identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B487A07F12FFC6FF30108A6AB3A20C.text	03B487A07F12FFC6FF30108A6AB3A20C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus Sharp 1882	<div><p>Chasmogenus Sharp, 1882</p> <p>Chasmogenus Sharp, 1882: 73.</p> </div>	http://treatment.plazi.org/id/03B487A07F12FFC6FF30108A6AB3A20C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
03B487A07F12FFC2FF301304698DA569.text	03B487A07F12FFC2FF301304698DA569.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus clinatus Glynn & Short 2021	<div><p>Chasmogenus clinatus sp. n.</p> <p>Figures 2A–B, 3A, 4A–D, 6</p> <p>Type Material. Holotype (male): “BRAZIL: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Amazonas</a>: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Manaus</a> / -2.93079, -59.97514, 75m / <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Ducke Reserve</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Igarape Barro</a> / Branco; Short &amp; team; forest/ pools; 6.vi.2018; BR18-0606-02C”, “[barcode]/ SEMC1623338 / KUNHM-ENT” (INPA). Paratypes (34): BRAZIL: Amazonas: Same data as holotype (8 exs., INPA, SEMC, including DNA Voucher is SLE1848); same data but “forest pools”, BR18-0606-02A (4 exs., SEMC); same data but stream margin and associated backwater swampy area, 9–10.vi.2018, BR18-0609-3A (2 exs., SEMC). VENEZUELA: Amazonas: Cerro de la Neblina, 1 km S. Basecamp, 17.ii.1985, leg. Faitoute &amp; Steiner, “along small whitewater stream, pools of dead leaves and sticks” (20 exs., USNM, MIZA, SEMC).</p> <p>Differential diagnosis. Chasmogenus clinatus is very similar to a cluster of species that have sinuate parameres and a well-developed accessory sclerite on the median lobe (Fig. 4A–D). The aedeagus of this species is most similar to that of C. pandus (Fig. 4F) in that both have a bent medial sclerite that lies dorsally to the median lobe and the outer margins of the parameres have a weak undulating appearance. However, the median lobe of C. clinatus is uniformly wide throughout its length until the apex acuminates past the parameres, whereas the median lobe of C. pandus is wider at the base and gradually acuminates throughout the entire length towards the apex. The outer margins of the parameres of C. clinatus have a single medial constriction (sinuate) between the base and the position of the gonopore and C. pandus has two lateral constrictions (bisinuate), one medial and one closer to the apex of the parameres. The two species may be differentiated by dorsal coloration as well: C. clinatus is dark brown, whereas C. pandus is a slightly paler reddish brown.</p> <p>Description. Size and color. Total body length 2.9–3.6 mm. Body form elongate oval with slightly curved lateral margins. Dorsum of head orange-brown to very dark brown, clypeus and labrum paler brown (Fig. 3A). Pronotum and elytra uniformly dark orange-brown (Fig. 2A). Venter uniformly dark orange-brown (Fig. 2B). Head. Ground punctation on head fine. Clypeus with anteromedial emargination, which exposes a narrow, rounded to angulate gap between the clypeus and labrum (Fig. 3A). Mentum deeply depressed in anterior two-thirds with a triangular to rounded anteromedial notch. Maxillary palps long, longer than width of head immediately posterior to eyes. Thorax. Ground punctation on pronotum fine (Fig. 2A). Prosternum very weakly to not at all tectiform. Mesoventrite with moderate elevation forming a thin posteromedial longitudinal carina. Metafemora densely pubescent in basal nine-tenths (Fig. 2B). Aedeagus. Aedeagus (Fig. 4A–D) with median lobe nearly parallel-sided and uniformly wide, then tapering gradually to a weakly acuminate apex, distinctly extending beyond the apex of the parameres. Sclerite of the median lobe expanded and developed into a long, narrow sliver, angled slightly to the left with a sharply acute apex that extends to or just beyond the apex of the parameres. Gonopore situated in the middle of the median lobe, ca. two to three gonopore widths below the apex. Parameres symmetrical, outer margins with a single medial constriction, apical half slightly narrower than basal half; apex slightly acuminate. Basal piece short, ca. one-third the length of the parameres.</p> <p>Etymology. The species name is derived from the Latin clinatus meaning “bent” or “sloped” after the condition of the medial sclerite on the median lobe of the aedeagus.</p> <p>Distribution. This species is known from the Ducke Reserve in Manaus, state of Amazonas, Brazil and from the extreme south of Venezuela at the foot of Cerro de la Neblina.</p> <p>Biology. This species was collected from a series of forest pools and riparian areas in northern Brazil (Amazonas) and along whitewater streams in southern Venezuela.</p></div> 	http://treatment.plazi.org/id/03B487A07F12FFC2FF301304698DA569	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
03B487A07F16FFC1FF30149F6E65A6A5.text	03B487A07F16FFC1FF30149F6E65A6A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus gironae Glynn & Short 2021	<div><p>Chasmogenus gironae sp. n.</p> <p>Figures 2C–D, 3B, 5C–D, 6</p> <p>Type Material. Holotype (male): “VENEZUELA, T.F.Amaz. / <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.166664&amp;materialsCitation.latitude=0.8333333" title="Search Plazi for locations around (long -66.166664/lat 0.8333333)">Cerro de la Neblina</a> / 1.5 km S Basecamp / 0°50’N, 66°10’W / 250 m., 15 Feb. 1985 ”, “In small ponds full/ of dead leaves;/ rainforest ridge/ P. J. &amp; P. M. Spangler,/ R. Faitoute, W. Steiner ” (MIZA). Paratypes (36): VENEZUELA: Amazonas: Same data as holotype (36 exs., SEMC, USNM).</p> <p>Differential Diagnosis. Among species with the median lobe of the aedeagus and the parameres approximately even in length and a long basal piece, this species is similar in morphology and general form of the aedeagus to C. brownsbergensis Smith &amp; Short, 2020, but may be distinguished by characters of the median lobe and dorsal coloration of the head. The median lobe of C. gironae bears a rounded triangular apex past the position of the gonopore and a constriction located medially, whereas the apex of C. brownsbergensis is acutely triangular with straight sides and the constriction is situated more distally closer to the gonopore. The coloration of the head of C. brownsbergensis has mesal dark patches on the clypeus and the labrum compared to the uniformly colored head of C. gironae.</p> <p>Description. Size and color. Total body length 3.4–3.7 mm. Body form elongate oval. Dorsum of head dark brown to very dark brown posteriorly, clypeus and labrum slightly paler (Fig. 3B). Pronotum and elytra uniformly very dark brown (Fig. 2C). Venter uniformly dark orange-brown to dark brown (Fig. 2D). Head. Ground punctation on head fine. Clypeus with anteromedial emargination, which exposes a broadly rounded gap between the clypeus and labrum (Fig. 3B). Mentum moderately depressed in anterior half with a triangular to rounded anteromedial notch. Maxillary palps long, longer than width of head immediately posterior to eyes. Thorax. Ground punctation on pronotum fine (Fig. 2C). Prosternum very weakly to not at all tectiform. Mesoventrite with moderate elevation forming a thin posteromedial longitudinal carina. Metafemora densely pubescent in basal nine-tenths (Fig. 2D).</p> <p>Aedeagus. Aedeagus (Fig. 5C,D) with median lobe that bears a single medial constriction situated halfway between base and apex, base and position of gonopore of equal width, tapering past position of gonopore narrowing abruptly to form a rounded triangular apex which is even with the apex of the parameres. Sclerite of the median lobe not expanded. Gonopore situated ca. half of one gonopore width below the apex of the median lobe. Parameres symmetrical, with outer margins straight, with apex slightly inwardly curved and bluntly rounded. Basal piece long, ca. half the length of the parameres.</p> <p>Etymology. This species is named in honor of Dr. Jennifer Girón, for her substantial contributions and dedication to the study of the taxonomy and morphology of the subfamily Acidocerinae and for her valuable mentorship of the first author.</p> <p>Distribution. Known only from the type locality near the base of Cerro de la Neblina in southern Venezuela.</p> <p>Biology. Chasmogenus gironae was collected in small ponds full of leaves in lowland rainforest.</p> </div>	http://treatment.plazi.org/id/03B487A07F16FFC1FF30149F6E65A6A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
03B487A07F15FFC0FF3017D36F75A7A7.text	03B487A07F15FFC0FF3017D36F75A7A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus inpa Glynn & Short 2021	<div><p>Chasmogenus inpa sp. n.</p> <p>Figures 3C, 5A–B, 6</p> <p>Type Material. Holotype (male): “BRAZIL: Amazonas: Manaus / -2.93079, -59.97514, 75 m / <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Ducke Reserve</a>, trail to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Igarape</a> / <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Acara</a>; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.97514&amp;materialsCitation.latitude=-2.93079" title="Search Plazi for locations around (long -59.97514/lat -2.93079)">Short</a> &amp; team; muddy/ area with emergent vegetation/ 7–8.vi.2018; BR18-0607-01A”, “[barcode]/ SEMC1623365 / KUNHM-ENT” (INPA). Paratypes (7): BRAZIL: Amazonas: Same data as holotype (4 exs., INPA, SEMC); same locality but Flight Intercept <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.9384&amp;materialsCitation.latitude=-2.68396" title="Search Plazi for locations around (long -60.9384/lat -2.68396)">Trap</a> (FIT) (2 exs., SEMC, DNA vouchers SLE1892 and SLE1894); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.9384&amp;materialsCitation.latitude=-2.68396" title="Search Plazi for locations around (long -60.9384/lat -2.68396)">Novo Airão Municipio</a>, -2.68396, -60.93840, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.9384&amp;materialsCitation.latitude=-2.68396" title="Search Plazi for locations around (long -60.9384/lat -2.68396)">Densely</a> vegetated margin of blackwater creek, BR17- 0609-04A (1 ex., SEMC, DNA voucher SLE1267)</p> <p>Differential diagnosis. This species is similar to a group that have an aedeagal form with relatively broad, straight parameres and a wide median lobe which is slightly shorter to slightly longer than the apex of the parameres. It is distinctly smaller in overall size than C. amplius Smith &amp; Short, 2020, C. berbicensis Smith &amp; Short, 2020, and C. clavijoi Smith &amp; Short, 2020. The apex of the parameres curve slightly inward like that of C. schmits Smith &amp; Short, 2020, but are not bluntly squared and the apex of the median lobe distinctly reaches the apex of the parameres or just slightly beyond. It is most similar to the comparably-sized C. guianensis Smith &amp; Short, 2020, but the median lobe is distinctly longer and even with the apex of the parameres.</p> <p>Description. Size and color. Total body length 3.7–3.9 mm. Body form elongate oval with slightly curved lateral margins. Dorsum of head orange brown to dark brown, clypeus and labrum distinctly paler (Fig. 3C). Pronotum and elytra uniformly orange-brown. Venter uniformly orange-brown to dark orange brown. Head. Ground punctation on head fine. Clypeus with anteromedial emargination, which exposes a broadly rounded gap between the clypeus and labrum (Fig. 3C). Mentum weakly depressed in anterior third with a triangular to rounded anteromedial notch. Maxillary palps long, longer than width of head immediately posterior to eyes. Thorax. Ground punctation on pronotum fine. Prosternum weakly tectiform. Mesoventrite with moderate elevation forming a thin posteromedial longitudinal carina. Metafemora densely pubescent in basal nine-tenths. Aedeagus. Aedeagus (Fig. 5A, B) with median lobe widest at base, and slightly tapering gradually until the apical quarter, then narrowing abruptly to form an acute triangular apex which is even with or extends slightly past the apex of the parameres. Sclerite of the median lobe not expanded. Gonopore situated less than half of one gonopore width below the apex of the median lobe. Parameres symmetrical, with outer margins slightly curved inwardly and apex slightly inwardly curved and bluntly rounded. Basal piece of medium to long length, ca. one half to two-thirds the length of the parameres.</p> <p>Etymology. Named after the Instituto Nacional de Pesquisas da Amazônia, an organization dedicated to advancing our understanding of the Amazon, and who also administers the Adolfo Ducke Forest Reserve from where this species was collected.</p> <p>Distribution. This species is known only from the Ducke Reserve in Amazonas State, northern Brazil.</p> <p>Biology. Collected in muddy vegetated areas along a trail to Igarapé Acará.</p></div> 	http://treatment.plazi.org/id/03B487A07F15FFC0FF3017D36F75A7A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
03B487A07F14FFCFFF3016D56EFBA669.text	03B487A07F14FFCFFF3016D56EFBA669.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus Sharp 1882	<div><p>Key to the species of Chasmogenus Sharp of Venezuela, Suriname, Guyana, French Guiana, and Brazil north of the Amazon River</p> <p>We have modified the key presented in Smith &amp; Short (2020) to insert the three species described here. All three species would key to couplet 13 in the original key, so we have not repeated those here. We emphasize that examination of the genitalia is crucial to accurate identifications and unassociated females of the three species described here cannot be identified with precision.</p> <p>13 Aedeagus with apex of median lobe long, distinctly extending beyond the apex of the parameres; basal piece short, one-third the total length of the parameres or less (e.g. Smith &amp; Short 2020: fig. 15A–E, 15G–I)............................. 14</p> <p>- Aedeagus with apex of median lobe shorter or not extending beyond the apex of the parameres; basal piece long, up to one-half the length of the entire aedeagus........................................................................ 17</p> <p>14 Parameres of aedeagus smooth along outer edges, not sinuate or undulate, apex of median lobe acuminates at the apex (Smith &amp; Short 2020: fig. 15H, I); gonopore situated toward the apex of the median lobe............ C. acuminatus Smith &amp; Short</p> <p>- Parameres moderately sinuate along outer margins; gonopore situated about in the middle of the median lobe............15</p> <p>15 Median lobe of aedeagus terminating in a broadly rounded apex past the apex of the parameres (e.g. Smith &amp; Short 2020: fig. 15E);.......................................................................... C. ligulatus Smith &amp; Short</p> <p>- Median lobe narrowing to a moderately to acutely pointed apex past the apex of the parameres...................... 16a</p> <p>16a Median lobe with apex acuminate (e.g. Smith &amp; Short 2020: fig. 15A–D); emargination of the clypeus narrow (e.g. Smith &amp; Short 2020: figs 5D–F, 10D).......................................................................... 16b</p> <p>- Median lobe with apex rounded (Smith &amp; Short 2020: fig. 15G); dorsal coloration dark brown, emargination of the clypeus rounded to angulate............................................................. C. undulatus Smith &amp; Short</p> <p>16b Outer margins of parameres with two constrictions (bisinuate) (e.g. Smith and Short 2020: fig. 15A–D); dorsal coloration dark reddish brown and translucent (e.g. Smith and Short 2020: figs. 5D–F, 10D)................... C. pandus Smith &amp; Short</p> <p>- Outer margins of parameres with a single constriction (sinuate) (fig. 4A–D); dorsal coloration uniformly dark brown (fig. 2A, B).................................................................................... C. clinatus sp. n.</p> <p>17 Median lobe of aedeagus shorter than length of parameres, aedeagus symmetrical................................. 18</p> <p>- Apex of median lobe of aedeagus approximately even with apex of parameres.................................... 19</p> <p>18 Apex of parameres bluntly squared (Smith &amp; Short 2020: fig. 14F).......................... C. schmits Smith &amp; Short</p> <p>- Apex of parameres rounded (Smith &amp; Short 2020: fig. 14G, H).......................... C. guianensis Smith &amp; Short</p> <p>19 Distinct dark patches on mesal portion of clypeus and labrum (Smith &amp; Short 2020: fig.10A).................................................................................................. C. brownsbergensis Smith &amp; Short</p> <p>- Clypeus and labrum without distinct dark patches........................................................... 20</p> <p>20 Aedeagus with parameres straight along outer margins, only curving inward past the position of the gonopore........... 21</p> <p>- Aedeagus with parameres that curve slightly inward along the outer margins throughout the entire length (figs. 5A, B)............................................................................................... C. inpa sp. n.</p> <p>21 Apex of median lobe rounded triangular in shape (figs. 5C, D).................................... C. gironae sp. n.</p> <p>- Apex of median lobe acutely triangular in shape............................................................ 22</p> <p>22 Total body size ≤ 3.8 mm, dorsal coloration very dark brown................................... C. gato Smith &amp; Short</p> <p>- Total body length ≥ 3.8 mm, dorsal coloration pale orange-brown..................... C. clavijoi Smith &amp; Short (in part)</p> </div>	http://treatment.plazi.org/id/03B487A07F14FFCFFF3016D56EFBA669	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
03B487A07F1BFFCFFF3016296996A7C2.text	03B487A07F1BFFCFFF3016296996A7C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chasmogenus acuminatus Smith & Short 2020	<div><p>Chasmogenus acuminatus Smith &amp; Short, 2020</p> <p>New material examined (21): BRAZIL: Amapá: Oiapoque, (c. 22 km S on BR-156), 3.65822°, -51.76958°, 17 m, 20.vii.2018, leg. Short, forested detrital pool, BR18-0720-01B (9 exs., INPA, SEMC). Pará: Alenquer, Vale do Paraíso, ca. 55 km N of Alenquer, -1.49292, -54.51566, 150 m, 3.ii.2018, leg. A. Short, isolated detrital pool in rocks next to river, BR18-0203-01B (9 exs., INPA, SEMC).</p></div> 	http://treatment.plazi.org/id/03B487A07F1BFFCFFF3016296996A7C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glynn, Rachel D.;Short, Andrew Edward Z.	Glynn, Rachel D., Short, Andrew Edward Z. (2021): New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048 (3): 435-443, DOI: https://doi.org/10.11646/zootaxa.5048.3.8
