taxonID	type	description	language	source
132B8783FFF6FFCFFCC8526E867DFA25.taxon	description	(Figs 1, 2)	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF6FFCFFCC8526E867DFA25.taxon	diagnosis	Diagnosis. Body size: 265 – 282 × 166 – 182. Rostrum pointed. Prolamella short, represented only by basal part. Translamella present, interrupted medially. Rostral, lamellar and interlamellar setae long, setiform, barbed; ro shortest. Bothridial seta long, clavate, barbed. Notogaster with one pair of short, setiform, smooth setae (p 1) and nine pairs of alveoli. Epimeral and anogenital setae short, setiform, roughened. Circumpedal carina of medium length. Leg tarsus I with 19 setae (l ” absent); genu II with two setae (v’ absent).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF6FFCFFCC8526E867DFA25.taxon	description	Description of adult. Measurements. Body length: 273 (holotype: female), 265 – 282 (three paratypes: all females); body width: 173 (holotype), 166 – 182 (three paratypes). Integument. Body color light brown. Cuticle microporose (visible under high magnification, × 1000). Lateral side of body with microgranulate cerotegument between bothridium and acetabula I – III. Prodorsum (Figs 1 A, C). Rostrum pointed. Lamella about 1 / 2 length of prodorsum. Prolamella short, represented only by basal part. Sublamella linear, similar to lamella in length. Sublamellar porose area (4 – 6) rounded. Translamella present, shortly interrupted medially. Lateral keel-shaped ridge distinct. Rostral (41 – 45), lamellar (57 – 61) and interlamellar (61 – 65) setae setiform, barbed. Exobothridial seta not observed. Bothridial seta (41 – 45) clavate, barbed. Dorsosejugal porose area not observed. Notogaster (Figs 1 A, C). One pair of setiform, smooth notogastral setae (p 1: 10 – 12) developed; other setae represented by alveoli. Four pairs of small sacculi. Opisthonotal gland opening and all lyrifissures distinct. Gnathosoma (Figs 2 A – D). Subcapitulum longer than wide (69 – 77 × 53 – 57). All subcapitular (a: 14 – 16; m: 10 – 12; h: 18 – 20) and adoral (10 – 12) setae setiform, barbed; m thinner than a and h. Palp (45 – 49) with typical setation: 0 - 2 - 1 - 3 - 9 (+ ω). Postpalpal seta (6) spiniform, roughened. Chelicera (77 – 82) with two setiform, barbed setae (cha: 24 – 26; chb: 16). Epimeral and lateral podosomal regions (Figs 1 B, C). Epimeral setal formula: 3 - 1 - 3 - 3. All setae (1 b, 3 b, 3 c, 4 c: 18 – 20; others: 10 – 12) setiform, roughened. Pedotectum II with small posterolateral tooth. Discidium triangular, rounded apically. Circumpedal carina of medium length, directed to pedotectum II but not reaching it. Anogenital region (Figs 1 B, C). Genital (g 1: 12; others: 8 – 10), aggenital (12), anal (8 – 10), and adanal (12) setae setiform, roughened. Adanal lyrifissure distinct. Marginal porose area band-like, complete. Legs (Figs 2 E – H). Bidactylous; all claws slightly barbed on dorsal side; paraxial claw (on tarsi I, II) and antiaxial claw (on tarsi III, IV) thinner than the accompanying claw, with small tooth distoventrally. Tibiae I and II with tubercle proximoventrally. Femur II with broadly rounded ledge distoventrally. Dorsoparaxial porose area on femora I-IV and on trochanters III, IV well visible; proximoventral porose area on tarsi I-IV and distoventral porose area on tibiae I-IV not observed. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 19) [1 - 2 - 2], II (1 - 5 - 2 - 4 - 15) [1 - 1 - 2], III (2 - 3 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 – 1 – 0]; homology of setae and solenidia indicated in Table 1. Famulus of tarsus I short, erect, slightly swollen distally, inserted between solenidion ω 2 and seta ft ”. Solenidion ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III bacilliform, other solenidia setiform. Seta s on tarsus I setiform, barbed (not eupathidial), inserted behind setae a’ and a ”. Note: Roman letters refer to normal setae, Greek letters to solenidia (except ɛ = famulus), Single quote (’) marks seta on anterior and double quote () seta on posterior side of the given leg segment. Parentheses refer to a pair of setae. * – Seta v’ on genu I present and l ” on femur II absent in P. elisabethae (versus v’ on genu I absent and l ” on femur II present S. (B.) eduardoi and S. (H.) dianae).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF6FFCFFCC8526E867DFA25.taxon	materials_examined	Type deposition. The holotype is deposited in the collection of the MUSM; three paratypes are deposited in the collection of the TSUMZ. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF6FFCFFCC8526E867DFA25.taxon	etymology	Etymology. The specific name eduardoi is given in honour of our friend and colleague Dr. Eduardo Ruiz (Complutense University, Madrid, Spain).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF6FFCFFCC8526E867DFA25.taxon	discussion	Remarks. Distinctive characters of the new species compared with other recent members of the genus can be found in the identification key below.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	description	(Figs 3, 4)	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	diagnosis	Diagnosis. Body size: 365 – 498 × 215 – 282. Rostrum rounded. Prolamella complete. Translamella absent. Rostral, lamellar and interlamellar setae long, setiform, barbed; ro shortest. Bothridial seta long, narrowly spindleform, with setiform apical tip, unilaterally ciliate. Anterolateral part of notogaster with triangular process. Notogastral setae setiform, smooth; c, la, lm, and lp long, other setae of medium length. Epimeral and anogenital setae short, setiform, barbed. Circumpedal carina short. Leg tarsus I with 19 setae (l ” absent); genu II with two setae (v’ absent).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	description	Description of adult. Measurements. Body length: 464 (holotype: female), 365 – 498 (17 paratypes: eight males and nine females); body width: 265 (holotype), 215 – 282 (17 paratypes). No differences between males and females in body size. Integument. Body color brown. Cuticle microporose (visible under high magnification, × 1000). Lateral side of body with microgranulate cerotegument between bothridium and acetabula I – IV. Prodorsum (Figs 3 A, C). Rostrum rounded. Lamella about 1 / 2 length of prodorsum. Prolamella complete. Sublamella linear, similar to lamella in length. Sublamellar porose area (4 – 6) rounded. Translamella absent. Lateral keel-shaped ridge distinct. Rostral (69 – 73), lamellar (102 – 106) and interlamellar (102 – 106) setae setiform, barbed. Exobothridial seta (32 – 36) setiform, slightly barbed. Bothridial seta (106 – 110) narrowly spindleform, with setiform apical tip, unilaterally shortly ciliate. Dorsosejugal porose area not observed. Notogaster (Figs 3 A, C). Anterolateral part of notogaster with triangular process. All notogastral setae (c, la, lm, lp: 61 – 73; others: 32 – 36) setiform, smooth. Four pairs of small sacculi. Opisthonotal gland opening and all lyrifissures distinct. Gnathosoma (Figs 4 A – D). Subcapitulum longer than wide (102 – 106 × 73 – 86). All subcapitular (a: 20; m: 14 – 16; h: 24) and adoral (10 – 12) setae setiform, barbed; m thinner than a and h. Palp (65 – 69) with typical setation: 0 - 2 - 1 - 3 - 9 (+ ω). Postpalpal seta (6) spiniform, roughened. Chelicera (106 – 110) with two setiform, barbed setae (cha: 32 – 36; chb: 20 – 24). Epimeral and lateral podosomal regions (Figs 3 B, C). Epimeral setal formula: 3 - 1 - 3 - 3. All setae setiform; 4 a and 4 b (16) slightly barbed, 1 a, 2 a and 3 a (20) heavily barbed, 1 b, 1 c, 3 b, 3 c, and 4 c (24 – 28) barbed. Pedotectum II with rounded lateral part. Discidium triangular, rounded apically, seta 4 c inserted on its dorsal side. Circumpedal carina comparatively short, posteriorly not reaching ventral plate, anteriorly not reaching pedotectum II. Anogenital region (Figs 3 B, C). Genital (g 1: 20; others: 14 – 16), aggenital (16), anal (14 – 16), and adanal (20) setae setiform, barbed. Adanal lyrifissure distinct. Marginal porose area band-like, complete. Legs (Figs 4 E – H). Tridactylous; all claws slightly barbed on dorsal side; lateral claws thinner median claw, with small tooth distoventrally. Femur II with broadly rounded ledge distoventrally. Dorsoparaxial porose area on femora I-IV and on trochanters III, IV, proximoventral porose area on tarsi I-IV and distoventral porose area on tibiae I-IV well visible. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 19) [1 - 2 - 2], II (1 - 5 - 2 - 4 - 15) [1 - 1 - 2], III (2 - 3 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 – 1 – 0]; homology of setae and solenidia indicated in Table 1. Famulus of tarsus I short, erect, slightly swollen distally, inserted between solenidion ω 2 and seta ft ”. Solenidion ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III bacilliform, other solenidia setiform. Seta s on tarsus I setiform, barbed (not eupathidial), inserted behind setae a’ and a ”.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	materials_examined	Type deposition. The holotype is deposited in the collection of the MUSM; 17 paratypes are deposited in the collection of the TSUMZ. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	etymology	Etymology. The specific name dianae is given in honour of Dr. Diana Silva Dávila, head of the arachnological department of the Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru for her always kind and valuable help and cooperation.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFF2FFC3FCC8502486FDFC3B.taxon	discussion	Remarks. In having narrowly spindleform bothridial seta with setiform apical tip, the new species is similar to Scheloribates (Hemileius) muscicola (Hammer, 1961) from Peru, but it differs from the latter by the smaller body length (365 – 498 versus 600), presence (versus absence) of anterolateral triangular notogastral processes and long (versus represented by alveoli) notogastral setae.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	description	(Figs 5, 6)	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	diagnosis	Diagnosis. Body size: 282 – 398 × 182 – 282. Prodorsum microfoveolate and slightly rugose; notogaster and ventral side micro- and macrofoveolate. Rostrum rounded. Prolamella and translamella absent. Rostral, lamellar and interlamellar setae long, setiform, barbed, in and le with apical swollen; ro shortest. Bothridial seta long, clavate, barbed. Notogastral setae setiform, barbed, with apical swollen; dm and h 2 of medium length, other setae long. Epimeral and anogenital setae short, setiform, barbed or roughened. Circumpedal carina long. Leg tarsus I with 19 setae (l ” absent); genu II with three setae (v’ present); femur II with four setae (l ” absent).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	description	Description of adult. Measurements. Body length: 398 (holotype: female), 282 (one male paratype), 332 – 398 (three female paratypes); body width: 265 (holotype), 182 (male paratype), 215 – 282 (female paratypes). Integument (Figs 1 A – C; 2 A, E, H). Body color brown. Prodorsum microfoveolate and slightly rugose. Notogaster and ventral side densely microfoveolate and sparsely macrofoveolate (diameter of foveola up to 8). Prodorsum (Figs 5 A, C). Rostrum rounded. Lamella about 1 / 2 length of prodorsum. Prolamella, translamella and sublamella absent. Sublamellar porose area (10 – 12) rounded. Tutorium with apical tooth. Rostral (49 – 53), lamellar (90 – 102) and interlamellar (94 – 110) setae setiform, barbed, le and in with apical swollen. Exobothridial seta (20 – 24) setiform, slightly barbed. Bothridial seta (77 – 82) clavate, barbed. Dorsosejugal porose area not observed. Notogaster (Figs 5 A, C). All notogastral setae (dm and h 2: 57 – 65; others: 106 – 147) setiform, barbed, with apically swollen tip. Four pairs of small sacculi. Opisthonotal gland opening and all lyrifissures distinct. Gnathosoma (Figs 6 A – D). Subcapitulum longer than wide (86 – 90 × 69 – 73). All subcapitular (a: 16; m: 20; h: 10) and adoral (10 – 12) setae setiform, barbed; h thinner than a and m. Palp (61 – 65) with typical setation: 0 - 2 - 1 - 3 - 9 (+ ω). Postpalpal seta (6) spiniform, roughened. Chelicera (98 – 102) with two setiform, barbed setae (cha: 32 – 34; chb: 20 – 22). Epimeral and lateral podosomal regions (Figs 5 B, C). Epimeral setal formula: 3 - 1 - 3 - 3. All setae setiform; 3 c (20 – 24) barbed, 1 b and 3 b (10 – 12) sparsely barbed, other setae (6 – 8) roughened. Pedotectum II with truncate lateral part. Discidium triangular, rounded apically. Custodium present. Circumpedal carina long, reaching anteriorly pedotectum II. Anogenital region (Figs 5 B, C). Genital (g 1: 10 – 12; others: 6 – 8), aggenital (6 – 8), anal (6 – 8), and adanal (6 – 8) setae setiform, roughened. Adanal lyrifissure distinct. Marginal porose represented by numerous rounded and oval parts. Legs (Figs 6 E – H). Tridactylous; all claws slightly barbed on dorsal side; lateral claws thinner median claw, with small tooth distoventrally. Tibiae I and II with tubercle proximoventrally. Femur II with broadly rounded ledge distoventrally. Dorsoparaxial porose area on femora I-IV and on trochanters III, IV, proximoventral porose area on tarsi I-IV and distoventral porose area on tibiae I-IV well visible. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 19) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 15) [1 - 1 - 2], III (2 - 3 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 – 1 – 0]; homology of setae and solenidia indicated in Table 1. Famulus of tarsus I short, erect, slightly swollen distally, inserted between solenidion ω 2 and seta ft ”. Solenidion ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III bacilliform, other solenidia setiform. Seta s on tarsus I eupathidial, inserted before setae a’ and a ”. Seta l ” on genu I inserted on tubercle.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	materials_examined	Type deposition. The holotype is deposited in the collection of the MUSM; four paratypes are deposited in the collection of the TSUMZ. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	etymology	Etymology. The specific name elisabethae is given in honour of Frau Elisabeth Hintelmann (Munich, Germany) for her effort and more than generous financial support of Panguana field station since many years.	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
132B8783FFFFFFC0FCC850AB81D6FCAC.taxon	discussion	Remarks. In having long notogastral setae with apically swollen tip, the new species is similar to P. buntotanus Corpuz-Raros, 1981, P. pilipinus Corpuz-Raros, 1981 and P. varisculptus Corpuz-Raros, 1981 from the Philippines, P. guttatoides Hammer, 1979 and P. guttatus Hammer, 1979 from the Oriental region, and P. moderatus Aoki, 1984 from Japan, but it differs from all above listed species by the presence of notogastral setae dm and h 2 distinctly shorter (versus not shorter) than other notogastral setae and lamellar seta with apical swollen (versus without swollen).	en	Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya., Friedrich, Stefan (2021): New sacculonotic Oripodoidea (Acari: Oribatida) from Peru. Zootaxa 5048 (3): 422-434, DOI: https://doi.org/10.11646/zootaxa.5048.3.7
