taxonID	type	description	language	source
03BD95561E317A78FF38FAF6F78AF86C.taxon	description	(Figs 1 A – B, 2 A, 3, 4 A, 5)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E317A78FF38FAF6F78AF86C.taxon	type_taxon	Type species: Argelodes magnificus sp. nov., by present designation.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E317A78FF38FAF6F78AF86C.taxon	diagnosis	Diagnosis. Large (TL 9.6 mm), not saltatorial, elongate, with long legs, elytra subglabrous, covered with very short and sparse setae, setae on elytra shorter than distance between two closest punctures. Subantennal grooves moderately deep, subgenal ridge without buttonhole configuration, eyes moderately protuberant. Pronotum small, distinctly narrower than basal portion of elytra; anterolateral pronotal angles acute, distinctly protruding anteriorly. Parameroids bifid, trigonium deeply divided; apices of parameres curved and hooked.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E317A78FF38FAF6F78AF86C.taxon	description	Description. Male. Body (Fig. 1 A) elongate, TL 9.6 mm, moderately convex. Body brown, head and pronotum covered with long, yellowish setae, elytra covered with sparse and very short setae, shorter than distance between two closest elytral punctures. Head and pronotum covered with sparse and shallow punctures and regular, polygonal microsculpture; elytra with larger, uniform punctures, separated by 1.5 – 2.0 × diameter of a puncture, without microsculpture, punctation irregular. Head (Fig. 1 B) relatively small, wider than long; eyes large and strongly protuberant, finely facetted; tempora moderately long, distance from posterior margin of eye to occipital ridge ca. 0.5 × diameter of the eye, slightly converging basally; distance from ventral margin of eye to subgenal ridge ca. 0.4 × diameter of the eye; supraantennal ridges slightly elevated, joined to margins of eyes; subtriangular area behind eyes devoid of setation, covered with microreticulation only; frons almost flat, with two shallow depressions behind antennal insertions; subantennal grooves slightly concave, moderately deep, glossy and impunctate, each subantennal groove separated from frons by a short ridge subparallel to supraantennal ridge, continued along margin of eye as sharp ridge, in lower portion connected with subgenal ridge. Frontoclypeal suture absent; clypeus subtrapezoidal, ca. 2.6 × wider than long, with straight anterior margin, lateral margins slightly converging posteriorly. Anteclypeus present, ca. 4 × shorter than clypeus. Ventral portion of head (Fig. 2 A) between subgenal ridges concave, gular area convex, rectangular, slightly longer than wide. Subgenal ridge well marked, without buttonhole configuration. Antennae filiform, antennomeres 6 – 11 missing in the only known specimen; antennomere 1 widest, subcylindrical, with subtle ridge marked on apical half of anterior margin; antennomere 2 subglobular and small, distinctly narrower and ca. 2 × shorter than antennomere 1; antennomere 3 very long and narrow, ca. 1.8 × longer than antennomere 1, slightly widening apically; antennomeres 4 – 5 similar to antennomere 3. Labrum covered with long setae, transverse, ca. 2.0 × wider than long, anterior margin slightly emarginated, anterolateral angles rounded. Mandibles (Fig. 3 A) symmetrical, ca. 2.3 × longer than width at bases, abruptly curved in apical 1 / 4, apices unidentate, outer margin covered with setae; inner margin of mandibles with big subtriangular denticle and two smaller ones; molar region covered with poorly visible microsetae. Maxillary palpi (Fig. 3 C): palpomere 1 shortest, subconical, palpomere 2 longest, ca. 3 × longer than palpomere 1, palpomere 3 shorter than 2, palpomere 4 fusiform, shorter than 3 and ca. 2 × shorter than 2. Galea (Fig. 3 D) with subparallel sides, much shorter than lacinia, with long fringe of irregularly arranged setae at apex; lacinia with straight or slightly curved setae on inner margin and stronger, hooked setae on apex. Mentum (Fig. 3 B) slightly transverse (W / L = 1.4), subtrapezoidal. Prementum subtriangular, strongly widening apically, apical margin straight, ligula covered with dense setae. Labial palpomeres: palpomere 1 long, narrow, ca. 2.5 × longer than wide, sparsely covered with setae; palpomere 2 wider, subconical, ca. 1.7 × longer than palpomere 1, covered with long setae; apical palpomere arising from apex of preapical one, narrow, slightly longer than palpomere 1, covered with setae. Pronotum (Fig. 1 B) transverse, distinctly narrower than base of elytra, ca. 1.7 × wider than long; lateral carinae almost straight, subtly curved in anterior portion; disc moderately convex; anterior margin rounded; anterolateral angles subtriangular, projecting; posterior angles obtuse; base curved, with distinct margination. Prosternum reduced anteriorly; prosternal process narrow, lanceolate, posterior apex pointed, ca. 3.0 × longer than wide, covered with setae. Scutellar shield equilaterally triangular, with slightly curved lateral sides, flat, with pointed apex; anterior margin straight, impunctate, covered with microsculpture similar to that on pronotum. Elytra elongated, distinctly wider at base than pronotal base; humeri marked, apices regularly rounded. Epipleura wide at base, slightly narrower than metanepisternum, distinctly narrowed behind metaventrite, present till apices. Hind wings (Fig. 4 A) ca. 2.3 × longer than its width; radial cell subtriangular; apical portion of r 3 well visible, straight; rp-mp 2 arcuate, forming acute angle with MP 1 + 2; MP 4 long, oblique, joining MP 3 and CuA + AA 1 + 2; anal field with well marked AP 3 and AP 4. Medial field with 3 free veins; pigmented areas in radial cell, radial, central and apical fields. Mesoventrite (Fig. 2 A) small; anterior margin with coxal rests; mesoventral process with well marked longitudinal suture; anterior portion forming deep, subtriangular fossa for reception of prosternal process; mesoventral process ca. 2 × longer than wide, sides subparallel, apex bilobed, touching metaventrite; mesocoxae completely separated; mesanepisternum rhomboidal, without setation, wider than long, forming coxal rest for procoxa; mesepimeron smaller, setose, subtriangular, strongly narrowed mesally; mesocoxae subconical, slightly projecting. Metaventrite (Fig. 2 A) relatively short, strongly transverse (W / L 3.4), convex; discrimen complete; metanepisternum subtrapezoidal, ca. 2 × longer than its width, slightly narrowing posteriorly. Metacoxae moderately large, strongly oblique, 2.7 × wider than long, extending laterally to epipleura; metacoxal plate obliquely narrowing. Legs long; femora extending beyond elytral margin, trochanters moderately long; trochantero-femoral joint oblique; metafemora longer than metatibiae, metatibiae slightly curved. Tibiae narrow, slightly widening towards apices, with paired carinae; metatibiae slightly sinuate; length of metatibial spurs equal largest width of metatibiae; mesotarsi only slightly shorter than mesotibiae [protarsi and metatarsi incomplete in the only known specimen]; tarsomere 1 elongated, sides subparallel, as long as tarsomeres 2 – 3 combined; tarsomere 4 emarginated at apex; apical tarsomere as long as tarsomeres 3 – 4 combined, with simple claws. Abdomen ca. 1.4 × longer than its width, convex; ventrites 1 – 4 of subequal length; ventrite 5 slightly longer, regularly rounded at apex. Abdominal ventrites regularly covered with setae. Genitalia (Fig. 5) of Microcara type (sensu: Zwick 2015), symmetrical, penis large, dorsoventrally flattened, trigonium bifid, parameroids moderately wide, bifid in apical portions, longer than trigonium. Tegmen large, parameres wide at bases, narrowed and hooked in apical portion, all terminal segments are present. Female. Unknown.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E317A78FF38FAF6F78AF86C.taxon	biology_ecology	Biology. Unknown.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E317A78FF38FAF6F78AF86C.taxon	etymology	Etymology. Combination of terra typica — Argentina and a genus name Elodes Latreille; reference to the geographical distribution of the genus. Gender masculine.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	description	(Figs 1 A – B, 2 A, 3, 4 A, 5)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	materials_examined	Type material. Holotype, male (ZMUC): “ ARGENTINA, Rio Negro \ 22: Lago Nahuel Huapi \ Puerto Blest, 770 m \ 25. - 27. x. 1981 \ Nielsen & Karsholt ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	diagnosis	Diagnosis. Large (TL 9.6 mm); pronotum distinctly narrower than base of elytra; elytra subglabrous. Parameroids of penis bifid at apices, parameres wide in basal portion, narrowed and hooked in apical half.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	description	Description. Male. Body large (TL 9.6 mm), elongate (TL / EW 2.0), widest in the middle of elytra, moderately convex. Colouration chestnut brown, head covered with relatively long and dense setae, setation of pronotum sparser, elytra with short and sparse setae, subglabrous. Head ca. 1.2 × wider than long, ca. 1.9 × wider than interocular space, convex, sparsely covered with weakly impressed punctation, punctures separated by 1.5 – 2.0 × diameter of a puncture, area between punctures microreticulate, covered with regular, polygonal microsculpture; eyes large, strongly protuberant. Pronotum transverse, narrow (PL 1.6 mm, PW 2.7 mm), with punctation and microsculpture similar to those on head. Elytra long (EL 8.3 mm), 5.2 × longer than pronotum, 1.8 × wider than pronotum; punctation irregular, sparse, separated by ca. 1.0 – 2.0 × diameter of a puncture. Aedeagus symmetrical, penis (Fig. 5 A) large (L 1.53 mm, W 0.53 mm), dorsoventrally flattened; trigonium bifid, each process of trigonium with two denticles at apex, a lateral one and an apical one; parameroids moderately wide, bifid in apical portions, longer than trigonium, outer apical process subtriangular, inner apical process longer than outer one, narrower, digitiform and subtly sinuate. Tegmen (Fig. 5 B) large (L 1.40 mm, W 0.80 mm), with semicircular basal portion; parameres wide at bases, with subparallel sides, narrowed and hooked in apical portion, with short, sparse setae. Sternite VIII (Fig. 5 C) (L 0.53 mm, W 0.83 mm) widely u-shaped, with sparse setation at apex; sternite IX (Fig. 5 D) (L 0.73 mm, W 0.68 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with setae. Tergite VIII (Fig. 5 E) (L 1.17 mm, W 1.17 mm) with apodemes shorter than apical plate, apical plate subtrapezoidal, apical margin rounded, with a row of dense, short setae, and longer, sparse ones. Tergite IX (Fig. 5 F) (L 1.0 mm, W 0.97 mm) with short apodemes, apical portion membranous, setation indistinct. Measurements and ratios. TL 9.6 mm, PW 2.7 mm, PL 1.6 mm, EL 8.3 mm, EW 4.9 mm, TL / EW 2.0, PW / PL 1.7, EL / EW 1.7, EL / PL 5.2, EW / PW 1.8.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	distribution	Distribution. Known only from the environs of Puerto Blest in the province Río Negro of Argentina (Fig. 23).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E337A7EFF38F938F418FE1C.taxon	etymology	Etymology. Lat. magnificus — splendid, magnificent, allusion to the remarkable body shape of the new species. Gender masculine.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	description	(Figs 1 C – E, 2 B, 4 B, 7 – 9)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	type_taxon	Type species: Solierodes rousseli (Solier, 1849), by present designation.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	diagnosis	Diagnosis. Large (TL 9.5 – 10.7 mm), non saltatorial, elongate, with subparallel sides of elytra, body covered with golden setae, setation of elytra in males patchy. Subantennal grooves shallow in males and moderately deep in females, subgenal ridge without buttonhole configuration. Pronotum short, in males distinctly narrower than basal portion of elytra, pronotum of females wider, slightly narrower than basal portion of elytra, with tubercles in posterior portion; anterolateral pronotal angles rounded, slightly protruding. Males: penis with bifid trigonium, parameres long, with subparallel sides and serrate margins. Females: coxites subtriangular, sclerotized, styli subapical; ventrite 4 with transverse patch of setae, ventrite 5 with elongate concavity.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	description	Description. Male. Body (Fig. 1 C) elongate, TL 9.5 – 10.7 mm, moderately convex. Colouration brown, head and pronotum covered with long, yellowish setae, setation of elytra a bit shorter, yellowish, with distinct brown patches, head and pronotum covered with dense but small punctures, elytra with slightly larger, dense punctures, punctation irregular. Head relatively small, wider than long; eyes large and protuberant, finely facetted; tempora long, distance from posterior margin of eye to occipital ridge ca. 0.6 × diameter of the eye, subparallel; distance from ventral margin of eye to subgenal ridge equal to ca. 0.3 × diameter of the eye; supraantennal ridges not elevated, joined to margins of eyes; frons with two shallow depressions behind antennal insertions; subantennal grooves slightly concave, relatively shallow, glossy and impunctate, each subantennal groove separated from frons by a short ridge near supraantennal ridge, continued along margin of eye as sharp ridge, in lower portion connected with subgenal ridge. Frontoclypeal suture absent; clypeus subtrapezoidal, ca. 2.3 × wider than long, with straight anterior margin, lateral margins converging posteriorly. Anteclypeus present, ca. 3 × shorter than clypeus. Ventral portion of head between subgenal ridges flat, gular area convex, rectangular, slightly longer than wide. Subgenal ridge well marked, without buttonhole configuration. Antennae filiform; antennomere 1 widest, subcylindrical, slightly curved; antennomere 2 subglobular and small, distinctly narrower and ca. 3 × shorter than antennomere 1; antennomere 3 long and narrow, as long as antennomere 1, slightly widening apically; antennomeres 4 – 10 similar to antennomere 3 but ca. 1.5 × longer, apical antennomere slightly longer than penultimate one, slightly widened in apical portion. Labrum covered with long setae, transverse, ca. 2.0 × wider than long, anterior margin slightly emarginated, anterolateral angles rounded. Mandibles (Fig. 7 A) symmetrical, ca. 2.0 × longer than width at bases, abruptly curved in apical 1 / 4, apices unidentate, outer margin covered with setae; inner margin of mandibles with big subtriangular denticle and two smaller ones; molar region with indistinct setation. Maxillary palpi (Figs 7 C): palpomere 1 shortest, subconical, 2 longest, ca. 2.5 × longer than palpomere 1, 3 ca. 3 × longer than palpomere 1, 4 narrow, digitiform, almost 2 × shorter than 3. Galea subrectangular, slightly longer than wide, much shorter than lacinia, with group of dense, irregularly arranged setae at apex; lacinia with dense, slightly curved setae on inner margin and apex. Mentum (Fig. 7 B) slightly transverse (W / L = 1.6), subtrapezoidal. Prementum subtriangular strongly widening apically, apical margin straight, ligula covered with dense setae. Labial palpomeres: palpomere 1 long, narrow, sinuate ca. 3 × longer than wide, sparsely covered with setae; palpomere 2 strongly widened, enlarged, subconical, as long as palpomere 1, ca. 2 × longer than wide, densely covered with long setae; apical palpomere arising from apex of preapical one, narrow, slightly shorter than palpomere 2, very sparsely covered with short setae. Pronotum small, transversely rectangular, distinctly narrower than base of elytra, ca. 2.1 × wider than long; lateral carinae subtly curved; disc convex, lateral portions flattened; anterior margin rounded; anterolateral angles rounded, slightly produced; posterior angles right-angled; base curved, with distinct margination ending laterally with small shiny area with neither setae nor punctures. Prosternum reduced anteriorly; prosternal process rhomboidal, posterior apex pointed and bent ventrally, ca. 1.8 × longer than wide, covered with setae. Scutellar shield triangular, slightly longer than wide, flat, with pointed apex; anterior margin straight, impunctate, covered with weakly impressed punctures. Elytra very long, with three subtle carinae, sides subparallel, distinctly wider at base than pronotal base; slightly widened in posterior half; humeri marked, apices regularly rounded. Epipleura wide at base, slightly narrower than metanepisternum, distinctly narrowed behind metaventrite, present till apices. Hind wings (Fig. 4 B) ca. 2.2 × longer than its width; radial cell subtriangular; apical portion of r 3 well visible, straight; rp-mp 2 arcuate, forming acute angle with MP 1 + 2; MP 4 long, oblique, joining MP 3 and CuA + AA 1 + 2; anal field with well marked AP 3 and AP 4. Medial field with 3 free veins; pigmented areas in radial cell, medial, radial, central and apical fields. Mesoventrite (Fig. 2 B) small; anterior margin with coxal rests; mesoventral process with well marked longitudinal suture; anterior portion forming deep, subtriangular fossa for reception of prosternal process; mesoventral process ca. 2.5 × longer than wide, sides subparallel, apex bilobed, touching metaventrite; mesocoxae completely separated; mesanepisternum rhomboidal, covered with setae, punctate, wider than long, forming coxal rest for procoxa; mesepimeron smaller, setose, subtriangular, strongly narrowed mesally; mesocoxae subconical, slightly projecting. Metaventrite (Fig. 2 B) moderately long, transverse (W / L 2.6), convex; discrimen complete; metanepisternum subtrapezoidal, ca. 2 × longer than its width, slightly narrowing posteriorly. Metacoxae moderately large, strongly oblique, 3.2 × wider than long, extending laterally to epipleura; metacoxal plate abruptly narrowing in outer half. Legs long; femora extending beyond elytral margin, trochanters moderately long; trochantero-femoral joint oblique; metafemora slightly shorter than metatibiae, metatibiae straight. Tibiae narrow, slightly widening towards apices, with paired carinae; length of metatibial spurs equal to largest width of metatibiae; tarsi distinctly shorter than tibiae; tarsomere 1 elongated, sides subparallel, as long as tarsomeres 2 – 3 combined; tarsomere 4 emarginated at apex; apical tarsomere as long as tarsomere 1, with simple claws. Abdomen ca. 1.5 × longer than its width, convex; ventrites 1 – 4 of subequal length; ventrite 5 slightly shorter; ventrite 5 regularly rounded at apex. Abdominal ventrites regularly covered with setae. Genitalia (Fig. 8) of Microcara type, symmetrical, penis large, dorsoventrally flattened, trigonium bifid, parameroids narrow, with subparallel sides, longer than trigonium. Tegmen large, parameres wide at bases, narrowing towards apices, apices pointed, margins serrate. Female (Figs 1 D, E). Body large (TL 10.4 – 10.6 mm), covered with longer and more erect setae; head more transverse, eyes less protuberant, subantennal grooves deeper than in males; pronotum wide (PW / PL 2.2), anterolateral angles rounded, slightly more produced than in males, with paired raised protuberances in lateral portion of posterior half; scutellar shield equilaterally triangular, elytra long, with subparallel sides; ovipositor long, branchlets absent; coxites subtriangular, sclerotized, styli subapical; bursal sclerites present; ventrite 4 with transverse patch of setae, ventrite 5 with elongate concavity.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	biology_ecology	Biology. According to the original description (Solier 1849), this species was found among scrub and on trees, together with Mucronotus velutinus (Solier). Specimens collected in recent decades were found in Valdivian forests, in a single case a specimen was fogged from Saxegothaea conspicua Lindley (Podocarpaceae).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E377A72FF38FF77F418FBA8.taxon	etymology	Etymology. Dedicated to Antoine Joseph Jean Solier (1792 – 1851), to commemorate his pioneering studies on Chilean marsh beetles. The genus name is a combination of Solier’s name and ending of a genus Elodes Latreille. Gender masculine.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E397A71FF38FD36F4DCFA88.taxon	description	(Figs 1 C – E, 2 B, 4 B, 6 A, 7 – 9)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E397A71FF38FD36F4DCFA88.taxon	materials_examined	Type material. Lectotype (MNHN, Fig. 6 A): “ Elodes \ rousseli Sol \ Calbuco ” [handwritten with brown ink, probably by Solier]; “ dans les broussailles, \ sur les arbres - \ contrefaît très bien \ le mort en applicant \ et tête et pattes sur le \ ventre. \ Calbuco. Mauldler ” [handwritten with brown ink, probably by Solier]; “ 9 \ 45 ” [handwritten on a green, round label]. The specimen is missing head and pronotum, and lateral portion of its right elytron is damaged. Additional material studied. 1 male (MNHN): “ 1362 ” [handwritten, without other labels]. 1 male (DBET): “ CHILE CW, reg. IX \ SE of NUEVA TOLTÉN \ BOROA CENTRAL \ 22. I. 2004 \ LGT M. SNIŽEK ”. 2 males (EMEC): “ 54 - CHILE X REGION \ Oncol Park, Fogging 14: 36 PM \ 300 cc / l, 08. I. 2007, 576 m 19 ° C \ S 39 ° 41.973 / W 73 ° 18.917 \ Saxegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 4 \ det. G. Arriagada 2007 ”. 1 female (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ D. Inward ”; “ Litter transect \ 3: 84 m. \ xi. 2003 ”; “ Scirt \ 2 ”. 1 female (CMNH): “ Prov. AYSEN \ 10 Km. N. Pto. Ibanez \ 14.17 Enero 1961 \ Coll: L. E. Pena ”. 1 male (FMNH): “ CUCAO \ Is. Chiloe \ 15 - 23 - II- 81 \ Coll. M. Marin ”; “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17 - 422 ”. 7 males (FMNH): “ Pto. Cisnes \ Aysen II 1961 \ Coll. L. E. Peña ”; “ FMNH, 1986 \ P. Peña Coll. \ Acc. # 17 - 422 ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E397A71FF38FD36F4DCFA88.taxon	diagnosis	Diagnosis. Large beetles with subparallel body; pronotum in males narrower than base of elytra; elytra covered with distinct setation. Parameroids of penis with subparallel sides, trigonium with two processes, parameres long, with serrate margins. Pronotum of females only slightly narrower than base of elytra, with tubercles in posterior portion.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E397A71FF38FD36F4DCFA88.taxon	description	Description. Male. Body large (TL 9.5 – 10.7 mm), elongate (TL / EW 2.0 – 2.2), widest in the posterior portion of elytra, moderately convex. Colouration chestnut brown, head covered with relatively long setae, pronotum covered with long, dense setae, elytra covered with shorter and sparser setae than pronotum, setation of elytra recumbent, patchy — yellowish with small black spots. Head ca. 1.3 × wider than long, ca. 1.6 × wider than interocular space, covered with coarse punctation, punctures separated by 0.2 – 0.3 × diameter of a puncture; frons with two elongated depressions; eyes large, protuberant. Pronotum transverse (PW / PL 1.9 – 2.2), punctation dense, punctures on disc separated by 0.3 – 0.5 × diameter of a puncture, punctures on lateral portion confluent. Elytra long (8.30 – 9.40 mm), punctures small, separated by ca. the diameter of a puncture, space between punctures matt. Aedeagus symmetrical, penis (Fig. 8 A) large (L 1.83 mm, W 0.43 mm), dorsoventrally flattened; trigonium as long as pala, bifid, processes of trigonium very narrow; parameroids longer than pala and trigonium, with subparallel sides, rounded at apices, outer margins serrate in apical portions. Tegmen (Fig. 8 B) large (L 1.70 mm, W 1.07 mm), widely u-shaped; parameres relatively wide at bases, regularly narrowing in basal halves and subparallel in apical halves, apices pointed on the outer margin and rounded on the inner margin, with few short setae. Sternite VIII (Fig. 8 C) (L 0.83 mm, W 1.03 mm) widely u-shaped, with sparse setation at apex; sternite IX (Fig. 8 D) (L 1.36 mm, W 0.73 mm) oval, consisting of two elongated, subtriangular hemisternites, apical 1 / 3 covered with setae. Tergite VIII (Fig. 8 E) (L 1.40 mm, W 1.00 mm) with apodemes shorter than apical plate, apical plate oval, covered with short setae, with rounded margin. Tergite IX (Fig. 8 F) (L 0.97 mm, W 0.76 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Body large (TL 10.4 – 10.6 mm), setation of dorsum uniformly yellowish, more erect than in males; head with coarse, confluent punctures, frons with two oblique elevated ridges, eyes less protuberant than in males; anterior portion of pronotal disc with dual punctation: larger punctures and small, weakly impressed punctures between them, lateral sides of pronotum with coarse, confluent, granulate punctures, posterior portion of pronotum with only very weakly impressed punctures; punctures on elytra separated by ca. the diameter of a puncture, area between punctures matt; pronotum wide (PW 3.5 mm), with elevated transverse ridge in posterior portion ending on sides with subconical tubercles; lateral sides of elytra subparallel; abdominal ventrite 4 with transverse patch of dense setae, ventrite 5 with subtriangular concavity (Fig. 9 D); ovipositor long (L 6.1 mm, Fig. 9 A), without branchlets, coxites subtriangular, elongated, apices of coxites with numerous short setae, styli subapical, eaxh stylus with single patch of sensillae (Fig. 9 B), bursal sclerite (L 1.34 mm, W 1.05 mm, width measured with membranous parts) subconical, widened in posterior portion and rounded in anterior part, surrounded with membranes (Fig. 9 C). Measurements and ratios. Males (n = 10) TL 9.50 – 10.70 mm (10.10 mm), PL 1.35 – 1.70 mm (1.52 mm), PW 2.90 – 3.50 mm (3.10 mm), EL 8.30 – 9.40 mm (8.81 mm), EW 4.50 – 5.10 mm (4.83 mm), TL / EW 2.0 – 2.2 (2.1), PW / PL 1.9 – 2.2 (2.1), EL / EW 1.8 – 1.9 (1.8), EL / PL 5.0 – 6.3 (5.8), EW / PW 1.4 – 1.6 (1.5). Females (n = 2) TL 10.40 – 10.60 mm (10.50 mm), PL 1.70 mm, PW 3.50 mm, EL 8.90 – 9.00 mm (8.95 mm), EW 4.60 – 4.70 mm (4.65 mm), TL / EW 2.3, PW / PL 2.1, EL / EW 1.9, EL / PL 5.2 – 5.3 (5.3), EW / PW 1.3.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E397A71FF38FD36F4DCFA88.taxon	distribution	Distribution. A rather rare species, known from scattered localities in the regions Los Ríos, Los Lagos, and Aysén of Chile (Fig. 23).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	description	(Figs 4 C, 10 – 22)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	type_taxon	Type species: Mucronotus velutinus (Solier, 1849), by present designation.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	diagnosis	Diagnosis. Moderately sized (TL 4.8 – 8.3 mm), non saltatorial beetles, covered with suberect vestiture. Subantennal grooves deep, subgenal ridge without buttonhole configuration, eyes moderately protuberant. Pronotum slightly narrower than base of elytra, anterolateral angles subtriangular, protruding, posterolateral angles obtuse, rounded. Male: penis with oval pala, distinct parameroids and bifid trigonium; tegmen with wide parameres forming two projections of variable shape in apical portion. Female: ovipositor with membranous coxites, styli apical; abdominal ventrites with regular setation.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	description	Description. Male. Body (Fig. 10) oblong oval, TL 4.8 – 8.3 mm, moderately convex. Body yellowish brown to brownish black, covered with dense, suberect setae, head and pronotum covered with granulate punctures, remaining portions of dorsum covered with normal (not granulate) punctures. Head moderately big, wider than long; eyes big, oval, moderately protuberant, finely facetted; tempora long, distance from posterior margin of eye to occipital ridge ca. diameter of the eye, converging basally; distance from ventral margin of eye to subgenal ridge equal to ca. 0.5 × diameter of the eye; supraantennal ridges slightly elevated, joined mesally to clypeal margin; frons flat to slightly convex; subantennal groove deep, ventrally reaching subgenal ridge. Frontoclypeal suture absent; clypeus transverse, ca. 3.6 × wider than long, with straight anterior margin, lateral margins converging posteriorly. Anteclypeus present, short, ca. 2 × shorter than clypeus. Ventral portion of head between subgenal ridges concave, gular area convex, square, gular ridges well developed. Subgenal ridge well marked, without buttonhole configuration. Antennae filiform, reaching basal 1 / 3 of elytra; antennomere 1 widest, subcylindrical, without ridge on anterior margin; antennomere 2 subconical, narrower and ca. 2.5 × shorter than antennomere 1; antennomere 3 very narrow, as long as antennomere 1, slightly widening apically; antennomeres 4 – 10 subtly widening apically, antennomere 4 slightly longer than antennomeres 5 – 10, which have similar length; apical antennomere slightly longer than preceding one. Labrum well visible, covered with relatively long setae, transverse, ca. 2.0 × wider than long, anterior margin slightly emarginated, anterolateral angles widely rounded. Mandibles (Fig. 12 A) symmetrical or subtly asymmetrical, ca. 2.0 × longer than width at bases, abruptly curved in apical 1 / 4, apices unidentate, outer margin covered with long setae; inner margin of mandibles with one well developed triangular denticle and two smaller subtriangular denticles, often fused and forming subrectangular lamina; molar region without setae. Maxillary palpi (Fig. 12 C): palpomere 1 short, subconical, 2 longest, ca. 2.0 × longer than palpomere 1, subcylindrical, 3 shorter than 2, subconical, 4 fusiform, narrow, slightly shorter than 3. Galea with rounded sides, shorter than lacinia, with long, irregularly arranged setae at apex; lacinia with straight or slightly curved, strong setae. Mentum (Fig. 12 B) transverse (W / L 1.4), subtrapezoidal. Ligula narrow at base, subtriangularly widening apically, apical margin membranous and bilobed, covered with dense setae. Labial palpomeres: palpomere 1 long, narrow, ca. 2.5 × longer than wide, slightly curved; palpomere 2 wide, subconical to axe-shaped; apical palpomere arising subapically from preapical one, digitiform, slightly curved, as long as palpomere 1. Pronotum transverse, distinctly narrower than base of elytra, ca. 2.0 × wider than long; lateral carinae slightly curved, converging anteriorly; disc moderately convex; anterolateral angles subtriangular, strongly projecting; posterior angles obtuse; base bisinuate. Prosternum reduced anteriorly; prosternal process narrow, rhomboidal, with elongated posterior portion that is slightly curved ventrad, ca. 2.5 – 3.0 × longer than wide, covered with subtle setae. Scutellar shield small, equilaterally triangular, with slightly curved lateral margins, flat, with pointed apex; anterior margin straight. Elytra elongated, distinctly wider at base than pronotal base; humeri well marked, apices regularly rounded, each elytron with three subtly marked longitudinal carinae. Epipleura wide at base, slightly narrower than metanepisternum, regularly narrowing till apex, present till apices. Punctation of elytra irregular, not granulate, uniform, sparse, punctures separated by ca. 1.0 – 1.5 × diameter of a puncture. Hind wings (Fig. 4 C) ca. 2.3 × longer than wide; radial cell subtriangular; apical portion of r 3 well visible, straight; rp-mp 2 arcuate, forming acute angle with MP 1 + 2; MP 4 long, oblique, joining MP 3 and CuA + AA 1 + 2; anal field with well marked AP 3 and AP 4. Medial field with 3 free veins; pigmented areas in radial cell, medial, radial, central and apical fields. Mesoventrite (Fig. 11) small; anterior margin with distinct coxal rests; mesoventral process with well marked longitudinal suture; anterior portion forming narrow but deep subtriangular fossa for reception of prosternal process; mesoventral process short, ca. 1.6 × longer than wide, sides subparallel to slightly converging posteriorly, apex bilobed, touching metaventrite; mesocoxae completely separated; mesanepisternum rhomboidal, wider than long, forming coxal rest for procoxa; mesepimeron smaller, impunctate, subtriangular, strongly narrowed mesally; mesocoxae triangular, slightly projecting. Metaventrite moderately long, transverse (W / L 3.2), convex; discrimen complete; metanepisternum subtrapezoidal, ca. 2 × longer than its width, slightly narrowing posteriorly. Metacoxae moderately large, strongly oblique, 3.0 × wider than long, extending laterally to epipleura; metacoxal plate obliquely narrowing, absent in outermost portion. Legs moderately long; femora slightly extending beyond elytral margin, trochanters moderately long; trochantero-femoral joint oblique; femora as long as or slightly longer than tibiae, widened in the middle of their length. Tibiae narrow, not widening towards apices, with paired carinae; metatibial spurs short, shorter than metatibial width; tarsi longer than half length of tibiae; tarsomere 1 elongated, narrow, slightly triangularly widened, as long as tarsomeres 2 – 4 combined; tarsomere 4 bilobed apically; apical tarsomere as long as tarsomeres 3 & 4 combined. Abdomen ca. 1.2 × longer than its width, moderately convex; ventrites 1 – 4 of subequal length; ventrite 5 slightly longer; ventrite 5 regularly rounded at apex. Abdominal ventrites regularly covered with setae. Genitalia (Figs 14, 16, 17, 19, 21) of Microcara type, symmetrical, penis large, dorsoventrally flattened, trigonium bifid, parameroids long, as long as trigonium. Tegmen large, subrectangular, parameres narrow at bases, additional ventral process present at base of each paramere, surrounding ventral portion of penis. Female (Figs 10 B, E, G, I). Body moderately large (TL 5.4 – 7.5 mm), in some species on average slightly smaller than male, in most species covered with longer, more erect setae; pronotum transverse, more convex, with subtriangular area of dense setae in posterior portion of central part of disc; scutellar shield equilaterally triangular, elytra long, with subparallel sides and three well marked longitudinal carinae, adscutellar portion of elytra slightly depressed in some species; abdominal ventrites with regular setation; ovipositor long, branchlets absent; coxites elongate, membranous, styli apical; bursal sclerite present, with small glands opening in its anterior half (Fig. 22 A); anterior portion of bursal sclerite connected with large gland (Fig. 22 A, B) and smaller, strongly folded structure, presumably bursella (Fig. 22 B, D).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	biology_ecology	Biology. Larvae possibly saproxylic (Ruta et al. 2018). Adults collected in temperate forests of southern Chile and Argentina.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3B7A74FF38FADEF6E6F9F9.taxon	etymology	Etymology. A hybrid of Latin words mucro = sword, sword point, sharp point and pronotum, a reference to sharp, pointing anterolateral angles of pronotum. Gender masculine.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	description	(Figs 10 A – B, 13 A – B, 14 – 15)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	materials_examined	Type material. Holotype, male (FMNH): “ CHILE: Arauco: Alto \ Llupehue, 4. I. 1977 \ L. E. Pena leg. \ FIELD MUS. NAT. HIST. ”. Paratypes, 1 male, 2 females (FMNH): same data as holotype. 1 male (CMNC): “ CHILE: Malleco; Puren \ Contulmo Natur Mon \ 11. XII. 84 - 13. II. 85 \ S & J Peck, FIT, 350 m \ mixed evergr. forest ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	diagnosis	Diagnosis. Posterolateral angles of pronotum well marked; elytra covered with dense but small punctures; processes of trigonium shorter than pala, bisinuate, with ca. 30 denticles on its outer margin and distinct basal lobe; anterior portion of bursal sclerite transversely oval.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	description	Description. Male. Body moderately large (TL 7.2 – 7.8 mm), elongate (TL / EW 1.8), widest in the middle of elytra, moderately convex. Colouration chestnut brown. Head ca. 1.5 × wider than interocular space, convex, densely covered with strong, granulate punctation, punctures separated by 0.3 – 0.5 × diameter of a puncture; eyes moderately large, protuberant. Pronotum transverse (PW / PL 2.14), covered with dense (separated by 0.3 – 0.5 × diameter of a puncture), strongly granulate punctures on sides and sparse punctures on central portion, lateral carinae of pronotum rounded, posterolateral angles well marked, anterolateral angles subtriangular. Elytra long, 4.4 × longer than pronotum, 1.4 × wider than pronotum; punctation irregular, sparse, separated by ca. the diameter of a puncture. Aedeagus symmetrical, penis (Fig. 14 B) large (L 1.32 mm, W 0.64 mm), dorsoventrally flattened; trigonium bifid, each process of trigonium sinuate, with curved lobe at base, outer margin with ca. 30 denticles; parameroids subtriangular, narrowed in apical portion, apices reaching apices of trigonium. Tegmen (Fig. 14 C) large (L 1.48 mm, W 1.14 mm), subrectangular, on each side with small, subtriangular lateral process; ventral processes wide, each with two lobes (subtriangular and suboval one); parameres digitiform, densely covered with short setae. Sternite VIII (Fig. 14 D) (L 0.64 mm, W 1.04 mm) widely v-shaped, with sparse setation at apex; sternite IX (Fig. 14 E) (L 0.74 mm, W 1.02 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with setae. Tergite VIII (Fig. 14 F) large (L 1.1 mm, W 1.06 mm), apodemes shorter than apical plate, apical plate subtrapezoidal, covered with microsetae, apical margin rounded, with a row of sparse setae. Tergite IX (Fig. 14 G) (L 1.06 mm, W 0.74 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Body slightly smaller (TL 6.5 – 7.5 mm), setation of dorsum dense, recumbent; head with coarse punctures separated by ca. 0.3 × diameter of a puncture; disc of pronotum more convex than in males; pronotal disc densely covered with small, not granulate punctures, lateral sides of pronotum with coarse, confluent, granulate punctures; punctures on elytra separated by ca. 0.5 – 1.0 × diameter of a puncture; ovipositor long (L 3.9 mm, Fig. 15 A), without branchlets, coxites subtriangular, elongated, apices of coxites with numerous short setae, styli apical, with two single patches of sensillae, bursal sclerite (L 1.6 mm, W 0.84 mm) elongate, with transversely oval sclerotization in anterior portion (Fig. 15 B). Measurements and ratios. Males (n = 3) TL 6.20 – 7.80 mm (7.1 mm), PL 1.20 – 1.50 mm (1.3 mm), PW 2.50 – 3.20 mm (2.8 mm), EL 5.10 – 6.40 mm (5.8 mm), EW 3.25 – 4.20 mm (3.9 mm), TL / EW 1.7 – 1.9 (1.8), PW / PL 2.1 – 2.2 (2.1), EL / EW 1.4 – 1.6 (1.5), EL / PL 4.3 – 4.5 (4.3), EW / PW 1.3 – 1.5 (1.4). Females (n = 2) TL 6.50 – 7.50 mm (7.00 mm), PL 1.20 – 1.30 mm (1.25 mm), PW 2.70 – 3.10 mm (2.90 mm), EL 5.20 – 6.30 mm (5.75 mm), EW 3.50 – 4.10 mm (3.80 mm), TL / EW 1.8 – 1.9 (1.8), PW / PL 2.3 – 2.4 (2.3), EL / EW 1.5 (1.5), EL / PL 4.3 – 4.9 (4.6), EW / PW 1.3 (1.3).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	distribution	Distribution. A very rare species, known from two localities in the northern part of the Araucanía region of Chile (Fig. 24 A, see Remarks below).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	biology_ecology	Biology. The species was collected in Valdivian temperate forest.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	etymology	Etymology. Greek αινιγματικός — mystery, riddle, a reference to the uncertain locality of the species (see Remarks below) and its extreme rarity.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E3F7A6AFF38FEC7F00DF8FF.taxon	discussion	Remarks. Locality of specimens from the collection of FMNH is probably misspelled and refer to the environs of Yupehue in Cautin province of Araucanía region (approximate coordinates: 38 ° 32 ’ 57.72 ” S, 73 ° 26 ’ 17.77 ” W).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	description	(Figs 10 C, 13 C, 16)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	materials_examined	Type material. Holotype, male (EMEC): “ 59 - CHILE X REGION \ Oncol Park, Quitaqui Way Fog- \ ging 12: 03 PM, 10. I. 2007, 526 m \ 19 ° C, S 39 ° 42.009 / W 73 ° 18.558 \ Sexegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 4 \ det. G. Arriagada 2007 ”. Paratypes (17 exx. in total, all males): 1 ex. (DBET): “ CHILE W, Reg. X \ W of Pargua \ 40 m 29. I. 2005 \ M. Snižek lgt. ”. 1 ex. (DBET): “ CHILE CW, reg. IX \ SE of NUEVA TOLTÉN \ BOROA CENTRAL \ 22. I. 2004 \ LGT. M. SNIŽEK ”. 1 ex. (EMEC): same data as holotype. 1 ex. (EMEC): “ 60 - CHILE X REGION \ Oncol Park, Quitaqui Way Fog- \ ging 15: 32 PM, 10. I. 2007, 519 m \ 13 ° C, S 39 ° 42.071 / W 73 ° 18.547 \ Saxegothaea conspicua 45 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx. (EMEC): “ 54 - CHILE X REGION \ Oncol Park, Fogging 14: 36 PM \ 300 cc / l, 08. I. 2007, 576 m 19 ° C \ S 39 ° 41.973 / W 73 ° 18.917 \ Sexegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 2 exx. (EMEC): “ 56 - CHILE X REGION \ Punta Curinanco, Fogging \ 12: 44 PM, 09. I. 2007, 177 m \ 17 ° C, S 39 ° 42.788 / W 73 ° 24.323 \ Aextoxicon punctatum 25 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 2 exx. (EMEC): “ 52 - CHILE X REGION \ Oncol Park, Calfuco Way Fog- \ ging 14: 13 PM, 150 cc / l, 07. I. 2007 \ 515 m 14 ° C, S 39 ° 42.114 / W 73 ° \ 19.244 Sexegothaea conspicua \ 35 m, Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ CHILE, X Region Los Rios, \ Oncol Park, 5 - 19.01.2007 \ Malaise trap T 1 \ Arias et Ruiz UC Berkeley ”; “ Scirtidae \ Genera \ sp. 1 \ det. J. E. Barriga-Tufion 2008 ”. 2 exx. (EUM): „ CHILI, Prov. Valdivia, \ Parque Oncol, 485 m, 9 I \ 2001 T. Cekalovic leg. ”. 1 ex. (EUM): “ CHILI, Prov. Valdivia, \ Parque Oncol, 485 m \ 6 I 2002 T. Cekalovic leg. ”. 1 ex. (EUM): “ CHILI, Region X, N of \ Valdivia, Parque Oncol 13. \ I. 2007 WD Shepard leg. ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ Malaise trap \ Week 1 Beach ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	diagnosis	Diagnosis. Posterolateral angles of pronotum well marked; elytra covered with dense but small punctures; processes of trigonium shorter than pala, curved, with ca. 15 denticles on its outer margin, gradually narrowing till apices.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	description	Description. Male. Body moderately large (TL 6.5 – 8.0 mm), elongate (TL / EW 1.7), widest in the middle of elytra, moderately convex. Colouration chestnut brown. Head ca. 1.5 × wider than interocular space, convex, densely covered with strong, granulate punctation, punctures separated by 0.3 – 0.5 × diameter of a puncture; eyes moderately large, protuberant. Pronotum transverse (PW / PL 2.0), covered with dense (separated by 0.3 – 0.5 × diameter of a puncture) and strongly granulate punctures on sides and more sparse punctures on central portion, lateral carinae of pronotum rounded, posterolateral angles well marked, anterolateral angles subtriangular. Elytra long, ca. 4 × longer than pronotum, 1.5 × wider than pronotum; punctation irregular, sparse, separated by ca. the diameter of a puncture. Aedeagus symmetrical, penis (Fig. 16 B) large (L 1.70 mm, W 0.67 mm), dorsoventrally flattened; trigonium bifid, each process of trigonium curved, with ca. 15 denticles on margin; parameroids subtriangular, narrowed in apical portion, as long as processes of trigonium. Tegmen (Fig. 16 C – D) large (L 1.77 mm, W 1.03 mm), subrectangular, with small, subtriangular lateral processes each with two denticles at apex; ventral processes wide, each with two lobes (large subtriangular and small suboval one); parameres digitiform, without distinct setation. Sternite VIII (Fig. 16 E) (L 0.70 mm, W 1.0 mm) widely v-shaped, with sparse setation at apex; sternite IX (Fig. 16 F) (L 1.0 mm, W 0.80 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with dense setae. Tergite VIII (Fig. 16 G) (L 1.17 mm, W 1.10 mm) with apodemes shorter than apical plate, apical plate subtrapezoidal, apical margin rounded, with sparse setae. Tergite IX (Fig. 16 H) (L 0.93 mm, W 0.73 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Unknown. Measurements and ratios. Males (n = 10) TL 6.50 – 8.0 mm (7.46 mm), PL 1.1 – 1.9 mm (1.44 mm), PW 2.5 – 3.3 mm (2.86 mm), EL 5.5 – 6.6 mm (6.24 mm), EW 3.8 – 4.7 mm (4.83 mm), TL / EW 1.70 – 1.83 (1.71), PW / PL 1.74 – 2.27 (2.0), EL / EW 1.38 – 1.55 (1.42), EL / PL 3.42 – 5.0 (4.21), EW / PW 1.42 – 1.52 (1.47).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	distribution	Distribution. A rare species, known from numerous collections from the Oncol Park (Los Ríos region) and two distant localities in regions Los Lagos and Aysén of Chile (Fig. 24).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	biology_ecology	Biology. Most specimens were collected in Valdivian temperate forests with predominating Aextoxicon punctatum Ruiz & Pav. and Sexegothaea conspicua.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E217A68FF38FAD4F693F8FC.taxon	etymology	Etymology. Referring to the area inhabited by the species — Patagonia.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	description	(Figs 10 D – E, 11 A, 13 D, 17 – 18)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	materials_examined	Type material. Lectotype (present designation), male (SDEI, Fig. 6 B): “ Puerto Puyuhuapi \ Süd-Chile \ leg. H. Schwabe ”; “ W 13 [illegible] \ 27.12.37 ” [handwritten]; “ Holotypus ” [red label, printed]; “ type ” [handwritten by Pic]; “ Pic det. ” [printed]; “ Cyphon \ schwabei \ n sp ” [handwritten by Pic]; “ Dtsch. Entomol. \ Institut Berlin ”; “ DEI-Müncheberg \ Col — 04805 ” [green label, printed]. Paralectotype (present designation), female (SDEI, Fig. 6 C): “ Puerto Puyuhuapi \ Süd-Chile \ leg. H. Schwabe ”; “ W 13 [illegible] \ 27.12.37 ” [handwritten]; “ Paratypus ” [red label, printed]; “ Pic det. ” [printed]; “ var ” [handwritten by Pic]; “ Dtsch. Entomol. \ Institut Berlin ”; “ DEI- Müncheberg \ Col — 04806 ” [green label, printed]. Paralectotype (present designation), 1 ex. (MNHN, Fig. 6 D): “ Puerto Puyuhuapi \ Süd-Chile \ leg. H. Schwabe ”; “ W 13 [illegible] \ 26.12.37 ” [handwritten]; “ Cyphon \ schwabei \ n sp ” [handwritten by Pic]; “ cotype ” [handwritten by Pic]. Additional material examined. 2 exx. (CMNC): “ CHILE: Valdivia \ 34 km WNW La Union \ 17. XII. 84 - 7. II. 85 \ S & J Peck, 700 m, FIT \ mixed evergr. forest ”. 1 ex. (CMNC): “ CHILE: Osorno: Puyehue \ Nat. Pk; Aquas Calientes \ 18. XII. 84 - 8. II. 85 \ S & J Peck, 600 m, malaise \ Nothofagus forest ”. 3 exx. (CMNC), 2 exx. (FMNH): “ CHILE: Malleco; Puren \ Contulmo Natur Mon \ 11. XII. 84 - 13. II. 85 \ S & J Peck, FIT, 350 m \ mixed evergr. forest ”. 1 ex. (CMNC): “ CHILE: Palena Prov. \ 37 km SE Chaiten \ 28. XII. 84 - 30. I. 85 \ S & J Peck, FIT, 60 m \ riverside 2 nd forest ”. 3 exx. (CMNC): “ CHILE: Cautin; 21 km NE \ Pucon, Lago Caburga \ 15. XII. 84 - 10. II. 85 \ S & J Peck, FIT, 600 m \ mixed forest remnant ”. 3 exx. (CMNC): “ CHILE: Lago Chapo \ 34 km E Pto. Montt \ 24. XII. 84 - 2. II. 85 \ S & J Peck, 300 m, FIT \ 2 nd growth Nothofagus ”. 2 exx. (CMNC): “ CHILE: Cautin, 10 kmS \ Pucon, Vol. Villarrica N. P. \ 15. XII. 84 - 10. II. 85 \ S & J Peck, FIT, 900 m \ Nothofagus grove on ash ”. 3 exx. (CMNC): “ CHILE: Malleco: Princesa \ 20 km W Curacautin \ 12. XII. 84 - 16. II. 85 \ S & J Peck, FIT, 1000 m \ Nothofagus forest ”. 2 exx. (CMNC): “ CHILE: Aisen; 16 km NW \ Cisnes Medio, Rio Grande \ 30. XII. 84 - 28. I. 85 \ S & J Peck, 200 m, FIT \ mature beech forest ”. 2 exx. (CMNH): “ Villarrica \ Cautin \ II-III- 1979 \ Coll. L. E. Pena ”; “ CMNH Acc. \ No. 31811 ”. 3 exx. (DBET): “ CHILE W, Reg. X \ Chiloe Isl. SE, 20 km \ NNW of Quellon \ 216 m, 42.58, 73.45 \ 31.1.2005, Snižek lgt. ”. 1 ex. (DBET): “ CHILE E, reg. X \ E of PUERTO MONTT \ SE of LAGO CHAPO \ 30.1.2004 \ LGT. M. SNIŽEK ”. 1 ex. (DBET): “ CHILE CW, reg. X \ W of LA UNION \ E of EL MIRADOR \ 27.1.2004 \ LGT. M. SNIŽEK ”. 2 exx. (DBET): “ CHILE W, Reg. X \ W of Pargua \ 40 m 29. I. 2005 \ M. Snižek lgt. ”. 1 ex. (DBET): “ CHILE \ Huerquehue NP. \ nr Tingika L \> = 780 m \ 20 - 26.12.2013 ”, “ S 39 ° 10 ’ \ W 71 ° 44 ’ \ leg. E. Lukasievich ”. 2 exx. (EMEC): “ 54 - CHILE X REGION \ Oncol Park, Fogging 14: 36 PM \ 300 cc / l, 08. I. 2007, 576 m 19 ° C \ S 39 ° 41.973 / W 73 ° 18.917 \ Sexegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 62 - CHILE X REGION \ Chaihuin, Hueicolla Way Fog- \ ging 12: 42 PM, 12. I. 2007 107 m \ 18 ° C, S 39 ° 59.926 / W 73 ° 38.976 \ Nothofagus dombeyi 20 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx. (EMEC): “ 61 - CHILE X REGION \ Chaihiun Hueicolla Way, Fogging \ 12: 23 PM, 12. I. 2007, 107 m, 18 ° CS \ 39 ° 51.926 / W 73 ° 38.976 \ Nothofagus dombeyi 20 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 56 - CHILE X REGION \ Punta Curinanco, Fogging \ 12: 44 PM, 09. I. 2007, 177 m \ 17 ° C, S 39 ° 42.788 / W 73 ° 24.323 \ Aextoxicon punctatum 25 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx. (EMEC): “ 70 - CHILE X REGION \ Chaihuin Res. Costera Valdivi- \ ana, Fogging 11: 44 AM, 14. I. 2007, \ 191 m 21 ° C, S 39 ° 59.986 / W 73 ° 38. \ 971, Nothofagus nitida 35 m, \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx (EMEC): “ 66. - Chile X Region, Chaihuin \ Res. Costera Valdiviana _ 362 m \ Las Garzas Way Fogging 11: 27 AM \ 12 / I / 2007 17 ° C Nothofagus nitida \ S 39 ° 59.700 / W 73 ° 34.663 \ Arias et al Berkeley ”. 1 ex. (EMEC): “ 59 - CHILE X REGION \ Oncol Park, Quitaqui Way Fog- \ ging 12: 03 PM, 10. I. 2007, 526 m \ 19 ° C, S 39 ° 42.009 / W 73 ° 18.558 \ Sexegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 53 - CHILE X REGION \ Oncol Park, Bonifacio Way \ Fogging 12: 43 PM 300 cc / l \ 08. I. 2007, 606 m 17.6 ° C \ S 39 ° 41.979 / W 73 ° 18.938 \ Saxegothaea conspicua 40 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx. (EMEC): “ 69 - CHILE X REGION \ Chaihuin Res. Costera Valdivi- \ ana, Las Garzas Way Fogging \ 09: 13 AM, 14. I. 2007, 210 m 19 ° C, \ S 40 ° 00.026 / W 73 ° 38.962, Nothofagus \ dombeyi 20 m, Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ CHILE, X Region Los Rios, \ Oncol Park, 19.01 / 1.02.2007 \ Malaise trap T 2 \ Arias et Ruiz UC Berkeley ”; “ Scirtidae \ Genera \ sp. 1 \ det. J. E. Barriga-Tuñon 2008 ”. 1 ex. (EMEC): “ CHILE, X Region Los Rios, \ Oncol Park, 6 - 20.03.2007 \ Malaise trap T 2 \ Arias et Ruiz UC Berkeley ”; “ Scirtidae \ Genera \ sp. 1 \ det. J. E. Barriga-Tuñon 2008 ”. 1 ex. (EMEC): “ 37 ° 48 ’ 12 ’’ S / 73 ° 01 ’ 28.5 ’’ W \ CHILE, IX Region, vic. Naheulb \ pitfall traps, 7 - 25: i: 2003 \ Colls. K. Will, S. Lew ”; “ Cyphon \ sp. \ det. J. E. Barriga ”; “ U. C. Berkeley \ EMEC 107091 ”. 1 ex. (FMNH): “ CHILE: Llanquihue Pr.: \ P. N. Vicente Perez Bos- \ ales, 9.2 km NE Ensen- \ ada on road to Petrohué, ”, “ 125 m, 41 ° 10.20 ’ S, \ 72 ° 27.10 ’ W, 2 - 28. \ 1997, Valdivian rainfor. \ with Nothofagus spp. ”, “ FMHD 97 - 9, carrion \ trap (squid), A. Newton \ & M. Thayer 987 \ FIELD MUS. NAT. HIST. ”. 2 exx. (FMNH): “ CHILE: Llanquihue \ Prov., Lenca, 45 km SE \ Pto Montt., 100 m \ 25. XII. 1984 ”, “ FMHD # 85 - 947, forest \ remnant, leaf stick \ litter, S. & J. Peck, \ P # 85 - 63, berlese \ FIELD MUS. NAT. HIST ”. 1 ex. (FMNH): “ DUHATAO \ Is. Chiloe W. \ 16 - 21 - I- 81 \ Coll. M. Marin ”, “ FMNH, 1986 \ L. E. Pena Coll. \ Acc. # 17 - 422 ”. 1 ex. (FMNH): “ CUCAO \ Is. Chiloe \ 23 - 15 - 11 - 81 \ Coll. M. Marin ”, “ FMNH, 1986 \ L. E. Pena Coll. \ Acc. # 17 - 422 ”. 5 exx. (FMNH): “ LLANQUIHUE \ Hornohuinco \ Lag. Chapo ”, “ Dec. 1968 \ L. E. Peña ”, “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17.422 ”. 1 ex. (FMNH): “ ENCO 1100 m \ Valdivia CH. \ II- 78 \ Coll: L. E. Pena ”, “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17.422 ”. 1 ex. (FMNH): “ Prov. Malleco \ Rio Blanco \ Curacautin ”, “ Febr. 1964 \ Coll. L. E. Pena ”, “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17.422 ”. 2 exx. (MHNG): “ CHILE: X Reg. Prov. Osorno \ PN Puyehue, Aguas \ Calientes, 400 - 500 m \ 31. XII. 1990 - 1. I. 1991 \ Agosti & Burckhardt # 25 b ”. 3 exx. (MHNG): “ CHILE: IX Reg. Prov. Cautin \ PN Huerquehue, 800 m \ 22 - 25. XII. 1990, Valdivian \ lauriphyllous forest \ Agosti & Burckhardt # 17 b ”. 1 ex. (MHNG): “ CHILE: X Reg. Prov. Chiloé \ Cucao, 30 km SW Castro \ PN Chiloé, 30 m, temp. rain \ forest, 4 - 6. I. 1991 \ Agosti & Burckhardt # 29 a ”. 6 exx. (MHNG): “ CHILE: X Reg. Prov. Chiloé \ Cucao, 30 km SW Castro \ PN Chiloé, 30 m, temp. rain \ forest, 4 - 6. I. 1991 \ Agosti & Burckhardt # 29 b ”. 1 female (MNHN): “ 1361 ” [handwritten, without other labels]. 1 male (MNHN): “ 9 \ 45 ” [round label, green on reverse], “ MUSEUM PARIS \ CHILI \ CL. GAY 1845 ” [printed], “ Elodes \ velutinus \ Chili ” [handwritten]. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Sweeping \ herbage ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 8 exx. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Glacier Path \ MT \ 25. xi. - 12. xii. 03 ”. 3 exx. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; „ Glacier Path \ MT \ 21. xi. - 12. xii. 03 ”; “ ridge \ epiphytes ”. 2 exx. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Glacier Path \ Hilltop FIT \ 30. xi. - 7. xii. 03 ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Litter transect \ 2: 21 m. \ xi. 2003 ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Litter transect \ 5: 14 m. \ xi. 2003 ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Litter transect \ 7: 7 m. \ xi. 2003 ”. 2 exx. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Below base \ 1 - 10. xii. 03 \ shrub flowers ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Below Base \ 21. xi- 12. xii. 03 \ beating ”. 3 exx. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Base area \ 21. xi. - 12. xii. 03 \ beating ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Base area \ 21. xi. - 12. xii. 03 ”; “ litter at \ forest edge ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ Base area, MT \ 21. xi. - 12. xii. 03 ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ New Path \ 21. xi. - 12. xii. 03 ”. 1 ex. (NHM): “ CHILE: Reg. XI \ Queulat N. P. \ P. M. Hammond ”; “ New path \ xi. 2003 ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ F. I. T. 1 / 2 \ in forest ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 1 ex. (NHM): “ Peulla. \ 12 - 13. xii. 1926. ”; „ S. Chile: \ Llanquihue prov. \ F. & M. Edwards. \ B. M. 1927 - 63. ”. 1 ex. (NHM): “ Chiloe I., \ Chile. \ C. Darwin. ”; „ Darwin Coll. \ 1885 - 119 ”; „ Chiloe \ 2309 ”; „ Cyphon \ velutinus \ Solier ” [written with pencil]. 2 exx. (NHMB): “ Chile Prov \ Valdivia \ E. Krahmer ”; “ Santo \ Domingo \ 27. XI. 1977 ”. 6 exx. (NHMF): “ CHILE, prov. Valdivia, \ Chaihuín, Camino a \ Huaicolla, 12 ene 2007, \ 107 mt, 39 ° 59.926 S 73 ° \ 38.976 W, fogging \ Nothofagus dombeyi, \ leg. J. E. Barriga-Tuñón ”. 7 exx. (NMPC): “ CHILE, IX. r. 1. - 3.2. \ 2005, Manzanar \ 38 ° 28.071 - 71 ° 41.873 \ 800 m, Sv. Bilý leg. ”. 1 ex. (NMPC): “ CHILE, IX. r., 9. - 11.2. \ 2005, P. N. Nahuelbuta \ 37 ° 48.530 - 73 ° 00.954 \ 1200 m, Sv. Bilý leg. ”. 1 ex. (NMPC): “ CHILE, IX. r., 4. - 5.2. \ 2005, Laguna Icalma \ 38 ° 49.148 - 71 ° 19.097 \ 1250 m, Sv. Bilý leg. ”. 2 exx. (SEMC): “ CHILE: Valdivia \ 30 km SE Panguipulli \ 39 ° 44 ’ S, 72 ° 20 ’ W, 200 m \ 15 Nov 1994. R. Les- \ chen & C. Carlton # 107 \ ex: sifting leaf litter ”. 1 ex. (SEMC): “ CHILE: Osorno \ 48.5 km W. Osorno \ 40 ° 37 ’ S, 73 ° 45 ’ W, 75 m \ 28 Nov. 1994, R. Les- \ chen & C. Carlton # 181 \ ex: sifting leaf litter ”. 1 ex. (SEMC): “ CHILE: Valdivia \ 10 km NW Choshuenco \ 39 ° 45 ’ S, 72 ° 20 ’ W, 250 m \ 15 Nov 1994, R. Les- \ chen & C. Carlton # 102 \ ex: beating vegetation ”. 3 exx. (SEMC): “ CHILE: Chiloe \ 33 km SW Chonchi \ Lago Tepuhueco, 25 m \ 42 ° 49 ’ S, 73 ° 55 ’ W \ 26 Nov 1994, R. Les- \ chen & C. Carlton # 163 \ ex: beating vegetation ”. 1 ex. (SEMC): “ CHILE: Chiloe \ 31.4 km SW Chonchi \ Lago Tepuhueco, 50 m \ 42 ° 51 ’ S, 73 ° 47 ’ W \ 26 Nov 1994, R. Leschen \ & C. Carlton # 172 \ ex: sifting leaf litter ”. 1 ex. (SEMC): “ CHILE: Valdivia \ 37 km SE Panguipulli \ 39 ° 45 ’ S, 72 ° 20 ’ W, 300 m \ 14 Nov 1994, R. Leschen \ & C. Carlton # 099 \ ex: sifting leaf litter ”. 1 ex. (SEMC): “ CHILE: Llanquihue \ Lago Llanquihue, 200 m \ 3.3 km NW Ensenada \ 41 ° 18 ’ S, 72 ° 29 ’ W \ 12 Nov 1994, R. Leschen \ & C. Carlton # 087 \ ex: beating vegetation ”. 1 ex. (ZMUC): “ ARGENTINA, Rio Negro \ 8: Lago Nahuel Huapi \ Puerto Blest, 770 m \ 23. xii. 1978 \ Mision Cientifica Danesa ”. 1 ex. (ZMUC): “ ARGENTINA, Rio Negro \ 8: Lago Nahuel Huapi \ Puerto Blest, 770 m \ 1. i. 1979 \ Mision Cientifica Danesa ”. 1 ex. (ZMUC): “ CHILE, Osorno \ 13: Parque Nacional \ Puyehue Anticura \ 15. xii., 350 m \ Nielsen & Karsholt ”. 1 ex. (ZMUC): “ CHILE, Valdivia \ 15: 20 km S Valdivia \ Rincon de la Piedra \ 14. xi. 1981, 180 m \ Nielsen & Karsholt ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	diagnosis	Diagnosis. Posterolateral angles of pronotum well marked; elytra covered with dense, large punctures; processes of trigonium longer than pala, bisinuate, with ca. 15 denticles on its dorsal edge, slightly widened at apex; anterior portion of bursal sclerite with transverse bar, hooked at sides.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	description	Description. Male. Body moderately large (TL 4.8 – 8.0 mm), elongate (TL / EW 1.7 – 1.9), widest in the middle of elytra, moderately convex. Colouration yellowish brown to chestnut brown. Head ca. 1.5 × wider than interocular space, convex, sparsely covered with granulate punctures, punctures separated by ca. 0.5 – 1.0 × diameter of a puncture, frons with two impunctate depressions; eyes moderately large, protuberant. Pronotum transverse (PW / PL 2.1 – 2.6), punctation on disc not granulate, sparse, punctures separated by ca. 2.0 × diameter of a puncture, punctation on sides granulate, as on head, lateral carinae of pronotum almost straight, posterolateral angles well marked, anterolateral angles subtriangular. Elytra long, 4.6 – 6.0 × longer than pronotum, 1.4 – 1.5 × wider than pronotum; punctation irregular, punctures large, dense, separated by ca. 0.5 – 1.0 × diameter of a puncture. Aedeagus symmetrical, penis (Fig. 17 A) large, (L 1.13 mm, W 0.53 mm) dorsoventrally flattened, pala shorter than parameroids and trigonium; trigonium bifid, each process of trigonium long, narrow, sinuate, slightly enlarged at apex, ventral edge with ca. 15 denticles; parameroids subtriangular, very long and narrow, apices subtly longer than processes of trigonium. Tegmen (Fig. 17 B) large (L 1.2 mm, W 0.6 mm), elongate, without lateral processes; ventral processes narrow, digitiform; parameres lyre-shaped, subtly denticulate on outer margins. Sternite VIII (Fig. 17 C) (L 0.38 mm, W 0.53 mm) widely v-shaped, with indistinct setation at apex; sternite IX (Fig. 17 D) (L 0.43 mm, W 0.73 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with sparse setae. Tergite VIII (Fig. 17 E) elongated (L 0.77 mm, W 0.67 mm), apodemes shorter than apical plate, apical plate subrectangular, covered with microsetae, apical margin tempered, with a row of sparse, relatively long setae and much more numerous, very short, dense setae. Tergite IX (Fig. 17 F) (L 0.50 mm, W 0.40 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Body on average smaller (TL 5.40 – 6.70), setation of dorsum similar to that on males, no external differences in punctation of dorsum, structure of pronotum or adscutellar portion of elytra noticeable; ovipositor long (L 3.6 mm, Fig. 18 A), without branchlets, coxites subtriangular, elongated, apices of coxites rounded, with numerous short setae, styli apical, with two patches of sensillae, bursal sclerite (L 1.38 mm, W 0.78 mm) with indistinct elongate sclerotizations and well developed transverse bar hooked at sides (Fig. 18 B). Measurements and ratios. Males (n = 10) TL 4.80 – 8.00 mm (6.3 mm), PL 0.80 – 1.40 mm (1.0 mm), PW 1.90 – 3.00 mm (2.4 mm), EL 4.20 – 7.20 mm (5.4 mm), EW 2.90 – 4.30 mm (3.5 mm), TL / EW 1.7 – 1.9 (1.7), PW / PL 2.1 – 2.6 (2.3), EL / EW 1.4 – 1.7 (1.5), EL / PL 4.6 – 6.0 (5.0), EW / PW 1.4 – 1.5 (1.5). Females (n = 10) TL 5.40 – 6.70 mm (6.0 mm), PL 0.90 – 1.10 mm (1.0 mm), PW 2.05 – 2.60 mm (2.32 mm), EL 4.40 – 5.70 mm (5.0 mm), EW 3.00 – 3.90 mm (3.4 mm), TL / EW 1.7 – 1.9 (1.8), PW / PL 2.1 – 2.4 (2.2), EL / EW 1.4 – 1.6 (1.5), EL / PL 4.4 – 5.2 (5.1), EW / PW 1.4 – 1.6 (1.5).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	distribution	Distribution. A common species, recorded from regions Bío Bío, Araucanía, Los Ríos, Los Lagos, and Aysénin of Chile and province Rio Negro in Argentina (Fig. 24 B).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	biology_ecology	Biology. Most specimens were collected in Valdivian temperate forests with Aextoxicon punctatum, Nothofagus dombeyi (Mirb.) Oerst., N. nitida (Phil.) Krasser, and Sexegothaea conspicua.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E237A6DFF38FDC3F6F8FEF9.taxon	discussion	Remarks. According to the original description (Pic 1938) type specimens were collected 27.12.1937, and were deposited in DEI and the collection of Pic (now in MNHN). The specimen from the collection of MNHN bears the date 26.12.1937 but it was collected in the same locality and was designated a paralectotype. The specimen from the collection of SDEI is designated here as the lectotype in order to preserve the stability of nomenclature by selecting one specimen as the sole, name-bearing type of the taxon.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	description	(Figs 10 F – G, 13 E, 19 – 20)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	materials_examined	Type material. Holotype, male (DBET): “ CHILE W, Reg. X \ Chiloe Isl. SE, 20 km \ NNW of Quellon \ 216 m, 42.58, 73.45 \ 31.1.2005, Snižek lgt. ”. Paratypes (97 exx. in total). 5 exx. (CMNC): “ CHILE: Chiloe Isl. \ 8 km S Ancud \ 1. II. 85, S & J Peck \ forest remnant litter ”. 4 exx. (CMNC): “ CHILE: Osorno: Puyehue \ Nat. Pk; Aquas Calientes \ 18. XII. 84 - 8. II. 85 \ S & J Peck, 600 m, malaise \ Nothofagus forest ”. 1 ex. (CMNC): “ CHILE: Osorno: Puyehue \ N. P.; Aguas Calientes \ 20. XII. 84, S & J Peck \ 500 m, sifted forest \ stick litter, Pionero Tr. ”. 1 ex. (CMNC): “ CHILE: Osorno: Puyehue \ N. P.; Aguas Calientes \ 20. XII. 84 - 8. II. 85 \ S & J Peck, 500 m, FIT \ Derumbes Forest Trail ”. 1 ex. (CMNC): “ CHILE: Osorno: Puyehue \ Nat. Park, Antillanca Rd. \ 18 - 20. XII. 84, S & J Peck \ 500 - 1000 m, carnetting ”. 1 ex. (CMNC): “ CHILE: Lago Chapo \ 34 km E Pto. Montt \ 24. XII. 84 - 2. II. 85 \ S & J Peck, 300 m, FIT \ 2 nd growth Nothofagus ”. 2 exx. (CMNH): „ Prov. AYSEN \ 10 Km. N. Pto. Ibanez \ 14.17 Enero 1961 \ Coll: L. E. Pena ”; „ CHILE ”. 31 exx. (DBET): same data as holotype. 2 exx. (DBET): “ CHILE W, Reg. X \ Fresia, WWN of Parga \ 480 m 27. I. 2005 \ M. Snižek lgt. ”. 3 exx. (DBET): “ CHILE W, Reg. X \ Purrangue 600 m \ W of Hueyusca \ 26 I 2005, \ M. Snižek ”. 5 exx. (DBET): “ CHILE W Reg. X \ W of Los Muermos \ 1 - 2 km E Oc. Pacifico \ 160 m 41.20, 73.49 \ 28 I 05, leg. M. Snižek ”. 1 ex. (DBET): “ CHILE W, Reg. X \ NC Chiloe, 20 km \ NNE Castro, Inia env. \ 180 m 42.15.72 73.40.50 \ 31. I. 2005 Snižek lgt. ”. 1 ex. (DBET): „ CHILE — reg. X, 50 kmSW \ Los Muermos, 30.1.2005 \ 41 ° 30.0 ’ S, 73 ° 35.7 ’ W \ leg. M. Halada, 150 m ”. 4 exx. (DBET): “ CHILE W, Reg. X \ NW Chiloe, SW of \ Aneud 250 m \ 4.11.2005 Snižek lgt. ”. 1 ex. (DBET): “ CHILE — Isla de Chiloe \ W of Castro, 9.2.2005 \ 42 ° 20 ’ S, 73 ° 48 ’ W \ leg. M. Halada 200 m ”. 1 male (EMEC): “ 62 - CHILE X REGION \ Chaihuin, Hueicolla Way Fog- \ ging 12: 42 PM, 12. I. 2007 107 m \ 18 ° C, S 39 ° 59.926 / W 73 ° 38.976 \ Nothofagus dombeyi 20 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 male (EMEC): “ 42 ° 36 ’ 50.3 ’’ S / 74 ° 06 ’ 14.7 ’’ \ CHILE, X Region, Chiloe Nat. Pk. \ under logs / rocks, 20: i: 2002 \ Colls. K. Will, S. Lew ”; “ Cyphon \ sp. \ det.: J. E. Barriga ”; “ U. C. Berkeley \ EMEC 107098 ”. 2 males (EMEC): “ 66. - Chile X Region, Chaihuin \ Res. Costera Valdiviana _ 362 m \ Las Garzas Way Fogging 1: 27 AM \ 12 / I / 2007 17 ° C Nothofagus nitida \ S 39 ° 59.700 / W 73 ° 34.663 \ Arias et al Berkeley ”; “ Elodes \ rouselii \ Solier 1849 \ det. J. E. Barriga-Tuñon 2008 ”; “ UC Berkeley \ EMEC 118524 ”. 1 male (EMEC): “ 70. - Chile X Region, Chaihuin \ Res. Costera Valdiviana \ Fogging 11: 44 AM 14 / I / 2007. \ 191 m 21 ° C Nothofagus nitida \ 35 m S 39 ° 59.986 / W 73 ° 38.971 \ Arias et al Berkeley ”; “ Elodes \ rouselii \ Solier 1849 \ det. J. E. Barriga-Tuñon 2008 ”; “ UC Berkeley \ EMEC 118160 ”. 3 exx. (EMEC): “ 60 - CHILE X REGION \ Oncol Park, Quitaqui Way Fog- \ ging 15: 32 PM, 10. I. 2007, 519 m \ 13 ° C, S 39 ° 42.071 / W 73 ° 18.547 \ Saxegothaea conspicua 45 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 61 - CHILE X REGION \ Chaihiun Hueicolla Way, Fogging \ 12: 23 PM, 12. I. 2007, 107 m, 18 ° CS \ 39 ° 51.926 / W 73 ° 38.976 \ Nothofagus dombeyi 20 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 3 exx. (EMEC): “ 67 - CHILE X REGION \ Chaihuin Reserva Costera Valdivi- \ ana, Las Garzas Way, Fogging \ 15: 27 PM, 13. I. 2007, 357 m 17 ° C, \ S 39 ° 59.669 / W 73 ° 34.721, Nothofagus \ dombeyi 35 m, Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EUM): “ CHILI, Prov. Valdivia, \ Pargue Oncol, Sendero \ Calfuco 17. I. 2007 \ T. Cekalovic leg TC- 831 ”. 1 ex. (EUM): “ CHILI, Prov. Valdivia, \ Pargue Oncol, 700 m, \ 14. I. 2001 T. \ Cekalovic leg. TC- 647 ”. 1 ex. (EUM): “ Prov. Llanquinve, \ Chinquihve Alto, \ 12. I. 1990 \ T. Cekalovic leg. ”. 1 ex. (FMNH): “ LLANQUIHUE \ Hornohuinco \ Lag. Chapo ”, “ Dec. 1968 \ L. E. Peña ”, “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17.422 ”. 1 ex. (FMNH): “ Pto Cisnes \ Aysen II- 1961 \ Coll. L. E. Pena ”, “ FMNH, 1986 \ L. Peña Coll. \ Acc. # 17.422 ”. 1 ex. (MHNG): “ CHILE: X Reg. Prov. Osorno \ PN Puyehue, Aguas \ Calientes, 400 - 500 m \ 31. XII. 1990 - 1. I. 1991 \ Agosti & Burckhardt # 25 b ”. 2 exx. (MHNG): “ CHILE: X Reg. Prov. Chiloé \ Cucao, 30 km SW Castro \ PN Chiloé, 30 m, temp. rain \ forest, 4 - 6. I. 1991 \ Agosti & Burckhardt # 29 a ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ Moss from \ tree ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ LEAF LITTER ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 2 exx. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ Forest Litter ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 1 ex. (NHM): “ 19 CHILE X Reg. \ PN Puyehue \ Anticura \ I. Ribera leg. 31. I. 1999 ”; “ BM 1999 - 195 \ 19. i. - 9. ii. 1999 \ I. Ribera ”. 1 ex. (NHM): “ 49685 ”; “ ex Mus \ Murray ”; “ Chili ”; “ Fry Coll. / 1905 - 100. ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Malaise \ Trap 2 ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 2 exx. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ Malaise trap \ Week 1 Beach ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ” [1 specimen with additional label “? gen. n. 1 \ sp. 1 \ P. M. Hammond det. ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ Base Camp \ 46 ° 44 ’ 44 ’’ S \ 73 ° 56 ’ 34 ’’ W ”; “ Moss from \ ground ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 1 ex. (NHM): “ CHILE: \ Laguna San Rafael \ Feb-March 1999 ”; “ BM- 1999 - 107 \ P. M. Hammond \ K. A. Jackson ”. 1 ex. (NHMF): „ CHILE, prov. Valdivia, \ Chaihuín, Camino a \ Huaicolla, 12 ene 2007, \ 107 mt, 39 ° 59.926 S 73 ° \ 38.976 W, fogging s / \ Nothofagus dombeyi, \ leg. J. E. Barriga-Tuñón ”. 1 ex. (SEMC): “ CHILE: Osorno \ 14.5 km W Puaucho \ 40 ° 37 ’ S, 73 ° 45 ’ W, 175 m \ 11 - 28 Nov 1994. R. Les- \ chen & C. Carlton # 170 \ ex flight intercept trap ”.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	diagnosis	Diagnosis. Posterolateral angles of pronotum well marked; elytra covered with dense but small punctures; processes of trigonium shorter than pala, curved, distinctly widened at apices, with ca. 20 denticles on its outer margin and 6 denticles on inner margin; anterior portion of bursal sclerite subtrapezoidal.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	description	Description. Male. Body moderately large (TL 6.2 – 8.3 mm), elongate (TL / EW 1.7 – 1.9), widest in the middle of elytra, moderately convex. Colouration chestnut brown. Head ca. 1.6 × wider than interocular space, convex, densely covered with strong granulate punctation, punctures separated by ca. 0.2 – 0.3 × diameter of a puncture; eyes moderately large, protuberant. Pronotum transverse (PW / PL 1.9 – 2.4), punctation on disc not granulate, sparse, punctures separated by ca. 2.0 × diameter of a puncture, punctation on sides dense, granulate, similar to that on head, lateral carinae of pronotum almost straight, posterolateral angles well marked, anterolateral angles long, subtriangular. Elytra long, 4.0 – 5.0 × longer than pronotum, 1.4 – 1.5 × wider than pronotum; punctation irregular, dense, punctures small, separated by ca. the diameter of a puncture. Aedeagus symmetrical, penis (Fig. 19 A) large (L 1.40 mm, W 0.57 mm), dorsoventrally flattened; trigonium bifid, each process of trigonium curved widened in apical portion, outer margin with ca. 20 denticles and ca. 6 denticles on inner margin of widened apical portion; parameroids subtriangular, with relatively wide apical portion, apices slightly longer than processes of trigonium. Tegmen (Fig. 19 B) large (L 1.40 mm, W 0.97 mm), subrectangular, with small, subtriangular lateral processes; ventral processes wide, each with two subtriangular lobes; parameres digitiform, without setation. Sternite VIII (Fig. 19 C) (L 0.60 mm, W 0.83 mm) widely v-shaped, with sparse setation at apex; sternite IX (Fig. 19 D) (L 0.80 mm, W 0.90 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with setae. Tergite VIII (Fig. 19 E) large (L 1.1 mm, W 1.07 mm), apodemes shorter than apical plate, apical plate subtrapezoidal, apical margin rounded, with a row of sparse setae. Tergite IX (Fig. 19 F) (L 0.80 mm, W 0.73 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Body slightly smaller (TL TL 5.40 – 7.40 mm), setation of dorsum dense, recumbent; head with coarse punctures separated by ca. 0.2 – 0.3 × diameter of a puncture; disc of pronotum more convex than in males; pronotal disc densely covered with small, not granulate punctures, lateral sides of pronotum with coarse, granulate punctures separated by ca. 0.2 × diameter of a puncture; punctures on elytra separated by ca. 1.0 – 1.5 × diameter of a puncture; ovipositor long (L 4.2 mm, Fig. 20 A), without branchlets, coxites subtriangular, elongated, apices of coxites with numerous short setae, styli apical, with two single patches of sensillae, bursal sclerite (L 2.1 mm, W 0.8 mm) elongate, with subtrapezoidal sclerotization in anterior portion (Fig. 20 B). Measurements and ratios. Males (n = 10) TL 6.20 – 8.30 mm (7.2 mm), PL 1.20 – 1.55 mm (1.34 mm), PW 2.50 – 3.00 mm (2.8 mm), EL 5.00 – 6.90 mm (5.9 mm), EW 3.70 – 4.50 mm (4.0 mm), TL / EW 1.7 – 1.9 (1.8), PW / PL 1.9 – 2.4 (2.2), EL / EW 1.3 – 1.6 (1.5), EL / PL 4.0 – 5.0 (4.6), EW / PW 1.4 – 1.5 (1.4). Females (n = 10) TL 5.40 – 7.40 mm (6.6 mm), PL 1.00 – 1.50 mm (1.3 mm), PW 2.10 – 3.10 mm (2.7 mm), EL 4.30 – 5.90 mm (5.4 mm), EW 2.90 – 4.20 mm (3.7 mm), TL / EW 1.7 – 1.9 (1.7), PW / PL 2.0 – 2.3 (2.1), EL / EW 1.3 – 1.5 (1.4), EL / PL 3.7 – 4.6 (3.9), EW / PW 1.3 – 1.4 (1.4).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	distribution	Distribution. Species known from numerous localities scattered in the Chilean regions Los Ríos, Los Lagos, and Aysén (Fig. 24 C).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	biology_ecology	Biology. Most specimens were collected in Valdivian temperate forests which included Nothofagus dombeyi and N. nitida.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E277A62FF38FF77F60DFA38.taxon	etymology	Etymology. After Valdivia, a region where the species occurs.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	description	(Figs 10 H – I, 11 B, 12, 13 F, 21 – 22)	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	materials_examined	Material examined. 1 male, 1 female (ANIC): “ CHILE: Osorno Prov \ Parque Nac. Puyehue \ 4.1 km E Anticura \ 430 m Site 662 19 ”, “ 26 Dec. 1982 Valdi- \ vian RF. Newton & \ Thayer ”, “ Malaise \ trap ”. 1 male (ANIC): “ Chile Chiloe \ Is. 21 km \ N Quellon \ II- 6 - 1968 ”, “ Collectors: L & \ C. W. O’Brien ”. 2 exx. (ANIC): “ CHILE: Llanquihue \ Pr. Lago Chapo \ 13.5 km E Correntoso \ Site 656.310 m 16 - 27 \ Dec. 1982 Valdivian \ RF A. Newton M. Thayer ”; 11 exx. (CMNC), 1 ex. (FMNH): “ CHILE: Osorno: Puyehue \ Nat. Pk; Aquas Calientes \ 18. XII. 84 - 8. II. 85 \ S & J Peck, 600 m, malaise \ Nothofagus forest ”. 1 ex. (CMNC): “ CHILE: Palena Prov. \ 37 km SE Chaiten \ 28. XII. 84 - 30. I. 85 \ S & J Peck, FIT, 60 m \ riverside 2 nd rainforest ”. 6 exx. (CMNC): “ CHILE: 40 km W Angol \ Nahuelbuta Nat. Pk. \ 9. XII. 84 - 17. II. 85 \ S & J Peck, 12 - 1500 m, FITS \ Nothofagus-Araucaria For. ”. 11 exx. (CMNC): “ CHILE: Malleco \ 17 km W Angol, 800 m \ 8. XII. 84 - 16. II. 85 \ S & J Peck, FIT \ mixed Nothofagus ”. 4 exx. (CMNC): “ CHILE: Valdivia \ 34 km WNW La Union \ 17. XII. 84 - 7. II. 85 \ S & J Peck, 700 m, FIT \ mixed evergr. forest ”. 28 exx. (CMNC), 30 exx. (FMNH): “ CHILE: Malleco; Puren \ Contulmo Natur. Mon. \ 11. XII. 84 - 13. II. 85 \ S & J Peck, FIT, 350 m \ mixed evergr. forest ”. 6 exx. (CMNC): “ CHILE: Valdivia \ 26 km SE Panguipulli \ 16. XII. 84 - 11. II. 85 \ S & J Peck, 300 m, cartp \ Nothofagus remnant ”. 1 ex. (CMNC): “ CHILE: Lago Chapo \ 34 km E Pto. Montt \ 24. XII. 84 - 2. II. 85 \ S & J Peck, 300 m, FIT \ 2 nd growth Nothofagus ”. 1 ex. (CMNC): “ CHILE: 11 km W Angol \ 9. XII. 84 - 16. II. 85 \ S & J Peck, 1000 m \ carrion trap, boggy \ mixed forest remnant ”. 5 exx. (CMNC): “ CHILE: Palena Prov. \ 37 km SE Chaiten \ 28. XII. 84 - 30. I. 85 \ S & J Peck, FIT, 60 m \ riverside 2 nd forest ”. 3 exx. (CMNC): “ CHILE: Cautin; 21 km NE \ Pucon, Lago Caburga \ 15. XII. 84 - 10. II. 85 \ S & J Peck, FIT, 600 m \ mixed forest remnant ”. 5 exx. (CMNC): “ CHILE: 45 km W Angol \ Nahuelbuta Nat. Pk. \ 9. XII. 84 - 16. II. 85 \ S & J Peck, 1400 m, car. tp. \ Nothofagus-Araucaria For. ”. 4 exx. (CMNH): “ Villarrica \ Cautin \ II-III- 1979 \ Coll. L. E. Pena ”; “ CHILE ”; “ CMNH Acc. \ No. 31811 ”. 1 ex. (CMNH): “ Nahuelbuta \ Malleco \ II-III- 1979 \ Coll. L. E. Pena ”; „ CHILE ”; “ CMNH Acc. \ No. 31811 ”. 2 exx. (DBET): “ Chile, N. P. Puyehue, Anticura Sector \ Nothofahus forest, \ in litter. 10.01.2014 \ K. Eskov, R. Rakitov ”. 10 exx. (DBET): “ Chile, Chiloe Is., Senda \ Darwin Biol. Station. \ Boggy Baccharis mator- \ ral, in Sphagnum, 27.12. \ 2013 - 4.01.14, K. Eskov ”. 28 exx. (DBET): “ CHILE, Reg. X Chiloe \ Is. SE, 20 km NNW of Quellon, 216 m, 42.58, \ 73.45, 31 I 05, leg. Snižek ”. 1 ex. (DBET): “ CHILE w, Reg. X \ SW of Los Muermos \ 50 m 41.30, 73.35 \ 28. I. 2005, Snižek lgt. ”. 6 exx. (DBET): “ CHILE CW, reg. X \ W of LA UNION \ E of EL MIRADOR \ 27.1.2004 \ LGT. M. SNIŽEK ”. 1 ex. (DBET): “ CHILE W., Reg. X \ Purrangue 600 m \ W of Hueyusca \ 26. I. 2005 M. Snižek ”. 3 exx. (DBET): “ CHILE W, reg. X \ SW of PUERTO MONTT \ W of PARGUA \ 29.1.2004 \ LGT. M. SNIŽEK ”. 1 ex. (DBET): “ CHILE, reg. X \ S of CARIRRINGUE \ PANGUIPULLI \ 20.1.2004, LGT. M. SNIŽEK ”. 4 exx. (DBET): “ CHILE W, Reg. X Chiloe C, NW Castro \ San Pedro env. \ 2 II 2005, leg. Snižek ”. 3 exx. (DBET): “ CHILE CW, reg. X \ NW of LA UNION \ St. ELISA env. \ 24. I. 2004 \ LGT. M. SNIŽEK ”. 1 ex. (DBET): “ CHILE E, reg. X \ E of PUERTO MONTT \ SE of LAGO CHAPO \ 30. I. 2004, LGT. M. SNIŽEK ”. 1 ex. (DBET): “ CHILE CW, reg. IX \ SE of NUEVA TOLTÉN \ BOROA CENTRAL \ 22. I. 2004 \ LGT. SNIŽEK ”. 7 exx. (EMEC): “ 37 ° 48 ’ 12 ’’ S / 73 ° 01 ’ 28.5 ’’ W \ CHILE, IX Region, vic. Nahuelbuta \ pitfall traps 7 - 25: i: 2003 \ Colls. K. Will, S. Lew ”; “ Cyphon \ sp. \ det. J. E. Barriga ”; “ U. C. Berkeley \ EMEC 107088 ” [each specimen with individual number]. 2 exx. (EMEC): “ 60 - CHILE X REGION \ Oncol Park, Quitaqui Way Fog- \ ging 15: 32 PM, 10. I. 2007, 519 m \ 13 ° C, S 39 ° 42.071 / W 73 ° 18.547 \ Saxegothaea conspicua 45 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 57 - CHILE X REGION \ Punta Curiñanco, Fogging \ 14: 41 PM, 09. I. 2007, 177 m \ 16 ° C, S 39 ° 42.791 / W 73 ° 24.328 \ Aextoxicon punctatum 45 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 58. - CHILE X REGION \ Punta Curiñanco, Fogging \ 16: 33 PM, 09. I. 2007, 115 m \ 19 ° C S 39 ° 42.942 / W 73 ° 24.375 \ Aextoxicon punctatum 20 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 72 - CHILE X REGION \ La Estancilla, Fogging 14: 02 PM, \ 15. I. 2007, 62 m 25 ° C, S 39 ° 50. \ 397 / W 73 ° 18.949 Nothofagus dom - \ beyi, 45 m, Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 1 ex. (EMEC): “ 46. - CHILE IX REGION \ Nahuelbuta NP exit, Pichinahuel \ Fogging 13: 53 PM, 05. I. 2007 \ 1238 m 22 ° C, S 37 ° 49.041 / W \ 73 ° 00.214, Araucaria araucana \ 15 m \ Arias et al UC Berkeley ”; “ Scirtidae \ Genero sp. 1 \ det. G. Arriagada 2007 ”. 2 males (FMNH): “ Pichinahuel \ Arauco Chile \ 3 - 11 - I- 1977 \ Coll. L. E. Pena ”, “ Collection \ Pena-Vidal ”. 1 male (FMNH): “ CHILE: Chiloé Pr.: Mira- \ flores, road to (0.6 km W \ Hwy 5), 130 m, 42 ° \ 46.73 ’ S, 73 ° 47.71 ’ W, \ 17. i. 1997, secondary \ Valdivian rainforest; ”, “ beating flowering Cald- \ cluvia paniculata (Cun- \ oniaceae), A. Newton & \ M. Thayer 994 \ FIELD MUS. NAT. HIST. ”. 4 males (FMHN): “ Chile ”, “ specimen orig. without \ labels in colln. but \ asociated with specimens \ bearing attached data ”, CNHM \ Karl Brancsik colln. \ ex Eduard Knirsch ”. 3 exx. (MHNG): “ CHILE: X Reg. Prov. Osorno \ PN Puyehue, Aguas \ Calientes, 400 - 500 m \ 31. XII. 1990 - 1. I. 1991 \ Agosti & Burckhardt # 25 b ”. 1 ex. (MHNG): “ CHILE: X Reg. Prov. Osorno \ PN Puyehue, Aguas \ Calientes, 400 - 500 m \ 31. XII. 1990 - 1. I. 1991 \ Agosti & Burckhardt # 25 a ”. 5 exx. (MHNG): “ CHILE: IX Reg. Prov. Cautin \ PN Huerquehue, 800 m \ 22 - 25. XII. 1990, Valdivian \ lauriphyllous forest \ Agosti & Burckhardt # 17 b ”. 1 male (MNHN): “ 1361 ” [handwritten, without other labels]. 1 male (MNHN): “ 1363 ” [handwritten], “ Elodes \? Rousselii Sol ” [handwritten]. 1 ex. (NHM): “ 30 CHILE Chiloé \ Cucao: PN Chiloé \ Stream in tepú forest \ I. Ribera leg. 4. II. 1999 ”; “ BM 1999 - 195 \ 19. i. - 9. ii. 1999 ”. 12 exx. (NMPC): “ CHILE, IX. r. 1. - 3.2. \ 2005, Manzanar \ 38 ° 28.071 - 71 ° 41.873 \ 800 m, Sv. Bilý leg. ”. 6 exx. (NMPC): “ CHILE, IX. r., 9. - 11.2. \ 2005, P. N. Nahuelbuta \ 37 ° 48.530 - 73 ° 00.954 \ 1200 m, Sv. Bilý leg. ”. 10 exx. (NMPC): “ CHILE, X. r., 6.2.2005 \ 45 km W of La Unión \ 40 ° 12.784 - 73 ° 23.462 \ 900 m, Sv. Bilý leg. ”. 3 exx. (NMPC): “ CHILE, IX. r., 25. - 31.1. \ 2005, P. N. Nahuelbuta \ 37 ° 48.530 - 73 ° 00.954 \ 1200 m, Sv. Bilý leg. ”. 2 exx. (NMPC): “ CHILE, IX. r., 12. - 14.2. \ 2005, Trm. de Tolhuaca \ 38 ° 14.162 - 71 ° 44.027 \ 900 m, Sv. Bilý leg. ”. 2 exx. (SDEI): “ Chile, coll. Kraatz ”, “ Pic det., Elodes var. velutina Sol. ”	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	diagnosis	Diagnosis. Posterolateral angles of pronotum broadly rounded; elytra covered with dense but small punctures; processes of trigonium shorter than pala, curved, with ca. 30 denticles on its margin; anterior portion of bursal sclerite with two crescent-like emarginations.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	description	Description. Male. Body moderately large (TL 5.4 – 6.9 mm), elongate (TL / EW 1.8 – 1.9), widest in the middle of elytra, moderately convex. Colouration chestnut brown. Head ca. 1.4 × wider than interocular space, convex, covered with dense, nearly confluent, granulate punctures, separated by ca. 0.1 × diameter of a puncture; eyes moderately large, protuberant. Pronotum transverse (PW / PL 2.1 – 2.2), punctation on disc not granulate, sparse, punctures separated by ca. the diameter of a puncture, punctation on sides granulate, very dense, similar to that on head, lateral carinae of pronotum rounded, posterolateral angles widely rounded, anterolateral angles subtriangular. Elytra long, 4.3 – 4.8 × longer than pronotum, 1.3 – 1.5 × wider than pronotum; punctation irregular, dense, separated by ca. 0.5 – 1.0 × diameter of a puncture. Aedeagus symmetrical, penis (Fig. 21 B) large (L 1.27 mm, W 0.33 mm), dorsoventrally flattened; trigonium bifid, each process of trigonium curved, with ca. 30 denticles; parameroids subtriangular, digitiform in apical portion, slightly longer than processes of trigonium. Tegmen (Fig. 21 C) large (L 1.2 mm, W 0.77 mm), subrectangular, with small, subtriangular lateral processes; ventral processes wide, each with two lobes (large suboval and small subtriangular one); parameres relatively wide, digitiform, densely covered with short setae. Sternite VIII (Fig. 21 D) (L 0.67 mm, W 0.43 mm) widely v-shaped, with sparse setation at apex; sternite IX (Fig. 21 E) (L 0.70 mm, W 0.50 mm) oval, consisting of two subtriangular hemisternites, apical portion covered with setae. Tergite VIII (Fig. 21 F) large (L 0.87 mm, W 0.77 mm), apodemes shorter than apical plate, apical plate subtrapezoidal, apical margin rounded, with a row of sparse setae. Tergite IX (Fig. 21 G) (L 0.63 mm, W 0.47 mm) with short apodemes, apical portion membranous, setation indistinct. Female. Body of similar size (TL 5.60 – 6.30 mm), setation of dorsum dense, recumbent; head with coarse, confluent punctures; disc of pronotum more convex than in males; pronotal disc densely covered with small, not granulate punctures, separated by ca. the diameter of a puncture, lateral sides of pronotum with coarse, confluent, granulate punctures, similar to those on head; punctures on elytra small, separated by ca. 1.0 – 2.0 × diameter of a puncture; adscutellar area of elytra slightly depressed; ovipositor long (L 3.7 mm, Fig. 22 A), without branchlets, coxites subtriangular, elongated, apices of coxites with numerous short setae, styli apical, with two single patches of sensillae, bursal sclerite (L 1.7 mm, W 0.5 mm) elongate, anterior portion with two crescent-like emarginations (Fig. 22 C). Measurements and ratios. Males (n = 10) TL 5.40 – 6.90 mm (6.10 mm), PL 1.00 – 1.30 mm (1.1 mm), PW 2.10 – 2.80 mm (2.4 mm), EL 4.60 – 5.80 mm (5.0 mm), EW 3.00 – 3.90 mm (3.3 mm), TL / EW 1.8 – 1.9 (1.8), PW / PL 2.1 – 2.2 (2.2), EL / EW 1.4 – 1.6 (1.5), EL / PL 4.3 – 4.8 (4.5), EW / PW 1.3 – 1.5 (1.4). Females (n = 10) TL 5.60 – 6.30 mm (6.0 mm), PL 1.00 – 1.25 mm (1.1 mm), PW 2.40 – 2.70 mm (2.5 mm), EL 4.60 – 5.20 mm (4.9 mm), EW 3.00 – 3.50 mm (3.3 mm), TL / EW 1.7 – 1.9 (1.8), PW / PL 2.1 – 2.5 (2.3), EL / EW 1.4 – 1.6 (1.6), EL / PL 4.0 – 4.8 (4.7), EW / PW 1.3 – 1.4 (1.3).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	distribution	Distribution. A very common species, recorded from the following regions of Chile: Bío Bío, Araucanía, Los Ríos, and Los Lagos (Fig. 24 D).	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
03BD95561E297A67FF38F94BF013FE76.taxon	biology_ecology	Biology. The species occurs in Valdivian temperate forests with Aextoxicon punctatum, Araucaria araucana, Nothofagus dombeyi, and Saxegothaea conspicua. One of the specimens was collected on a flowering Caldcluvia paniculata (Cav.) D. Don. A description of a plausible larva of M. velutinus was published by Ruta et al. (2018). In the original description, Solier (1849) noted that M. velutinus was collected together with Solierodes rousseli.	en	Ruta, Rafał (2021): Three new genera of large marsh beetles (Coleoptera: Scirtidae) from Valdivian temperate rain forests of southern South America. Zootaxa 5048 (4): 451-485, DOI: https://doi.org/10.11646/zootaxa.5048.4.1
