin slightly concave, beaded in median line; fore angles slightly protruding; widest at beginning of posterior third, gently and gradually narrowed anteriorly, strongly contracted towards posterior angles; posterior angles with obtuse teeth, lateral margin between posterior angles and basal constriction with two conspicuously toothed projections; with two pairs of lateral setiferous pores, anterior one at about anterior quarter, posterior one at basal angle and distinctly removed from channel; reflexed lateral margin with slight notches, lateral channel narrow before anterior seta, distinctly widened between anterior and posterior lateral setae, ending before posterior seta; basal constriction wide, basal carina narrow; median line engraved, deep, wide, distinctly joining basal constriction, surpassing anterior transverse line without joining. Peduncle short, scutellum small. Prosternum and propleura smooth, with dense and isodiametric abdominal punctuation. ABDOMEN. Ventrites also densely punctured, ventrites four to six each with a pair of setae; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, wider than pronotum, EW / PW = 1.27, much longer than wide, EL / EW = 1.86; strongly convex; widest behind middle, gently contracted anteriad and posteriad; base finely bisinuate, with a pair of additional tubercles followed by a basal setiferous puncture located at base of interval two; shoulders broadly obtuse, with sharp tooth; apex of elytra pointed; sides distinctly crenulated from base to apical quarter; elytral striae punctate-striate, wide and deep, with large and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals distinctly convex; intervals one to four unbordered at base; intervals seven and eight joined near base, then joined to interval six at base, intervals six to eight carinated throughout, intervals four and five carinated near base; arrangement of elytral setiferous chaetotaxy as in Fig. 4 D, five foveolate setiferous pores on interval three; marginal channel with uninterrupted series of small setiferous pores and several large pores bearing much longer setae. Hind wings reduced. LEGS. Moderately elongated; proleg stout, profemur distinctly dilated, smooth, with a long seta and two shorter ones; protibia well developed, with distinct and complete carina dorsally, sulcus indistinct, quadridentate; lateral teeth blunt at apex, lowest lateral one much shorter than others, upper two stoutensiform; subapical spur elongate-ensiform, blunt at apex, shorter and more slender than uppermost lateral teeth; protarsi slender, tarsomere one longer than tarsomeres two to four combined; meso- and meta-legs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with an elongated, tuber-like subapical spur, longer than wide and furnished with an isometric seta. GENITALIA (Fig. 5 C). Moderately sclerotized; aedeagus similar to those of G. deharvengi Tian, 2014 (Fig. 5 A) and G. cavicola Tian, 2013 (Fig. 5 B), median lobe gently arcuated ventrally, whereas strongly arcuated in G. deharvengi, or evidently bisinuate in G. cavicola, blunt at apex; parameres asymmetrical, one much longer and broader than the other, both parameres with three long setae at apex. Female Unknown.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFBCFF9CFDF1A2CBFB71FD9C.taxon	distribution	Distribution China (Guangxi: Huanjiang County) (Fig. 1). Known only from Dapo Dong cave. This cave opens at the bottom of a hill near a country road in the village of Jiale. The entrance is narrow, but accessible (Fig. 6 A). It is about 90 m long with a short and narrow side passage inside. A large part of the passage is dry, but there are some moist places. The single blind beetle specimen was discovered on the moist wall about 30 m from the entrance at the left side of the main passage. Other syntopic cave animals were observed during the investigation in the cave, for instance, a flatworm and a bat. (Fig. 6 B, D).	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	description	urn: lsid: zoobank. org: act: 54 A 7 DD 47 - 8 EAB- 4 A 9 A- 9 C 49 - 2 E 6383 B 4842 A Figs 1, 3 E, 4 E, 5 D, 7 – 8	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	diagnosis	Diagnosis This new species belongs to the group with distinctly projecting clypeal wings and is close to G. deharvengi Tian, 2014, which also occurs in the same karst area in Huanjiang County, by the similar character configuration of head and elytra, but it is easily separated from the latter species by the missing central pore on the frons of the head and the flattened neck constriction. In addition, the pronotum, with the anterior margin almost not emarginate, is much longer, its median line is much narrower, and the anterior transverse line is finely developed.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	etymology	Etymology The species epithet is derived from Yueliang Shan (Moon Mountain in Chinese), a famous karst landscape with a naturally-formed moon-like cavern passing through it (Fig. 8 A). The type locality, Huang Dong, is one of the caves of this landscape.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	materials_examined	Type material Holotype CHINA • ♂, broken on the left supra-antennal plate; Guangxi, Huanjiang, Shuiyuan, Sancai, Neiwen, Huang Dong; 24 ° 50 ′ 05.74 ″ N, 108 ° 05 ′ 1.86 ″ E; alt. 354 m; 6 Jun. 2017; S. Huang leg.; SCAU.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	description	Description Male MEASUREMENTS. Length: 9.3 mm; width: 2.9 mm. Habitus as in Fig. 7. BODY. Well-pigmented, concolorous dark brown. HEAD. Stout (Fig. 3 E), from apex of mandible slightly longer than wide, HLm / HW = 1.06, or shorter than wide when measured from apex of labrum, HLl / HW = 0.82; clypeus strongly transverse and moderately emarginate, surface of median part moderately convex, bisetose at base, with flattened clypeofrontal sulcus, clypeal wings fused, divided from supra-antennal plates by distinct notches; supra-antennal plates rounded and well developed, reflexed margined, strongly convex, smooth and glabrous; frons and vertex fairly convex; laterally with two setiferous pores, situated at posterior end of supra-antennal plates and at level of neck constriction, respectively; frontal furrows deep, conspicuously widened, subparallel at middle of frons, divergent posteriorly; frontal carinae blunt but well marked, not parallelsided; eyes completely lacking; genae well developed, anteriorly conically raised laterally then sinuated, sides nearly vertically truncated; neck constriction distinct, relatively wide; labrum transverse, slightly emarginate, slightly wider than clypeus, seven-setose, ciliate laterally; mandibles of moderate size. Palpomeres ensiform, glabrous, apical segments of both maxillary and labial palpomeres much longer than penultimate ones, respectively; labial palpomere two bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pair situated beneath mental tooth, posterior one localized in centre of two large, deep concavities restricted at base near lateral margin; median tooth simple, pointed at apex, lateral lobes wide, obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, elongate, reaching over base of pronotum; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; pedicellus slightly longer than antennomere three, then gradually decreasing in length towards penultimate segment, antennomeres five to ten subelongate, penultimate segment slightly shorter than terminal one. Underside of head rough. THORAX. Pronotum peltate (Fig. 3 E), much wider than head, PW / HW = 1.25, slightly shorter than wide, PL / PW = 0.96; disc strongly convex and with noticeable rugosity; anterior margin unbordered, slightly convex in median portion, margin barely emarginate, anterior transverse line fine and traceable, fore angles protruding; widest slightly behind middle, gently curved and gradually narrowed anteriorly, more evidently contracted towards posterior angles; with distinct teeth at posterior angles, lateral margin between posterior angles and basal constriction with one conspicuous and one minute toothed projections before basal constriction; with two lateral setiferous pores, anterior one at about apical quarter, posterior one at basal angle, just before tooth at posterior angle; reflexed lateral margin nearly smooth, lateral channel with uneven surface, narrower before anterior seta, wider between anterior and posterior lateral setae, interrupted by tooth of posterior angle; basal constriction wide, with noticeable transverse carina; median line deeply furrowed but faint when joined with transverse line in front. Peduncle short, scutellum small. Prosternum and propleura smooth, with dense and isodiametric punctuation. ABDOMEN. Abdominal ventrites also densely punctured; ventrites four to six each with a pair of setae; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, wider than pronotum, EW / PW = 1.29, much longer than wide, EL / EW = 1.84; strongly convex, widest at middle, gently contracted anteriad and posteriad; base with pair of basal setiferous pores located at beginning of stria one, joined with an additional tubercle at base of interval two; shoulders broadly obtuse, with two teeth on each side; apex pointed; sides distinctly crenulated from base to apical third, then very feebly marked towards apex; elytral striae punctatestriate, narrow and deep, with small and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals distinctly convex; intervals seven and eight joined near base, then joined to interval six at base, intervals six to eight carinated near base; arrangement of elytral setiferous chaetotaxy as in Fig. 4 E: five foveolate setiferous pores present on interval three; marginal channel with uninterrupted series of small setiferous pores and several large pores bearing much longer setae. Hind wings reduced. LEGS. Legs moderately elongated; proleg stout, profemur moderately dilated, smooth, with several sparsely distributed setae; protibia well developed, with distinct and complete carina dorsally, sulcus indistinct, quadridentate; lateral teeth blunt at apex, lowest lateral tooth unobtrusive, upper three stoutensiform and moderately protruded anteriorly toward apex; subapical spur elongate-ensiform, blunt at apex, much longer and more slender than uppermost lateral teeth; protarsi slender, tarsomere one very long, much longer than tarsomeres two to four combined; meso- and meta-legs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with elongated tuber-like subapical spur, distinctly longer than wide and furnished with isometric seta. GENITALIA (Fig. 5 D). Moderately sclerotized; aedeagus similar to that of G. deharvengi Tian, 2014 (Fig. 5 A); median lobe strongly arcuated ventrally, elongated and blunt at apex; copulatory piece irregular, elongated, strongly sclerotized in basal and frontal part; parameres asymmetrical, large one with three setae, small one with four setae. Female Unknown.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB7FF98FE3DA2EEFE48FD52.taxon	distribution	Distribution China (Guangxi: Huanjiang County) (Fig. 1). Known only from Huang Dong cave (Fig. 8 B). The cave is still preserved as a natural environment and the entrance opens at the middle of the hill. The cave is not well explored and its length remains unknown. The passages in the cave are complicated, and most of them are moist and wet. The single beetle specimen was collected in a wet place about 100 m from the entrance (Fig. 8 C).	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	description	urn: lsid: zoobank. org: act: EA 3 D 7117 - F 8 F 3 - 4 F 45 - 848 A- 9 E 3879 ABCBF 8 Figs 1, 3 F, 4 F, 9, 10 D, 11	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	diagnosis	Diagnosis This new species is most similar to G. bedosae Tian, 2014, which also occurs in Longzhou County, by the similar character configuration of the head and elytra and the seven-setose labrum present in both species. It is easily separated from the latter species by its much larger body, conspicuously emarginate labrum, the strongly protruding fore angles of the pronotum and the special elytral setiferous chaetotaxy.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	etymology	Etymology The species epithet refers to the unexpected discovery of this new species. The single specimen was encountered by Jiajun Zhou (the second author) after diving in the cave. It is his first discovery of a new cave beetle.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	materials_examined	Type material Holotype CHINA • ♀, broken on the left supra-antennal plate; Guangxi, Longzhou, Xiadong, Zhuzhu Dong; 22 ° 24 ′ 10.75 ″ N, 106 ° 38 ′ 52.36 ″ E; alt. 170 m; 18 Oct. 2020; J. Zhou leg.; SCAU.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	description	Description Female MEASUREMENTS. Length: 7.5 mm; width: 2.4 mm. Habitus as in Fig. 9. BODY. Moderately depigmented, concolorous brown. HEAD. Stout (Fig. 3 F), much longer than wide from apex of mandible, HLm / HW = 1.44, or slightly shorter than wide when measured from apex of labrum, HLl / HW = 0.97; clypeus strongly transverse and with anterior margin slightly, median part moderately concave, bisetose at base, frontoclypeal sulcus well marked, clypeal wings not projecting, nearly at same level as clypeus, distinctly separated from clypeus and supra-antennal plates by notches; supra-antennal plates well developed, reflexed margined, strongly convex, smooth and glabrous; frons and vertex moderately convex, with transverse rugosity; with two setiferous pores, situated at posterior end of supra-antennal plates and before neck constriction, respectively, frontal furrows distinct, wide, slightly divergent towards middle of frons, strongly divergent posteriorly; frontal carinae blunt, wide, parallel; eyes completely lacking; genae well developed, not parallel-sided, sides tumid; neck constriction distinct, wide; labrum transverse, frontal margin conspicuously emarginate as a semicircular incision, seven-setose along frontal part, with six cilia on both sides; mandibles short and stout. Palpomeres ensiform, glabrous, apical segments of maxillary palpomere about twice as long as penultimate one, labial palpomeres subequal; labial palpomere two bisetose on inner margin; ligula unisetose at apex; labial suture well marked; mentum well developed, with two pairs of setae, one pair situated beneath median tooth, the other at base near lateral margin, with two large and deep concavities near base; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; basal foveae slightly convex; submentum narrow, quadrisetose. Antennae filiform, elongated, reaching elytral base; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; antennomere three slightly longer than pedicellus, then gradually decreasing in length towards penultimate segment, which is slightly shorter than terminal segment. Underside of head rough. THORAX. Pronotum peltate (Fig. 3 F), much wider than head, PW / HW = 1.47, slightly shorter than wide, PL / PW = 0.90; disc moderately convex; anterior margin unbordered, indistinctly convex in median portion, margin with a row of cilia throughout, anterior transverse line marked with punctation, fore angles strongly protruding; widest at end of second third, gently and gradually narrowed anteriorly, with conspicuous notch before posterior angle, posterior angle large, with two conspicuously toothed projections between posterior angle and basal constriction; with two lateral setiferous pores, anterior seta at about anterior fifth, posterior one slightly before notch at posterior angle; reflexed lateral margin slightly crenulate, lateral channel broad throughout, narrower before anterior seta, wider between anterior and posterior lateral setae, ending at notch of posterior angle; basal constriction wide, unbordered, with narrow transverse carina; median line engraved, wide, not joining basal constriction. Peduncle short, scutellum small. Prosternum and propleura smooth, with sparse punctures. ABDOMEN. Abdominal ventrites more or less punctured; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, strongly convex, wider than pronotum, EW / PW = 1.29, much longer than wide, EL / EW = 1.67; widest at about one third from apex, slightly contracted anteriad and moderately and regularly contracted posteriad; base with a pair of setiferous punctures at base of stria one, intervals two and three each with a pair of small tubercles at base; shoulders broadly obtuse, with two large teeth on each side; apex pointed; sides distinctly crenulate from base to apical third, then indistinctly continued towards apex; elytral striae broad and well marked, with large and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals slightly convex; intervals six and seven carinate at base, joining and continuing into humeral tooth, interval eight carinate throughout, joining with seven before reaching base; arrangement of elytral setiferous chaetotaxy as in Fig. 3 F: nine and six foveolate setiferous dorsal pores on interval three and five, respectively, pores nearly regularly arranged; with series of setiferous pores in marginal channel, uninterrupted and small, with several large pores bearing much longer setae. Hind wings reduced. LEGS. Moderately elongated; proleg stout, profemur moderately dilated, smooth, with a long seta and several short ones; protibia well developed, with distinct and complete carina dorsally, sulcus distinct, quadridentate; lateral teeth blunt at apex, lowest lateral much shorter than others, upper two stoutensiform; subapical spur elongate-ensiform, blunt at apex, shorter and more slender than uppermost lateral teeth; protarsi slender, tarsomere longer than tarsomeres two to four combined; meso- and metalegs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with an elongated tuber-like subapical spur, longer than wide and furnished with an isometric seta. GENITALIA (Fig. 10 D). Similar to those of G. cavicola (Fig. 10 B), gonosubcoxite IX stout, with one long seta and two short setae at median portion of outer margin; gonocoxite IX curved and slender, apex very sharp, with three dorsal and three ventral setae, plus two short and sturdy ensiform setae on outer margin. Male Unknown.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
03CC663BFFB3FF84FDCCA2A8FD81FB54.taxon	distribution	Distribution China (Guangxi: Longzhou County) (Fig. 1). Known only from Zhuzhu Dong cave (Fig. 11). The cave is not easy to access due to the extremely narrow entrance as well as the vertical depth of three meters after the entrance. Then, it reaches a relatively wider and flat passage. But the ventilation of the passage is inadequate, causing the collector to experience symptoms of shortness of breath and chest distress. The single specimen was found on the muddy surface near the bank of an underground river, which is 50 meters from the entrance.	en	Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi, Faille, Arnaud (2021): Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135-154, DOI: https://doi.org/10.5852/ejt.2021.774.1537, URL: http://dx.doi.org/10.5852/ejt.2021.774.1537
